2339:
325:
113:
2345:
2895:
131:
2925:
507:
2901:
1070:
904:
was unearthed from the
Sheeregeen Gashoon beds, which are part of the Upper Bayan Shireh. The presence of caliche, fluvial and lacustrine sediments, indicate a semiarid climate with rivers and large lakes around the zone. Fossilized fruits have also been recovered from the upper and lower parts of
528:
are not fused, which indicates that this specimen was not fully grown when it died, probably a juvenile individual. Its size is estimated at 60 cm (2.0 ft) long with a weight between 2.27 to 9.1 kg (5.0 to 20.1 lb). However, due to the immature nature of the specimen, the adult
543:
is recognised by the fragile frill and characteristic tibial-femoral ratio (1.2:1); the frill is also briefly elongated with well developed squamosal processes. Seven sacral vertebrae were identified and not fused. The scapula is very gracile in constitution but thicker at the
1420:
Kurumada, Y.; Aoki, S.; Aoki, K.; Kato, D.; Saneyoshi, M.; Tsogtbaatar, K.; Windley, B. F.; Ishigaki, S. (2020). "Calcite U–Pb age of the
Cretaceous vertebrate-bearing Bayn Shire Formation in the Eastern Gobi Desert of Mongolia: usefulness of caliche for age determination".
355:. The discoveries were made during field exploration by the Polish-Mongolian Palaeontological Expedition, in 1971. Four years later, in 1975, the specimen was described by
3018:
548:, with a relatively large coracoid; the humerus is also very slender. The femur measures 9.5 cm (95 mm), it is lightly curved and has a large head; the
1329:
Sochava, A. V. (1975). "Stratigraphy and lithology of the Upper
Cretaceous sediments in southern Mongolia. In Stratigraphy of Mesozoic sediments of Mongolia".
1566:
3072:
552:
is fragile and place above the midlength of the femoral end. Being larger than the femur, the tibia measures 11 cm (110 mm) and its proximal
3005:
3092:
3087:
1383:"Paleomagnetic and palynologic analyses of Albian to Santonian strata at Bayn Shireh, Burkhant, and Khuren Dukh, eastern Gobi Desert, Mongolia"
2338:
1660:
3097:
2344:
324:
2918:
1352:
1083:
130:
3102:
3082:
2894:
1183:
Sereno, P. C. (2000). "The fossil record, systematics and evolution of pachycephalosaurs and ceratopsians from Asia".
535:. The frill has large fenestrae bounded by very slender struts. This structure is very similar to that of the later
1226:"A Ceratopsian Dinosaur from the Lower Cretaceous of Western North America, and the Biogeography of Neoceratopsia"
1534:"The Description and Phylogenetic Position of a New Nanhsiungchelyid Turtle from the Late Cretaceous of Mongolia"
1184:
1589:
1751:
1653:
1367:
1143:
Mateus, O. (2008). "Two ornithischian dinosaurs renamed: Microceratops Bohlin 1953 and
Diceratops Lull 1905".
920:
371:
noted that the referral for this specimen was injustified and overall, the genus lacked diagnosis, therefore,
112:
1869:
1209:
556:
is more developed than distally. The right pes is virtually complete, only lacking the distal end of the IV
491:
2945:
924:
344:
1286:
Jerzykiewicz, T.; Russell, D. A. (1991). "Late
Mesozoic stratigraphy and vertebrates of the Gobi Basin".
