471:, the rear part of the left lower jaw; forty single teeth; five neck vertebrae; the right shoulder joint; the left ulna; the left radius; the proximal and distal left wrist bones; two fourth metacarpals; phalanges from the first to third fingers of the left hand; and the first phalanx of the fourth finger. It represents a fully-grown but not yet mature animal. The skeleton has largely been preserved three-dimensionally due to the ironstone, but some bones however, have been crushed. It was probably fossilized in articulation but got some time prior to the discovery dispersed by erosion and cattle.
379:
7035:
5802:
7129:
4604:
143:
4610:
555:
484:
3987:
5818:
7042:
3992:
7141:
3446:
7048:
3454:
3461:
5810:
517:
lower jaws are not expanded sideways. The rear teeth are directed vertically, gradually declining in size. The tooth sockets are swollen relative to the outer wall of the jaw bones. The snout bears a premaxillary crest, the front edge of which continues the line of the snout tip, steeply rising under an angle of 60 degrees, and ending in a rounded crest top.
526:
115:
542:(0.16 in) thick. It is hollow inside, the smooth bone walls being connected by small struts. The crest has a base length of 131 millimeters (5.2 in) and a height of 128 millimeters (5.0 in). The symphysis of the lower jaws, their front fused area, probably extended to below in a second crest.
516:
Additionally, a unique combination is present of traits that in themselves are not unique. The front edge of the premaxilla is flattened and triangular. The first tooth pair in the premaxillae is directed vertically and is slightly set-off to above from the jawline. The front parts of the upper and
545:
There is an estimated total of twelve teeth in the upper jaw and thirteen teeth in the lower jaw for a total of fifty in the head as a whole. The teeth are formed as conical spikes with an oval cross-section, transversely flattened. The swollen tooth sockets cause an undulating profile of the
541:
The length of the skull has been estimated to be about 60 centimeters (2.0 ft). The snout bears a relatively high crest, and it probably had a triangular profile in side view; the rear edge has not been preserved. The crest is very thin transversely however, only up to 4 millimeters
420:
was named and described by Adele H. Pentland, Stephen
Francis Poropat, Travis R. Tischler, Trish Sloan, Robert A. Elliott, Harry A. Elliott, Judy A. Elliott and David A. Elliott. It was subsequently X-ray CT scanned at high-resolution using the Imaging and Medical Beamline at the
409:
specimen was initially discovered in April 2017 when cattle farmer Robert A. Elliott was spraying herbicide near
Belmont Station. It was excavated by a team led by Adele H. Pentland. Nicknamed 'Butch', it was further prepared by volunteer Ali Calvey.
694:(particularly the post-cranial skeleton) and a renewed phylogenetic appraisal that unequivocally demonstrated that it belongs in the family Anhangueridae as proposed by Holgado and Pêgas. This study suggested that the precise position of
2261:
Pentland, Adele H.; Poropat, Stephen F.; White, Matt A.; Rigby, Samantha L.; Bevitt, Joseph J.; Duncan, Ruairidh J.; Sloan, Trish; Elliott, Robert A.; Elliott, Harry A.; Elliott, Judy A.; Elliott, David A. (2022-03-30).
512:
had. The first tooth pair in both the premaxillae of the snout and in the front lower jaws is smaller than the other front teeth. The fourth to seventh tooth pairs are smaller than the third and eighth pair.
467:, about ninety-six million years old. It consists of a partial skeleton with skull and lower jaws. It contains the front part of the head with the premaxillae, the maxillae and the dentaries; the left
340:, consisting primarily of the anterior portion of the skull and dentary, cervical vertebral centra and a partial wing. Its wingspan was estimated to be about 4 meters (13 ft).
2197:
Pentland, Adele H.; Poropat, Stephen F.; Tischler, Travis R.; Sloan, Trish; Elliott, Robert A.; Elliott, Harry A.; Elliott, Judy A.; Elliott, David A. (December 2019).
475:
is the only pterosaur fossil known from the Winton
Formation, and is the most complete pterosaur ever found in Australia, a continent where such finds are rare.
595:. In 2010 however, an ornithocheirid jaw fragment that includes two teeth (specimen WAM 68.5.11) had been reported from Australia, the same place where
702:
within
Anhangueridae still remains uncertain, and that the diversity of Australian pterosaur fauna has been greatly underestimated. In 2023, Richards
2395:"A second specimen of the pterosaur Thapunngaka shawi from the Lower Cretaceous (upper Albian) Toolebuc Formation of North West Queensland, Australia"
7161:
2447:
2343:
Kear, B.P., Deacon, G.L. & Siverson, M. 2010. "Remains of a Late
Cretaceous pterosaur from the Molecap Greensand of Western Australia".
7034:
2320:
448:
honors the late mayor of Winton Shire, Graham Thomas ‘Butch’ Lenton, for his work for the local community and his support for the
7166:
2203:
gen. et sp. nov., a new ornithocheirid pterosaur from the Winton
Formation (Cenomanian–lower Turonian) of Queensland, Australia"
2158:
2362:"A taxonomic and phylogenetic review of the anhanguerid pterosaur group Coloborhynchinae and the new clade Tropeognathinae"
378:
5801:
4609:
7119:
4603:
449:
142:
3986:
2440:
7171:
2544:
3445:
4022:
445:
3991:
3453:
7067:
6942:
5100:
1912:
1543:
904:
658:
422:
7041:
7099:
7047:
5817:
2394:
5195:
3634:
3544:
3460:
2433:
5809:
2264:"The osteology of Ferrodraco lentoni, an anhanguerid pterosaur from the mid-Cretaceous of Australia"
7133:
6699:
6388:
6075:
5374:
5301:
3069:
2789:
336:. It is the most complete pterosaur fossil from Australia, being known from the holotype specimen
6228:
4702:
4545:
3688:
3491:
3478:
3392:
3234:
2865:
2840:
2326:
578:
329:
137:
7104:
6654:
5469:
5398:
4640:
4627:
4570:
4333:
4231:
4133:
4051:
3829:
3806:
3772:
3702:
3243:
3159:
3111:
3085:
2798:
2703:
2316:
2285:
2240:
2222:
600:
7145:
6883:
6766:
6758:
6550:
6479:
6404:
5946:
5849:
5836:
5768:
5648:
5637:
4887:
4744:
4681:
4456:
4438:
4276:
4100:
3916:
3902:
3860:
3366:
3352:
3284:
2963:
2513:
2402:
2373:
2308:
2275:
2230:
2214:
1618:
1269:
953:
926:
800:
635:
487:
456:
345:
311:
200:
6972:
6815:
6664:
6449:
6353:
6328:
6321:
6308:
6241:
6127:
6025:
5991:
5887:
5867:
5721:
5579:
5520:
5507:
5478:
5354:
5265:
5124:
4997:
4952:
4824:
4753:
4732:
4558:
4219:
4199:
4171:
4147:
4083:
4075:
4068:
4060:
3815:
3795:
3625:
3601:
3551:
3359:
3342:
3332:
3309:
3265:
3257:
3225:
3214:
3193:
3147:
3139:
3078:
2994:
2934:
2853:
2822:
2712:
2691:
2644:
2529:
2504:
1968:
1841:
1476:
1150:
887:
683:
592:
367:
308:
6949:
6922:
6900:
6724:
6638:
6623:
6575:
6514:
6499:
6465:
6290:
6274:
6262:
6248:
6201:
6158:
6118:
6032:
5978:
5706:
5699:
5692:
5664:
5571:
5492:
5419:
5330:
5086:
4978:
4913:
4854:
4794:
4760:
4480:
4407:
4379:
4298:
4248:
4187:
4156:
3756:
3732:
3710:
3656:
3614:
3558:
3526:
3207:
3124:
3101:
2980:
2906:
2874:
2727:
2676:
2624:
2235:
2198:
1782:
1697:
1656:
1409:
1348:
1307:
991:
967:
863:
717:
670:
664:
639:
631:
554:
505:
483:
648:, another Australian pterosaur from somewhat older layers. The second analysis placed
7155:
6986:
6957:
6933:
6914:
6872:
6857:
6806:
6795:
6688:
6680:
6630:
6567:
6538:
6522:
6506:
6492:
6433:
6411:
6362:
6255:
6016:
5959:
5878:
5713:
5685:
5657:
5528:
5411:
5202:
5132:
5004:
4987:
4843:
4780:
4724:
4489:
4445:
4326:
4319:
4312:
4305:
4290:
4240:
4164:
4121:
4114:
4040:
3887:
3880:
3873:
3722:
3664:
3649:
3642:
3590:
3373:
3042:
3002:
2952:
2921:
2882:
2777:
2769:
2747:
2651:
2635:
2330:
2006:
1989:
1875:
1858:
1737:
1601:
1517:
1500:
1388:
1188:
1171:
1140:
940:
873:
850:
679:
573:
567:
363:
213:
128:
72:
6979:
6907:
6778:
6673:
6607:
6559:
6369:
6297:
6175:
6143:
5911:
5678:
5606:
5547:
5442:
5079:
5064:
5048:
5039:
5028:
4868:
4831:
4717:
4667:
4510:
4423:
4416:
4396:
4368:
4361:
4347:
4340:
4255:
4212:
4179:
3895:
3837:
3780:
3765:
3745:
3570:
3301:
3293:
3273:
3200:
3179:
3172:
3027:
2815:
2761:
2720:
1758:
1680:
1443:
1331:
817:
619:
468:
414:
333:
256:
2407:
2280:
7090:
6865:
6825:
6744:
6708:
6442:
6217:
6150:
6093:
6039:
5896:
5564:
5435:
5427:
5387:
5346:
5338:
5213:
5180:
5148:
5140:
5111:
5072:
5056:
5015:
4944:
4936:
4929:
4875:
4816:
4808:
4469:
4431:
4354:
4009:
3942:
3909:
3845:
3186:
3055:
3010:
2987:
2662:
2263:
2076:
2023:
747:
708:
588:
397:
301:
47:
2218:
6788:
6716:
6583:
6418:
6397:
6342:
6210:
6185:
6135:
6100:
6046:
5934:
5926:
5919:
5671:
5594:
5586:
5485:
5449:
5287:
5188:
5167:
5156:
5093:
4710:
4503:
4496:
3931:
3822:
2829:
2486:
2045:
713:
460:
315:
226:
92:
57:
30:
2289:
2226:
674:. A more recent study made in 2020 by Borja Holgado and Rodrigo Pêgas placed
501:
had a wingspan of 4 meters (13 ft) and weighed 10 kg (22 lb).
