1138:
1491:
1753:
1363:
377:
1330:
2037:
537:
316:
3457:
1241:
1976:
3418:
1085:
1940:
384:
700:
2389:
1300:
3959:
Escaso, Fernando; Ortega, Francisco; Dantas, Pedro; Malafaia, Elisabete; Pimentel, Nuno L.; Pereda-Suberbiola, Xabier; Sanz, José Luis; Kullberg, José Carlos; Kullberg, María Carla (2006-12-23). "New evidence of shared dinosaur across Upper
Jurassic Proto-North Atlantic: Stegosaurus from Portugal".
1133:"...one tooth, five cervical vertebrae (including the axis), five dorsal vertebrae, cervical and dorsal ribs, three caudal vertebrae, chevrons, preacetabular process of the left ilium, right tibia, fibula, astragalus and , a cervical plate, and several fragments of plates."
429:. It is an important source of information on the diversity of Late Jurassic mammals. Many of the fossils were collected from the now disused and flooded Camadas de Guimarota coal mine.
3682:"Life at the continental–marine interface: palaeoenvironments and biota of the Alcobaça Formation (Late Jurassic, Central Portugal), with a formal definition of the unit appended"
3609:
4164:
3922:"M ARTIN , T. & K REBS , B. (eds) 2000. Guimarota. A Jurassic Ecosystem . 155 pp. München: Verlag Dr. Friedrich Pfeil. Price DM 120.00 (hard covers). ISBN 3 931516 80 6"
4010:"16.1 Distrito do Leiria, Portugal; 3. Camadas de Alcobaça," and "16.4 Distrito do Lisboa, Portugal; 1. Camadas de Alcobaça," in Weishampel, et al. (2004). Pages 548-549.
833:
are one of the most numerous faunal components of the
Guimarota mine with around 9000 isolated remains, including parts of the skull and limbs. Remains of indeterminate
4109:
3604:
4154:
3631:"The Lourinhã Formation: the Upper Jurassic to lower most Cretaceous of the Lusitanian Basin, Portugal – landscapes where dinosaurs walked"
4114:
376:
4169:
4085:
295:
4029:
Malafaia, E.; Ortega, F.; Escaso, F.; Dantas, P.; Pimentel, N.; Gasulla, J. M.; Ribeiro, B.; Barriga, F.; Sanz, J. L. (2010).
4159:
1021:
Indeterminate euornithopod remains located in Lisbon
District. Indeterminate stegosaurid remains present in Lisbon District.
3921:
3413:
This genus is regarded as an indicator for brachyhaline water conditions and it was likely buried in its natural habitat
214:
260:
221:
4124:
4144:
4119:
4139:
3746:
3721:
4134:
4129:
3538:
4149:
3583:
3528:
1137:
3680:
Fürsich, Franz T.; Schneider, Simon; Werner, Winfried; Lopez-Mir, Berta; Pierce, Colm S. (2021-09-13).
3248:
346:
151:
2722:
4030:
3969:
161:
141:
131:
3843:"16.1 Distrito do Leiria, Portugal; 3. Camadas de Alcobaça," in Weishampel, et al. (2004). Page 548.
3827:"16.4 Distrito do Lisboa, Portugal; 1. Camadas de Alcobaça," in Weishampel, et al. (2004). Page 548.
3164:
1490:
3192:
2455:
1763:
414:
3337:
3220:
2275:
1752:
734:
655:
3993:
3563:
3291:
1588:
1563:
590:
545:
3309:
3266:
3238:
3210:
3182:
1362:
3115:
3097:
2831:
1818:
4081:
4052:
3985:
3941:
3703:
3662:
1641:
896:
830:
4031:"Vertebrate fauna at the Allosaurus fossil-site of Andrés (Upper Jurassic), Pombal, Portugal"
4042:
3977:
3933:
3693:
3652:
3642:
2939:
2917:
2850:
2507:
2504:
2445:
2442:
2356:
2353:
2265:
2262:
2232:
2151:
2140:
2111:
2108:
1950:
1692:
1498:
1329:
568:
331:
3587:
2779:
2696:
2573:
2089:
2036:
2030:
1933:
1881:
1862:
1610:
1470:
1287:
1228:
1072:
442:
438:
303:
299:
3973:
3427:
3120:
2366:
1373:
1340:
930:
458:
253:
4103:
3630:
2892:
2882:
2805:
2769:
2644:
2547:
2170:
2011:
1969:
1703:
1028:
907:
803:
756:
514:
426:
81:
38:
3997:
3647:
4047:
2879:
2741:
2243:
1986:
1398:
1310:
1251:
961:
623:
536:
422:
356:
42:
3456:
1240:
4075:
3410:
Dozens of well preserved shells sometimes making up entire stratigraphic layers
3017:
2670:
2618:
2213:
2173:
2121:
1975:
1914:
1792:
1721:
1186:
1148:
1116:
1061:
993:
679:
56:
3698:
3681:
3586:
on the shells of this taxon indicate a marine lifestyle, modern relatives like
3981:
3937:
3510:
3475:
3417:
3124:
3003:
2599:
2536:
2485:
2474:
2423:
2334:
2305:
2294:
2202:
2183:
1732:
1674:
1219:
1094:
1084:
834:
101:
66:
4056:
3945:
3707:
3666:
275:
262:
236:
223:
3394:
3319:
2752:
2517:
2324:
1939:
1845:
1434:
1276:
880:
847:
727:
Two species of different size can be distinguished from the fossil material
709:
202:
198:
194:
106:
50:
3989:
724:
isolated lateral teeth, lower jaws, ganoid scales and isolated skull bones
3062:
2397:
418:
321:
186:
96:
91:
76:
71:
61:
17:
699:
3657:
3010:
2862:
779:
490:
111:
86:
3590:
also venture onto land, which can also be expected from M. jurassicus
2388:
1751:
1299:
2492:
K. guimarotensis, K. dietrichi, K. simpsoni, K. uniradiculatus, K. sp
2056:
1530:
2976:
2981:
1782:
838:
190:
176:
4074:
Weishampel, David B.; Dodson, Peter; Osmólska, Halszka (2004),
1179:
Indeterminate sauropod remains located in Leiria and Lisboa.
