533:
throughout the lower jaw, with the exception of the region between the sixth and ninth dentary teeth. The sixth and seventh tooth are very closely spaced, as are the eight and ninth, with prominent diastemas separating them from each other as well as the fifth and tenth dentary teeth. The eight tooth also happens to be the smallest tooth in the lower jaw, whereas the 11th and 13th are the largest. All teeth are laterally compressed, but unlike other peirosaurids they lack the serrations or crenulations that would make them
122:
1984:
104:
2556:
2003:
2549:
2563:
2571:
1996:
1990:
1171:, which may suggest a similar semi-aquatic lifestyle. Pepesuchines are however not perfect anatomical analogues to neosuchians, as they still have some adaptations seen in more terrestrial groups. Rather than having eyes located atop the head, as would be useful for laying in ambush with the body submerged, the eyes of
518:
had a long and narrow snout, with the preserved portion alone measuring 35 cm (14 in). It's roughly constant in height safe for a small increase towards the back of the skull. The dentaries are also not compressed mediolaterally but have a convex outer surface, which supports the placement
611:. Within Peirosauridae, Ruiz and colleagues recovered a clear split between two distinct lineages, peirosaurines and pepesuchines. While the former are described as semi-aquatic to terrestrial animals with oreinirostral snouts (meaning their snouts appeared tall and domed) and includes taxa like
532:
The lower jaw preserves a total of 14 teeth of varying size and diameter, with the individual teeth and alveoli separated by prominent gaps that likely serve to allow the teeth of the upper and lower jaw to interlock. The distance between the individual teeth or alveoli is relatively consistent
523:
within
Pepesuchinae. The edges of the lower jaw seem to run subparallel to the length of the skull as a whole and don't display noticeable narrowing towards the tip nor widening towards the back. However, there is a noticeable notch in the lower jaw close to the fourth dentary tooth, which might
1255:
Ruiz, Juan V.; Queiroz, Marcos V. L.; Martins, Kawan C.; Godoy, Pedro L.; Iori, Fabiano V.; Langer, Max C.; Montefeltro, Felipe C.; Bronzati, Mario (2024-08-29). "A new
Peirosauridae (Crocodyliformes, Notosuchia) from the Adamantina Formation (Bauru Group, Late Cretaceous), with a revised
1147:, which consists of peirosaurids, itasuchids and mahajangasuchids. Within Peirosauria, Peirosauridae and Itasuchidae are roughly equivalent to the clades Peirosaurinae and Pepesuchinae as recovered by Ruiz and colleagues.
1314:
Fachini, T.S.; Godoy, P.L.; Marsola, J.C.A.; Montefeltro, F.C.; Langer, M.C. (2022). "A large-sized mesoeucrocodylian from the Late
Cretaceous of Brazil with possible neosuchian affinities".
2623:
1209:. Though the Bauru Group is renown for its diverse notosuchian fauna, only a single neosuchian has been recovered from any of the formations that are part of the unit. Said neosuchian is
1159:
had elongated and narrow jaws that are wildly different from the tall, deep skulls seen in peirosaurines. Such a skull shape is unique among notosuchians, but much more widespread among
396:
had long and gracile jaws, which may indicate that they were semi-aquatic animals more similar to modern crocodilians. The genus is monotypic, meaning it only contains a single species,
426:. The specimen; which consists of a nearly tubular snout, a partial skull roof, parts of the mandible and a single rib; was recovered from a road cut and subsequently sent to the
1281:
Leardi, J. M.; Pol, D.; Montefeltro, F.; Marinho, T. S.; Ruiz, J. V.; Bravo, G. G.; Pinheiro, A. E. P.; Godoy, P. L.; Nicholl, C. S. C.; Lecuona, A.; Larsson, H. C. E. (2024).
