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Grandidier's trident bat

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of western Madagascar, which have nonoverlapping ranges. Their paper reported the first new material of the species since its original description. They described a third species of this group in 2008—
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Monadjem, A.; Andriafidison, D.; Cardiff, S.G.; Goodman, S.; Hutson, A.M.; Jenkins, R.K.B.; Kofoky, A.; Racey, P.A.; Ranivo, J.; Ratrimomanarivo, F.H.; Razafimanahaka, J. (2017).
342:, who reviewed the genus in 1982. In their 1995 study of Madagascar bats, however, R.L. Peterson and colleagues reinstated it as a species. They also changed the 1431: 467:
is a large-eared, short-tailed bat. The fur is reddish gold and is darker above than below. Both males and females are significantly larger than individuals of
351: 1562: 1143: 913: 1508: 1366: 1521: 1392: 845: 506:. It is relatively abundant in its range and occurs in dry forests. It roosts in caves, and the largest colony contains an estimated 2000 bats. 874: 780: 547: 761: 1572: 834: 1418: 1567: 1539: 1526: 1397: 766:(Chiroptera: Hipposideridae) with description of a new species from southern Arabia and definitions of a new genus and tribe 1136: 906: 551: 65: 1534: 856: 1436: 355: 1331: 985: 1129: 899: 558:. The IUCN recommends that conservation efforts be focused on monitoring and protecting the known cave roosts. 806:(Chiroptera: Hipposideridae) reputed to be from Madagascar and the description of a replacement species name 503: 394:
in the nearby Seychelles. In 2007 and 2008, Amy Russell and colleagues investigated the relationships among
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Working at the interface of phylogenetics and population genetics: a biogeographical analysis of
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Coalescent analyses support multiple mainland-to-island dispersals in the evolution of Malagasy
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and their status as distinct species. In view of the significant differences between the
1246: 1189: 1086: 959: 543: 272: 1556: 1405: 1200: 1042: 1000: 860: 852: 791:(Mammalia, Chiroptera, Hipposideridae) from Aldabra Atoll, Picard Island (Seychelles) 589: 580: 515: 420: 293: 137: 50: 312:, on the basis of a single poorly preserved specimen collected at Diégo-Suarez (now 1500: 1358: 399: 1486: 772: 1423: 1379: 1325: 1153: 822:
Russell, A.L., Ranivo, J., Palkovacs, E.P., Goodman, S.M. and Yoder, A.D. 2007.
313: 1316: 1121: 1183: 965: 279: 418:, Petr Benda and Peter Villa removed the former in 2009 to a separate genus, 1236: 1066: 884: 287: 77: 797: 812: 1480: 1310: 1171: 976: 953: 947: 941: 935: 472: 97: 846:
Late Pleistocene bat fossils from Anjohibe Cave, northwestern Madagascar
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Goodman and Ranivo, 2008, p. 686; Benda and Vallo, 2009, p. 33
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has a small range in far northern Madagascar, perhaps south to the
452:, an extinct species described in 2007 that may also be related to 316:) in northernmost Madagascar. In his 1939 list of African mammals, 283: 866:
Mammal Species of the World: A Taxonomic and Geographic Reference