6893:
6335:
6283:
6005:
5998:
5966:
5904:
5630:
5623:
5615:
5535:
5313:
5279:
5229:
5222:
4959:
4897:
4801:
4389:
3788:
3505:
3132:
2972:
2807:
2684:
2606:
2593:
2556:
2495:
2456:
1632:
1283:
441:
426:
319:
304:
290:
187:
154:
97:
41:
2378:
2361:
2244:
2312:
7084:
6965:
6849:
6841:
6615:
6599:
6530:
6107:
5457:
5362:
5294:
4966:
4921:
4694:
3923:
3583:
2480:
2093:
1799:
1426:
1221:
1032:
762:
653:
644:
530:
464:
406:
391:
382:
350:
174:
87:
82:
67:
62:
52:
34:
525:
294:
114:
102:
77:
2393:
Richards, Timothy; Stumkat, Paul; Salisbury, Steven (6 October 2023).
5556:
5323:
4905:
3513:
3380:
2474:
558:
164:
7061:
440:, "dragon", in reference to the fact that the skeleton was found in
4654:
3250:
553:
524:
482:
430:
377:
297:
7065:
5834:
4625:
4007:
3476:
2591:
2467:
2429:
690:. In 2022, Pentland et al. published a detailed description of
2425:
652:
as a basal member of the
Anhangueria, and sister taxon to the
591:, and proves that the clade did not become extinct during the
727:
429:
described in detail. The generic name is derived from the
537:
in different angles (left), and schematic diagram (right)
16:
Genus of anhanguerid pterosaur from the Late
Cretaceous
7117:
603:, layers that possibly have an even younger age than
455:
The holotype, AODF 876, was found in a layer of the
263:
7074:
6932:
6882:
6814:
6805:
6777:
6742:
6698:
6663:
6653:
6549:
6478:
6463:
6432:
6387:
6352:
6307:
6273:
6227:
6200:
6173:
6117:
6073:
6015:
5977:
5945:
5886:
5877:
5847:
5767:
5647:
5605:
5546:
5506:
5468:
5397:
5373:
5312:
5263:
5212:
5166:
5110:
5038:
5014:
4986:
4977:
4886:
4852:
4841:
4778:
4743:
4680:
4665:
4638:
4569:
4544:
4479:
4455:
4406:
4378:
4274:
4230:
4198:
4146:
4131:
4098:
4050:
4020:
3941:
3859:
3805:
3755:
3731:
3720:
3686:
3624:
3600:
3569:
3524:
3489:
3391:
3341:
3331:
3283:
3233:
3224:
3158:
3110:
3099:
3068:
3053:
3025:
2962:
2932:
2904:
2864:
2839:
2797:
2788:
2745:
2702:
2661:
2634:
2604:
2555:
2528:
626:in the evolutionary tree. The first analysis found
599:was found, and this specimen was unearthed in the
328:. The species was named after the former mayor of
2192:
2190:
2188:
2186:
2184:
2182:
2180:
2178:
2176:
2174:
622:analyses, trying to determine the position of
2441:
126:showing known material on a diagram based on
8:
344:was found to have been within the subfamily
2307:. Princeton University Press. p. 166.
587:was the youngest known member of the clade
7062:
6811:
6750:
6660:
6484:
6475:
6471:
6429:
6425:
6313:
6233:
6197:
6193:
6085:
6081:
5983:
5951:
5883:
5859:
5855:
5844:
5831:
5764:
5602:
5543:
5512:
5403:
5379:
5271:
5172:
5116:
5035:
5020:
4983:
4860:
4849:
4786:
4686:
4677:
4673:
4646:
4635:
4622:
4550:
4541:
4461:
4282:
4204:
4143:
4139:
4106:
4032:
4028:
4017:
4004:
3865:
3737:
3728:
3694:
3606:
3575:
3536:
3532:
3497:
3486:
3473:
3338:
3230:
3164:
3116:
3107:
3065:
3061:
3034:
2944:
2940:
2913:
2845:
2794:
2753:
2668:
2616:
2612:
2601:
2588:
2534:
2525:
2464:
2448:
2434:
2426:
113:
20:
2406:
2377:
2279:
2234:
682:, more specifically within the subfamily
366:, more specifically within the subfamily
7124:
2305:The Princeton Field Guide to Pterosaurs
2170:
634:, more precisely within the subfamily
716:. The clade comprises all Australian
686:, although still the sister taxon to
504:The describing authors indicated two
7:
2256:
2254:
362:, but both placed within the family
2051:
1981:
1974:
1964:
1891:
1833:
1774:
1750:
1743:
1672:
1648:
1624:
1614:
1607:
1597:
1492:
1468:
1401:
1394:
1323:
1299:
1275:
1213:
1163:
1156:
1146:
1136:
1007:
983:
959:
932:
879:
869:
842:
792:
785:
778:
754:
742:
706:. published a revised diagnosis of
2268:Journal of Vertebrate Paleontology
712:where they also erected the clade
14:
2360:Holgado, B.; Pêgas, R.V. (2020).
389:(A) compared to the holotypes of
7139:
7127:
7046:
7040:
7033:
5816:
5808:
5800:
4608:
4602:
3990:
3985:
3459:
3452:
3444:
354:. A recent study also recovered
322:, containing the single species
141:
565:(A) compared to the palates of
293:the fossil was found in) is an
2159:Timeline of pterosaur research
494:Size and distinguishing traits
1:
7162:Fossil taxa described in 2019
2408:10.1016/j.cretres.2023.105740
2366:Acta Palaeontologica Polonica
2281:10.1080/02724634.2021.2038182
1133:Holgado & Pêgas, (2020).
618:contained the results of two
508:(distinguishing traits) that
334:Graham Thomas ‘Butch’ Lenton
735:First analysis by Pentland
450:Australian Age of Dinosaurs
7188:
2219:10.1038/s41598-019-49789-4
7030:
6753:
6487:
6474:
6428:
6316:
6236:
6196:
6088:
6084:
5986:
5954:
5862:
5858:
5843:
5830:
5797:
5763:
5515:
5406:
5382:
5274:
5175:
5119:
5023:
4863:
4789:
4689:
4676:
4649:
4634:
4621:
4599:
4553:
4540:
4464:
4285:
4207:
4142:
4109:
4035:
4031:
4016:
4003:
3983:
3868:
3740:
3697:
3609:
3578:
3539:
3535:
3500:
3485:
3472:
3441:
3167:
3119:
3064:
3037:
2947:
2943:
2916:
2848:
2756:
2671:
2619:
2615:
2600:
2587:
2537:
2524:
2463:
2303:Paul, Gregory S. (2022).
2090:
2073:
2056:
2049:
2020:
2003:
1986:
1979:
1972:
1962:
1909:
1896:
1889:
1872:
1855:
1838:
1831:
1796:
1779:
1772:
1755:
1748:
1741:
1694:
1677:
1670:
1653:
1646:
1629:
1622:
1612:
1605:
1540:
1514:
1497:
1490:
1473:
1466:
1440:
1423:
1406:
1399:
1392:
1345:
1328:
1321:
1304:
1297:
1280:
1273:
1235:
1218:
1211:
1185:
1168:
1161:
1154:
1144:
1029:
1012:
1005:
988:
981:
964:
957:
937:
930:
901:
884:
877:
867:
847:
840:
814:
797:
790:
783:
776:
759:
752:
289:("Iron Dragon" after the
262:
255:
138:Scientific classification
136:
121:
112:
23:
577:(C), all of them are in
7167:Pterosaurs of Australia
4023:Archaeopterodactyloidea
2379:10.4202/app.00751.2020
581:
546:jawlines in top view.
538:
490:
423:Australian Synchrotron
402:
7100:Paleobiology Database
2313:10.1515/9780691232218
557:
528:
486:
381:
348:, as sister taxon to
3635:Archaeoistiodactylus
3545:Archaeoistiodactylus
374:Discovery and naming
6700:Targaryendraconidae
6389:Ornithocheiriformes
6076:Ornithocheiromorpha
5375:Neopterodactyloidea
5302:Dsungaripteromorpha
3070:Campylognathoididae
2790:Eudimorphodontoidea
2399:Cretaceous Research
452:. He died in 2017.
