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Glyphoglossus smithi

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is present but hidden under skin. The finger tips are not expanded but the toe tips are expanded into small discs. The toes are about one-fourth webbed. Skin is smooth. Colouration is dark brown above with dark markings edged with pink or red. A conspicuous red or pink arc may be present above the
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occurs in lowland rainforests. It is a secretive, burrowing species that is rarely observed. The eggs are probably laid in pools of standing water on the forest floor. It is threatened by
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in Sarawak has experienced severe habitat loss and the species might no longer persist there. The Sabah locality is, however, within the
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honours Dr Harrison Willard Smith, an American scientist who collected the
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Amphibian Species of the World: an Online Reference. Version 6.0
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Beolens, Bo; Watkins, Michael & Grayson, Michael (2013).
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Adult females measure up to 39 mm (1.5 in) in
825: 693: 546: 438: 436: 434: 443:Haas, A.; Das, I. & Hertwig, S.T. (2019). 379:10.2305/IUCN.UK.2018-1.RLTS.T57726A95676801.en 8: 354:IUCN SSC Amphibian Specialist Group (2018). 534: 41: 20: 522: 377: 402: 400: 398: 396: 333: 905:IUCN Red List near threatened species 199: 7: 940:Taxa named by Gladwyn Kingsley Noble 419:. American Museum of Natural History 484:. Pelagic Publishing. p. 200. 481:The Eponym Dictionary of Amphibians 365:IUCN Red List of Threatened Species 14: 66: 323:Danum Valley Conservation Area 1: 317:caused by clear-cutting. The 935:Taxa named by Thomas Barbour 930:Amphibians described in 1916 213:(Barbour and Noble, 1916) 325:, which is well protected. 961: 413:(Barbour and Noble, 1916)" 221:(Barbour and Noble, 1916) 920:Endemic fauna of Malaysia 449:(Smith's Burrowing Frog)" 407:Frost, Darrel R. (2019). 205:Barbour and Noble, 1916 198: 191: 168: 161: 63:Scientific classification 61: 39: 30: 23: 306:Habitat and conservation 925:Endemic fauna of Borneo 915:Amphibians of Malaysia 524:10.5962/bhl.title.3147 241:Smith's burrowing frog 505:Inger, R. F. (1966). 372:: e.T57726A95676801. 945:Amphibians of Borneo 737:Glyphoglossus-smithi 695:Glyphoglossus smithi 578:Glyphoglossus smithi 459:on 21 September 2019 447:Glyphoglossus smithi 411:Glyphoglossus smithi 358:Glyphoglossus smithi 311:Glyphoglossus smithi 232:Glyphoglossus smithi 172:Glyphoglossus smithi 25:Glyphoglossus smithi 210:Colpoglossus smithi 33:Conservation status 827:Calliglutus smithi 243:, is a species of 237:Smith's squat frog 202:Calliglutus smithi 892: 891: 678:Open Tree of Life 540:Taxon identifiers 511:Fieldiana Zoology 491:978-1-907807-42-8 295:snout–vent length 228: 227: 222: 214: 206: 56: 952: 885: 884: 872: 871: 859: 858: 846: 845: 844: 818: 817: 805: 804: 792: 791: 779: 778: 766: 765: 753: 752: 740: 739: 727: 726: 714: 713: 712: 686: 685: 673: 672: 660: 659: 647: 646: 634: 633: 621: 620: 608: 607: 595: 594: 592:Calluella_smithi 582: 581: 580: 567: 566: 565: 548:Calluella smithi 535: 529: 528: 526: 502: 496: 495: 475: 469: 468: 466: 464: 455:. Archived from 440: 429: 428: 426: 424: 404: 391: 390: 388: 386: 381: 351: 235:, also known as 220: 218:Calluella smithi 212: 204: 174: 71: 70: 50: 45: 44: 21: 960: 959: 955: 954: 953: 951: 950: 949: 895: 894: 893: 888: 880: 875: 867: 862: 854: 849: 840: 839: 834: 821: 813: 810:Observation.org 808: 800: 795: 787: 782: 774: 769: 761: 756: 748: 743: 735: 730: 722: 717: 708: 707: 702: 689: 681: 676: 668: 665:Observation.org 663: 655: 650: 642: 637: 629: 624: 616: 611: 603: 598: 590: 585: 576: 575: 570: 561: 560: 555: 542: 532: 504: 503: 499: 492: 477: 476: 472: 462: 460: 453:Frogs of Borneo 442: 441: 432: 422: 420: 406: 405: 