650:
758:
570:
500:
709:
737:
521:
824:
147:
421:
120:
671:
591:
2003:
1154:
472:
841:
by the way in which their wings develop. Endopterygota (literally "internal winged forms") develop wings inside the body and undergo an elaborate metamorphosis involving a pupal stage. Exopterygota ("external winged forms") develop wings on the outside their bodies and do not go through a pupal
948:
A. Nel; P. Roques; P. Nel; J. Prokop; J. S. Steyer (2007). "The earliest holometabolous insect from the
Carboniferous: a "crucial" innovation with delayed success (Insecta Protomeropina Protomeropidae)".
951:
2246:
2195:
1057:
2182:
2074:
887:
Wang, Y.-h. et al. Fossil record of stem groups employed in evaluating the chronogram of insects (Arthropoda: Hexapoda). Sci. Rep. 6, 38939;
649:
1050:
2187:
2079:
269:
146:
2241:
338:
The
Holometabola constitute the most diverse insect superorder, with over 1 million living species divided between 11
757:
285:
1043:
244:
2200:
569:
499:
279:
1687:
987:
319:
108:
2032:
1069:
263:
2236:
2208:
2087:
911:
1938:
708:
2169:
2143:
1880:
1860:
1953:
2251:
1968:
736:
1978:
1900:
1875:
968:
930:
234:
213:
141:
2213:
2092:
2134:
1895:
823:
1928:
1021:
863:
851:
1870:
1721:
1407:
1011:
1003:
960:
920:
907:"Endopterygote systematics – where do we stand and what is the goal (Hexapoda, Arthropoda)?"
888:
631:
520:
128:
1948:
1865:
1855:
1832:
1399:
339:
2066:
326:
and adult stages differing considerably in their structure and behaviour. This is called
2007:
1963:
1958:
1943:
1890:
1493:
1016:
991:
868:
827:
443:
420:
385:
327:
2230:
2148:
1918:
1910:
1136:
1128:
925:
906:
843:
466:
433:
374:
259:
70:
972:
934:
1850:
1623:
1600:
1592:
1489:
1321:
1264:
1093:
964:
838:
752:
585:
551:
453:
395:
Molecular analysis has clarified the group's phylogeny, as shown in the cladogram.
381:
670:
119:
2055:
2014:
2128:
1973:
1905:
1837:
1792:
1738:
1639:
1569:
1420:
1358:
1234:
831:
665:
644:
590:
494:
415:
133:
45:
1923:
1647:
1457:
1366:
1350:
1334:
1162:
992:"Progress, pitfalls and parallel universes: a history of insect phylogenetics"
564:
515:
299:
198:
90:
55:
2119:
17:
1885:
1729:
1689:
1664:
1517:
1428:
1387:
1374:
1205:
1192:
1153:
1087:
731:
621:
389:
343:
290:
178:
158:
95:
1025:
1007:
471:
294:
2156:
2113:
2049:
1933:
1842:
1631:
1501:
1293:
1099:
307:
85:
80:
65:
60:
50:
2161:
2002:
1272:
847:
703:
384:(Neuroptera, Megaloptera, Raphidioptera, Strepsiptera and Coleoptera),
100:
75:
892:
1609:
1509:
1436:
1342:
1251:
1243:
1081:
1066:
370:
363:
323:
303:
188:
168:
2026:
392:(Siphonaptera, Diptera, Trichoptera, Lepidoptera and Mecoptera).
311:
2174:
1697:
1035:
850:(dragonflies and damselflies), which are not Neoptera, but more
351:
315:
2030:
1817:
1112:
1039:
380:
The
Holometabola are sometimes divided into three assemblages:
1707:
359:
355:
347:
1650:(net-winged insects: lacewings, mantidflies, antlions)
990:; Yavorskaya, Margarita & Beutel, Rolf G. (2016).
2103:
2039:
1720:
1678:
1663:
1622:
1591:
1582:
1533:
1487:
1456:
1419:
1386:
1319:
1292:
1263:
1242:
1233:
1204:
1161:
1127:
846:, however, as it is found also in groups such as
1051:
952:Annales de la Société Entomologique de France
837:The Endopterygota are distinguished from the
8:
318:, and adult stages. They undergo a radical
2027:
1825:
1814:
1675:
1588:
1561:
1484:
1480:
1327:
1316:
1312:
1239:
1230:
1226:
1185:
1124:
1120:
1109:
1058:
1044:
1036:
118:
31:
1015:
924:
1779:Four most speciose orders are marked in
1369:(crickets, wetas, grasshoppers, locusts)
822:
880:
2247:Extant Pennsylvanian first appearances
996:Journal of the Royal Society Interface
107:Molecular clock indicates a possible
7:
1642:(alderflies, dobsonflies, fishflies)
724:
696:
637:
627:
557:
487:
459:
449:
439:
408:
398:
905:Rolf G. Beutel; Hans Pohl (2006).