358:as sister taxon to
132:, scale bar = 50mm
6229:Istiodactyliformes
4703:Lonchognathosaurus
4546:Eupterodactyloidea
3689:Pterodactyliformes
3492:Pterodactylomorpha
3479:Pterodactylomorpha
3393:Pterodactylomorpha
3235:Angustinaripterini
2866:Eudimorphodontinae
2841:Eudimorphodontidae
2207:Scientific Reports
2201:Ferrodraco lentoni
678:within the family
610:The 2019 study of
582:
539:
491:
418:Ferrodraco lentoni
403:
268:Ferrodraco lentoni
124:Ferrodraco lentoni
122:Reconstruction of
7115:
7114:
7068:Taxon identifiers
7059:
7058:
7055:
7054:
7028:
7027:
7024:
7023:
7020:
7019:
7016:
7015:
7012:
7011:
7008:
7007:
7004:
7003:
7000:
6999:
6996:
6995:
6738:
6737:
6734:
6733:
6655:Targaryendraconia
6649:
6648:
6459:
6458:
6383:
6382:
6379:
6378:
6169:
6168:
6069:
6068:
6065:
6064:
6061:
6060:
6057:
6056:
5826:
5825:
5795:
5794:
5791:
5790:
5787:
5786:
5759:
5758:
5755:
5754:
5751:
5750:
5747:
5746:
5743:
5742:
5739:
5738:
5735:
5734:
5731:
5730:
5502:
5501:
5470:Chaoyangopterinae
5399:Chaoyangopteridae
5259:
5258:
5255:
5254:
5251:
5250:
5247:
5246:
5243:
5242:
5239:
5238:
4774:
4773:
4770:
4769:
4641:Ornithocheiroidea
4628:Ornithocheiroidea
4617:
4616:
4597:
4596:
4593:
4592:
4589:
4588:
4571:Ornithocheiroidea
4536:
4535:
4532:
4531:
4528:
4527:
4524:
4523:
4520:
4519:
4334:Gladocephaloideus
4270:
4269:
4266:
4265:
4232:Aurorazhdarchidae
4134:Ctenochasmatoidea
4094:
4093:
4052:Germanodactylidae
3999:
3998:
3981:
3980:
3977:
3976:
3973:
3972:
3969:
3968:
3965:
3964:
3961:
3960:
3855:
3854:
3830:Dendrorhynchoides
3807:Batrachognathinae
3773:Dendrorhynchoides
3703:Changchengopterus
3682:
3681:
3678:
3677:
3674:
3673:
3468:
3467:
3439:
3438:
3435:
3434:
3431:
3430:
3427:
3426:
3423:
3422:
3419:
3418:
3415:
3414:
3411:
3410:
3327:
3326:
3323:
3322:
3319:
3318:
3244:Angustinaripterus
3160:Rhamphorhynchinae
3112:Rhamphorhynchidae
3095:
3094:
3086:Campylognathoides
3021:
3020:
2900:
2899:
2896:
2895:
2892:
2891:
2799:Raeticodactylidae
2741:
2740:
2737:
2736:
2704:Austriadraconidae
2583:
2582:
2579:
2578:
2575:
2574:
2149:
2148:
2141:
2140:
2132:
2131:
2123:
2122:
2114:
2113:
2105:
2104:
2035:
2034:
1951:
1950:
1942:
1941:
1933:
1932:
1924:
1923:
1820:
1819:
1811:
1810:
1727:
1726:
1718:
1717:
1709:
1708:
1582:
1581:
1573:
1572:
1564:
1563:
1555:
1554:
1529:
1528:
1455:
1454:
1378:
1377:
1369:
1368:
1360:
1359:
1259:
1258:
1250:
1249:
1200:
1199:
1125:
1124:
1116:
1115:
1107:
1106:
1098:
1097:
1089:
1088:
1080:
1079:
1071:
1070:
1062:
1061:
1053:
1052:
1044:
1043:
916:
915:
829:
828:
630:within the clade
614:made by Pentland
601:Molecap Greensand
282:
281:
251:
7179:
7144:
7143:
7142:
7132:
7131:
7130:
7123:
7108:
7107:
7095:
7094:
7093:
7063:
7050:
7044:
7038:
7037:
6884:Coloborhynchinae
6812:
6767:Ornithocheiridae
6759:Brasileodactylus
6751:
6661:
6551:Ornithocheirinae
6485:
6480:Ornithocheiridae
6476:
6472:
6430:
6426:
6405:Brasileodactylus
6314:
6234:
6202:Lonchodraconidae
6198:
6194:
6086:
6082:
5984:
5952:
5947:Nyctosauromorpha
5884:
5860:
5856:
5850:Pteranodontoidea
5845:
5837:Pteranodontoidea
5832:
5821:
5820:
5813:
5812:
5805:
5804:
5769:Pteranodontoidea
5765:
5649:Quetzalcoatlinae
5638:Mistralazhdarcho
5603:
5544:
5513:
5404:
5380:
5272:
5173:
5117:
5036:
5021:
4984:
4888:Thalassodromidae
4861:
4850:
4787:
4745:Dsungaripterinae
4687:
4682:Dsungaripteridae
4678:
4674:
4647:
4636:
4623:
4612:
4607:
4606:
4551:
4542:
4462:
4457:Ctenochasmatinae
4439:Plataleorhynchus
4283:
4277:Ctenochasmatidae
4205:
4144:
4140:
4107:
4101:Euctenochasmatia
4033:
4029:
4018:
4005:
3994:
3989:
3917:Propterodactylus
3903:Ningchengopterus
3866:
3861:Pterodactyloidea
3738:
3729:
3695:
3607:
3576:
3537:
3533:
3498:
3487:
3474:
3463:
3457:
3456:
3449:
3448:
3367:Jianchangopterus
3353:Fenghuangopterus
3339:
3285:Rhamphorhynchini
3231:
3165:
3117:
3108:
3066:
3062:
3035:
2964:Dimorphodontidae
2945:
2941:
2914:
2846:
2795:
2754:
2669:
2617:
2613:
2602:
2589:
2535:
2526:
2519:
2518:
2514:Pterosauromorpha
2465:
2450:
2443:
2436:
2427:
2420:
2419:
2417:
2415:
2410:
2390:
2384:
2383:
2381:
2357:
2351:
2341:
2335:
2334:
2300:
2294:
2293:
2283:
2258:
2249:
2248:
2238:
2194:
2052:
1982:
1975:
1965:
1892:
1834:
1775:
1751:
1744:
1673:
1649:
1625:
1619:Coloborhynchinae
1615:
1608:
1598:
1493:
1469:
1402:
1395:
1324:
1300:
1276:
1270:Coloborhynchinae
1214:
1164:
1157:
1147:
1137:
1008:
984:
960:
954:Ornithocheirinae
933:
927:Ornithocheiridae
880:
870:
843:
801:Brasileodactylus
793:
786:
779:
755:
743:
728:
636:Ornithocheirinae
488:Life restoration
459:dating from the
457:Winton Formation
346:Ornithocheirinae
312:Winton Formation
265:
245:
238:
225:
212:
201:Pterodactyloidea
199:
186:
146:
145:
117:
107:
44:
29:Temporal range:
21:
7187:
7186:
7182:
7181:
7180:
7178:
7177:
7176:
7172:Pteranodontoids
7152:
7151:
7150:
7140:
7138:
7128:
7126:
7118:
7116:
7111:
7103:
7098:
7089:
7088:
7083:
7070:
7060:
7051:
7032:
6992:
6973:Liaoningopterus
6928:
6878:
6816:Tropeognathinae
6801:
6773:
6747:
6730:
6694:
6665:Cimoliopteridae
6645:
6545:
6468:
6455:
6450:Zhenyuanopterus
6424:
6375:
6354:Istiodactylinae
6348:
6329:Longchengpterus
6322:Lingyuanopterus
6309:Istiodactylidae
6303:
6269:
6242:Hongshanopterus
6223:
6192:
6178:
6165:
6128:Hongshanopterus
6113:
6078:
6053:
6026:Barbaridactylus
6011:
5973:
5941:
5888:Pteranodontidae
5873:
5868:Santanadactylus
5852:
5839:
5822:
5815:
5807:
5799:
5783:
5727:
5722:Zhejiangopterus
5643:
5601:
5580:Palaeocursornis
5542:
5521:Montanazhdarcho
5508:Azhdarchiformes
5498:
5479:Chaoyangopterus
5464:
5393:
5369:
5355:Montanazhdarcho
5308:
5268:
5266:Azhdarchomorpha
5235:
5208:
5162:
5125:Aymberedactylus
5106:
5034:
5010:
4998:Aymberedactylus
4973:
4953:Thalassodromeus
4882:
4857:
4846:
4837:
4825:Montanazhdarcho
4783:
4766:
4754:Domeykodactylus
4739:
4733:Tendaguripterus
4670:
4661:
4643:
4630:
4613:
4601:
4585:
4565:
4559:Altmuehlopterus
4516:
4475:
4451:
4402:
4374:
4279:
4262:
4226:
4220:Gallodactylidae
4200:Aurorazhdarchia
4194:
4172:Normannognathus
4148:Gallodactylidae
4136:
4127:
4103:
4090:
4084:Tendaguripterus
4076:Normannognathus
4069:Germanodactylus
4061:Altmuehlopterus
4046:
4025:
4012:
3995:
3957:
3937:
3851:
3816:Batrachognathus
3801:
3796:Vesperopterylus
3751:
3725:
3716:
3691:
3670:
3626:Wukongopterinae
3620:
3602:Wukongopteridae
3596:
3565:
3552:Normannognathus
3529:
3520:
3494:
3481:
3464:
3451:
3443:
3407:
3387:
3360:Jianchangnathus
3343:Scaphognathidae
3333:Digibrevisauria
3315:
3310:Rhamphorhynchus
3279:
3266:Qinglongopterus
3258:Harpactognathus
3226:Rhamphorhynchae
3220:
3215:Qinglongopterus
3194:Harpactognathus
3154:
3148:Scaphognathinae
3140:Parapsicephalus
3104:
3091:
3079:Bergamodactylus
3058:
3049:
3031:
3017:
2995:Parapsicephalus
2958:
2937:
2935:Macronychoptera
2928:
2910:
2888:
2860:
2854:Arcticodactylus
2835:
2823:Raeticodactylus
2784:
2750:
2733:
2713:Arcticodactylus
2698:
2692:Raeticodactylus
2657:
2645:Austriadactylus
2630:
2609:
2596:
2571:
2551:
2545:Avemetatarsalia
2530:Avemetatarsalia
2520:
2505:Avemetatarsalia
2470:
2469:
2459:
2454:
2424:
2423:
2413:
2411:
2392:
2391:
2387:
2359:
2358:
2354:
2342:
2338:
2323:
2302:
2301:
2297:
2274:(5): e2038182.