1175:
faced towards the side of the skull. Considering both these similarities and differences might suggest that though more semi-aquatic than their relatives, pepesuchines such as
1179:
were still less dependent on water than modern crocodilians. Semi-aquatic habits have also been reported from mahajangasuchine notosuchians, which includes
453:. The use of "epoide" is a reference to the city of Catanduva, which is also known as the "Magic Spell City". The species name on the other hand honors
434:. The animal was eventually fully described in 2024 and recognized as a distinct species and genus by Juan V. Ruiz and colleagues, who coined the name
2633:
1127:
2018 Peirosauridae would be restricted to the clade they dub
Peirosaurinae. Their version of Pepesuchinae meanwhile would be separated into the clade
486:, with the distance between it and the eyesocket being longer than the fossa itself is high. The skull table is dominated by two large openings, the
427:
2638:
1359:
2598:
1123:
Though the name
Pepesuchinae is used by Ruiz and colleagues to describe the clade, they do note that following the terminology of Pinheiro
629:, the later consists primarily of animals that are hypothesized to be more aquatic, possessing elongated or flattened snouts as seen in
482:
possessed a small opening in the skull before the eyes, known as the antorbital fossa. The fossa is separated from the eyesocket by the
1197:, with the latter being much closer in anatomy to pepesuchines. This might suggest some degree of convergence between pepesuchines and
490:, the rims of which are on level with the rest of the skull roof rather than raised. The skull roof extends the furthest back with the
595:
recovers a tree that alligns with the so-called "Sebecia hypothesis", meaning that peirosaurids are found to clade with the families
478:, a piece of a lacrimal and a skull fragment that preserves parts of the region that sits before the eyes. Unlike modern crocodiles,
1139:
would not be regarded as a pepesuchine nor an itasuchid, eventhough its position would not change. Furthermore, the description of
449:"epoide" meaning "enchanted" and "souchus", a commonly used suffix among crocodilians that harkons back to the Egyptian god
2628:
1201:
specifically. Ruiz and colleagues note that pepesuchines often occur in units were neosuchians are rare or even absent.
121:
2555:
2643:
1983:
1143:
does not take into account the 2024 nomenclature published by Leardi and colleagues, in which they coin the clade
1352:
603:. This hypothesis stands opposed to the "Sebecosuchia hypothesis", in which sebecids are most closely related to
498:. Notably, beyond this point the bones lack the ornamentation that is present across most the rest of the skull.
2002:
2548:
454:
2562:
487:
2570:
2648:
1444:
1258:
559:
was at least 50% larger than other peirosaurids that inhabited the
Adamantina Formation, which includes
334:
1995:
1345:
415:
385:
674:
431:
392:. Like other members of the Pepesuchinae and unlike the closely related members of Peirosaurinae,
2298:
1144:
116:
1989:
1135:
and the recovered polytomy) and various more basal taxa. Subsequently, following this taxonomy
1858:
1588:
1216:
723:
600:
2442:
1404:
1323:
1294:
1263:
2501:
2476:
2314:
2287:
2232:
2207:
2200:
2154:
2107:
2081:
1893:
1865:
1604:
1181:
1164:
625:
608:
579:
215:
202:
2517:
2467:
2429:
2375:
2353:
2239:
2177:
2124:
1851:
1775:
1754:
1652:
1569:
892:
820:
543:
491:
103:
2617:
2510:
2483:
2449:
2410:
2395:
2368:
2361:
2307:
2273:
2266:
2257:
2219:
2184:
2096:
2066:
2059:
1928:
1844:
1837:
1821:
1747:
1719:
1211:
770:
757:
619:
604:
495:
483:
446:
381:
373:
276:
61:
2490:
2386:
2340:
2131:
2074:
1900:
1872:
1830:
1782:
1766:
1739:
1732:
1694:
1631:
1429:
1168:
957:
940:
782:
631:
613:
511:
370:
289:
247:
189:
1327:
494:, which project back and upward even after their lateral processes connect to the
457:, a paleontologist and the director of the museum that houses the type specimen.
2403:
2321:
2280:
2192:
2164:
2051:
1921:
1907:
1810:
1708:
1659:
1645:
1638:
1624:
1597:
1555:
1524:
1506:
1493:
1456:
1368:
1206:
1187:
1128:
991:
844:
637:
573:
475:
36:
2456:
2418:
2138:
2117:
2089:
2033:
2020:
1951:
1914:
1886:
1879:
1803:
1687:
1680:
1673:
1577:
1413:
1398:
1205:
for example was native to Brazil's
Adamantina Formation, which is part of the
1193:
916:
796:
661:
643:
567:
534:
231:
81:
46:
2246:
1796:
1666:
1615:
1562:
1283:"Phylogenetic nomenclature of Notosuchia (Crocodylomorpha; Crocodyliformes)"
1160:
974:
745:
700:
596:
561:
423:
377:
176:
133:
86:
1392:
153:
76:
71:
56:
51:
41:
1701:
1541:
1299:
1282:
686:
430:. The material was first identified as a peirosaurid that same year by
263:
163:
91:
66:
1386:
1267:
419:
389:
143:
524:
accommodate an enlarged maxillary tooth when the jaws were closed.