25:
2001:
1152:
926:10.1111/j.1365-3113.2006.00341.x
756:
735:
707:
669:
648:
589:
568:
519:
498:
470:
419:
145:
2064:Cockroach Species File (new):
965:10.1080/00379271.2007.10697531
418:(sawflies, wasps, ants, bees)
1:
1572:(sawflies, wasps, ants, bees)
342:, containing insects such as
1520:(cicadas, aphids, true bugs)
330:, or complete metamorphism.
310:that go through distinctive
842:stage. The latter trait is
369:The earliest holometabolan
2268:
1710:(gnats, mosquitoes, flies)
1603:(twisted-winged parasites)
1275:(dragonflies, damselflies)
1995:
1828:
1824:
1813:
1775:
1564:
1483:
1330:
1315:
1229:
1188:
1150:
1123:
1119:
1108:
1076:
750:
729:
722:
701:
694:
663:
642:
635:
625:
588:(twisted-wing parasites)
583:
562:
555:
513:
492:
485:
464:
457:
447:
437:
413:
406:
273:"change"), also known as
240:
233:
226:
221:
142:Scientific classification
140:
126:
117:
34:
1361:(stick and leaf insects)
497:(alderflies and allies)
1788:are paraphyletic groups
1431:(cockroaches, termites)
1195:(silverfish, firebrats)
518:(Lacewings and allies)
284:
278:
262:
1139:(jumping bristletails)
1008:10.1098/rsif.2016.0363
834:
306:within the infraclass
293:
268:
2209:Paleobiology Database
2088:Paleobiology Database
912:Systematic Entomology
826:
668:(butterflies, moths)
27:Superorder of insects
1741:(moths, butterflies)
41:Pennsylvanian–Recent
388:(Hymenoptera), and
2242:Insect superorders
2008:Insects portal
1979:Triadophlebioptera
835:
136:(order Mecoptera)
2224:
2223:
2033:Taxon identifiers
2024:
2023:
1991:
1990:
1987:
1986:
1929:Palaeodictyoptera
1914:
1846:
1809:
1808:
1805:
1804:
1771:
1770:
1767:
1766:
1763:
1762:
1759:
1758:
1755:
1754:
1751:
1750:
1742:
1733:
1711:
1701:
1693:
1671:
1659:
1658:
1651:
1643:
1635:
1613:
1604:
1573:
1529:
1528:
1521:
1513:
1505:
1452:
1451:
1448:
1447:
1440:
1432:
1411:
1408:Mantophasmatodea
1403:
1392:
1378:
1370:
1362:
1354:
1346:
1338:
1288:
1287:
1284:
1283:
1276:
1255:
1244:Ephemeropteroidea
1196:
1148:
1147:
1140:
893:10.1038/srep38939
864:Insect morphology
815:
814:
805:
804:
796:
795:
787:
786:
778:
777:
769:
768:
734:(scorpionflies)
682:
681:
611:
610:
602:
601:
541:
540:
532:
531:
298:"-having"), is a
253:
252:
248:
217:
112:
16:(Redirected from
2259:
2217:
2216:
2204:
2203:
2191:
2190:
2178:
2177:
2165:
2164:
2152:
2151:
2139:
2138:
2137:
2124:
2123:
2122:
2096:
2095:
2083:
2082:
2070:
2069:
2060:
2059:
2058:
2028:
2006:
2005:
1939:Permoplectoptera
1912:
1871:Diaphanopterodea
1844:
1826:
1815:
1740:
1731:
1722:Amphiesmenoptera
1709:
1703:
1699:
1691:
1681:
1676:
1670:
1667:
1649:
1641:
1633:
1611:
1602:
1589:
1585:
1571:
1562:
1519:
1511:
1504:(barklice, lice)
1503:
1495:
1485:
1481:
1438:
1430:
1409:
1401:
1400:Grylloblattodea
1390:
1376:
1368:
1360:
1352:
1344:
1336:
1328:
1324:
1317:
1313:
1274:
1253:
1240:
1231:
1227:
1194:
1186:
1156:
1138:
1125:
1121:
1110:
1060:
1053:
1046:
1037:
1030:
1029:
1019:
983:
977:
976:
945:
939:
938:
928:
902:
896:
885:
811:(Endopterygota)
760:
739:
725:
711:
697:
673:
652:
638:
632:Amphiesmenoptera
628:
593:
572:
558:
523:
502:
488:
474:
460:
450:
440:
423:
409:
399:
243:
212:
150:
149:
129:Panorpa communis
122:
106:
105:
42:
38:Temporal range:
32:
21:
2267:
2266:
2262:
2261:
2260:
2258:
2257:
2256:
2227:
2226:
2225:
2220:
2212:
2207:
2199:
2194:
2186:
2181:
2173:
2168:
2160:
2155:
2147:
2142:
2133:
2132:
2127:
2118:
2117:
2112:
2099:
2091:
2086:
2078:
2073:
2065:
2063:
2054:
2053:
2048:
2035:
2025:
2020:
2000:
1983:
1949:Protelytroptera
1881:Eudiaphanoptera
1866:Coxoplectoptera
1861:Carbotriplurida
1856:Campylopteridae
1833:Aethiocarenodea
1820:
1801:
1747:
1716:
1692:(scorpionflies)
1679:
1668:
1666:
1655:
1618:
1583:
1578:
1556:
1554:
1552:
1550:
1548:
1546:
1544:
1542:
1540:
1538:
1536:
1525:
1475:
1473:
1471:
1469:
1467:
1465:
1463:
1461:
1459:
1444:
1415:
1389:
1382:
1377:(angel insects)
1320:
1307:
1305:
1303:
1301:
1299:
1297:
1295:
1280:
1259:
1221:
1219:
1217:
1215:
1213:
1211:
1209:
1207:
1200:
1180:
1178:
1176:
1174:
1172:
1170:
1168:
1166:
1164:
1157:
1144:
1115:
1104:
1072:
1064:
1034:
1033:
986:Kjer, Karl M.;
985:
984:
980:
947:
946:
942:
904:
903:
899:
886:
882:
877:
860:
854:among insects.
821:
816:
806:
797:
788:
779:
770:
683:
612:
603:
542:
533:
336:
211:
144:
113:
104:
103:
98:
93:
88:
83:
78:
73:
68:
63:
58:
53:
48:
40:
39:
36:
28:
23:
22:
15:
12:
11:
5:
2265:
2263:
2255:
2254:
2249:
2244:
2239:
2229:
2228:
2222:
2221:
2219:
2218:
2205:
2192:
2179:
2166:
2153:
2140:
2125:
2109:
2107:
2101:
2100:
2098:
2097:
2084:
2071:
2061:
2045:
2043:
2037:
2036:
2031:
2022:
2021:
2019:
2018:
2011:
1996:
1993:
1992:
1989:
1988:
1985:
1984:
1982:
1981:
1976:
1971:
1966:
1964:Protozygoptera
1961:
1959:Protorthoptera
1956:
1954:Protephemerida
1951:
1946:
1944:Protanisoptera
1941:
1936:
1931:
1926:
1921:
1916:
1913:(griffinflies)
1911:Meganisoptera
1908:
1903:
1898:
1893:
1891:Glosselytrodea
1888:
1883:
1878:
1873:
1868:
1863:
1858:
1853:
1848:
1840:
1835:
1829:
1822:
1821:
1818:
1811:
1810:
1807:
1806:
1803:
1802:
1800:
1799:
1789:
1783:
1776:
1773:
1772:
1769:
1768:
1765:
1764:
1761:
1760:
1757:
1756:
1753:
1752:
1749:
1748:
1746:
1745:
1735:
1726:
1724:
1718:
1717:
1715:
1714:
1704:
1684:
1682:
1673:
1661:
1660:
1657:
1656:
1654:
1653:
1645:
1637:
1632:Raphidioptera
1628:
1626:
1620:
1619:
1617:
1616:
1606:
1597:
1595:
1586:
1584:Neuropteroidea
1580:
1579:
1577:
1576:
1565:
1559:
1531:
1530:
1527:
1526:
1524:
1523:
1515:
1507:
1498:
1496:
1478:
1454:
1453:
1450:
1449:
1446:
1445:
1443:
1442:
1434:
1425:
1423:
1417:
1416:
1414:
1413:
1405:
1402:(ice-crawlers)
1396:
1394:
1384:
1383:
1381:
1380:
1372:
1364:
1356:
1348:
1340:
1331:
1325:
1310:
1290:
1289:
1286:
1285:
1282:
1281:
1279:
1278:
1269:
1267:
1261:
1260:
1258:
1257:
1252:Ephemeroptera
1248:
1246:
1237:
1224:
1202:
1201:
1199:
1198:
1189:
1183:
1159:
1158:
1151:
1149:
1146:
1145:
1143:
1142:
1137:Archaeognatha
1133:
1131:
1117:
1116:
1113:
1106:
1105:
1103:
1102:
1096:
1090:
1084:
1077:
1074:
1073:
1065:
1063:
1062:
1055:
1048:
1040:
1032:
1031:
978:
959:(3): 349–355.