3077:
3044:
2928:
1778:
2900:
2992:
2983:
1486:
1430:
1394:
1295:
1237:
913:
2034:
1646:
1475:"A Review of Shamosuchus and Paralligator (Crocodyliformes, Neosuchia) from the Cretaceous of Asia"
360:
356:
1038:
2754:
2704:
2136:
1581:
1160:
1118:
401:
397:
125:
3049:
1607:
3031:
3023:
2837:
2775:
2738:
2212:
2099:
1567:"Correlation of Late Cretaceous continental vertebrate assemblages in Middle and Central Asia"
1514:
1311:
1265:
1212:
932:
630:
586:
was found to be a basal neoceratopsian. Below are the results obtained for the
Neoceratopsia:
549:
453:
3036:
2867:
2665:
2490:
2192:
1887:
1622:
1545:
1504:
1494:
1438:
1402:
1363:
1303:
1255:
1245:
1152:
1075:
973:
2761:
2713:
2672:
2608:
2511:
2462:
2164:
2043:
1917:
1903:
1862:
1736:
1720:
313:
30:
1490:
1434:
1398:
1299:
1241:
1058:
have been reported from at least two locations, compromising mainly cervical vertebrae.
2768:
2727:
2658:
2645:
2616:
2601:
2571:
2532:
2518:
2435:
2396:
2275:
2185:
2150:
2143:
2023:
1973:
1896:
1876:
1509:
1474:
1260:
1225:
948:
810:
724:
487:
478:
207:
516:
Although very damaged, the skull measures approximately 20 cm (200 mm), the
3066:
2846:
2825:
2795:
2786:
2686:
2580:
2525:
2504:
2476:
2469:
2455:
2448:
2254:
2243:
2236:
2178:
2171:
2157:
2115:
2050:
2016:
1987:
1940:
1457:
1307:
1205:
Dinosaurs: The Most
Complete, Up-to-Date Encyclopedia for Dinosaur Lovers of All Ages
952:
939:
685:
645:
531:
449:
421:
239:
70:
1585:
1331:
Transactions of Joint Soviet–Mongolian
Scientific Research and Geological Expedition
1164:
2860:
2811:
2804:
2720:
2693:
2679:
2587:
2560:
2553:
2546:
2497:
2483:
2375:
2362:
2293:
2282:
2267:
2229:
2221:
2203:
2108:
2074:
2057:
2009:
2002:
1958:
1763:
1711:
1089:
1032:
1022:
982:
963:
958:
943:
827:
760:
738:
700:
662:
594:
433:
429:
382:
377:
265:
191:
1353:"Erketu ellisoni, a long-necked sauropod from Bor Guvé (Dornogov Aimag, Mongolia)"
506:
385:. The referred specimen was redescribed by him, creating a new genus and species:
1499:
1250:
2977:
2853:
2747:
2630:
2594:
2417:
2129:
1994:
1052:
1030:. Additional paleofauna has been recovered, expanding the aquatic biodiversity:
1008:
990:
986:
977:
968:
777:
368:
282:
45:
938:
It lived alongside other dinosaurs from the upper part, most notably the large
2623:
2539:
2389:
2122:
1910:
1844:
1831:
1790:
1669:
1065:
1055:
1026:
999:
906:
579:
557:
328:
302:
223:
90:
55:
2968:
1456:
Weishampel, D. B.; Dodson, P.; Osmolska, H. (2004). "Dinosaur
Distribution".
1315:
994:
928:
352:
305:
142:
95:
39:
1518:
1406:
1269:
1627:
1550:
1533:
2962:
2424:
2410:
2403:
1966:
1702:
1693:
1608:"The first discovery of pterosaurs from the Upper Cretaceous of Mongolia"
1013:
1003:
574:
521:
517:
510:
413:
348:
175:
162:
85:
80:
65:
60:
50:
3010:
2818:
1980:
1924:
1047:
917:
561:
545:
437:
417:
409:
100:
75:
1442:
428:; proximal and distal end of left humerus; proximal fragments of both
1687:
1156:
1017:
909:
525:
405:
152:
2939:
1382:
1224:
Farke, A. A.; Maxwell, W. D.; Cifelli, R. L.; Wedel, M. J. (2014).
1043:
553:
505:
469:
461:
445:
441:
393:
323:
298:
2997:
457:
425:
309:
120:
Skeletal diagram featuring the known elements from the holotype
2943:
2360:
1829:
1680:
1642:
1532:
Sukhanov, V. B.; Danilov, I. G.; Syromyatnikova, E. V. (2008).