450:
474:
is known from various incomplete remains, including much of the
2587:
2018:
1491:
1379:
1341:
1215:, which has so far only been found within the sediments of the
428:
1337:
591:
The phylogenetic analysis conducted during the description of
510:
is the left hemimandible, which preserves most of the
2500:
2466:
2428:
2385:
2339:
2297:
2256:
2218:
2163:
2152:
2106:
2031:
1950:
1820:
1765:
1718:
1614:
1587:
1539:
1504:
1455:
1428:
607:while peirosaurids clade with mahajangasuchids and
514:. The size and shape of the mandible suggests that
2624:Late Cretaceous crocodylomorphs of South America
1353:
8:
1155:Like several other members of Pepesuchinae,
2584:
2434:
2345:
2224:
2169:
2160:
2043:
2039:
2028:
2015:
1947:
1724:
1547:
1516:
1512:
1501:
1488:
1434:
1425:
1376:
1360:
1346:
1338:
102:
20:
1298:
555:The size of the lower jaw indicates that
1228:
414:were found in 2011 in outcrops of the
1287:Bulletin of Phylogenetic Nomenclature
1250:
1248:
1246:
1244:
1242:
1240:
1238:
1236:
1234:
1232:
7:
537:or pseudo-ziphodont, something that
506:Another largely complete element of
1256:phylogenetic analysis of Sebecia".
932:
908:
884:
860:
836:
812:
788:
763:
738:
716:
692:
667:
657:
650:
14:
2634:Prehistoric pseudosuchian genera
2569:
2561:
2554:
2547:
2001:
1994:
1988:
1982:
120:
1:
2639:Fossil taxa described in 2024
1328:10.1080/08912963.2022.2122822
1131:(which would be composed of
2599:Terrestrial crocodylomorphs
2665:
376:from the Late Cretaceous (
2594:
2583:
2545:
2437:
2348:
2227:
2172:
2046:
2042:
2027:
2014:
1980:
1946:
1727:
1550:
1519:
1515:
1500:
1487:
1437:
1424:
1375:
1191:, as well as the sebecid
988:
971:
954:
937:
930:
913:
906:
889:
882:
865:
858:
841:
834:
817:
810:
793:
786:
768:
761:
743:
736:
721:
714:
697:
690:
672:
665:
655:
410:The holotype specimen of
340:
333:
117:Scientific classification
115:
110:
101:
23:
541:shares with the African
455:Sandra Simionato Tavares
445:is a combination of the
16:Extinct genus of reptile
32:Campanian–Maastrichtian
701:Doratodon carcharidens
488:supratemporal fenestra
436:Epoidesuchus tavaresae
399:Epoidesuchus tavaresae
347:Epoidesuchus tavaresae
1259:The Anatomical Record
2629:Adamantina Formation
416:Adamantina Formation
386:Adamantina Formation
2589:Related categories
2299:Pissarrachampsinae
1316:Historical Biology
1300:10.11646/bpn.1.3.2
406:History and naming
2644:Fossils of Brazil
2611:
2610:
2607:
2606:
2579:
2578:
2543:
2542:
2539:
2538:
2535:
2534:
2531:
2530:
2527:
2526:
2335:
2334:
2331:
2330:
2148:
2147:
2010:
2009:
1978:
1977:
1974:
1973:
1970:
1969:
1942:
1941:
1938:
1937:
1859:Colhuehuapisuchus
1589:Mahajangasuchidae
1535:
1534:
1483:
1482:
1479:
1478:
1475:
1474:
1322:(10): 1817–1830.