4080:, Berkeley: University of California Press, pp. 1–880,
2035:
1974:
1938:
1361:
1328:
1298:
1239:
1166:
Later determined to be indeterminate ornithischian remains.
1136:
1083:
441:, Portugal. Dinosaur tracks are geographically located in
3531:
on the shells of this taxon indicate a marine lifestyle
3449:
Dozens of well preserved shells with articulated valves
2051:
1876:
1636:
1525:
1416:
Later determined to be indeterminate theropod remains.
1181:
1023:
957:
843:
4019:"Table 13.1," in Weishampel, et al. (2004). Page 265.
3610:
List of fossiliferous stratigraphic units in
Portugal
362:
352:
342:
337:
327:
309:
291:
252:
213:
208:
182:
172:
167:
157:
147:
137:
127:
32:
3629:Mateus, O.; Dinis, J.; Cunha, P. P. (2017-09-28).
1356:" cervicals, dorsals, ribs, limb elements."
8:
437:Dinosaur eggs are geographically located in
4165:Fossiliferous stratigraphic units of Europe
530:isolated teeth, fin-spine and a head-spine
3635:Ciências da Terra / Earth Sciences Journal
3455:
3416:
2797:A multituberculate of uncertain placement
2387:
1489:
698:
535:
383:
29:
4046:
3823:
3821:
3819:
3817:
3815:
3813:
3811:
3809:
3807:
3805:
3803:
3801:
3799:
3797:
3795:
3793:
3791:
3789:
3787:
3697:
3686:Palaeobiodiversity and Palaeoenvironments
3656:
3646:
3839:
3837:
3835:
3833:
3785:
3783:
3781:
3779:
3777:
3775:
3773:
3771:
3769:
3767:
3605:List of dinosaur-bearing rock formations
3469:
3356:
3119:is a typical component of Late Jurassic
3056:
2054:
1879:
1639:
1528:
1428:
1184:
1026:
960:
846:
617:
452:
3621:
3123:environments in Europe. Member of the
3915:
3913:
3911:
3909:
3907:
3905:
3903:
3901:
3899:
3897:
3895:
3893:
3891:
3889:
3887:
3885:
3883:
3881:
3879:
3877:
3875:
3873:
3871:
3869:
7:
3867:
3865:
3863:
3861:
3859:
3857:
3855:
3853:
3851:
3849:
3033:Uncertain or tentative taxa are in
794:Isolated Teeth and a partial skull
3920:Palmer, Douglas (September 2001).
2910:A paulchoffatiid multituberculate
2823:A paulchoffatiid multituberculate
2714:A paulchoffatiid multituberculate
2688:A pinheirodontid multituberculate
2662:A paulchoffatiid multituberculate
2591:A paulchoffatiid multituberculate
2565:A paulchoffatiid multituberculate
2415:A pinheirodontid multituberculate
25:
2993:Taxon falsely reported as present
2932:4 distinct genera with 7 species
693:Isolated teeth and prearticulars
841:are also present but far rarer.
382:
375:
314:
27:Geological formation in Portugal
4110:Geologic formations of Portugal
2998:Dubious taxon or junior synonym
559:Isolated teeth and a fin-spine
4048:10.5209/rev_JIGE.2010.v36.n2.7
1:
4155:Ichnofossiliferous formations
3648:10.21695/cterra/esj.v19i1.355
2677:P. sp, P. vastus, P. pygmaeus
391:Alcobaça Formation (Portugal)
2812:P. dryas, P krebsi, P. oreas
254:Approximate paleocoordinates
4077:The Dinosauria, 2nd edition
2960:
4186:
4035:Journal of Iberian Geology
3747:"Porto Pinheiro, Lourinha"
3699:10.1007/s12549-021-00496-x
964:of the Alcobaça Formation
850:of the Alcobaça Formation
626:of the Alcobaça Formation
461:of the Alcobaça Formation
405:, previously known as the
4115:Jurassic System of Europe
3982:10.1007/s00114-006-0209-8
3938:10.1017/S0016756801316084
3474:
3361:
3264:
3246:
3113:
3095:
3061:
2967:
2963:
2616:
2597:
2059:of the Alcobaça Formation
1884:of the Alcobaça Formation
1644:of the Alcobaça Formation
1533:of the Alcobaça Formation
1433:
1189:of the Alcobaça Formation
1031:of the Alcobaça Formation
923:Indeterminate salamander
622:
457:
370:
37:
4170:Paleontology in Portugal
2965:
769:Isolated grasping tooth
3495:Stratigraphic Position
3379:Stratigraphic Position
3082:Stratigraphic Position
3048:Invertebrate Paleofauna
2703:P. thoas, P. peperethos
1825:S. henkeli, cf. obtusus
1748:Basal Lepidosauromorph
1454:Stratigraphic Position
640:Stratigraphic Position
475:Stratigraphic Position
421:. It dates back to the
3041:taxa are discredited.