940:
919:(2): 202–219.
897:
879:
878:
876:
873:
872:
871:
869:Holometabolism
866:
859:
856:
828:Holometabolism
820:
817:
813:
812:
808:
807:
803:
802:
799:
798:
794:
793:
790:
789:
785:
784:
781:
780:
776:
775:
772:
771:
767:
766:
763:
762:
749:
746:
745:
742:
741:
728:
723:
721:
718:
717:
714:
713:
700:
695:
693:
689:
688:
685:
684:
680:
679:
676:
675:
662:
659:
658:
655:
654:
647:(caddisflies)
641:
636:
634:
626:
624:
618:
617:
614:
613:
609:
608:
605:
604:
600:
599:
596:
595:
582:
579:
578:
575:
574:
561:
556:
554:
548:
547:
544:
543:
539:
538:
535:
534:
530:
529:
526:
525:
512:
509:
508:
505:
504:
491:
486:
484:
481:
480:
477:
476:
463:
458:
456:
448:
446:
444:Neuropteroidea
438:
436:
430:
429:
426:
425:
412:
407:
405:
397:
386:Hymenopteroida
373:date from the
335:
332:
328:holometabolism
251:
250:
242:Endopterygota
238:
237:
231:
230:
224:
223:
219:
218:
206:
202:
201:
196:
192:
191:
186:
182:
181:
176:
172:
171:
166:
162:
161:
156:
152:
151:
138:
137:
124:
123:
115:
114:
99:
94:
89:
84:
79:
74:
69:
64:
59:
54:
49:
44:
43:
37:
26:
24:
14:
13:
10:
9:
6:
4:
3:
2:
2264:
2253:
2250:
2248:
2245:
2243:
2240:
2238:
2235:
2234:
2232:
2215:
2210:
2206:
2202:
2197:
2193:
2189:
2184:
2180:
2176:
2171:
2167:
2163:
2158:
2154:
2150:
2145:
2141:
2136:
2130:
2126:
2121:
2115:
2111:
2110:
2108:
2106:
2105:Endopterygota
2102:
2094:
2089:
2085:
2081:
2076:
2072:
2068:
2062:
2057:
2051:
2047:
2046:
2044:
2042:
2038:
2034:
2029:
2017:
2016:
2012:
2010:
2009:
2004:
1998:
1997:
1994:
1980:
1977:
1975:
1972:
1970:
1967:
1965:
1962:
1960:
1957:
1955:
1952:
1950:
1947:
1945:
1942:
1940:
1937:
1935:
1932:
1930:
1927:
1925:
1922:
1920:
1919:Megasecoptera
1917:
1915:
1909:
1907:
1904:
1902:
1899:
1897:
1894:
1892:
1889:
1887:
1884:
1882:
1879:
1877:
1874:
1872:
1869:
1867:
1864:
1862:
1859:
1857:
1854:
1852:
1849:
1847:
1841:
1839:
1836:
1834:
1831:
1830:
1827:
1823:
1816:
1812:
1798:
1796:
1790:
1787:
1784:
1782:
1778:
1777:
1774:
1744:
1743:
1736:
1734:
1732:(caddisflies)
1728:
1727:
1725:
1723:
1719:
1713:
1712:
1705:
1702:
1698:Siphonaptera
1695:
1694:
1686:
1685:
1683:
1677:
1674:
1672:
1669:(Mecopterida)
1662:
1652:
1646:
1644:
1638:
1636:
1630:
1629:
1627:
1625:
1621:
1615:
1614:
1607:
1605:
1601:Strepsiptera
1599:
1598:
1596:
1594:
1590:
1587:
1581:
1575:
1574:
1567:
1566:
1563:
1560:
1558:
1532:
1522:
1516:
1514:
1510:Thysanoptera
1508:
1506:
1500:
1499:
1497:
1492:
1491:
1486:
1482:
1479:
1477:
1455:
1441:
1435:
1433:
1427:
1426:
1424:
1422:
1418:
1412:
1406:
1404:
1398:
1397:
1395:
1393:
1385:
1379:
1373:
1371:
1365:
1363:
1357:
1355:
1353:(webspinners)
1349:
1347:
1341:
1339:
1333:
1332:
1329:
1326:
1323:
1318:
1314:
1311:
1309:
1291:
1277:
1271:
1270:
1268:
1266:
1262:
1256:
1250:
1249:
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1238:
1236:
1232:
1228:
1225:
1223:
1203:
1197:
1191:
1190:
1187:
1184:
1182:
1160:
1155:
1141:
1135:
1134:
1132:
1130:
1126:
1122:
1118:
1111:
1107:
1101:
1097:
1095:
1091:
1089:
1085:
1083:
1079:
1078:
1075:
1071:
1068:
1061:
1056:
1054:
1049:
1047:
1042:
1041:
1038:
1027:
1023:
1018:
1013:
1009:
1005:
1001:
997:
993:
989:
982:
979:
974:
970:
966:
962:
958:
954:
953:
944:
941:
936:
932:
927:
922:
918:
914:
913:
908:
901:
898:
894:
890:
884:
881:
874:
870:
867:
865:
862:
861:
857:
855:
853:
849:
845:
844:plesiomorphic
840:
833:
829:
825:
818:
810:
809:
801:
800:
792:
791:
783:
782:
774:
773:
765:
764:
761:
759:
754:
748:
747:
744:
743:
740:
738:
733:
727:
726:
720:
719:
716:
715:
712:
710:
706:(true flies)
705:
699:
698:
691:
690:
687:
686:
678:
677:
674:
672:
667:
661:
660:
657:
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633:
630:
629:
623:
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619:
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598:
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587:
581:
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566:
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553:
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537:
536:
528:
527:
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522:
517:
511:
510:
507:
506:
503:
501:
496:
490:
489:
483:
482:
479:
478:
475:
473:
469:(snakeflies)
468:
467:Raphidioptera
462:
461:
455:
452:
451:
445:
442:
441:
435:
434:Aparaglossata
432:
431:
428:
427:
424:
422:
417:
411:
410:
404:
401:
400:
396:
393:
391:
387:
383:
378:
376:
375:Carboniferous
372:
367:
365:
361:
357:
353:
349:
345:
341:
333:
331:
329:
325:
321:
320:metamorphosis
317:
313:
309:
305:
301:
297:
296:
292:
288:
287:
282:
281:
276:
275:Endopterygota
272:
271:
267:"complete" +
266:
265:
261:
260:Ancient Greek
257:
249:
246:
239:
236:
232:
229:
225:
220:
215:
210:
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204:
203:
200:
197:
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190:
187:
184:
183:
180:
177:
174:
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170:
167:
164:
163:
160:
157:
154:
153:
148:
143:
139:
135:
131:
130:
125:
121:
116:
110:
109:Mississippian
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
33:
30:
19:
18:Endopterygota
2237:Holometabola
2149:Holometabola
2135:Holometabola
2104:
2041:Holometabola
2040:
2013:
1999:
1969:Syntonoptera
1851:Caloneurodea
1843:Blattoptera
1794:
1785:
1780:
1739:Lepidoptera
1737:
1730:Trichoptera
1706:
1688:
1640:Megaloptera
1634:(snakeflies)
1624:Neuropterida
1608:
1593:Coleopterida
1570:Hymenoptera
1568:
1534:
1490:Paraneoptera
1488:
1410:(gladiators)
1359:Phasmatodea
1337:(stoneflies)
1322:Polyneoptera
1265:Odonatoptera
1129:Monocondylia
1094:Pancrustacea
1092:(unranked):
1002:(121): 121.