1291: 1125: 895: 487:, the three lancets are straight and about equal in length; in 438:), which is widespread in western Madagascar and overlaps with 574: 572: 570: 268: 117: 869:(3rd ed.). Johns Hopkins University Press. p. 378. 334:) of western Madagascar, and in his 1948 review of the genus 491:, the middle lancet is longer and the outer two are curved. 471:. On its face are the three lancets at the back of the 534:. Agricultural activities in particular are causing 1470: 1300: 1234: 1198: 1064: 1040: 998: 974: 530:" because of its small, fragmented, and declining 833:Russell, A.L., Goodman, S.M. and Cox, M.P. 2008. 798:The geographical origin of the type specimens of 817:malgaches (Mammalia, Chiroptera, Hipposideridae) 376:of northern Madagascar and the closely related 604:10.2305/IUCN.UK.2017-2.RLTS.T40025A22064746.en 402:data; they confirmed the relationship between 372:in 2006 and clarified the distinction between 1137: 907: 775:. London: A & C Black, 304 pp.  8: 808:(subscription required). Mammalia 73:47–55. 1288: 1144: 1130: 1122: 914: 900: 892: 744:Ranivo and Goodman, 2006, pp. 969–970 624:Ranivo and Goodman, 2006, pp. 964–965 291:, but is now placed in the separate genus 186: 39: 20: 602: 768:. Folia Zoologica 58 (Monograph 1):1–45. 308:first described the species in 1912, as 773:Mammals of Madagascar: A Complete Guide 681:Russell et al., 2007, fig. 2; 2008 641: 639: 566: 841:. Journal of Biogeography 35:995–1003. 735:Goodman and Ranivo, 2008, p. 688 672:Goodman and Ranivo, 2008, p. 681 663:Ranivo and Goodman, 2006, p. 965 645:Ranivo and Goodman, 2006, p. 964 7: 428:still includes the Malagasy species 848:. Acta Chiropterologica 9(1):39–65. 796:Goodman, S.M. and Ranivo, J. 2009. 785:Goodman, S.M. and Ranivo, J. 2008. 590:IUCN Red List of Threatened Species 811:Ranivo, J. and Goodman S.M. 2006. 717:Russell et al., 2008, p. 1001 282:. It was formerly assigned to the 14: 1028:Paulian's triple leaf-nosed bat ( 839:bats (Chiroptera: Hipposideridae) 828:spp. (Chiroptera: Hipposideridae) 699:Russell et al., 2007, fig. 1 690:Benda and Villa, 2009, p. 34 1563:IUCN Red List vulnerable species 762:Taxonomic revision of the genus 64: 830:. Molecular Ecology 16:839–851. 338:, Jean Dorst concurred, as did 760:Benda, P. and Vallo, P. 2009. 552:RĂ©serve SpĂ©ciale d'Analamerana 1: 24:Grandidier's trident bat 819:. Zoosystema 28(4):963–985. 793:. Zoosystema 30(3):681–693. 548:RĂ©serve SpĂ©ciale d'Ankarana 475:that are characteristic of 414:group and other species of 1594: 1019:Trouessart's trident bat ( 1010:Grandidier's trident bat ( 654:Simmons, 2005, p. 378 1573:Mammals described in 1912 1160: 930: 922:Extant species of family 813:RĂ©vision taxinomique des 726:Garbutt, 2007, p. 72 708:Samonds, 2007, p. 46 224: 217: 194: 185: 166: 159: 61:Scientific classification 59: 37: 28: 23: 986:Percival's trident bat ( 495:Distribution and ecology 320:placed the species as a 259:Grandidier's trident bat 1052:Orange leaf-nosed bat ( 633:Allen, 1939, p. 82 863:; Reeder, D.M (eds.). 504:Andrafiamena Mountains 387:Paratriaenops pauliani 1568:Mammals of Madagascar 1332:Paratriaenops auritus 1302:Paratriaenops auritus 1107:Persian trident bat ( 597:: e.T40025A22064746. 583:Paratriaenops auritus 465:Paratriaenops auritus 331:Paratriaenops furcula 264:Paratriaenops auritus 249:Benda and Vallo, 2009 245:Paratriaenops auritus 240:Peterson et al., 1995 170:Paratriaenops auritus 844:Samonds, K.E. 2007. 318:Glover Morrill Allen 306:Guillaume Grandidier 536:habitat destruction 510:Conservation status 368:revised Madagascar 31:Conservation status 857:"Order Chiroptera" 771:Garbutt, N. 2007. 