2074:Stratigraphic position
2040:
1979:
1943:
1899:Stratigraphic position
1756:
1659:Stratigraphic position
1548:Stratigraphic position
1366:
1333:
1303:
1244:
1204:Stratigraphic position
1141:
1088:
1046:Stratigraphic position
978:Stratigraphic position
864:Stratigraphic position
409:and also known as the
4160:Ooliferous formations
3751:Paleobiology Database
3726:Paleobiology Database
2404:I. quadrituberculatus
2039:
1978:
1942:
1755:
1365:
1332:
1302:
1243:
1140:
1087:
433:Vertebrate paleofauna
2128:B. sp, B. atlanticus
162:Montejunto Formation
142:Camadas de Guimarota
132:Geological formation
3974:2007NW.....94..367E
3962:Naturwissenschaften
3926:Geological Magazine
3471:
3358:
3058:
1484:Isolated Fragments
1430:
945:Indeterminate frog
619:
581:One isolated tooth
454:
407:Guimarota Formation
272: /
233: /
39:Stratigraphic range
4125:Kimmeridgian Stage
3480:Alcobaça Formation
3470:
3364:Alcobaça Formation
3357:
3067:Alcobaça Formation
3057:
2988:Reclassified taxon
2041:
1980:
1944:
1757:
1439:Alcobaça Formation
1429:
1367:
1334:
1304:
1245:
1142:
1089:
618:
453:
403:Alcobaça Formation
347:Alcobaça, Portugal
152:Lourinhã Formation
33:Alcobaça Formation
4145:Lagoonal deposits
4120:Jurassic Portugal
3596:
3595:
3584:clionidid sponges
3582:Boring traces of
3529:clionidid sponges
3527:Boring traces of
3463:
3462:
3345:
3344:
3045:
3044:
3036:
3024:
3023:
3020:
3013:
3006:
2999:
2994:
2989:
2984:
2959:
2958:
2853:multituberculate
2580:M. krausei, M. sp
2235:multituberculate
2143:multituberculate
2045:
2044:
1870:
1869:
1642:Lepidosauromorphs
1633:Lepidosauromorphs
1630:
1629:
1519:
1518:
1422:
1421:
1347:D. lourinhanensis
1172:
1171:
1014:
1013:
951:
950:
823:
822:
611:
610:
399:
398:
49:156.9–152.5
16:(Redirected from
4177:
4140:Coal in Portugal
4096:
4095:
4094:
4061:
4060:
4050:
4026:
4020:
4017:
4011:
4008:
4002:
4001:
3956:
3950:
3949:
3917:
3844:
3841:
3828:
3825:
3762:
3761:
3759:
3757:
3743:
3737:
3736:
3734:
3732:
3718:
3712:
3711:
3701:
3677:
3671:
3670:
3660:
3650:
3626:
3472:
3459:
3452:freshwater clam
3420:
3362:Bivalves of the
3359:
3227:T. guimarotensis
3059:
3034:
3016:
3009:
3002:
2997:
2992:
2987:
2980:
2977:
2961:
2940:Pinheirodontidae
2918:Paulchoffatiidae
2723:Portopinheirodon
2508:multituberculate
2446:multituberculate
2391:
2357:multituberculate
2266:multituberculate
2152:Drescheratherium
2112:multituberculate
2052:
1993:L. mitracostatus
1951:Knoetschkesuchus
1921:G. baryglyphaeus
1877:
1863:rhynchocephalian
1637:
1526:
1499:Pterodactyloidea
1493:
1431:
1380:L. alenquerensis
1182:
1123:S. cf. ungulatus
1024:
958:
844:
831:Albanerpetontids
702:
620:
569:Orectolobiformes
539:
455:
413:is a geological
411:Consolação Unit,
386:
385:
379:
332:Lusitanian Basin
320:
318:
317:
287:
286:
284:
283:
282:
277:
273:
270:
269:
268:
265:
248:
247:
245:
244:
243:
238:
234:
231:
230:
229:
226:
123:
53:
30:
21:
4185:
4184:
4180:
4179:
4178:
4176:
4175:
4174:
4135:Coal formations
4130:Marl formations
4100:
4099:
4092:
4090:
4088:
4073:
4070:
4065:
4064:
4028:
4027:
4023:
4018:
4014:
4009:
4005:
3958:
3957:
3953:
3919:
3918:
3847:
3842:
3831:
3826:
3765:
3755:
3753:
3745:
3744:
3740:
3730:
3728:
3720:
3719:
3715:
3679:
3678:
3674:
3628:
3627:
3623:
3618:
3601:
3468:
3355:
3350:
3199:B. pahasapensis
3055:
3050:
3032:
2975:
2947:Porto Pinheiro
2871:Porto Pinheiro
2786:P. plagiocyrtus
2780:Proalbionbaatar
2761:Porto Pinheiro
2733:Porto Pinheiro
2729:P. asymmetricus
2697:Plesiochoffatia
2681:Porto Pinheiro
2610:Porto Pinheiro
2574:Meketichoffatia
2528:Porto Pinheiro
2408:Porto Pinheiro
2384:A dryolestidan
2341:H. naias, H. sp
2224:Porto Pinheiro
2132:Porto Pinheiro
2090:Bathmochoffatia
2050:
1882:Crocodyliformes
1875:
1873:Crocodyliformes
1854:Andrés, Pombal
1710:D. purbeckensis
1681:B. hoffstetteri
1635:
1611:Pleurosternidae
1524:
1471:Rhamphorhynchus
1427:
1177:
1019:
956:
897:Albanerpetontid
889:Porto Pinheiro
828:
747:Isolated teeth
669:Isolated teeth
616:
603:Isolated teeth
504:Isolated Teeth
451:
443:Leiria District
439:Lisbon District
435:
395:
394:
393:
392:
389:
388:
387:
315:
313:
280:
278:
274:
271:
266:
263:
261:
259:
258:
241:
239:
235:
232:
227:
224:
222:
220:
219:
122:
121:
120:
119:
114:
109:
104:
99:
94:
89:
84:
79:
74:
69:
64:
59:
48:
47:
45:
28:
23:
22:
15:
12:
11:
5:
4183:
4181:
4173:
4172:
4167:
4162:
4157:
4152:
4147:
4142:
4137:
4132:
4127:
4122:
4117:
4112:
4102:
4101:
4098:
4097:
4086:
4069:
4066:
4063:
4062:
4041:(2): 193–204.
4021:
4012:
4003:
3968:(5): 367–374.
3951:
3932:(5): 619–630.
3845:
3829:
3763:
3738:
3713:
3692:(2): 265–329.