2260:
2259:
2252:
2196:
2195:
2172:
2167:
2155:
2150:
2142:
2133:
2124:
2115:
2106:
2036:
1969:Tropeognathinae
1952:
1943:
1934:
1925:
1842:Liaoningopterus
1821:
1812:
1728:
1719:
1710:
1583:
1574:
1565:
1556:
1530:
1477:Liaoningopterus
1456:
1379:
1370:
1361:
1260:
1251:
1201:
1151:Tropeognathinae
1126:
1117:
1108:
1099:
1090:
1081:
1072:
1063:
1054:
1045:
917:
888:Liaoningopterus
830:
718:tropeognathines
684:Tropeognathinae
656:that comprises
593:Late Cretaceous
552:
523:
496:
481:
376:
368:Tropeognathinae
309:Late Cretaceous
307:known from the
278:
271:
244:
236:
223:
210:
197:
184:
140:
108:
106:
105:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
39:
38:
27:
17:
12:
11:
5:
7185:
7183:
7175:
7174:
7169:
7164:
7154:
7153:
7149:
7148:
7136:
7113:
7112:
7110:
7109:
7096:
7080:
7078:
7072:
7071:
7066:
7057:
7056:
7053:
7052:
7031:
7029:
7026:
7025:
7022:
7021:
7018:
7017:
7014:
7013:
7010:
7009:
7006:
7005:
7002:
7001:
6998:
6997:
6994:
6993:
6991:
6990:
6983:
6976:
6969:
6962:
6954:
6950:Caulkicephalus
6946:
6938:
6936:
6930:
6929:
6927:
6926:
6923:Uktenadactylus
6919:
6911:
6904:
6901:Coloborhynchus
6897:
6889:
6887:
6880:
6879:
6877:
6876:
6869:
6862:
6854:
6846:
6838:
6830:
6821:
6819:
6809:
6803:
6802:
6800:
6799:
6792:
6784:
6782:
6775:
6774:
6772:
6771:
6763:
6754:
6748:
6743:
6740:
6739:
6736:
6735:
6732:
6731:
6729:
6728:
6725:Targaryendraco
6721:
6713:
6704:
6702:
6696:
6695:
6693:
6692:
6685:
6677:
6669:
6667:
6658:
6651:
6650:
6647:
6646:
6644:
6643:
6639:Uktenadactylus
6635:
6627:
6624:Ornithocheirus
6620:
6612:
6604:
6596:
6588:
6580:
6576:Coloborhynchus
6572:
6564:
6555:
6553:
6547:
6546:
6544:
6543:
6535:
6527:
6519:
6515:Caulkicephalus
6511:
6503:
6500:Arthurdactylus
6496:
6488:
6482:
6469:
6466:Ornithocheirae
6464:
6461:
6460:
6457:
6456:
6454:
6453:
6446:
6438:
6436:
6423:
6422:
6415:
6408:
6401:
6393:
6391:
6385:
6384:
6381:
6380:
6377:
6376:
6374:
6373:
6366:
6358:
6356:
6350:
6349:
6347:
6346:
6339:
6332:
6325:
6317:
6311:
6305:
6304:
6302:
6301:
6294:
6291:Linlongopterus
6287:
6279:
6277:
6275:Mimodactylidae
6271:
6270:
6268:
6267:
6263:Lonchodectidae
6259:
6252:
6249:Linlongopterus
6245:
6237:
6231:
6225:
6224:
6222:
6221:
6214:
6206:
6204:
6191:
6190:
6181:
6179:
6174:
6171:
6170:
6167:
6166:
6164:
6163:
6159:Targaryendraco
6155:
6147:
6140:
6132:
6123:
6121:
6119:Lonchodectidae
6115:
6114:
6112:
6111:
6104:
6097:
6089:
6079:
6074:
6071:
6070:
6067:
6066:
6063:
6062:
6059:
6058:
6055:
6054:
6052:
6051:
6043:
6036:
6033:Muzquizopteryx
6029:
6021:
6019:
6013:
6012:
6010:
6009:
6002:
5995:
5987:
5981:
5979:Aponyctosauria
5975:
5974:
5972:
5971:
5963:
5955:
5949:
5943:
5942:
5940:
5939:
5931:
5923:
5916:
5908:
5901:
5892:
5890:
5881:
5875:
5874:
5872:
5871:
5863:
5853:
5848:
5841:
5840:
5835:
5828:
5827:
5824:
5823:
5798:
5796:
5793:
5792:
5789:
5788:
5785:
5784:
5782:
5781:
5780:
5779:
5773:
5771:
5761:
5760:
5757:
5756:
5753:
5752:
5749:
5748:
5745:
5744:
5741:
5740:
5737:
5736:
5733:
5732:
5729:
5728:
5726:
5725:
5718:
5710:
5707:Thanatosdrakon
5703:
5700:Quetzalcoatlus
5696:
5693:Phosphatodraco
5689:
5682:
5675:
5668:
5665:Arambourgiania
5661:
5653:
5651:
5645:
5644:
5642:
5641:
5634:
5627:
5620:
5611:
5609:
5600:
5599:
5591:
5583:
5576:
5572:Navajodactylus
5568:
5561:
5552:
5550:
5541:
5540:
5532:
5525:
5516:
5510:
5504:
5503:
5500:
5499:
5497:
5496:
5493:Shenzhoupterus
5489:
5482:
5474:
5472:
5466:
5465:
5463:
5462:
5454:
5446:
5439:
5432:
5424:
5420:Argentinadraco
5416:
5407:
5401:
5395:
5394:
5392:
5391:
5383:
5377:
5371:
5370:
5368:
5367:
5359:
5351:
5343:
5335:
5331:Argentinadraco
5327:
5319:
5317:
5310:
5309:
5307:
5306:
5298:
5291:
5284:
5275:
5269:
5264:
5261:
5260:
5257:
5256:
5253:
5252:
5249:
5248:
5245:
5244:
5241:
5240:
5237:
5236:
5234:
5233:
5226:
5218:
5216:
5210:
5209:
5207:
5206:
5199:
5192:
5185:
5176:
5170:
5164:
5163:
5161:
5160:
5153:
5145:
5137:
5129:
5120:
5114:
5108:
5107:
5105:
5104:
5097:
5090:
5087:Nemicolopterus
5083:
5076:
5069:
5061:
5053:
5044:
5042:
5033:
5032:
5024:
5018:
5012:
5011:
5009:
5008:
5001:
4993:
4991:
4981:
4979:Tapejariformes
4975:
4974:
4972:
4971:
4963:
4956:
4949:
4941:
4933:
4926:
4918:
4914:Argentinadraco
4910:
4902:
4893:
4891:
4884:
4883:
4881:
4880:
4872:
4864:
4858:
4855:Tapejaromorpha
4853:
4847:
4842:
4839:
4838:
4836:
4835:
4828:
4821:
4813:
4805:
4798:
4795:Argentinadraco
4790:
4784:
4779:
4776:
4775:
4772:
4771:
4768:
4767:
4765:
4764:
4761:Dsungaripterus
4757:
4749:
4747:
4741:
4740:
4738:
4737:
4729:
4721:
4714:
4707:
4699:
4690:
4684:
4671:
4666:
4663:
4662:
4660:
4659:
4650:
4644:
4639:
4632:
4631:
4626:
4619:
4618:
4615:
4614:
4600:
4598:
4595:
4594:
4591:
4590:
4587:
4586:
4584:
4583:
4582:
4581:
4575:
4573:
4567:
4566:
4564:
4563:
4554:
4548:
4538:
4537:
4534:
4533:
4530:
4529:
4526:
4525:
4522:
4521:
4518:
4517:
4515:
4514:
4507:
4500:
4493:
4485:
4483:
4481:Pterodaustrini
4477:
4476:
4474:
4473:
4465:
4459:
4453:
4452:
4450:
4449:
4442:
4435:
4428:
4420:
4412:
4410:
4408:Gnathosaurinae
4404:
4403:
4401:
4400:
4393:
4385:
4383:
4380:Moganopterinae
4376:
4375:
4373:
4372:
4365:
4358:
4351:
4344:
4337:
4330:
4323:
4316:
4309:
4302:
4299:Balaenognathus
4295:
4286:
4280:
4275:
4272:
4271:
4268:
4267:
4264:
4263:
4261:
4260:
4252:
4249:Aurorazhdarcho
4245:
4236:
4234:
4228:
4227:
4225:
4224:
4216:
4208:
4202:
4196:
4195:
4193:
4192:
4188:Moganopterinae
4184:
4176:
4168:
4161:
4157:Aurorazhdarcho
4152:
4150:
4137:
4132:
4129:
4128:
4126:
4125:
4118:
4110:
4104:
4099:
4096:
4095:
4092:
4091:
4089:
4088:
4080:
4072:
4065:
4056:
4054:
4048:
4047:
4045:
4044:
4036:
4026:
4021:
4014:
4013:
4008:
4001:
4000:
3997:
3996:
3984:
3982:
3979:
3978:
3975:
3974:
3971:
3970:
3967:
3966:
3963:
3962:
3959:
3958:
3956:
3955:
3954:
3953:
3947:
3945:
3939:
3938:
3936:
3935:
3928:
3920:
3913:
3906:
3899:
3892:
3884:
3877:
3869:
3863:
3857:
3856:
3853:
3852:
3850:
3849:
3842:
3834:
3826:
3819:
3811:
3809:
3803:
3802:
3800:
3799:
3792:
3785:
3777:
3769:
3761:
3759:
3757:Anurognathinae
3753:
3752:
3750:
3749:
3741:
3735:
3733:Anurognathidae
3726:
3721:
3718:
3717:
3715:
3714:
3711:Douzhanopterus
3707:
3698:
3692:
3687:
3684:
3683:
3680:
3679:
3676:
3675:
3672:
3671:
3669:
3668:
3661:
3657:Douzhanopterus
3653:
3646:
3639:
3630:
3628:
3622:
3621:
3619:
3618:
3615:Kunpengopterus
3610:
3604:
3598:
3597:
3595:
3594:
3587:
3579:
3573:
3567:
3566:
3564:
3563:
3559:Anurognathidae
3555:
3548:
3540:
3530:
3527:Monofenestrata
3525:
3522:
3521:
3519:
3518:
3510:
3501:
3495:
3490:
3483:
3482:
3477:
3470:
3469:
3466:
3465:
3442:
3440:
3437:
3436:
3433:
3432:
3429:
3428:
3425:
3424:
3421:
3420:
3417:
3416:
3413:
3412:
3409:
3408:
3406:
3405:
3404:
3403:
3397:
3395:
3389:
3388:
3386:
3385:
3377:
3370:
3363:
3356:
3348:
3346:
3336:
3329:
3328:
3325:
3324:
3321:
3320:
3317:
3316:
3314:
3313:
3306:
3298:
3289:
3287:
3281:
3280:
3278:
3277:
3270:
3262:
3254:
3247:
3239:
3237:
3228:
3222:
3221:
3219:
3218:
3211:
3208:Orientognathus
3204:
3197:
3190:
3183:
3176:
3168:
3162:
3156:
3155:
3153:
3152:
3144:
3136:
3129:
3125:Dolicorhamphus
3120:
3114:
3105:
3102:Breviquartossa
3100:
3097:
3096:
3093:
3092:
3090:
3089:
3082:
3074:
3072:
3059:
3054:
3051:
3050:
3048:
3047:
3038:
3032:
3026:
3023:
3022:
3019:
3018:
3016:
3015:
3007:
2999:
2991:
2984:
2981:Caelestiventus
2977:
2968:
2966:
2960:
2959:
2957:
2956:
2948:
2938:
2933:
2930:
2929:
2927:
2926:
2917:
2911:
2907:Zambellisauria
2905:
2902:
2901:
2898:
2897:
2894:
2893:
2890:
2889:
2887:
2886:
2879:
2875:Carniadactylus
2870:
2868:
2862:
2861:
2859:
2858:
2849:
2843:
2837:
2836:
2834:
2833:
2826:
2819:
2812:
2803:
2801:
2792:
2786:
2785:
2783:
2782:
2774:
2766:
2757:
2751:
2746:
2743:
2742:
2739:
2738:
2735:
2734:
2732:
2731:
2728:Seazzadactylus
2724:
2717:
2708:
2706:
2700:
2699:
2697:
2696:
2688:
2681:
2677:Carniadactylus
2672:
2666:
2659:
2658:
2656:
2655:
2648:
2640:
2638:
2632:
2631:
2629:
2628:
2625:Daohugoupterus
2620:
2610:
2605:
2598:
2597:
2592:
2585:
2584:
2581:
2580:
2577:
2576:
2573:
2572:
2570:
2569:
2568:
2567:
2561:
2559:
2553:
2552:
2550:
2549:
2548:
2547:
2538:
2532:
2522:
2521:
2517:
2516:
2507:
2498:
2489:
2483:
2477:
2468:
2461:
2460:
2455:
2453:
2452:
2445:
2438:
2430:
2422:
2421:
2385:
2352:
2336:
2321:
2295:
2250:
2169:
2168:
2166:
2163:
2162:
2161:
2154:
2151:
2147:
2146:
2139:
2138:
2135:
2134:
2130:
2129:
2126:
2125:
2121:
2120:
2117:
2116:
2112:
2111:
2108:
2107:
2103:
2102:
2099:
2098:
2089:
2086:
2085:
2082:
2081:
2072:
2069:
2068:
2065:
2064:
2055:
2050:
2048:
2042:
2041:
2038:
2037:
2033:
2032:
2029:
2028:
2019:
2016:
2015:
2012:
2011:
2002:
1999:
1998:
1995:
1994:
1985:
1980:
1978:
1973:
1971:
1963:
1961:
1958:
1957:
1954:
1953:
1949:
1948:
1945:
1944:
1940:
1939:
1936:
1935:
1931:
1930:
1927:
1926:
1922:
1921:
1918:
1917:
1908:
1905:
1904:
1901:
1900:
1895:
1890:
1888:
1885:
1884:
1881:
1880:
1871:
1868:
1867:
1864:
1863:
1854:
1851:
1850:
1847:
1846:
1837:
1832:
1830:
1827:
1826:
1823:
1822:
1818:
1817:
1814:
1813:
1809:
1808:
1805:
1804:
1795:
1792:
1791:
1788:
1787:
1783:Caulkicephalus
1778:
1773:
1771:
1768:
1767:
1764:
1763:
1754:
1749:
1747:
1742:
1740:
1734:
1733:
1730:
1729:
1725:
1724:
1721:
1720:
1716:
1715:
1712:
1711:
1707:
1706:
1703:
1702:
1698:Uktenadactylus
1693:
1690:
1689:
1686:
1685:
1676:
1671:
1669:
1666:
1665:
1662:
1661:
1657:Coloborhynchus
1652:
1647:
1645:
1642:
1641:
1638:
1637:
1628:
1623:
1621:
1613:
1611:
1606:
1604:
1596:
1584:
1580:
1579:
1576:
1575:
1571:
1570:
1567:
1566:
1562:
1561:
1558:
1557:
1553:
1552:
1549:
1548:
1539:
1536:
1535:
1532:
1531:
1527:
1526:
1523:
1522:
1513:
1510:
1509:
1506:
1505:
1496:
1491:
1489:
1486:
1485:
1482:
1481:
1472:
1467:
1465:
1462:
1461:
1458:
1457:
1453:
1452:
1449:
1448:
1439:
1436:
1435:
1432:
1431:
1422:
1419:
1418:
1415:
1414:
1410:Caulkicephalus
1405:
1400:
1398:
1393:
1391:
1385:
1384:
1381:
1380:
1376:
1375:
1372:
1371:
1367:
1366:
1363:
1362:
1358:
1357:
1354:
1353:
1349:Uktenadactylus
1344:
1341:
1340:
1337:
1336:
1327:
1322:
1320:
1317:
1316:
1313:
1312:
1308:Coloborhynchus
1303:
1298:
1296:
1293:
1292:
1289:
1288:
1279:
1274:
1272:
1266:
1265:
1262:
1261:
1257:
1256:
1253:
1252:
1248:
1247:
1244:
1243:
1234:
1231:
1230:
1227:
1226:
1217:
1212:
1210:
1207:
1206:
1203:
1202:
1198:
1197:
1194:
1193:
1184:
1181:
1180:
1177:
1176:
1167:
1162:
1160:
1155:
1153:
1145:
1143:
1135:
1127:
1123:
1122:
1119:
1118:
1114:
1113:
1110:
1109:
1105:
1104:
1101:
1100:
1096:
1095:
1092:
1091:
1087:
1086:
1083:
1082:
1078:
1077:
1074:
1073:
1069:
1068:
1065:
1064:
1060:
1059:
1056:
1055:
1051:
1050:
1047:
1046:
1042:
1041:
1038:
1037:
1028:
1025:
1024:
1021:
1020:
1011:
1006:
1004:
1001:
1000:
997:
996:
992:Ornithocheirus
987:
982:
980:
977:
976:
973:
972:
968:Coloborhynchus
963:
958:
956:
950:
949:
946:
945:
936:
931:
929:
923:
922:
919:
918:
914:
913:
910:
909:
900:
897:
896:
893:
892:
883:
878:
876:
868:
866:
864:Ornithocheirae
860:
859:
856:
855:
846:
841:
839:
836:
835:
832:
831:
827:
826:
823:
822:
813:
810:
809:
806:
805:
796:
791:
789:
784:
782:
777:
775:
772:
771:
768:
767:
758:
753:
751:
741:
726:
671:Ornithocheirus
665:Coloborhynchus
640:sister species
632:Ornithocheirae
551:
550:Classification
548:
522:
519:
506:autapomorphies
495:
492:
480:
477:
436:, "iron", and
375:
372:
280:
279:
272:
260:
259:
253:
252:
234:
230:
229:
221:
217:
216:
208:
204:
203:
195:
191:
190:
182:
178:
177:
172:
168:
167:
162:
158:
157:
152:
148:
147:
134:
133:
119:
118:
110:
109:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
45:
28:
15:
13:
10:
9:
6:
4:
3:
2:
7184:
7173:
7170:
7168:
7165:
7163:
7160:
7159:
7157:
7147:
7137:
7135:
7125:
7121:
7106:
7101:
7097:
7092:
7086:
7082:
7081:
7079:
7077:
7073:
7069:
7064:
7049:
7045:
7043:
7036:
6989:
6988:
6987:Maaradactylus
6984:
6982:
6981:
6977:
6975:
6974:
6970:
6968:
6967:
6963:
6960:
6959:
6958:Cearadactylus
6955:
6952:
6951:
6947:
6945:
6944:
6940:
6939:
6937:
6935:
6934:Anhanguerinae
6931:
6925:
6924:
6920:
6917:
6916:
6915:Siroccopteryx
6912:
6910:
6909:
6905:
6903:
6902:
6898:
6896:
6895:
6891:
6890:
6888:
6885:
6881:
6875:
6874:
6873:Tropeognathus
6870:
6868:
6867:
6863:
6860:
6859:
6858:Siroccopteryx
6855:
6852:
6851:
6847:
6844:
6843:
6839:
6836:
6835:
6831:
6828:
6827:
6823:
6822:
6820:
6817:
6813:
6810:
6808:
6807:Anhangueridae
6804:
6798:
6797:
6796:Iberodactylus
6793:
6791:
6790:
6786:
6785:
6783:
6780:
6776:
6769:
6768:
6764:
6761:
6760:
6756:
6755:
6752:
6749:
6746:
6741:
6727:
6726:
6722:
6719:
6718:
6714:
6711:
6710:
6706:
6705:
6703:
6701:
6697:
6691:
6690:
6689:Cimoliopterus
6686:
6683:
6682:
6681:Camposipterus
6678:
6676:
6675:
6671:
6670:
6668:
6666:
6662:
6659:
6656:
6652:
6641:
6640:
6636:
6633:
6632:
6631:Siroccopteryx
6628:
6626:
6625:
6621:
6618:
6617:
6613:
6610:
6609:
6605:
6602:
6601:
6597:
6594:
6593:
6589:
6586:
6585:
6581:
6578:
6577:
6573:
6570:
6569:
6568:Camposipterus
6565:
6562:
6561:
6557:
6556:
6554:
6552:
6548:
6541:
6540:
6539:Tropeognathus
6536:
6533:
6532:
6528:
6525:
6524:
6523:Cimoliopterus
6520:
6517:
6516:
6512:
6509:
6508:
6507:Camposipterus
6504:
6502:
6501:
6497:
6495:
6494:
6493:Araripesaurus
6490:
6489:
6486:
6483:
6481:
6477:
6473:
6470:
6467:
6462:
6452:
6451:
6447:
6445:
6444:
6440:
6439:
6437:
6435:
6434:Boreopteridae
6431:
6427:
6421:
6420:
6416:
6414:
6413:
6412:Cearadactylus
6409:
6407:
6406:
6402:
6400:
6399:
6395:
6394:
6392:
6390:
6386:
6372:
6371:
6367:
6365:
6364:
6363:Istiodactylus
6360:
6359:
6357:
6355:
6351:
6345:
6344:
6340:
6338:
6337:
6333:
6331:
6330:
6326:
6324:
6323:
6319:
6318:
6315:
6312:
6310:
6306:
6300:
6299:
6295:
6293:
6292:
6288:
6286:
6285:
6281:
6280:
6278:
6276:
6272:
6265:
6264:
6260:
6258:
6257:
6256:Yixianopterus
6253:
6251:
6250:
6246:
6244:
6243:
6239:
6238:
6235:
6232:
6230:
6226:
6220:
6219:
6215:
6213:
6212:
6208:
6207:
6205:
6203:
6199:
6195:
6188:
6187:
6183:
6182:
6180:
6177:
6172:
6161:
6160:
6156:
6153:
6152:
6148:
6146:
6145:
6141:
6138:
6137:
6133:
6130:
6129:
6125:
6124:
6122:
6120:
6116:
6110:
6109:
6105:
6103:
6102:
6098:
6096:
6095:
6091:
6090:
6087:
6083:
6080:
6077:
6072:
6049:
6048:
6044:
6042:
6041:
6037:
6035:
6034:
6030:
6028:
6027:
6023:
6022:
6020:
6018:
6017:Nyctosauridae
6014:
6008:
6007:
6003:
6001:
6000:
5996:
5994:
5993:
5989:
5988:
5985:
5982:
5980:
5976:
5969:
5968:
5964:
5962:
5961:
5960:Alamodactylus
5957:
5956:
5953:
5950:
5948:
5944:
5937:
5936:
5932:
5929:
5928:
5924:
5922:
5921:
5917:
5914:
5913:
5909:
5907:
5906:
5902:
5899:
5898:
5894:
5893:
5891:
5889:
5885:
5882:
5880:
5879:Pteranodontia
5876:
5870:
5869:
5865:
5864:
5861:
5857:
5854:
5851:
5846:
5842:
5838:
5833:
5829:
5819:
5814:
5811:
5803:
5777:
5776:
5775:
5774:
5772:
5770:
5766:
5762:
5724:
5723:
5719:
5716:
5715:
5714:Wellnhopterus
5711:
5709:
5708:
5704:
5702:
5701:
5697:
5695:
5694:
5690:
5688:
5687:
5686:Hatzegopteryx
5683:
5681:
5680:
5676:
5674:
5673:
5669:
5667:
5666:
5662:
5660:
5659:
5658:Aralazhdarcho
5655:
5654:
5652:
5650:
5646:
5640:
5639:
5635:
5633:
5632:
5628:
5626:
5625:
5621:
5618:
5617:
5613:
5612:
5610:
5608:
5604:
5597:
5596:
5592:
5589:
5588:
5584:
5582:
5581:
5577:
5574:
5573:
5569:
5567:
5566:
5562:
5559:
5558:
5554:
5553:
5551:
5549:
5545:
5538:
5537:
5533:
5531:
5530:
5529:Radiodactylus
5526:
5523:
5522:
5518:
5517:
5514:
5511:
5509:
5505:
5495:
5494:
5490:
5488:
5487:
5483:
5481:
5480:
5476:
5475:
5473:
5471:
5467:
5460:
5459:
5455:
5452:
5451:
5447:
5445:
5444:
5440:
5438:
5437:
5433:
5430:
5429:
5425:
5422:
5421:
5417:
5414:
5413:
5412:Apatorhamphus
5409:
5408:
5405:
5402:
5400:
5396:
5390:
5389:
5385:
5384:
5381:
5378:
5376:
5372:
5365:
5364:
5360:
5357:
5356:
5352:
5349:
5348:
5344:
5341:
5340:
5336:
5333:
5332:
5328:
5326:
5325:
5321:
5320:
5318:
5315:
5311:
5304:
5303:
5299:
5297:
5296:
5292:
5290:
5289:
5285:
5282:
5281:
5277:
5276:
5273:
5270:
5267:
5262:
5232:
5231:
5227:
5225:
5224:
5220:
5219:
5217:
5215:
5211:
5205:
5204:
5203:Tupandactylus
5200:
5198:
5197:
5193:
5191:
5190:
5186:
5183:
5182:
5178:
5177:
5174:
5171:
5169:
5165:
5159:
5158:
5154:
5151:
5150:
5146:
5143:
5142:
5138:
5135:
5134:
5133:Caupedactylus
5130:
5127:
5126:
5122:
5121:
5118:
5115:
5113:
5109:
5103:
5102:
5098:
5096:
5095:
5091:
5089:
5088:
5084:
5082:
5081:
5077:
5075:
5074:
5070:
5067:
5066:
5062:
5059:
5058:
5054:
5051:
5050:
5046:
5045:
5043:
5041:
5037:
5031:
5030:
5026:
5025:
5022:
5019:
5017:
5013:
5007:
5006:
5005:Caupedactylus
5002:
5000:
4999:
4995:
4994:
4992:
4989:
4988:Caupedactylia
4985:
4982:
4980:
4976:
4969:
4968:
4964:
4962:
4961:
4957:
4955:
4954:
4950:
4947:
4946:
4942:
4939:
4938:
4934:
4932:
4931:
4927:
4924:
4923:
4919:
4916:
4915:
4911:
4908:
4907:
4903:
4900:
4899:
4895:
4894:
4892:
4889:
4885:
4878:
4877:
4873:
4871:
4870:
4866:
4865:
4862:
4859:
4856:
4851:
4848:
4845:
4844:Neoazhdarchia
4840:
4834:
4833:
4829:
4827:
4826:
4822:
4819:
4818:
4814:
4811:
4810:
4806:
4804:
4803:
4799:
4797:
4796:
4792:
4791:
4788:
4785:
4782:
4781:Azhdarchoidea
4777:
4763:
4762:
4758:
4756:
4755:
4751:
4750:
4748:
4746:
4742:
4735:
4734:
4730:
4727:
4726:
4725:Puntanipterus
4722:
4720:
4719:
4715:
4713:
4712:
4708:
4705:
4704:
4700:
4697:
4696:
4692:
4691:
4688:
4685:
4683:
4679:
4675:
4672:
4669:
4664:
4657:
4656:
4652:
4651:
4648:
4645:
4642:
4637:
4633:
4629:
4624:
4620:
4611:
4605:
4579:
4578:
4577:
4576:
4574:
4572:
4568:
4561:
4560:
4556:
4555:
4552:
4549:
4547:
4543:
4539:
4513:
4512:
4508:
4506:
4505:
4501:
4499:
4498:
4494:
4492:
4491:
4490:Beipiaopterus
4487:
4486:
4484:
4482:
4478:
4472:
4471:
4467:
4466:
4463:
4460:
4458:
4454:
4448:
4447:
4446:Tacuadactylus
4443:
4441:
4440:
4436:
4434:
4433:
4429:
4426:
4425:
4421:
4419:
4418:
4414:
4413:
4411:
4409:
4405:
4399:
4398:
4394:
4392:
4391:
4387:
4386:
4384:
4381:
4377:
4371:
4370:
4366:
4364:
4363:
4359:
4357:
4356:
4352:
4350:
4349:
4345:
4343:
4342:
4338:
4336:
4335:
4331:
4329:
4328:
4327:Forfexopterus
4324:
4322:
4321:
4320:Elanodactylus
4317:
4315:
4314:
4313:Cratonopterus
4310:
4308:
4307:
4306:Cathayopterus
4303:
4301:
4300:
4296:
4293:
4292:
4291:Ardeadactylus
4288:
4287:
4284:
4281:
4278:
4273:
4258:
4257:
4253:
4251:
4250:
4246:
4243:
4242:
4241:Ardeadactylus
4238:
4237:
4235:
4233:
4229:
4222:
4221:
4217:
4215:
4214:
4210:
4209:
4206:
4203:
4201:
4197:
4190:
4189:
4185:
4182:
4181:
4177:
4174:
4173:
4169:
4167:
4166:
4165:Cycnorhamphus
4162:
4159:
4158:
4154:
4153:
4151:
4149:
4145:
4141:
4138:
4135:
4130:
4124:
4123:
4122:Pterodactylus
4119:
4117:
4116:
4115:Diopecephalus
4112:
4111:
4108:
4105:
4102:
4097:
4086:
4085:
4081:
4078:
4077:
4073:
4071:
4070:
4066:
4063:
4062:
4058:
4057:
4055:
4053:
4049:
4043:
4042:
4041:Prejanopterus
4038:
4037:
4034:
4030:
4027:
4024:
4019:
4015:
4011:
4006:
4002:
3993:
3988:
3951:
3950:
3949:
3948:
3946:
3944:
3940:
3934:
3933:
3929:
3926:
3925:
3921:
3919:
3918:
3914:
3912:
3911:
3907:
3905:
3904:
3900:
3898:
3897:
3893:
3890:
3889:
3888:Herbstosaurus
3885:
3883:
3882:
3881:Eurolimnornis
3878:
3876:
3875:
3874:Dermodactylus
3871:
3870:
3867:
3864:
3862:
3858:
3848:
3847:
3843:
3840:
3839:
3835:
3832:
3831:
3827:
3825:
3824:
3820:
3818:
3817:
3813:
3812:
3810:
3808:
3804:
3798:
3797:
3793:
3791:
3790:
3786:
3783:
3782:
3778:
3775:
3774:
3770:
3768:
3767:
3763:
3762:
3760:
3758:
3754:
3748:
3747:
3743:
3742:
3739:
3736:
3734:
3730:
3727:
3724:
3723:Caelidracones
3719:
3713:
3712:
3708:
3705:
3704:
3700:
3699:
3696:
3693:
3690:
3685:
3667:
3666:
3665:Wukongopterus
3662:
3659:
3658:
3654:
3652:
3651:
3650:Darwinopterus
3647:
3645:
3644:
3643:Cuspicephalus
3640:
3637:
3636:
3632:
3631:
3629:
3627:
3623:
3617:
3616:
3612:
3611:
3608:
3605:
3603:
3599:
3593:
3592:
3591:Pterorhynchus
3588:
3586:
3585:
3581:
3580:
3577:
3574:
3572:
3568:
3561:
3560:
3556:
3554:
3553:
3549:
3547:
3546:
3542:
3541:
3538:
3534:
3531:
3528:
3523:
3516:
3515:
3511:
3508:
3507:
3503:
3502:
3499:
3496:
3493:
3488:
3484:
3480:
3475:
3471:
3462:
3458:
3455:
3447:
3401:
3400:
3399:
3398:
3396:
3394:
3390:
3383:
3382:
3378:
3376:
3375:
3374:Scaphognathus
3371:
3369:
3368:
3364:
3362:
3361:
3357:
3355:
3354:
3350:
3349:
3347:
3344:
3340:
3337:
3334:
3330:
3312:
3311:
3307:
3304:
3303:
3299:
3296:
3295:
3291:
3290:
3288:
3286:
3282:
3276:
3275:
3271:
3268:
3267:
3263:
3260:
3259:
3255:
3253:
3252:
3248:
3246:
3245:
3241:
3240:
3238:
3236:
3232:
3229:
3227:
3223:
3217:
3216:
3212:
3210:
3209:
3205:
3203:
3202:
3198:
3196:
3195:
3191:
3189:
3188:
3184:
3182:
3181:
3177:
3175:
3174:
3170:
3169:
3166:
3163:
3161:
3157:
3150:
3149:
3145:
3142:
3141:
3137:
3135:
3134:
3130:
3127:
3126:
3122:
3121:
3118:
3115:
3113:
3109:
3106:
3103:
3098:
3088:
3087:
3083:
3081:
3080:
3076:
3075:
3073:
3071:
3067:
3063:
3060:
3057:
3052:
3045:
3044:
3043:Eudimorphodon
3040:
3039:
3036:
3033:
3029:
3024:
3013:
3012:
3008:
3005:
3004:
3003:Peteinosaurus
3000:
2997:
2996:
2992:
2990:
2989:
2985:
2983:
2982:
2978:
2975:
2974:
2970:
2969:
2967:
2965:
2961:
2955:
2954:
2953:Herbstosaurus
2950:
2949:
2946:
2942:
2939:
2936:
2931:
2924:
2923:
2922:Peteinosaurus
2919:
2918:
2915:
2912:
2908:
2903:
2885:
2884:
2883:Eudimorphodon
2880:
2877:
2876:
2872:
2871:
2869:
2867:
2863:
2856:
2855:
2851:
2850:
2847:
2844:
2842:
2838:
2832:
2831:
2827:
2825:
2824:
2820:
2818:
2817:
2813:
2810:
2809:
2805:
2804:
2802:
2800:
2796:
2793:
2791:
2787:
2780:
2779:
2778:Preondactylia
2775:
2772:
2771:
2770:Peteinosaurus
2767:
2764:
2763:
2759:
2758:
2755:
2752:
2749:
2748:Eopterosauria
2744:
2730:
2729:
2725:
2723:
2722:
2718:
2715:
2714:
2710:
2709:
2707:
2705:
2701:
2694:
2693:
2689:
2687:
2686:
2682:
2679:
2678:
2674:
2673:
2670:
2667:
2664:
2660:
2654:
2653:
2652:Preondactylus
2649:
2647:
2646:
2642:
2641:
2639:
2637:
2636:Preondactylia
2633:
2627:
2626:
2622:
2621:
2618:
2614:
2611:
2608:
2603:
2599:
2595:
2590:
2586:
2565:
2564:
2563:
2562:
2560:
2558:
2554:
2546:
2542:
2541:
2540:
2539:
2536:
2533:
2531:
2527:
2523:
2515:
2511:
2508:
2506:
2502:
2499:
2497:
2493:
2490:
2488:
2484:
2482:
2478:
2476:
2472:
2471:
2466:
2462:
2458:
2451:
2446:
2444:
2439:
2437:
2432:
2431:
2428:
2409:
2404:
2400:
2396:
2389:
2386:
2380:
2375:
2371:
2367:
2363:
2356:
2353:
2349:
2346:
2340:
2337:
2332:
2328:
2324:
2322:9780691232218
2318:
2314:
2310:
2306:
2299:
2296:
2291:
2287:
2282:
2277:
2273:
2269:
2265:
2257:
2255:
2251:
2246:
2242:
2237:
2232:
2228:
2224:
2220:
2216:
2212:
2208:
2204:
2202:
2193:
2191:
2189:
2187:
2185:
2183:
2181:
2179:
2177:
2175:
2171:
2164:
2160:
2157:
2156:
2152:
2145:
2137:
2136:
2128:
2127:
2119:
2118:
2110:
2109:
2101:
2100:
2097:
2096:
2095:
2088:
2087:
2084:
2083:
2080:
2079:
2078:
2071:
2070:
2067:
2066:
2063:
2062:
2061:
2054:
2053:
2047:
2044:
2043:
2040:
2039:
2031:
2030:
2027:
2026:
2025:
2018:
2017:
2014:
2013:
2010:
2009:
2008:
2007:Tropeognathus
2001:
2000:
1997:
1996:
1993:
1992:
1991:
1990:Siroccopteryx
1984:
1983:
1977:
1976:
1970:
1967:
1966:
1960:
1959:
1956:
1955:
1947:
1946:
1938:
1937:
1929:
1928:
1920:
1919:
1916:
1915:
1914:
1907:
1906:
1903:
1902:
1899:
1894:
1893:
1887:
1886:
1883:
1882:
1879:
1878:
1877:
1876:Maaradactylus
1870:
1869:
1866:
1865:
1862:
1861:
1860:
1859:Cearadactylus
1853:
1852:
1849:
1848:
1845:
1844:
1843:
1836:
1835:
1829:
1828:
1825:
1824:
1816:
1815:
1807:
1806:
1803:
1802:
1801:
1794:
1793:
1790:
1789:
1786:
1785:
1784:
1777:
1776:
1770:
1769:
1766:
1765:
1762:
1761:
1760:
1753:
1752:
1746:
1745:
1739:
1738:Anhanguerinae
1736:
1735:
1732:
1731:
1723:
1722:
1714:
1713:
1705:
1704:
1701:
1700:
1699:
1692:
1691:
1688:
1687:
1684:
1683:
1682:
1675:
1674:
1668:
1667:
1664:
1663:
1660:
1659:
1658:
1651:
1650:
1644:
1643:
1640:
1639:
1636:
1635:
1634:
1627:
1626:
1620:
1617:
1616:
1610:
1609:
1603:
1602:Anhangueridae
1600:
1599:
1595:
1593:
1589:
1585:
1578:
1577:
1569:
1568:
1560:
1559:
1551:
1550:
1547:
1546:
1545:
1538:
1537:
1534:
1533:
1525:
1524:
1521:
1520:
1519:
1518:Maaradactylus
1512:
1511:
1508:
1507:
1504:
1503:
1502:
1501:Cearadactylus
1495:
1494:
1488:
1487:
1484:
1483:
1480:
1479:
1478:
1471:
1470:
1464:
1463:
1460:
1459:
1451:
1450:
1447:
1446:
1445:
1438:
1437:
1434:
1433:
1430:
1429:
1428:
1421:
1420:
1417:
1416:
1413:
1412:
1411:
1404:
1403:
1397:
1396:
1390:
1389:Anhanguerinae
1387:
1386:
1383:
1382:
1374:
1373:
1365:
1364:
1356:
1355:
1352:
1351:
1350:
1343:
1342:
1339:
1338:
1335:
1334:
1333:
1326:
1325:
1319:
1318:
1315:
1314:
1311:
1310:
1309:
1302:
1301:
1295:
1294:
1291:
1290:
1287:
1286:
1285:
1278:
1277:
1271:
1268:
1267:
1264:
1263:
1255:
1254:
1246:
1245:
1242:
1241:
1240:
1233:
1232:
1229:
1228:
1225:
1224:
1223:
1216:
1215:
1209:
1208:
1205:
1204:
1196:
1195:
1192:
1191:
1190:
1189:Tropeognathus
1183:
1182:
1179:
1178:
1175:
1174:
1173:
1172:Siroccopteryx
1166:
1165:
1159:
1158:
1152:
1149:
1148:
1142:
1141:Anhangueridae
1139:
1138:
1134:
1132:
1128:
1121:
1120:
1112:
1111:
1103:
1102:
1094:
1093:
1085:
1084:
1076:
1075:
1067:
1066:
1058:
1057:
1049:
1048:
1040:
1039:
1036:
1035:
1034:
1027:
1026:
1023:
1022:
1019:
1018:
1017:
1010:
1009:
1003:
1002:
999:
998:
995:
994:
993:
986:
985:
979:
978:
975:
974:
971:
970:
969:
962:
961:
955:
952:
951:
948:
947:
944:
943:
942:
941:Tropeognathus
935:
934:
928:
925:
924:
921:
920:
912:
911:
908:
907:
906:
899:
898:
895:
894:
891:
890:
889:
882:
881:
875:
874:Anhangueridae
872:
871:
865:
862:
861:
858:
857:
854:
853:
852:
851:Cearadactylus
845:
844:
838:
837:
834:
833:
825:
824:
821:
820:
819:
812:
811:
808:
807:
804:
803:
802:
795:
794:
788:
787:
781:
780:
774:
773:
770:
769:
766:
765:
764:
757:
756:
749:
745:
744:
740:
738:
734:
730:
729:
725:
723:
719:
715:
711:
710:
705:
701:
697:
693:
689:
685:
681:
680:Anhangueridae
677:
673:
672:
667:
666:
661:
660:
655:
651:
647:
646:
641:
637:
633:
629:
625:
621:
617:
613:
608:
606:
602:
598:
594:
590:
586:
580:
579:occlusal view
576:
575:
574:Siroccopteryx
570:
569:
568:Tropeognathus
564:
560:
556:
549:
547:
543:
536:
532:
527:
520:
518:
514:
511:
507:
502:
500:
493:
489:
485:
478:
476:
474:
470:
466:
462:
458:
453:
451:
447:
446:specific name
443:
439:
435:
432:
428:
424:
419:
416:
413:In 2019, the
411:
408:
400:
399:
394:
393:
388:
384:
380:
373:
371:
369:
365:
364:Anhangueridae
361:
357:
353:
352:
347:
343:
339:
335:
331:
327:
326:
321:
317:
313:
310:
306:
303:
299:
296:
292:
288:
287:
276:
270:
269:
261:
258:
254:
249:
243:
242:
235:
232:
231:
228:
222:
219:
218:
215:
214:Anhangueridae
209:
206:
205:
202:
196:
193:
192:
189:
183:
180:
179:
176:
173:
170:
169:
166:
163:
160:
159:
156:
153:
150:
149:
144:
139:
135:
131:
130:
129:Tropeognathus
125:
120:
116:
111:
104:
99:
94:
89:
84:
79:
74:
69:
64:
59:
54:
49:
43:
36:
32:
26:
22:
19:
7134:Paleontology
7075:
7039:
6985:
6980:Ludodactylus
6978:
6971:
6964:
6956:
6948:
6941:
6921:
6913:
6908:Nicorhynchus
6906:
6899:
6892:
6871:
6864:
6856:
6848:
6840:
6833:
6832:
6824:
6794:
6787:
6779:Hamipteridae
6765:
6757:
6723:
6715:
6707:
6687:
6679:
6674:Aetodactylus
6672:
6637:
6629:
6622:
6614:
6608:Ludodactylus
6606:
6598:
6591:
6590:
6582:
6574:
6566:
6560:Aetodactylus
6558:
6537:
6529:
6521:
6513:
6505:
6498:
6491:
6448:
6441:
6417:
6410:
6403:
6396:
6370:Liaoxipterus
6368:
6361:
6341:
6334:
6327:
6320:
6298:Mimodactylus
6296:
6289:
6282:
6261:
6254:
6247:
6240:
6216:
6209:
6184:
6176:Lanceodontia
6157:
6149:
6144:Lonchodectes
6142:
6134:
6126:
6106:
6099:
6092:
6045:
6038:
6031:
6024:
6004:
5997:
5990:
5965:
5958:
5933:
5925:
5918:
5912:Ornithostoma
5910:
5903:
5895:
5866:
5806:
5720:
5712:
5705:
5698:
5691:
5684:
5679:Eurazhdarcho
5677:
5670:
5663:
5656:
5636:
5629:
5622:
5614:
5607:Azhdarchinae
5593:
5585:
5578:
5570:
5563:
5555:
5548:Azhdarchidae
5534:
5527:
5519:
5491:
5484:
5477:
5456:
5448:
5443:Meilifeilong
5441:
5434:
5426:
5418:
5410:
5386:
5361:
5353:
5345:
5337:
5329:
5322:
5300:
5293:
5286:
5278:
5228:
5221:
5201:
5194:
5187:
5179:
5155:
5147:
5139:
5131:
5123:
5099:
5092:
5085:
5080:Huaxiapterus
5078:
5071:
5065:Eopteranodon
5063:
5055:
5049:Afrotapejara
5047:
5040:Sinopterinae
5029:Afrotapejara
5027:
5003:
4996:
4965:
4958:
4951:
4943:
4935:
4928:
4920:
4912:
4904:
4896:
4874:
4869:Bennettazhia
4867:
4832:Ornithostoma
4830:
4823:
4815:
4807:
4800:
4793:
4759:
4752:
4731:
4723:
4718:Ordosipterus
4716:
4709:
4701:
4693:
4668:Tapejaroidea
4653:
4557:
4511:Pterodaustro
4509:
4502:
4495:
4488:
4468:
4444:
4437:
4430:
4424:Huanhepterus
4422:
4417:Gnathosaurus
4415:
4397:Moganopterus
4395:
4388:
4369:Pterofiltrus
4367:
4362:Petrodactyle
4360:
4353:
4348:Liaodactylus
4346:
4341:Kepodactylus
4339:
4332:
4325:
4318:
4311:
4304:
4297:
4289:
4256:Huanhepterus
4254:
4247:
4239:
4218:
4213:Aerodactylus
4211:
4186:
4180:Petrodactyle
4178:
4170:
4163:
4155:
4120:
4113:
4082:
4074:
4067:
4059:
4039:
3930:
3922:
3915:
3908:
3901:
3896:Kryptodrakon
3894:
3886:
3879:
3872:
3844:
3838:Jeholopterus
3836:
3828:
3821:
3814:
3794:
3787:
3781:Jeholopterus
3779:
3771:
3766:Anurognathus
3764:
3746:Mesadactylus
3744:
3709:
3701:
3663:
3655:
3648:
3641:
3633:
3613:
3589:
3582:
3571:Darwinoptera
3557:
3550:
3543:
3512:
3504:
3450:
3379:
3372:
3365:
3358:
3351:
3308:
3302:Nesodactylus
3300:
3294:Cacibupteryx
3292:
3274:Sericipterus
3272:
3264:
3256:
3249:
3242:
3213:
3206:
3201:Nesodactylus
3199:
3192:
3185:
3180:Cacibupteryx
3178:
3173:Bellubrunnus
3171:
3146:
3138:
3131:
3123:
3084:
3077:
3041:
3028:Lonchognatha
3009:
3001:
2993:
2986:
2979:
2971:
2951:
2920:
2881:
2873:
2852:
2828:
2821:
2816:Pachagnathus
2814:
2806:
2776:
2768:
2762:Austriadraco
2760:
2726:
2721:Austriadraco
2719:
2711:
2690:
2683:
2675:
2650:
2643:
2623:
2509:
2500:
2491:
2412:. Retrieved
2398:
2388:
2369:
2365:
2355:
2347:
2344:
2339:
2304:
2298:
2271:
2267:
2213:(1): 13454.