554:, and a forest at 449:Triaenops goodmani 431:Triaenops menamena 382:Triaenops pauliani 267:) is a species of 1550: 1549: 1472:Triaenops auritus 1445:Open Tree of Life 1346:Triaenops_auritus 1294:Taxon identifiers 1285: 1284: 1152:Species of tribe 1119: 1118: 1079: 876:978-0-8018-8221-0 787:A new species of 781:978-0-7136-7043-1 524:Triaenops auritus 364:Julie Ranivo and 350:, presumably for 340:John Edwards Hill 326:Triaenops furcula 256: 255: 250: 241: 236:Triaenops auritus 232: 54: 1585: 1543: 1542: 1530: 1529: 1517: 1516: 1504: 1503: 1491: 1490: 1489: 1463: 1462: 1453: 1452: 1440: 1439: 1427: 1426: 1414: 1413: 1401: 1400: 1388: 1387: 1375: 1374: 1362: 1361: 1349: 1348: 1336: 1335: 1334: 1321: 1320: 1319: 1289: 1146: 1139: 1132: 1123: 1078: 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649: 644: 637: 632: 628: 623: 619: 609: 607: 578: 577: 568: 564: 544:protected areas 512: 497: 485:P. furcula 469:P. furcula 462: 444:P. furcula 440:P. auritus 436:Triaenops rufus 404:T. auritus 374:T. auritus 303: 181: 174: 168: 155: 152:P. auritus 128:Rhinonycteridae 63: 55: 44: 40: 33: 17: 12: 11: 5: 1591: 1589: 1581: 1580: 1575: 1570: 1565: 1555: 1554: 1548: 1547: 1545: 1544: 1531: 1518: 1505: 1492: 1476: 1474: 1468: 1467: 1465: 1464: 1454: 1441: 1428: 1415: 1402: 1389: 1376: 1363: 1350: 1337: 1322: 1306: 1304: 1298: 1297: 1292: 1283: 1282: 1280: 1279: 1272: 1265: 1258: 1250: 1242: 1240: 1232: 1231: 1229: 1228: 1221: 1214: 1206: 1204: 1196: 1195: 1193: 1192: 1190:Hipposideridae 1186: 1180: 1174: 1168: 1161: 1158: 1157: 1151: 1149: 1148: 1141: 1134: 1126: 1117: 1116: 1114: 1113: 1104: 1097: 1090: 1082: 1080: 1062: 1061: 1059: 1058: 1048: 1046: 1038: 1037: 1035: 1034: 1025: 1016: 1006: 1004: 996: 995: 993: 992: 982: 980: 972: 971: 969: 968: 962: 960:Laurasiatheria 956: 950: 944: 938: 931: 928: 927: 921: 919: 918: 911: 904: 896: 890: 889: 875: 849: 842: 831: 820: 809: 794: 783: 769: 756: 755: 746: 737: 728: 719: 710: 701: 692: 683: 674: 665: 656: 647: 635: 626: 617: 565: 563: 560: 511: 508: 496: 493: 461: 458: 446:in range, and 398:species using 366:Steven Goodman 302: 299: 273:Hipposideridae 271:in the family 254: 253: 252: 251: 242: 233: 222: 221: 215: 214: 192: 191: 183: 182: 175: 164: 163: 157: 156: 149: 147: 143: 142: 135: 131: 130: 125: 121: 120: 115: 111: 110: 105: 101: 100: 95: 91: 90: 85: 81: 80: 75: 71: 70: 57: 56: 38: 35: 34: 29: 26: 25: 16:Species of bat 15: 13: 10: 9: 6: 4: 3: 2: 1590: 1579: 1578:Paratriaenops 1576: 1574: 1571: 1569: 1566: 1564: 1561: 1560: 1558: 1541: 1536: 1532: 1528: 1523: 1519: 1515: 1510: 1506: 1502: 1497: 1493: 1488: 1482: 1478: 1477: 1475: 1473: 1469: 1461: 1455: 1451: 1446: 1442: 1438: 1433: 1429: 1425: 1420: 1416: 1412: 1407: 1403: 1399: 1394: 1390: 1386: 1381: 1377: 1373: 1368: 1364: 1360: 1355: 1351: 1347: 1342: 1338: 1333: 1327: 1323: 1318: 1312: 1308: 1307: 1305: 1303: 1299: 1295: 1290: 1278: 1277: 1273: 1271: 1270: 1266: 1264: 1263: 1259: 1257: 1256: 1251: 1249: 1248: 1244: 1243: 1241: 1239: 1238: 1233: 1227: 1226: 1222: 1220: 1219: 1215: 1213: 1212: 1208: 1207: 1205: 1203: 1202: 1201:Paratriaenops 1197: 1191: 1187: 1185: 1181: 1179: 1175: 1173: 1169: 1167: 1163: 1162: 1159: 1155: 1147: 1142: 1140: 1135: 1133: 1128: 1127: 1124: 1112: 1110: 1105: 1103: 1102: 1098: 1096: 1095: 1091: 1089: 1088: 1084: 1083: 1081: 1074:(triple leaf- 1069: 1068: 1063: 1057: 1055: 1050: 1049: 1047: 1045: 1044: 1043:Rhinonicteris 1039: 1033: 1031: 1026: 1024: 1022: 1017: 1015: 1013: 1008: 1007: 1005: 1003: 1002: 1001:Paratriaenops 997: 991: 989: 984: 983: 981: 979: 978: 973: 967: 963: 961: 957: 955: 951: 949: 945: 943: 939: 937: 933: 932: 929: 925: 917: 912: 910: 905: 903: 898: 897: 894: 886: 882: 878: 872: 868: 867: 862: 858: 854: 853:Simmons, N.B. 850: 847: 843: 840: 838: 832: 829: 827: 821: 818: 816: 810: 807: 805: 801: 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139:Paratriaenops 