3672:
3620:
3619:
3617:
3614:
3613:
3612:
3607:
3600:
3597:
3594:
3593:
3591:
3580:
3577:
3575:
3572:
3567:
3559:
3558:
3556:
3554:
3551:
3549:
3546:
3545:Indeterminate
3543:
3535:
3534:
3532:
3525:
3522:
3520:
3517:
3516:Indeterminate
3514:
3506:
3505:
3502:
3499:
3496:
3493:
3490:
3487:
3483:
3482:
3467:
3464:
3461:
3460:
3453:
3450:
3447:
3445:
3442:
3432:
3422:
3421:
3414:
3411:
3408:
3406:
3403:
3398:
3390:
3389:
3386:
3383:
3380:
3377:
3374:
3371:
3367:
3366:
3354:
3351:
3349:
3346:
3343:
3342:
3340:
3336:Member of the
3334:
3332:
3330:
3328:
3326:D. leguminella
3323:
3315:
3314:
3312:
3308:Member of the
3306:
3304:
3302:
3300:
3295:
3287:
3286:
3284:
3282:
3280:
3278:
3272:
3271:
3269:
3265:Member of the
3263:
3261:
3259:
3257:
3252:
3244:
3243:
3241:
3237:Member of the
3235:
3233:
3231:
3229:
3224:
3216:
3215:
3213:
3209:Member of the
3207:
3205:
3203:
3201:
3196:
3188:
3187:
3185:
3181:Member of the
3179:
3177:
3175:
3173:
3171:T. wyomingense
3168:
3165:Theriosynoecum
3160:
3159:
3157:
3155:
3153:
3151:
3145:
3144:
3142:
3140:
3138:
3136:
3130:
3129:
3127:
3121:brackish water
3112:
3110:
3108:
3106:
3101:
3093:
3092:
3089:
3086:
3083:
3080:
3077:
3074:
3070:
3069:
3054:
3051:
3049:
3046:
3043:
3042:
3027:
3025:
3022:
3021:
3014:
3007:
3000:
2995:
2990:
2985:
2969:
2968:
2966:
2964:
2957:
2956:
2954:
2952:
2950:
2948:
2945:
2944:Indeterminate
2942:
2936:
2935:
2933:
2930:
2928:
2926:
2923:
2922:Indeterminate
2920:
2914:
2913:
2911:
2908:
2906:
2904:
2901:
2896:
2888:
2887:
2885:
2876:
2874:
2872:
2869:
2868:Indeterminate
2866:
2857:
2856:
2854:
2851:paulchoffatiid
2847:
2845:
2843:
2840:
2835:
2827:
2826:
2824:
2821:
2819:
2817:
2814:
2809:
2801:
2800:
2798:
2795:
2793:
2791:
2788:
2783:
2775:
2774:
2772:
2766:
2764:
2762:
2759:
2758:Indeterminate
2756:
2748:
2747:
2745:
2738:
2736:
2734:
2731:
2726:
2718:
2717:
2715:
2712:
2710:
2708:
2705:
2700:
2692:
2691:
2689:
2686:
2684:
2682:
2679:
2674:
2666:
2665:
2663:
2660:
2658:
2656:
2653:
2648:
2640:
2639:
2637:
2635:
2633:
2630:
2624:
2623:
2621:
2615:
2613:
2611:
2608:
2603:
2595:
2594:
2592:
2589:
2587:
2585:
2582:
2577:
2569:
2568:
2566:
2563:
2561:
2559:
2556:
2551:
2543:
2542:
2540:
2533:
2531:
2529:
2526:
2521:
2513:
2512:
2510:
2505:paulchoffatiid
2501:
2499:
2497:
2494:
2489:
2481:
2480:
2478:
2471:
2469:
2467:
2464:
2462:K. lusitanicum
2459:
2451:
2450:
2448:
2443:paulchoffatiid
2439:
2437:
2435:
2432:
2427:
2419:
2418:
2416:
2413:
2411:
2409:
2406:
2401:
2393:
2392:
2385:
2382:
2380:
2378:
2375:
2370:
2367:Henkelotherium
2362:
2361:
2359:
2354:paulchoffatiid
2350:
2348:
2346:
2343:
2338:
2330:
2329:
2327:
2321:
2319:
2317:
2314:
2312:H. exspectatus
2309:
2301:
2300:
2298:
2291:
2289:
2287:
2284:
2279:
2271:
2270:
2268:
2263:paulchoffatiid
2259:
2257:
2255:
2252:
2247:
2239:
2238:
2236:
2233:pinheirodontid
2229:
2227:
2225:
2222:
2217:
2209:
2208:
2206:
2199:
2197:
2195:
2192:
2187:
2179:
2178:
2176:
2167:
2165:
2163:
2160:
2155:
2147:
2146:
2144:
2141:pinheirodontid
2137:
2135:
2133:
2130:
2125:
2117:
2116:
2114:
2109:paulchoffatiid
2105:
2103:
2101:
2098:
2093:
2085:
2084:
2081:
2078:
2075:
2072:
2069:
2066:
2062:
2061:
2049:
2046:
2043:
2042:
2033:
2027:
2025:
2023:
2020:
2015:
2007:
2006:
2004:
2002:
2000:
1998:
1995:
1990:
1982:
1981:
1972:
1966:
1964:
1962:
1959:
1954:
1946:
1945:
1936:
1930:
1928:
1926:
1923:
1918:
1910:
1909:
1906:
1903:
1900:
1897:
1894:
1891:
1887:
1886:
1874:
1871:
1868:
1867:
1865:
1859:
1857:
1855:
1852:
1851:Indeterminate
1849:
1840:
1839:
1837:
1834:
1832:
1830:
1827:
1822:
1814:
1813:
1811:
1808:
1806:
1804:
1801:
1799:S. proraformis
1796:
1788:
1787:
1785:
1779:
1777:
1775:
1772:
1767:
1759:
1758:
1749:
1746:
1744:
1742:
1739:
1738:Indeterminate
1736:
1728:
1727:
1725:
1719:
1717:
1715:
1712:
1707:
1699:
1698:
1696:
1693:Paramacellodid
1690:
1688:
1686:
1683:
1678:
1670:
1669:
1666:
1663:
1660:
1657:
1654:
1651:
1647:
1646:
1634:
1631:
1628:
1627:
1625:
1623:
1621:
1619:
1616:
1615:Indeterminate
1613:
1607:
1606:
1604:
1602:
1600:
1598:
1595:
1594:Indeterminate
1592:
1584:
1583:
1581:
1579:
1577:
1575:
1572:
1570:P. duchastelli
1567:
1559:
1558:
1555:
1552:
1549:
1546:
1543:
1540:
1536:
1535:
1523:
1520:
1517:
1516:
1514:
1512:
1509:
1507:
1504:
1503:Indeterminate
1501:
1495:
1494:
1487:
1485:
1482:
1480:
1477:
1476:Indeterminate
1474:
1465:
1464:
1461:
1458:
1455:
1452:
1449:
1446:
1442:
1441:
1426:
1423:
1420:
1419:
1417:
1414:
1412:
1410:
1407:
1402:
1394:
1393:
1391:
1389:
1387:
1385:
1382:
1377:
1374:Lourinhasaurus
1369:
1368:
1359:
1357:
1354:
1352:
1349:
1344:
1341:Dinheirosaurus
1336:
1335:
1326:
1324:
1322:
1320:
1317:
1316:Indeterminate
1314:
1306:
1305:
1296:
1294:
1292:
1290:
1285:
1283:L. atalaiensis
1280:
1272:
1271:
1269:
1267:
1265:
1263:
1260:
1255:
1247:
1246:
1237:
1235:
1233:
1231:
1226:
1225:Indeterminate
1223:
1215:
1214:
1211:
1208:
1205:
1202:
1199:
1196:
1192:
1191:
1176:
1173:
1170:
1169:
1167:
1164:
1162:
1160:
1157:
1152:
1144:
1143:
1134:
1131:
1129:
1127:
1125:
1120:
1112:
1111:
1109:
1107:
1105:
1103:
1100:
1099:Indeterminate
1097:
1091:
1090:
1081:
1079:
1077:
1075:
1070:
1065:
1057:
1056:
1053:
1050:
1047:
1044:
1041:
1038:
1034:
1033:
1029:Ornithischians
1018:
1017:Ornithischians
1015:
1012:
1011:
1009:
1007:
1005:
1003:
1000:
999:Indeterminate
997:
989:
988:
985:
982:
979:
976:
973:
970:
966:
965:
955:
952:
949:
948:
946:
943:
941:
939:
936:
935:Indeterminate
933:
931:Discoglossidae
927:
926:
924:
921:
919:
917:
914:
913:Indeterminate
911:
902:
901:
899:
894:
892:
890:
887:
886:Indeterminate
884:
875:
874:
871:
868:
865:
862:
859:
856:
852:
851:
827:
824:
821:
820:
818:
816:
814:
812:
809:
808:Indeterminate
806:
800:
799:
797:
795:
792:
790:
787:
786:Indeterminate
784:
775:
774:
772:
770:
767:
765:
762:
761:Indeterminate
759:
753:
752:
750:
748:
745:
743:
740:
739:Indeterminate
737:
731:
730:
728:
725:
722:
720:
717:
716:Indeterminate
714:
704:
703:
696:
694:
691:
689:
686:
685:Indeterminate
683:
675:
674:
672:
670:
667:
665:
662:
661:Indeterminate
659:
651:
650:
647:
644:
641:
638:
635:
632:
628:
627:
615:
612:
609:
608:
606:
604:
601:
599:
596:
595:Indeterminate
593:
587:
586:
584:
582:
579:
577:
574:
573:Indeterminate
571:
565:
564:
562:
560:
557:
555:
552:
551:Indeterminate
549:
541:
540:
533:
531:
528:
526:
523:
518:
510:
509:
507:
505:
502:
500:
497:
496:Indeterminate
494:
486:
485:
482:
479:
476:
473:
470:
467:
463:
462:
459:Chondrichthyes
450:
449:Chondrichthyes
447:
434:
431:
397:
396:
390:
381:
380:
374:
373:
372:
371:
368:
367:
364:
360:
359:
354:
350:
349:
344:
340:
339:
335:
334:
329:
325:
324:
311:
307:
306:
293:
289:
288:
256:
250:
249:
217:
211:
210:
206:
205:
184:
180:
179:
174:
170:
169:
165:
164:
159:
155:
154:
149:
145:
144:
139:
135:
134:
129:
125:
124:
117:
116:
115:
110:
105:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
54:
35:
34:
26:
24:
14:
13:
10:
9:
6:
4:
3:
2:
4182:
4171:
4168:
4166:
4163:
4161:
4158:
4156:
4153:
4151:
4150:Reef deposits
4148:
4146:
4143:
4141:
4138:
4136:
4133:
4131:
4128:
4126:
4123:
4121:
4118:
4116:
4113:
4111:
4108:
4107:
4105:
4089:
4087:0-520-24209-2
4083:
4079:
4078:
4072:
4071:
4067:
4058:
4054:
4049:
4044:
4040:
4036:
4032:
4025:
4022:
4016:
4013:
4007:
4004:
3999:
3995:
3991:
3987:
3983:
3979:
3975:
3971:
3967:
3963:
3955:
3952:
3947:
3943:
3939:
3935:
3931:
3927:
3923:
3916:
3914:
3912:
3910:
3908:
3906:
3904:
3902:
3900:
3898:
3896:
3894:
3892:
3890:
3888:
3886:
3884:
3882:
3880:
3878:
3876:
3874:
3872:
3870:
3868:
3866:
3864:
3862:
3860:
3858:
3856:
3854:
3852:
3850:
3846:
3840:
3838:
3836:
3834:
3830:
3824:
3822:
3820:
3818:
3816:
3814:
3812:
3810:
3808:
3806:
3804:
3802:
3800:
3798:
3796:
3794:
3792:
3790:
3788:
3786:
3784:
3782:
3780:
3778:
3776:
3774:
3772:
3770:
3768:
3764:
3752:
3748:
3742:
3739:
3727:
3723:
3717:
3714:
3709:
3705:
3700:
3695:
3691:
3687:
3683:
3676:
3673:
3668:
3664:
3659:
3654:
3649:
3644:
3640:
3636:
3632:
3625:
3622:
3615:
3611:
3608:
3606:
3603:
3602:
3598:
3592:
3589:
3585:
3581:
3578:
3576:
3573:
3571:
3570:M. jurassicus
3568:
3566:
3565:
3561:
3560:
3557:
3555:
3552:
3550:
3547:
3544:
3542:
3541:
3537:
3536:
3533:
3530:
3526:
3523:
3521:
3518:
3515:
3513:
3512:
3508:
3507:
3503:
3500:
3497:
3494:
3491:
3488:
3485:
3484:
3481:
3477:
3473:
3465:
3458:
3454:
3451:
3448:
3446:
3443:
3441:
3437:
3433:
3430:
3429:
3424:
3423:
3419:
3415:
3412:
3409:
3407:
3404:
3402:
3399:
3397:
3396:
3392:
3391:
3387:
3384:
3381:
3378:
3375:
3372:
3369:
3368:
3365:
3360:
3352:
3347:
3341:
3339:
3335:
3333:
3331:
3329:
3327:
3324:
3322:
3321:
3317:
3316:
3313:
3311:
3307:
3305:
3303:
3301:
3299:
3298:D. reticulata
3296:
3294:
3293:
3289:
3288:
3285:
3283:
3281:
3279:
3277:
3274:
3273:
3270:
3268:
3262:
3260:
3258:
3256:
3253:
3251:
3250:
3245:
3242:
3240:
3236:
3234:
3232:
3230:
3228:
3225:
3223:
3222:
3218:
3217:
3214:
3212:
3208:
3206:
3204:
3202:
3200:
3197:
3195:
3194:
3193:Bisulcocypris
3190:
3189:
3186:
3184:
3180:
3178:
3176:
3174:
3172:
3169:
3167:
3166:
3162:
3161:
3158:
3156:
3154:
3152:
3150:
3147:
3146:
3143:
3141:
3139:
3137:
3135:
3132:
3131:
3128:
3126:
3122:
3118:
3117:
3111:
3109:
3107:
3105:
3102:
3100:
3099:
3094:
3090:
3087:
3084:
3081:
3078:
3075:
3072:
3071:
3068:
3064:
3060:
3052:
3047:
3040:
3031:
3028:
3026:
3019:
3015:
3012:
3008:
3005:
3001:
2996:
2991:
2986:
2983:
2979:
2978:
2974:
2971:
2970:
2962:
2955:
2953:
2951:
2949:
2946:
2943:
2941:
2938:
2937:
2934:
2931:
2929:
2927:
2924:
2921:
2919:
2916:
2915:
2912:
2909:
2907:
2905:
2902:
2900:
2897:
2895:
2894:
2893:Xenachoffatia
2890:
2889:
2886:
2884:
2881:
2877:
2875:
2873:
2870:
2867:
2865:
2864:
2859:
2858:
2855:
2852:
2848:
2846:
2844:
2841:
2839:
2836:
2834:
2833:
2829:
2828:
2825:
2822:
2820:
2818:
2815:
2813:
2810:
2808:
2807:
2806:Pseudobolodon
2803:
2802:
2799:
2796:
2794:
2792:
2789:
2787:
2784:
2782:
2781:
2777:
2776:
2773:
2771:
2770:triconodontid
2767:
2765:
2763:
2760:
2757:
2755:
2754:
2750:
2749:
2746:
2744:dryolestidan
2743:
2739:
2737:
2735:
2732:
2730:
2727:
2725:
2724:
2720:
2719:
2716:
2713:
2711:
2709:
2706:
2704:
2701:
2699:
2698:
2694:
2693:
2690:
2687:
2685:
2683:
2680:
2678:
2675:
2673:
2672:
2668:
2667:
2664:
2661:
2659:
2657:
2654:
2652:
2649:
2647:
2646:
2645:Paulchoffatia
2642:
2641:
2638:
2636:
2634:
2631:
2629:
2628:N. drescherae
2626:
2625:
2622:
2620:
2614:
2612:
2609:
2607:
2604:
2602:
2601:
2596:
2593:
2590:
2588:
2586:
2583:
2581:
2578:
2576:
2575:
2571:
2570:
2567:
2564:
2562:
2560:
2557:
2555:
2552:
2550:
2549:
2548:Meketibolodon
2545:
2544:
2541:
2539:dryolestidan
2538:
2534:
2532:
2530:
2527:
2525:
2522:
2520:
2519:
2515:
2514:
2511:
2509:
2506:
2502:
2500:
2498:
2495:
2493:
2490:
2488:
2487:
2483:
2482:
2479:
2477:dryolestidan
2476:
2472:
2470:
2468:
2465:
2463:
2460:
2458:
2457:
2456:Krebsotherium
2453:
2452:
2449:
2447:
2444:
2440:
2438:
2436:
2433:
2431:
2428:
2426:
2425:
2421:
2420:
2417:
2414:
2412:
2410:
2407:
2405:
2402:
2400:
2399:
2395:
2394:
2390:
2386:
2383:
2381:
2379:
2376:
2374:
2373:H. guimarotae
2371:
2369:
2368:
2364:
2363:
2360:
2358:
2355:
2351:
2349:
2347:
2344:
2342:
2339:
2337:
2336:
2332:
2331:
2328:
2326:
2322:
2320:
2318:
2315:
2313:
2310:
2308:
2307:
2303:
2302:
2299:
2297:dryolestidan
2296:
2292:
2290:
2288:
2285:
2283:
2280:
2278:
2277:
2273:
2272:
2269:
2267:
2264:
2260:
2258:
2256:
2253:
2251:
2250:G. leiriensis
2248:
2246:
2245:
2241:
2240:
2237:
2234:
2230:
2228:
2226:
2223:
2221:
2218:
2216:
2215:
2211:
2210:
2207:
2205:dryolestidan
2204:
2200:
2198:
2196:
2193:
2191:
2188:
2186:
2185:
2181:
2180:
2177:
2175:
2172:
2168:
2166:
2164:
2161:
2159:
2156:
2154:
2153:
2149:
2148:
2145:
2142:
2138:
2136:
2134:
2131:
2129:
2126:
2124:
2123:
2119:
2118:
2115:
2113:
2110:
2106:
2104:
2102:
2099:
2097:
2094:
2092:
2091:
2087:
2086:
2082:
2079:
2076:
2073:
2070:
2067:
2064:
2063:
2060:
2058:
2053:
2047:
2038:
2034:
2032:
2031:machimosaurid
2028:
2026:
2024:
2021:
2019:
2016:
2014:
2013:
2012:Machimosaurus
2009:
2008:
2005:
2003:
2001:
1999:
1996:
1994:
1991:
1989:
1988:
1984:
1983:
1977:
1973:
1971:
1967:
1965:
1963:
1960:
1958:
1957:K. guimarotae
1955:
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1948:
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1941:
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1934:goniopholidid
1931:
1929:
1927:
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1764:Portugalophis
1761:
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1704:Dorsetisaurus
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986:
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977:
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962:Choristoderes
959:
954:Choristoderes
953:
947:
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937:
934:
932:
929:
928:
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922:
920:
918:
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908:Marmorerpeton
904:
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895:
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836:
832:
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819:
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804:Pachycormidae
802:
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788:
785:
783:
781:
777:
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773:
771:
768:
766:
763:
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757:Macrosemiidae
755:
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744:
741:
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733:
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723:
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556:
553:
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548:
547:
543:
542:
538:
534:
532:
529:
527:
524:
522:
521:A. biformatus
519:
517:
516:
515:Asteracanthus
512:
511:
508:
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501:
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471:
468:
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448:
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444:
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432:
430:
428:
427:Late Jurassic
425:stage of the
424:
420:
416:
412:
408:
404:
378:
369:
365:
361:
358:
355:
351:
348:
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73:
68:
63:
58:
52:
44:
40:
36:
31:
19:
4091:, retrieved
4076:
4068:Bibliography
4038:
4034:
4024:
4015:
4006:
3965:
3961:
3954:
3929:
3925:
3754:. Retrieved
3750:
3741:
3729:. Retrieved
3725:
3716:
3689:
3685:
3675:
3641:(1): 75–97.
3638:
3634:
3624:
3569:
3562:
3539:
3509:
3479:
3440:alcobacensis
3439:
3435:
3426:
3400:
3393:
3363:
3338:Darwinulacea
3325:
3318:
3297:
3290:
3275:
3255:P. bicostata
3254:
3247:
3226:
3221:Timiriasevia
3219:
3198:
3191:
3170:
3163:
3148:
3133:
3114:
3103:
3096:
3066:
3038:
3029:
2972:
2898:
2891:
2883:symmetrodont
2860:
2837:
2830:
2811:
2804:
2785:
2778:
2751:
2728:
2721:
2702:
2695:
2676:
2669:
2650:
2643:
2627:
2619:cladotherian
2605:
2598:
2579:
2572:
2553:
2546:
2523:
2516:
2491:
2484:
2461:
2454:
2429:
2422:
2403:
2396:
2372:
2365:
2340:
2333:
2311:
2304:
2281:
2276:Guimarotodus
2274:
2249:
2244:Guimarotodon
2242:
2219:
2212:
2190:D. leirensis
2189:
2182:
2174:dryolestidan
2157:
2150:
2127:
2120:
2095:
2088:
2055:
2017:
2010:
1992:
1987:Lisboasaurus
1985:
1956:
1949:
1920:
1913:
1880:
1844:
1824:
1817:
1798:
1791:
1769:
1762:
1731:
1709:
1702:
1680:
1673:
1640:
1587:
1569:
1562:
1529:
1469:
1438:
1404:
1399:Megalosaurus
1397:
1379:
1372:
1346:
1339:
1311:Ceratosaurus
1309:
1282:
1275:
1258:A. jurassica
1257:
1252:Aviatyrannis
1250:
1218:
1187:Saurischians
1185:
1178:
1175:Saurischians
1154:
1147:
1122:
1115:
1067:
1060:
1027:
1020:
992:
906:
879:
829:
778:
735:Ionoscopidae
707:
678:
656:Macromesodon
654:
624:Osteichthyes
614:Osteichthyes
544:
520:
513:
489:
445:, Portugal.
436:
423:Kimmeridgian
410:
406:
402:
400:
363:Year defined
338:Type section
276:33.7°N 1.6°E
237:39.6°N 9.0°W
43:Kimmeridgian
3722:"Guimarota"
3658:10316/79879
3564:Melampoides
3292:Dicrorygima
3039:crossed out
3018:Morphotaxon
2899:X. oinopion
2838:R. amalthea
2671:Pinheirodon
2651:P. delgadoi
2554:M. robustus
2282:G. inflatus
2214:Ecprepaulax
2171:paurodontid
2122:Bernardodon
1970:atoposaurid
1915:Goniopholis
1793:Saurillodon
1770:P. lignites
1589:Platychelys
1564:Peltochelys
1405:M. insignis
1155:T. cuneatus
1149:Trimucrodon
1117:Stegosaurus
1095:Ornithopoda
1062:Dacentrurus
994:Cteniogenys
835:salamanders
680:Proscinetes
591:Batomorphii
546:Polyacrodus
296:Estremadura
279: /
240: /
215:Coordinates
4104:Categories
4093:2019-02-21
3616:References
3574:Guimarota
3548:Guimarota
3540:Cryptaulax
3519:Guimarota
3511:Teinostoma
3476:Gastropods
3466:Gastropods
3444:Guimarota
3405:Guimarota
3401:I. rugosus
3310:Cytheracea
3276:P. clivosa
3267:Cytheracea
3239:Cytheracea
3211:Cytheracea
3183:Cytheracea
3149:C. striata
3134:C. inermis
3125:Cypridacea
3035:small text
3004:Ichnotaxon
2925:Guimarota
2903:Guimarota
2880:tinodontid
2842:Guimarota
2816:Guimarota
2790:Guimarota
2742:dryolestid
2707:Guimarota
2655:Guimarota
2632:Guimarota
2606:N. krusati
2600:Nanolestes
2584:Guimarota
2558:Guimarota