394: 384: 382: 353: 352: 335: 331: 308: 291: 187: 176: 170: 157: 65: 57: 48:Near Threatened 46: 42: 35: 17: 16:Species of frog 12: 11: 5: 958: 956: 948: 947: 942: 937: 932: 927: 922: 917: 912: 907: 897: 896: 890: 889: 887: 886: 873: 860: 847: 831: 829: 823: 822: 820: 819: 806: 793: 780: 767: 754: 741: 728: 715: 699: 697: 691: 690: 688: 687: 674: 661: 648: 635: 622: 609: 596: 583: 568: 552: 550: 544: 543: 538: 531: 530: 497: 490: 470: 430: 392: 332: 330: 327: 307: 304: 290: 287: 247:in the family 226: 225: 224: 223: 215: 207: 196: 195: 189: 188: 177: 166: 165: 159: 158: 154:G. smithi 151: 149: 145: 144: 137: 133: 132: 127: 123: 122: 117: 113: 112: 107: 103: 102: 97: 93: 92: 87: 83: 82: 77: 73: 72: 59: 58: 40: 37: 36: 31: 28: 27: 15: 13: 10: 9: 6: 4: 3: 2: 957: 946: 943: 941: 938: 936: 933: 931: 928: 926: 923: 921: 918: 916: 913: 911: 910:Glyphoglossus 908: 906: 903: 902: 900: 883: 878: 874: 870: 865: 861: 857: 852: 848: 843: 837: 833: 832: 830: 828: 824: 816: 811: 807: 803: 798: 794: 790: 785: 781: 777: 772: 768: 764: 759: 755: 751: 746: 742: 738: 733: 729: 725: 720: 716: 711: 705: 701: 700: 698: 696: 692: 684: 679: 675: 671: 666: 662: 658: 653: 649: 645: 640: 636: 632: 627: 623: 619: 614: 610: 606: 601: 597: 593: 588: 584: 579: 573: 569: 564: 558: 554: 553: 551: 549: 545: 541: 536: 525: 520: 516: 512: 508: 501: 498: 493: 487: 483: 482: 474: 471: 458: 454: 450: 448: 439: 437: 435: 431: 418: 414: 412: 403: 401: 399: 397: 393: 380: 375: 371: 367: 366: 361: 359: 350: 348: 346: 344: 342: 340: 338: 334: 328: 326: 324: 320: 319:type locality 316: 312: 305: 303: 300: 296: 288: 286: 284: 283:Limbang River 280: 276: 273: 272:specific name 269: 266: 262: 258: 254: 250: 246: 242: 238: 234: 233: 219: 216: 211: 208: 203: 200: 197: 194: 190: 185: 181: 175: 173: 167: 164: 163:Binomial name 160: 156: 155: 150: 147: 146: 143: 142: 141:Glyphoglossus 138: 135: 134: 131: 128: 125: 124: 121: 118: 115: 114: 111: 108: 105: 104: 101: 98: 95: 94: 91: 88: 85: 84: 81: 78: 75: 74: 69: 64: 60: 54: 49: 38: 34: 29: 26: 22: 19: 826: 694: 547: 514: 510: 500: 480: 473: 463:21 September 461:. Retrieved 457:the original 452: 446: 423:21 September 421:. Retrieved 416: 410: 383:. Retrieved 369: 363: 357: 315:habitat loss 310: 309: 292: 274: 249:Microhylidae 240: 236: 231: 230: 229: 217: 209: 201: 171: 169: 153: 152: 140: 130:Microhylidae 24: 18: 771:iNaturalist 719:AmphibiaWeb 572:Wikispecies 385:15 November 289:Description 899:Categories 329:References 285:district. 842:Q27149056 710:Q27149073 517:: 1–402. 281:from the 265:Malaysian 148:Species: 86:Kingdom: 80:Eukaryota 869:10976855 836:Wikidata 763:10861690 704:Wikidata 644:11335202 563:Q2241496 557:Wikidata 299:tympanum 279:holotype 251:. It is 193:Synonyms 126:Family: 110:Amphibia 100:Chordata 96:Phylum: 90:Animalia 76:Domain: 53:IUCN 3.1 882:1097599 789:1097598 683:3618705 631:2422128 618:1019319 261:Sarawak 253:endemic 186:, 1916) 180:Barbour 136:Genus: 116:Order: 106:Class: 51: ( 815:955932 776:476451 670:202173 657:664378 488:  302:anus. 275:smithi 268:Borneo 802:57726 750:3GGT3 639:IRMNG 257:Sabah 184:Noble 120:Anura 877:ITIS 864:GBIF 856:PQ3P 797:IUCN 784:ITIS 758:GBIF 724:5372 652:ITIS 626:GBIF 605:PW8V 486:ISBN 465:2019 425:2019 387:2021 370:2018 259:and 245:frog 182:and 851:CoL 745:CoL 732:ASW 613:EoL 600:CoL 587:ADW 519:doi 374:doi 263:in 255:to 239:or 901:: 879:: 866:: 853:: 838:: 812:: 799:: 786:: 773:: 760:: 747:: 734:: 721:: 706:: 680:: 667:: 654:: 641:: 628:: 615:: 602:: 589:: 574:: 559:: 515:52 513:. 509:. 451:. 433:^ 415:. 395:^ 368:. 362:. 336:^ 527:. 521:: 494:. 467:. 445:" 427:. 409:" 389:. 376:: 360:" 356:" 178:( 55:)

Index

Conservation status
Near Threatened
IUCN 3.1
Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Amphibia
Anura
Microhylidae
Glyphoglossus
Binomial name
Barbour
Noble
Synonyms
frog
Microhylidae
endemic
Sabah
Sarawak
Malaysian
Borneo
specific name
holotype
Limbang River
snout–vent length
tympanum
habitat loss
type locality

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