1381:
Hicks, J.F.; Brinkman, D.L.; Nichols, D.J.; Watabe, M. (1999).
529:
size is estimated around 2 m (6.6 ft), or similar to
1638:
564:
are dorsoventrally flattened and somewhat sharply-developed.
392:
The holotype is fragmentary, consisting of a very fragmented
335:
fossils have been collected near
Sainshand at area D (right)
1368:
10.1206/0003-0082(2006)3508[1:EEALSF]2.0.CO;2
498:, is to emphasize the place of its discovery: Mongolia.
923:
analyses indicate that the formation lies within the
274:
2952:
2836:
2794:
2785:
2737:
2703:
2644:
2570:
2434:
2373:
2292:
2253:
2211:
2202:
2098:
2072:
2033:
1938:
1886:
1842:
1789:
1762:
1735:
1462:. University of California Press. pp. 596–597.
452:; distal part of left tibia, fragmentary left pes;
351:, coming from the Sheeregeen Gashoon locality near
490:) in reference to its fragile build. Lastly, the
1606:Watabe, M.; Suzuki, D.; Tsogtbaatar, K. (2009).
1192:. Cambridge University Press. pp. 489–492.
1112:
1110:
1108:
1106:
1281:
1279:
331:-aged dinosaur fossil localities of Mongolia;
1654:
8:
1346:
1344:
1178:
1176:
1174:
1186:The Age of Dinosaurs in Russia and Mongolia
927:, which was deposited until the end of the
905:the formation, suggesting the existence of
2940:
2791:
2650:
2440:
2381:
2370:
2357:
2259:
2208:
2080:
1950:
1946:
1854:
1850:
1839:
1826:
1768:
1741:
1732:
1677:
1661:
1647:
1639:
111:
20:
1626:
1549:
1508:
1498:
1259:
1249:
1119:"Protoceratopsidae (Dinosauria) of Asia"
931:around 95.9 ± 6.0 million to 89.6 ± 4.0
1102:
1351:Ksepka, D. T.; Norell, M. A. (2006).
582:phylogenetic analysis was conducted.
412:; proximal end of left scapula; left
7:
1117:Maryańska, T.; Osmólska, H. (1975).
802:
752:
745:
716:
692:
637:
625:
601:
589:
14:
3073:Late Cretaceous dinosaurs of Asia
1203:Holtz, T. R.; Rey, L. V. (2007).
2924:
2923:
2899:
2893:
2343:
2337:
1565:Averianov, A.; Sues, H. (2012).
1084:Timeline of ceratopsian research
1068:
396:with mandibles; vertebrae, four
129:
1459:The Dinosauria, Second Edition
297:(meaning "slender horn") is a
1:
3093:Fossil taxa described in 2000
3088:Fossil taxa described in 1975
1615:Acta Palaeontologica Polonica
1538:Acta Palaeontologica Polonica
1500:10.1371/journal.pone.0118116
1308:10.1016/0195-6671(91)90015-5
1251:10.1371/journal.pone.0112055
387:Graciliceratops mongoliensis
277:Graciliceratops mongoliensis
3119:
572:During the description of
477:(meaning slender) and the
363:and referred to the genus
16:Extinct genus of dinosaurs
3098:Taxa named by Paul Sereno
2913:
2891:
2653:
2443:
2384:
2369:
2356:
2334:
2262:
2083:
1953:
1949:
1857:
1853:
1838:
1825:
1771:
1744:
1731:
1676:
1360:American Museum Novitates
1210:Genus List for Holtz 2012
824:
807:
800:
774:
757:
750:
743:
721:
714:
697:
690:
659:
642:
635:
623:
606:
599:
271:
264:
126:Scientific classification
124:
119:
110:
23:
343:, was discovered at the
1870:Micropachycephalosaurus
1574:Journal of Stratigraphy
1145:Journal of Paleontology
1126:Palaeontologia Polonica
1473:Turner, A. H. (2015).