2537:dryolestid
2524:L. andresi
2496:Guimarota
2486:Kuehneodon
2475:dryolestid
2466:Guimarota
2434:Guimarota
2430:K. hopsoni
2424:Kielanodon
2377:Guimarota
2345:Guimarota
2335:Henkelodon
2316:Guimarota
2306:Haldanodon
2295:dryolestid
2286:Guimarota
2254:Guimarota
2220:E. anomala
2203:dryolestid
2194:Guimarota
2184:Dryolestes
2162:Guimarota
2100:Guimarota
2022:Guimarota
1997:Guimarota
1961:Guimarota
1925:Guimarota
1829:Guimarota
1803:Guimarota
1774:Guimarota
1741:Guimarota
1733:Marmoretta
1722:Anguimorph
1714:Guimarota
1685:Guimarota
1675:Becklesius
1618:Guimarota
1597:Guimarota
1574:Guimarota
1506:Guimarota
1479:Guimarota
1435:Pterosaurs
1425:Pterosaurs
1262:Guimarota
1220:Allosaurus
1068:D. armatus
1002:Guimarota
938:Guimarota
916:Guimarota
848:Amphibians
826:Amphibians
811:Guimarota
789:Guimarota
764:Guimarota
742:Guimarota
719:Guimarota
688:Guimarota
664:Guimarota
598:Guimarota
576:Guimarota
554:Guimarota
525:Guimarota
499:Guimarota
242:39.6; -9.0
4057:1698-6180
3946:0016-7568
3756:31 August
3731:31 August
3708:1867-1608
3667:2183-4431
3498:Material
3492:Location
3395:Isognomon
3382:Material
3376:Location
3320:Darwinula
3116:Cetacella
3104:C. armata
3098:Cetacella
3085:Material
3079:Location
3063:Ostracods
3053:Ostracods
2973:Color key
2832:Renatodon
2753:Priacodon
2518:Laolestes
2158:D. acutum
2029:A marine
1846:Opisthias
1836:A lizard
1819:Saurillus
1810:A lizard
1457:Material
1451:Location
1277:Lusotitan
1207:Abundance
1049:Abundance
881:Celtedens
710:Lepidotes
643:Material
637:Location
478:Material
472:Location
415:formation
343:Named for
281:33.7; 1.6
203:sandstone
199:limestone
195:siltstone
168:Lithology
148:Underlies
138:Sub-units
18:Guimarota
3998:10930309
3990:17187254
3599:See also
3588:Melampus
3489:Species
3373:Species
3353:Bivalves
3348:Mollusca
3076:Species
2398:Iberodon
2325:docodont
2096:B. hapax
2077:Material
2071:Location
2018:M. hugii
1902:Material
1896:Location
1662:Material
1656:Location
1551:Material
1545:Location
1448:Species
1201:District
1043:District
981:Material
975:Location
867:Material
861:Location
634:Species
469:Species
419:Portugal
353:Named by
322:Portugal
209:Location
187:Mudstone
158:Overlies
3970:Bibcode
3579:Shells
3553:Shells
3524:Shells
3504:Images
3478:of the
3438:." cf.
3388:Images
3091:Images
3065:of the
3011:Ootaxon
2863:Tinodon
2083:Images
2068:Species
2057:Mammals
2048:Mammals
1908:Images
1893:Species
1724:lizard
1695:lizard
1668:Images
1653:Species
1557:Images
1542:Species
1531:Turtles
1522:Turtles
1463:Images
1437:of the
1409:Lisboa
1384:Lisboa
1351:Lisboa
1319:Lisboa
1213:Images
1198:Species
1159:Lisboa
1102:Lisboa
1055:Images
1040:Species
987:Images
972:Species
873:Images
858:Species
780:Caturus
649:Images
491:Hybodus
484:Images
357:Choffat
310:Country
264:33°42′N
225:39°36′N
173:Primary
118:↓
4084:
4055:
3996:
3988:
3944:
3706:
3665:
3501:Notes
3486:Genus
3385:Notes
3370:Genus
3249:Poisia
3088:Notes
3073:Genus
1511:Teeth
1460:Notes
1445:Genus
1288:Lisboa
1229:Leiria
1073:Lisboa
646:Notes
631:Genus
481:Notes
466:Genus
328:Extent
319:
304:Lisboa
300:Leiria
292:Region
267:1°36′E
228:9°00′W
3994:S2CID
3030:Notes
2982:Taxon
2080:Notes
2065:Genus
1905:Notes
1890:Genus
1783:snake
1781:Stem-
1665:Notes
1650:Genus
1554:Notes
1539:Genus
1468:aff.
1210:Notes
1195:Genus
1052:Notes
1037:Genus
984:Notes
969:Genus
870:Notes
855:Genus
839:frogs
183:Other
4082:ISBN
4053:ISSN
3986:PMID
3942:ISSN
3758:2018
3733:2018
3704:ISSN
3663:ISSN
3428:Unio
1843:cf.
905:cf.
878:cf.
837:and
401:The
366:1885
191:coal
177:Marl
128:Type
57:PreꞒ
4043:doi
3978:doi
3934:doi
3930:138
3694:doi
3690:102
3653:hdl
3643:doi
1968:An
417:in
4106::
4051:.
4039:36
4037:.
4033:.
3992:.
3984:.
3976:.
3966:94
3964:.
3940:.
3928:.
3924:.
3848:^
3832:^
3766:^
3749:.
3724:.
3702:.
3688:.
3684:.
3661:.
3651:.
3639:19
3637:.
3633:.
3431:"
3037:;
2878:A
2861:?
2849:A
2768:A
2740:A
2617:A
2535:A
2503:A
2473:A
2441:A
2352:A
2323:A
2293:A
2261:A
2231:A
2201:A
2169:A
2139:A
2107:A
1932:A
1861:A
302:,
298:,
201:,
197:,
193:,
189:,
107:Pg
51:Ma
41::
4059:.
4045::
4000:.
3980::
3972::
3948:.
3936::
3760:.
3735:.
3710:.
3696::
3669:.
3655::
3645::
3436:U
3434:"
3425:"
782:?
712:"
708:"
112:N
102:K
97:J
92:T
87:P
82:C
77:D
72:S
67:O
62:Ꞓ
46:~
20:)
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.