999:
995:
988:Simon, Chris
981:
956:
950:
943:
916:
910:
900:
883:
839:Exopterygota
836:
753:Siphonaptera
751:
730:
702:
692:Antliophora
664:
643:
586:Strepsiptera
584:
563:
552:Coleopterida
514:
493:
465:
454:Neuropterida
414:
403:Holometabola
402:
394:
382:Neuropterida
379:
368:
337:
274:
256:Holometabola
255:
254:
241:
227:
209:Holometabola
208:
205:(unranked):
195:(unranked):
127:
35:Holometabola
29:
2129:Wikispecies
2015:Wikispecies
1974:Titanoptera
1906:Lapeyriidae
1901:Hypoperlida
1876:Eoblattodea
1845:(roachoids)
1838:Archodonata
1680:Antliophora
1648:Neuroptera
1610:Coleoptera
1421:Dictyoptera
1391:(Xenonomia)
1367:Orthoptera
1351:Embioptera
1343:Dermaptera
1335:Plecoptera
1235:Palaeoptera
1098:Subphylum:
832:Hymenoptera
819:Description
666:Lepidoptera
645:Trichoptera
495:Megaloptera
416:Hymenoptera
344:butterflies
322:, with the
134:scorpionfly
2252:Eumetabola
2231:Categories
1924:Miomoptera
1690:Mecoptera
1518:Hemiptera
1439:(mantises)
1429:Blattodea
1375:Zoraptera
1254:(mayflies)
1193:Zygentoma
1088:Arthropoda
875:References
830:stages in
567:(beetles)
565:Coleoptera
516:Neuroptera
300:superorder
283:"inner" +
214:Burmeister
199:Eumetabola
179:Arthropoda
2056:Q37140800
1896:Heraridea
1886:Geroptera
1791:Based on
1665:Panorpida
1612:(beetles)
1502:Psocodea
1437:Mantodea
1388:Notoptera
1345:(earwigs)
1080:Kingdom:
732:Mecoptera
622:Panorpida
390:Panorpida
334:Evolution
291:Neo-Latin
289:"wing" +
228:See text
165:Kingdom:
159:Eukaryota
2157:BugGuide
2114:Wikidata
2050:Wikidata
1934:Paoliida
1708:Diptera
1512:(thrips)
1273:Odonata
1100:Hexapoda
1086:Phylum:
1082:Animalia
1026:27558853
973:86235521
935:83714402
858:See also
755:(fleas)
308:Neoptera
270:metabolḗ
235:Synonyms
175:Phylum:
169:Animalia
155:Domain:
2175:3016961
2120:Q304358
1819:Extinct
1793:Sasaki
1700:(fleas)
1494: *
1017:5014063
895:(2016).
848:Odonata
704:Diptera
371:fossils
364:beetles
304:insects
286:ptéryg-
222:Orders
189:Insecta
185:Class:
2214:133932
2188:109215
2162:172251
2093:198428
2080:914213
2067:856736
1797:(2013)
1795:et al.
1786:Italic
1114:Extant
1070:orders
1067:Insect
1024:
1014:
971:
933:
362:, and
340:orders
324:larval
312:larval
277:(from
258:(from
247:, 1898
216:, 1835
111:origin
2201:33392
969:S2CID
931:S2CID
852:basal
352:fleas
348:flies
316:pupal
280:endo-
264:holo-
245:Sharp
2196:NCBI
2183:ITIS
2075:ITIS
1781:bold
1022:PMID
360:ants
356:bees
295:-ota
132:, a
46:PreꞒ
2170:EoL
2144:ADW
1012:PMC
1004:doi
961:doi
921:doi
889:doi
302:of
2233::
2211::
2198::
2185::
2172::
2159::
2146::
2131::
2116::
2090::
2077::
2052::
1696:+
1020:.
1010:.
1000:13
998:.
994:.
967:.
957:43
955:.
929:.
917:31
915:.
909:.
377:.
366:.
358:,
354:,
350:,
346:,
314:,
96:Pg
1557:a
1555:l
1553:o
1551:b
1549:a
1547:t
1545:e
1543:m
1541:o
1539:l
1537:o
1535:H
1476:a
1474:l
1472:o
1470:b
1468:a
1466:t
1464:e
1462:m
1460:u
1458:E
1308:a
1306:r
1304:e
1302:t
1300:p
1298:o
1296:e
1294:N
1222:a
1220:t
1218:o
1216:g
1214:y
1212:r
1210:e
1208:t
1206:P
1181:a
1179:i
1177:l
1175:y
1173:d
1171:n
1169:o
1167:c
1165:i
1163:D
1059:e
1052:t
1045:v
1028:.
1006::
975:.
963::
937:.
923::
891::
101:N
91:K
86:J
81:T
76:P
71:C
66:D
61:S
56:O
51:Ꞓ
20:)
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