1217:Marilia Formation
1120:
1119:
1111:
1110:
1102:
1101:
1093:
1092:
1084:
1083:
1075:
1074:
1066:
1065:
1057:
1056:
1048:
1047:
1039:
1038:
1030:
1029:
1021:
1020:
1012:
1011:
1003:
1002:
724:Mahajangasuchidae
601:Mahajangasuchidae
362:
361:
326:E. tavaresae
314:
111:Life Restoration
2656:
2585:
2574:
2573:
2566:
2565:
2559:
2558:
2551:
2443:Adamantinasuchus
2435:
2346:
2225:
2170:
2161:
2044:
2040:
2029:
2016:
2005:
1999:
1998:
1992:
1986:
1948:
1725:
1548:
1517:
1513:
1502:
1489:
1435:
1426:
1419:
1418:
1405:Mesoeucrocodylia
1377:
1362:
1355:
1348:
1339:
1332:
1331:
1311:
1305:
1304:
1302:
1278:
1272:
1271:
1268:10.1002/ar.25559
1252:
933:
909:
885:
861:
837:
813:
789:
764:
739:
717:
693:
668:
658:
651:
422:municipality of
349:
345:
308:
301:
288:
275:
262:
246:
230:
125:
124:
106:
96:
33:
29:Temporal range:
21:
2664:
2663:
2659:
2658:
2657:
2655:
2654:
2653:
2614:
2613:
2612:
2603:
2590:
2575:
2568:
2560:
2553:
2523:
2502:Caipirasuchinae
2496:
2477:Armadillosuchus
2462:
2424:
2381:
2327:
2315:Pissarrachampsa
2293:
2288:Stratiotosuchus
2252:
2233:Cynodontosuchus
2214:
2208:Razanandrongobe
2201:Pehuenchesuchus
2157:
2155:Xenodontosuchia
2144:
2108:Candidodontidae
2102:
2082:Razanandrongobe
2036:
2023:
2006:
1993:
1966:
1934:
1894:Montealtosuchus
1866:Gasparinisuchus
1816:
1761:
1714:
1610:
1605:Mahajangasuchus
1583:
1544:
1531:
1509:
1496:
1471:
1451:
1420:
1382:
1381:
1371:
1366:
1336:
1335:
1313:
1312:
1308:
1280:
1279:
1275:
1254:
1253:
1230:
1225:
1182:Mahajangasuchus
1153:
1121:
1112:
1103:
1094:
1085:
1076:
1067:
1058:
1049:
1040:
1031:
1022:
1013:
1004:
626:Montealtosuchus
589:
580:Montealtosuchus
553:
530:
504:
492:squamosal bones
468:
463:
408:
358:
351:
343:
342:
329:
307:
299:
286:
273:
260:
244:
228:
216:Crocodyliformes
203:Crocodylomorpha
119:
97:
95:
94:
89:
84:
79:
74:
69:
64:
59:
54:
49:
44:
39:
31:
30:
27:
17:
12:
11:
5:
2662:
2660:
2652:
2651:
2646:
2641:
2636:
2631:
2626:
2616:
2615:
2609:
2608:
2605:
2604:
2602:
2601:
2595:
2592:
2591:
2588:
2581:
2580:
2577:
2576:
2546:
2544:
2541:
2540:
2537:
2536:
2533:
2532:
2529:
2528:
2525:
2524:
2522:
2521:
2518:Morrinhosuchus
2514:
2506:
2504:
2498:
2497:
2495:
2494:
2487:
2480:
2472:
2470:
2468:Sphagesaurinae
2464:
2463:
2461:
2460:
2453:
2446:
2438:
2432:
2430:Sphagesauridae
2426:
2425:
2423:
2422:
2415:
2407:
2400:
2391:
2389:
2383:
2382:
2380:
2379:
2376:Eptalofosuchus
2372:
2365:
2358:
2354:Brasileosaurus
2349:
2343:
2337:
2336:
2333:
2332:
2329:
2328:
2326:
2325:
2318:
2311:
2303:
2301:
2295:
2294:
2292:
2291:
2284:
2277:
2270:
2262:
2260:
2254:
2253:
2251:
2250:
2243:
2240:Gondwanasuchus
2236:
2228:
2222:
2216:
2215:
2213:
2212:
2204:
2197:
2189:
2181:
2178:Chimaerasuchus
2173:
2167:
2158:
2153:
2150:
2149:
2146:
2145:
2143:
2142:
2135:
2128:
2125:Lavocatchampsa
2121:
2113:
2111:
2104:
2103:
2101:
2100:
2093:
2086:
2078:
2071:
2063:
2056:
2047:
2037:
2032:
2025:
2024:
2019:
2012:
2011:
2008:
2007:
1981:
1979:
1976:
1975:
1972:
1971:
1968:
1967:
1965:
1964:
1963:
1962:
1956:
1954:
1944:
1943:
1940:
1939:
1936:
1935:
1933:
1932:
1925:
1918:
1911:
1904:
1897:
1890:
1883:
1876:
1869:
1862:
1855:
1852:Bayomesasuchus
1848:
1841:
1834:
1826:
1824:
1818:
1817:
1815:
1814:
1807:
1800:
1793:
1786:
1779:
1776:Barreirosuchus
1771:
1769:
1763:
1762:
1760:
1759:
1755:Stolokrosuchus
1751:
1744:
1736:
1728:
1722:
1716:
1715:
1713:
1712:
1705:
1698:
1691:
1684:
1677:
1670:
1663:
1656:
1653:Dentaneosuchus
1649:
1642:
1635:
1628:
1620:
1618:
1612:
1611:
1609:
1608:
1601:
1593:
1591:
1585:
1584:
1582:
1581:
1574:
1570:Trematochampsa
1566:
1559:
1551:
1545:
1540:
1537:
1536:
1533:
1532:
1530:
1529:
1520:
1510:
1505:
1498:
1497:
1492:
1485:
1484:
1481:
1480:
1477:
1476:
1473:
1472:
1470:
1469:
1468:
1467:
1461:
1459:
1453:
1452:
1450:
1449:
1448:
1447:
1438:
1432:
1422:
1421:
1417:
1416:
1407:
1401:
1395:
1389:
1380:
1373:
1372:
1367:
1365:
1364:
1357:
1350:
1342:
1334:
1333:
1306:
1273:
1227:
1226:
1224:
1221:
1165:goniopholidids
1152:
1149:
1118:
1117:
1114:
1113:
1109:
1108:
1105:
1104:
1100:
1099:
1096:
1095:
1091:
1090:
1087:
1086:
1082:
1081:
1078:
1077:
1073:
1072:
1069:
1068:
1064:
1063:
1060:
1059:
1055:
1054:
1051:
1050:
1046:
1045:
1042:
1041:
1037:
1036:
1033:
1032:
1028:
1027:
1024:
1023:
1019:
1018:
1015:
1014:
1010:
1009:
1006:
1005:
1001:
1000:
997:
996:
987:
984:
983:
980:
979:
970:
967:
966:
963:
962:
953:
950:
949:
946:
945:
936:
931:
929:
926:
925:
922:
921:
912:
907:
905:
902:
901:
898:
897:
893:Barreirosuchus
888:
883:
881:
878:
877:
874:
873:
864:
859:
857:
854:
853:
850:
849:
840:
835:
833:
830:
829:
826:
825:
821:Stolokrosuchus
816:
811:
809:
806:
805:
802:
801:
792:
787:
785:
779:
778:
775:
774:
767:
762:
760:
754:
753:
750:
749:
742:
737:
735:
732:
731:
728:
727:
720:
715:
713:
710:
709:
706:
705:
696:
691:
689:
683:
682:
679:
678:
671:
666:
664:
656:
654:
649:
609:uruguaysuchids
588:
585:
552:
549:
544:Stolokrosuchus
529:
526:
503:
500:
467:
464:
462:
459:
407:
404:
369:is an extinct
360:
359:
352:
338:
337:
331:
330:
322:
320:
316:
315:
297:
293:
292:
284:
280:
279:
271:
267:
266:
258:
251:
250:
242:
235:
234:
226:
219:
218:
213:
206:
205:
200:
193:
192:
187:
180:
179:
174:
167:
166:
161:
157:
156:
151:
147:
146:
141:
137:
136:
131:
127:
126:
113:
112:
108:
107:
99:
98:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
40:
35:
34:
28:
15:
13:
10:
9:
6:
4:
3:
2:
2661:
2650:
2647:
2645:
2642:
2640:
2637:
2635:
2632:
2630:
2627:
2625:
2622:
2621:
2619:
2600:
2597:
2596:
2593:
2586:
2582:
2572:
2567:
2564:
2557:
2550:
2520:
2519:
2515:
2513:
2512:
2511:Caipirasuchus
2508:
2507:
2505:
2503:
2499:
2493:
2492:
2488:
2486:
2485:
2484:Caryonosuchus
2481:
2479:
2478:
2474:
2473:
2471:
2469:
2465:
2459:
2458:
2454:
2452:
2451:
2450:Labidiosuchus
2447:
2445:
2444:
2440:
2439:
2436:
2433:
2431:
2427:
2421:
2420:
2416:
2413:
2412:
2411:Mariliasuchus
2408:
2406:
2405:
2401:
2398:
2397:
2396:Coronelsuchus
2393:
2392:
2390:
2388:
2384:
2378:
2377:
2373:
2371:
2370:
2369:Coronelsuchus
2366:
2364:
2363:
2362:Coringasuchus
2359:
2356:
2355:
2351:
2350:
2347:
2344:
2342:
2338:
2324:
2323:
2319:
2317:
2316:
2312:
2310:
2309:
2308:Campinasuchus
2305:
2304:
2302:
2300:
2296:
2290:
2289:
2285:
2283:
2282:
2278:
2276:
2275:
2274:Aplestosuchus
2271:
2269:
2268:
2267:Aphaurosuchus
2264:
2263:
2261:
2259:
2258:Baurusuchinae
2255:
2249:
2248:
2244:
2242:
2241:
2237:
2235:
2234:
2230:
2229:
2226:
2223:
2221:
2220:Baurusuchidae
2217:
2210:
2209:
2205:
2203:
2202:
2198:
2195:
2194:
2190:
2187:
2186:
2185:Comahuesuchus
2182:
2180:
2179:
2175:
2174:
2171:
2168:
2166:
2162:
2159:
2156:
2151:
2141:
2140:
2136:
2134:
2133:
2129:
2127:
2126:
2122:
2120:
2119:
2115:
2114:
2112:
2109:
2105:
2099:
2098:
2097:Uruguaysuchus
2094:
2092:
2091:
2087:
2084:
2083:
2079:
2077:
2076:
2072:
2069:
2068:
2067:Comahuesuchus
2064:
2062:
2061:
2060:Araripesuchus
2057:
2054:
2053:
2049:
2048:
2045:
2041:
2038:
2035:
2030:
2026:
2022:
2017:
2013:
2004:
2000:
1997:
1991:
1985:
1960:
1959:
1958:
1957:
1955:
1953:
1949:
1945:
1931:
1930:
1929:Uberabasuchus
1926:
1924:
1923:
1919:
1917:
1916:
1912:
1910:
1909:
1905:
1903:
1902:
1898:
1896:
1895:
1891:
1889:
1888:
1884:
1882:
1881:
1877:
1875:
1874:
1870:
1868:
1867:
1863:
1861:
1860:
1856:
1854:
1853:
1849:
1847:
1846:
1845:Barrosasuchus
1842:
1840:
1839:
1838:Barcinosuchus
1835:
1833:
1832:
1828:
1827:
1825:
1823:
1822:Peirosaurinae
1819:
1813:
1812:
1808:
1806:
1805:
1801:
1799:
1798:
1794:
1792:
1791:
1787:
1785:
1784:
1780:
1778:
1777:
1773:
1772:
1770:
1768:
1764:
1757:
1756:
1752:
1750:
1749:
1748:Miadanasuchus
1745:
1742:
1741:
1737:
1735:
1734:
1730:
1729:
1726:
1723:
1721:
1720:Peirosauridae
1717:
1711:
1710:
1706:
1704:
1703:
1699:
1697:
1696:
1692:
1690:
1689:
1685:
1683:
1682:
1678:
1676:
1675:
1671:
1669:
1668:
1664:
1662:
1661:
1657:
1655:
1654:
1650:
1648:
1647:
1643:
1641:
1640:
1636:
1634:
1633:
1629:
1627:
1626:
1622:
1621:
1619:
1617:
1613:
1607:
1606:
1602:
1600:
1599:
1595:
1594:
1592:
1590:
1586:
1580:
1579:
1575:
1572:
1571:
1567:
1565:
1564:
1560:
1558:
1557:
1553:
1552:
1549:
1546:
1543:
1538:
1527:
1526:
1522:
1521:
1518:
1514:
1511:
1508:
1503:
1499:
1495:
1490:
1486:
1465:
1464:
1463:
1462:
1460:
1458:
1454:
1446:
1442:
1441:
1440:
1439:
1436:
1433:
1431:
1427:
1423:
1415:
1411:
1408:
1406:
1402:
1400:
1396:
1394:
1390:
1388:
1384:
1383:
1378:
1374:
1370:
1363:
1358:
1356:
1351:
1349:
1344:
1343:
1340:
1329:
1325:
1321:
1317:
1310:
1307:
1301:
1296:
1292:
1288:
1284:
1277:
1274:
1269:
1265:
1261:
1260:
1251:
1249:
1247:
1245:
1243:
1241:
1239:
1237:
1235:
1233:
1229:
1222:
1220:
1218:
1214:
1213:
1212:Titanochampsa
1208:
1204:
1200:
1196:
1195:
1190:
1189:
1184:
1183:
1178:
1174:
1170:
1166:
1162:
1158:
1150:
1148:
1146:
1142:
1138:
1134:
1130:
1126:
1116:
1115:
1107:
1106:
1098:
1097:
1089:
1088:
1080:
1079:
1071:
1070:
1062:
1061:
1053:
1052:
1044:
1043:
1035:
1034:
1026:
1025:
1017:
1016:
1008:
1007:
999:
998:
995:
994:
993:
986:
985:
982:
981:
978:
977:
976:
969:
968:
965:
964:
961:
960:
959:
952:
951:
948:
947:
944:
943:
942:
935:
934:
928:
927:
924:
923:
920:
919:
918:
911:
910:
904:
903:
900:
899:
896:
895:
894:
887:
886:
880:
879:
876:
875:
872:
871:
870:
863:
862:
856:
855:
852:
851:
848:
847:
846:
839:
838:
832:
831:
828:
827:
824:
823:
822:
815:
814:
808:
807:
804:
803:
800:
799:
798:
791:
790:
784:
781:
780:
777:
776:
773:
772:
771:Peirosaurinae
766:
765:
759:
758:Peirosauridae
756:
755:
752:
751:
748:
747:
741:
740:
734:
733:
730:
729:
726:
725:
719:
718:
712:
711:
708:
707:
704:
703:
702:
695:
694:
688:
685:
684:
681:
680:
677:
676:
670:
669:
663:
660:
659:
653:
652:
648:
646:
645:
640:
639:
634:
633:
628:
627:
622:
621:
620:Uberabasuchus
616:
615:
610:
606:
602:
598:
594:
586:
584:
582:
581:
576:
575:
570:
569:
564:
563:
558:
550:
548:
546:
545:
540:
536:
527:
525:
522:
517:
513:
509:
501:
499:
497:
496:parietal bone
493:
489:
485:
484:lacrimal bone
481:
477:
473:
470:The skull of
465:
460:
458:
456:
452:
448:
447:Ancient Greek
444:
439:
437:
433:
429:
425:
421:
417:
413:
405:
403:
401:
400:
395:
391:
387:
383:
382:Maastrichtian
379:
375:
372:
368:
367:
356:
350:
348:
339:
336:
335:Binomial name
332:
328:
327:
321:
318:
317:
312:
306:
305:
298:
295:
294:
291:
285:
282:
281:
278:
277:Peirosauridae
272:
269:
268:
265:
259:
256:
253:
252:
249:
243:
240:
237:
236:
233:
227:
224:
221:
220:
217:
214:
211:
208:
207:
204:
201:
198:
195:
194:
191:
188:
185:
182:
181:
178:
175:
172:
169:
168:
165:
162:
159:
158:
155:
152:
149:
148:
145:
142:
139:
138:
135:
132:
129:
128:
123:
118:
114:
109:
105:
100:
93:
88:
83:
78:
73:
68:
63:
58:
53:
48:
43:
38:
26:
22:
19:
2649:Peirosaurids
2552:
2516:
2509:
2491:Sphagesaurus
2489:
2482:
2475:
2455:
2448:
2441:
2417:
2409:
2402:
2394:
2387:Notosuchidae
2374:
2367:
2360:
2352:
2341:Sphagesauria
2320:
2313:
2306:
2286:
2279:
2272:
2265:
2245:
2238:
2231:
2206:
2199:
2191:
2183:
2176:
2137:
2132:Malawisuchus
2130:
2123:
2116:
2095:
2088:
2080:
2075:Libycosuchus
2073:
2065:
2058:
2050:
1987:
1927:
1920:
1913:
1906:
1901:Patagosuchus
1899:
1892:
1885:
1878:
1873:Hamadasuchus
1871:
1864:
1857:
1850:
1843:
1836:
1831:Antaeusuchus
1829:
1809:
1802:
1795:
1790:Epoidesuchus
1789:
1788:
1783:Caririsuchus
1781:
1774:
1767:Pepesuchinae
1753:
1746:
1740:Fortignathus
1738:
1733:Amargasuchus
1731:
1707:
1700:
1695:Sahitisuchus
1693:
1686:
1679:
1672:
1665:
1658:
1651:
1644:
1637:
1632:Barinasuchus
1630:
1623:
1603:
1596:
1576:
1568:
1561:
1554:
1523:
1445:Pseudosuchia
1430:Pseudosuchia
1409:
1319:
1315:
1309:
1293:(3): 44–82.