2210:
2206:
2200:
2143:
2092:
2091:
2075:
2074:
2059:
2058:
2057:
2022:
2021:
2005:
2004:
1988:
1987:
1911:
1910:
1897:
1874:
1873:
1857:
1856:
1840:
1839:
1798:
1797:
1781:
1780:
1759:Ludodactylus
1757:
1756:
1696:
1695:
1681:Nicorhynchus
1679:
1678:
1655:
1654:
1631:
1630:
1591:
1587:
1586:
1542:
1541:
1516:
1515:
1499:
1498:
1475:
1474:
1444:Ludodactylus
1442:
1441:
1425:
1424:
1408:
1407:
1347:
1346:
1332:Nicorhynchus
1330:
1329:
1306:
1305:
1282:
1281:
1238:
1237:
1236:
1220:
1219:
1187:
1186:
1170:
1169:
1130:
1129:
1031:
1030:
1015:
1014:
1013:
990:
989:
966:
965:
939:
938:
903:
902:
886:
885:
849:
848:
818:Ludodactylus
816:
815:
799:
798:
761:
760:
736:
732:
731:
721:
720:, including
707:
703:
699:
695:
691:
687:
675:
669:
663:
657:
649:
643:
627:
623:
620:phylogenetic
615:
611:
609:
604:
596:
584:
583:
572:
566:
562:
544:
540:
534:
515:
509:
503:
498:
497:
472:
469:frontal bone
454:
437:
433:
417:
415:type species
412:
404:
396:
390:
386:
385:specimen of
359:
355:
349:
341:
337:
324:
323:
285:
284:
283:
274:
267:
266:
257:Type species
247:
240:
239:
127:
123:
24:
18:
6866:Thapunngaka
6826:Amblydectes
6745:Anhangueria
6709:Aussiedraco
6443:Boreopterus
6218:Lonchodraco
6151:Lonchodraco
6094:Aussiedraco
6040:Nyctosaurus
5897:Bogolubovia
5565:Bogolubovia
5436:Lacusovagus
5428:Eoazhdarcho
5388:Eoazhdarcho
5347:Leptostomia
5339:Keresdrakon
5214:Caiuajarina
5181:Bakonydraco
5149:Lacusovagus
5141:Keresdrakon
5112:Tapejarinae
5073:Huaxiadraco
5057:Bakonydraco
5016:Tapejaridae
4945:Leptostomia
4937:Lacusovagus
4930:Kariridraco
4876:Keresdrakon
4817:Leptostomia
4809:Keresdrakon
4470:Ctenochasma
4432:Lusognathus
4355:Otogopterus
4010:Lophocratia
3943:Lophocratia
3910:Pangupterus
3846:Sinomacrops
3187:Dorygnathus
3056:Novialoidea
3011:Rhamphinion
2988:Dimorphodon
2663:Caviramidae
2607:Pterosauria
2594:Pterosauria
2557:Pterosauria
2496:Archosauria
2457:Pterosauria
2077:Thapunngaka
2024:Amblydectes
1898:AMNH 22555
1594:., (2023).
1588:Topology 3:
1131:Topology 2:
748:Anhangueria
739:., (2019).
733:Topology 1:
709:Thapunngaka
589:Anhangueria
479:Description
398:Aussiedraco
302:anhanguerid
188:Pterosauria
40:96–90
7156:Categories
7076:Ferrodraco
6943:Anhanguera
6834:Ferrodraco
6789:Hamipterus
6717:Barbosania
6592:Ferrodraco
6584:Draigwenia
6419:Hamipterus
6398:Barbosania
6343:Nurhachius
6211:Ikrandraco
6186:Draigwenia
6136:Ikrandraco
6101:Serradraco
6047:Volgadraco
5935:Volgadraco
5927:Tethydraco
5920:Pteranodon
5778:see below↓
5672:Cryodrakon
5595:Volgadraco
5587:Tethydraco
5486:Jidapterus
5450:Microtuban
5288:Microtuban
5189:Europejara
5168:Tapejarini
5157:Vectidraco
5094:Sinopterus
4711:Noripterus
4580:see below↓
4504:Gegepterus
4497:Eosipterus
3952:see below↓
3932:Wenupteryx
3823:Cascocauda
3402:see below↓
2830:Yelaphomte
2566:see below↓
2487:Sauropsida
2345:Alcheringa
2165:References
2060:Ferrodraco
2046:Mythungini
1913:Anhanguera
1544:Anhanguera
1239:Ferrodraco
1016:Ferrodraco
905:Anhanguera
722:Ferrodraco
714:Mythungini
696:Ferrodraco
692:Ferrodraco
676:Ferrodraco
659:Anhanguera
650:Ferrodraco
628:Ferrodraco
624:Ferrodraco
612:Ferrodraco
605:Ferrodraco
597:Ferrodraco
585:Ferrodraco
563:Ferrodraco
535:Ferrodraco
529:Skull and
510:Ferrodraco
499:Ferrodraco
473:Ferrodraco
461:Cenomanian
387:Ferrodraco
356:Ferrodraco
342:Ferrodraco
325:F. lentoni
316:Queensland
286:Ferrodraco
241:Ferrodraco
227:Mythungini
194:Suborder:
31:Cenomanian
25:Ferrodraco
7146:Australia
7091:Q69739391
6894:Aerodraco
6336:Luchibang
6284:Haopterus
6006:Simurghia
5999:Epapatelo
5967:Cretornis
5905:Dawndraco
5631:Azhdarcho
5624:Albadraco
5616:Aerotitan
5536:Alanqidae
5314:Alanqidae
5280:Cretornis
5230:Torukjara
5223:Caiuajara
4960:Tupuxuara
4898:Aerotitan
4802:Inabtanin
4390:Feilongus
3789:Luopterus
3506:Allkaruen
3133:Klobiodon
2973:Allkaruen
2808:Caviramus
2685:Caviramus
2473:Kingdom:
2414:7 October
2350:: 273–279
2331:249332375
2290:0272-4634
2227:2045-2322
1633:Aerodraco
1590:Richards
1284:Aerodraco
442:ironstone
427:osteology
320:Australia
305:pterosaur
291:ironstone
273:Pentland
246:Pentland
161:Kingdom:
155:Eukaryota
7085:Wikidata
6966:Guidraco
6850:Mythunga
6842:Haliskia
6616:Mythunga
6600:Guidraco
6531:Haliskia
6108:Unwindia
5458:Xericeps
5363:Xericeps
5295:Xericeps
5196:Tapejara
4967:Xericeps
4922:Banguela
4695:Banguela
3924:Samrukia
3584:Ceoptera
2481:Chordata
2479:Phylum:
2475:Animalia
2245:31582757
2153:See also
2094:Mythunga
1800:Guidraco
1427:Guidraco
1222:Mythunga
1033:Mythunga
763:Guidraco
700:Mythunga
688:Mythunga
654:polytomy
645:Mythunga
571:(B) and
531:mandible
521:Skeleton
465:Turonian
463:- lower
407:holotype
395:(B) and
392:Mythunga
383:Holotype
360:Mythunga
351:Mythunga
338:AODF 876
207:Family:
175:Chordata
171:Phylum:
165:Animalia
151:Domain:
35:Turonian
7120:Portals
5992:Alcione
5101:Wightia
2485:Class:
2236:6776501
750:
638:as the
295:extinct
233:Genus:
220:Tribe:
181:Order:
7105:399803
5557:Alanqa
5324:Alanqa
4906:Alanqa
3514:Sordes
3381:Sordes
2329:
2319:
2288:
2243:
2233:
2225:
2144:
746:
616:et al.
559:Palate
444:. The
434:ferrum
330:Winton
277:, 2019
275:et al.
250:, 2019
248:et al.
4655:Piksi
3251:Dearc
2510:Clade
2501:Clade
2492:Clade
2327:S2CID
1592:et al
737:et al
704:et al
438:draco
431:Latin
298:genus
2543:see
2416:2023
2317:ISBN
2286:ISSN
2241:PMID
2223:ISSN
698:and
668:and
425:and
405:The
48:PreꞒ
2403:doi
2374:doi
2309:doi
2276:doi
2231:PMC
2215:doi
642:of
561:of
533:of
401:(C)
314:of
300:of
7158::
7102::
7087::
2512::
2503::
2494::
2401:.
2397:.
2372:.
2370:65
2368:.
2364:.
2348:34
2325:.
2315:.
2284:.
2272:41
2270:.
2266:.
2253:^
2239:.
2229:.
2221:.
2209:.
2205:.
2173:^
724:.
662:,
607:.
370:.
332:,
318:,
98:Pg
42:Ma
37:,
7122::
6961:?
6953:?
6918:?
6886:?
6861:?
6853:?
6845:?
6837:?
6829:?
6818:?
6781:?
6770:?
6762:?
6720:?
6712:?
6684:?
6657:?
6642:?
6634:?
6619:?
6611:?
6603:?
6595:?
6587:?
6579:?
6571:?
6563:?
6542:?
6534:?
6526:?
6518:?
6510:?
6266:?
6189:?
6162:?
6154:?
6139:?
6131:?
6050:?
5970:?
5938:?
5930:?
5915:?
5900:?
5717:?
5619:?
5598:?
5590:?
5575:?
5560:?
5539:?
5524:?
5461:?
5453:?
5431:?
5423:?
5415:?
5366:?
5358:?
5350:?
5342:?
5334:?
5316:?
5305:?
5283:?
5184:?
5152:?
5144:?
5136:?
5128:?
5068:?
5060:?
5052:?
4990:?
4970:?
4948:?
4940:?
4925:?
4917:?
4909:?
4901:?
4890:?
4879:?
4820:?
4812:?
4736:?
4728:?
4706:?
4698:?
4658:?
4562:?
4427:?
4382:?
4294:?
4259:?
4244:?
4223:?
4191:?
4183:?
4175:?
4160:?
4087:?
4079:?
4064:?
3927:?
3891:?
3841:?
3833:?
3784:?
3776:?
3706:?
3660:?
3638:?
3562:?
3517:?
3509:?
3384:?
3345:?
3335:?
3305:?
3297:?
3269:?
3261:?
3151:?
3143:?
3128:?
3046:?
3030:?
3014:?
3006:?
2998:?
2976:?
2925:?
2909:?
2878:?
2857:?
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2781:?
2773:?
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2716:?
2695:?
2680:?
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2449:e
2442:t
2435:v
2418:.
2405::
2382:.
2376::
2333:.
2311::
2292:.
2278::
2247:.
2217::
2211:9
2199:"
264:†
237:†
224:†
211:†
198:†
185:†
103:N
93:K
88:J
83:T
78:P
73:C
68:D
63:S
58:O
53:Ꞓ
33:-
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