136: 133: 132: 129: 126: 123: 122: 119: 116: 113: 112: 109: 106: 103: 102: 99: 96: 93: 92: 89: 86: 83: 82: 79: 76: 73: 72: 67: 62: 58: 52: 47: 36: 32: 27: 22: 19: 1471: 1301: 1274: 1267: 1260: 1253: 1245: 1235: 1223: 1216: 1210: 1209: 1199: 1108: 1099: 1092: 1085: 1065: 1053: 1041: 1029: 1020: 1011: 1009: 999: 988:C. percivali 987: 975: 958:Superorder: 952:Infraclass: 865: 861:Wilson, D.E. 836: 825: 814: 803: 799: 788: 763: 749: 740: 731: 722: 713: 704: 695: 686: 677: 668: 659: 650: 629: 620: 608:. Retrieved 594: 588: 582: 523: 519: 513: 499: 498: 488: 484: 480: 476: 468: 464: 463: 453: 447: 443: 439: 435: 429: 425: 419: 415: 411: 407: 403: 400:DNA sequence 395: 385: 381: 377: 373: 369: 363: 358: 356:generic name 347: 335: 329: 325: 309: 304: 292: 286: 263: 262: 258: 257: 244: 235: 227: 208: 202: 197: 196: 169: 167: 151: 150: 138: 18: 1380:iNaturalist 1326:Wikispecies 1276:T. persicus 1262:T. menamena 1255:T. goodmani 1225:P. pauliani 1218:P. furculua 1154:Triaenopini 1109:T. persicus 1094:T. menamena 1076:nosed bats) 1054:R. aurantia 1030:P. pauliani 804:T. humbloti 610:12 November 460:Description 314:Antsiranana 210:P. pauliani 1557:Categories 1211:P. auritus 1184:Chiroptera 1021:P. furcula 1012:P. auritus 966:Chiroptera 562:References 528:Vulnerable 520:P. auritus 500:P. auritus 434:(formerly 280:Madagascar 207:(red) and 204:P. furcula 198:P. auritus 178:Grandidier 118:Chiroptera 46:Vulnerable 1487:Q27440604 1269:T. parvus 1237:Triaenops 1164:Kingdom: 1101:T. parvus 1067:Triaenops 934:Kingdom: 837:Triaenops 826:Triaenops 815:Triaenops 789:Triaenops 764:Triaenops 489:Triaenops 477:Triaenops 426:Triaenops 416:Triaenops 396:Triaenops 370:Triaenops 359:Triaenops 354:with the 336:Triaenops 288:Triaenops 146:Species: 84:Kingdom: 78:Eukaryota 1540:13800854 1481:Wikidata 1317:Q1833289 1311:Wikidata 1188:Family: 1178:Mammalia 1172:Chordata 1170:Phylum: 1166:Animalia 977:Cloeotis 954:Eutheria 948:Mammalia 942:Chordata 940:Phylum: 936:Animalia 885:62265494 855:(2005). 483:. As in 473:noseleaf 301:Taxonomy 219:Synonyms 213:(green) 201:(blue), 124:Family: 108:Mammalia 98:Chordata 94:Phylum: 88:Animalia 74:Domain: 51:IUCN 3.1 1514:4265487 1460:7140078 1450:1071075 1424:1004762 1372:7568662 1247:T. afer 1182:Order: 1176:Class: 1087:T. afer 964:Order: 946:Class: 556:Daraina 392:Aldabra 390:) from 348:auritus 322:synonym 276:endemic 180:, 1912) 134:Genus: 114:Order: 104:Class: 49: ( 1527:946029 1457:uBio: 1437:329872 1398:947352 1385:154715 883:  873:  779:  526:) as " 518:lists 1501:5856T 1411:40025 1359:4DGHH 859:. In 532:range 384:(now 328:(now 284:genus 1522:ITIS 1509:GBIF 1432:NCBI 1406:IUCN 1393:ITIS 1367:GBIF 881:OCLC 871:ISBN 802:and 777:ISBN 612:2021 595:2017 538:and 522:(as 514:The 479:and 442:and 406:and 1535:MSW 1496:CoL 1419:MDD 1354:CoL 1341:ADW 599:doi 346:to 324:of 278:to 269:bat 1559:: 1537:: 1524:: 1511:: 1498:: 1483:: 1447:: 1434:: 1421:: 1408:: 1395:: 1382:: 1369:: 1356:: 1343:: 1328:: 1313:: 879:. 638:^ 593:. 587:. 569:^ 550:, 456:. 424:. 361:. 297:. 247:: 238:: 1252:† 1145:e 1138:t 1131:v 1111:) 1056:) 1032:) 1023:) 1014:) 990:) 915:e 908:t 901:v 887:. 614:. 601:: 585:" 581:" 546:— 261:( 176:( 53:)

Index

Conservation status
Vulnerable
IUCN 3.1
Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Mammalia
Chiroptera
Rhinonycteridae
Paratriaenops
Binomial name
Grandidier
Map.
P. furcula
P. pauliani
Synonyms
bat
Hipposideridae
endemic
Madagascar
genus
Triaenops
Paratriaenops
Guillaume Grandidier
Antsiranana
Glover Morrill Allen
synonym
Paratriaenops furcula

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