1290:
1286:
1276:
1257:
1210:
1203:Epoidesuchus
1202:
1198:
1192:
1186:
1180:
1177:Epoidesuchus
1176:
1173:Epoidesuchus
1172:
1169:crocodilians
1157:Epoidesuchus
1156:
1154:
1151:Paleobiology
1141:Epoidesuchus
1140:
1137:Epoidesuchus
1136:
1132:
1124:
1122:
990:
989:
973:
972:
958:Caririsuchus
956:
955:
941:Amargasuchus
939:
938:
915:
914:
891:
890:
869:Epoidesuchus
868:
867:
866:
843:
842:
819:
818:
795:
794:
783:Pepesuchinae
769:
744:
722:
699:
698:
675:Eunotosuchia
673:
642:
636:
632:Caririsuchus
630:
624:
618:
614:Hamadasuchus
612:
605:baurusuchids
593:Epoidesuchus
592:
590:
578:
572:
566:
560:
557:Epoidesuchus
556:
554:
542:
539:Epoidesuchus
538:
531:
521:Epoidesuchus
520:
516:Epoidesuchus
515:
512:dentary bone
508:Epoidesuchus
507:
505:
480:Epoidesuchus
479:
472:Epoidesuchus
471:
469:
443:Epoidesuchus
442:
440:
435:
432:Fabiano Iori
412:Epoidesuchus
411:
409:
398:
397:
394:Epoidesuchus
393:
366:Epoidesuchus
365:
364:
363:
354:
346:
341:
325:
324:
310:
304:Epoidesuchus
303:
302:
290:Pepesuchinae
254:
248:Sebecosuchia
238:
222:
209:
196:
190:Pseudosuchia
183:
170:
25:Epoidesuchus
24:
18:
2404:Llanosuchus
2322:Wargosuchus
2281:Baurusuchus
2193:Eremosuchus
2165:Baurusuchia
2052:Anatosuchus
1922:Rukwasuchus
1908:Peirosaurus
1811:Roxochampsa
1709:Zulmasuchus
1660:Iberosuchus
1646:Bretesuchus
1639:Bergisuchus
1625:Ayllusuchus
1598:Kaprosuchus
1556:Baharijodon
1525:Microsuchus
1507:Ziphosuchia
1494:Ziphosuchia
1457:Ziphosuchia
1369:Ziphosuchia
1207:Bauru Group
1188:Kaprosuchus
1167:and modern
1161:neosuchians
1145:Peirosauria
1129:Itasuchidae
992:Roxochampsa
845:Rukwasuchus
638:Roxochampsa
574:Roxochampsa
476:skull table
461:Description
374:peirosaurid
371:pepesuchine
283:Subfamily:
177:Archosauria
2618:Categories
2457:Yacarerani
2419:Notosuchus
2139:Pakasuchus
2118:Candidodon
2090:Simosuchus
2034:Notosuchia
2021:Notosuchia
1961:see below↓
1952:Notosuchia
1915:Pepesuchus
1887:Lomasuchus
1880:Kinesuchus
1804:Pepesuchus
1688:Ogresuchus
1681:Lorosuchus
1674:Langstonia
1578:Wanosuchus
1466:see below↓
1414:Metasuchia
1399:Sauropsida
1223:References
1199:Lorosuchus
1194:Lorosuchus
1133:Pepesuchus
917:Pepesuchus
797:Kinesuchus
662:Notosuchia
644:Pepesuchus
568:Pepesuchus
232:Notosuchia
2247:Pabwehshi
1797:Itasuchus
1667:Ilchunaia
1616:Sebecidae
1563:Doratodon
1385:Kingdom:
975:Itasuchus
746:Sebecidae
597:Sebecidae
587:Phylogeny
562:Itasuchus
535:ziphodont
528:Dentition
502:Lower jaw
441:The name
424:Catanduva
420:Brazilian
378:Campanian
319:Species:
140:Kingdom:
134:Eukaryota
1393:Chordata
1391:Phylum:
1387:Animalia
270:Family:
164:Reptilia
154:Chordata
150:Phylum:
144:Animalia
130:Domain:
1702:Sebecus
1542:Sebecia
1403:Order:
1397:Class:
687:Sebecia
466:Cranium
418:in the
344:†
323:†
296:Genus:
264:Sebecia
160:Class:
1125:et al.
390:Brazil
357:, 2024
355:et al.
313:, 2024
311:et al.
1410:Clade
1163:like
451:Sobek
353:Ruiz
309:Ruiz
255:Clade
239:Clade
223:Clade
210:Clade
197:Clade
184:Clade
171:Clade
1443:see
1185:and
641:and
623:and
599:and
577:and
551:Size
37:PreꞒ
1324:doi
1295:doi
1264:doi
519:of
388:of
380:to
2620::
1412::
1320:35
1318:.
1289:.
1285:.
1262:.
1231:^
1219:.
647:.
635:,
617:,
583:.
571:,
565:,
547:.
438:.
402:.
384:)
257::
241::
225::
212::
199::
186::
173::
87:Pg
2414:?
2399:?
2357:?
2211:?
2196:?
2188:?
2110:?
2085:?
2070:?
2055:?
1758:?
1743:?
1573:?
1528:?
1361:e
1354:t
1347:v
1330:.
1326::
1303:.
1297::
1291:1
1270:.
1266::
300:†
287:†
274:†
261:†
245:†
229:†
92:N
82:K
77:J
72:T
67:P
62:C
57:D
52:S
47:O
42:Ꞓ
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