1407:10.1006/cres.1999.0188
925:Cretaceous Long Normal
578:in 2014, an extensive
513:
468:, is derived from the
436:and fragment of right
345:Bayan Shireh Formation
336:
3045:Paleobiology Database
1628:10.4202/app.2006.0068
1551:10.4202/app.2008.0405
1051:. The discoveries of
509:
327:
3103:Ornithischian genera
914:Magnetostratigraphic
448:and nearly complete
432:; fragments of both
320:Discovery and naming
3083:Fossils of Mongolia
2035:Archaeoceratopsidae
1491:2015PLoSO..1018116T
1435:2020TeNov..32..246K
1399:1999CrRes..20..829H
1387:Cretaceous Research
1300:1991CrRes..12..345J
1288:Cretaceous Research
1242:2014PLoSO...9k2055F
381:) was considered a
2755:Crittendenceratops
2705:Albertaceratopsini
1213:Weight Information
514:
337:
3060:
3059:
3032:Open Tree of Life
2946:Taxon identifiers
2937:
2936:
2909:
2908:
2889:
2888:
2885:
2884:
2881:
2880:
2877:
2876:
2838:Pachyrhinosaurini
2776:Yehuecauhceratops
2739:Nasutoceratopsini
2640:
2639:
2352:
2351:
2332:
2331:
2328:
2327:
2324:
2323:
2320:
2319:
2316:
2315:
2312:
2311:
2213:Protoceratopsidae
2100:Leptoceratopsidae
2068:
2067:
1934:
1933:
1821:
1820:
1817:
1816:
1813:
1812:
1809:
1808:
1443:10.1111/ter.12456
956:, therizinosaurs
933:million years ago
893:
892:
884:
883:
875:
874:
866:
865:
857:
856:
848:
847:
839:
838:
789:
788:
674:
673:
631:Protoceratopsidae
550:fourth trochanter
290:
289:
260:
3110:
3053:
3052:
3040:
3039:
3027:
3026:
3014:
3013:
3001:
3000:
2988:
2987:
2986:
2973:
2972:
2971:
2941:
2927:
2926:
2904:
2903:
2897:
2868:Pachyrhinosaurus
2792:
2666:Machairoceratops
2651:
2491:Coahuilaceratops
2441:
2382:
2371:
2358:
2347:
2342:
2341:
2260:
2209:
2193:Zhuchengceratops
2081:
1951:
1947:
1888:Chaoyangsauridae
1855:
1851:
1840:
1827:
1769:
1742:
1733:
1726:
1725:
1678:
1663:
1656:
1649:
1640:
1633:
1632:
1630:
1612:
1603:
1597:
1596:
1594:
1588:. Archived from
1571:
1562:
1556:
1555:
1553:
1529:
1523:
1522:
1512:
1502:
1470:
1464:
1463:
1453:
1447:
1446:
1417:
1411:
1410:
1378:
1372:
1371:
1357:
1348:
1339:
1338:
1326:
1320:
1319:
1283:
1274:
1273:
1263:
1253:
1221:
1215:
1208:
1200:
1194:
1193:
1191:
1180:
1169:
1168:
1157:10.1666/07-069.1
1140:
1134:
1133:
1123:
1114:
1078:
1076:Dinosaurs portal
1073:
1072:
1071:
974:pachycephalosaur
803:
753:
746:
717:
693:
638:
626:
602:
590:
526:sacral vertebrae
511:Life restoration
486:(kéras, meaning
424:and fragmentary
361:Halszka Osmólska
357:Teresa Maryańska
276:
258:
251:
238:
222:
206:
190:
134:
133:
115:
105:
42:
29:Temporal range:
21:
3118:
3117:
3113:
3112:
3111:
3109:
3108:
3107:
3063:
3062:
3061:
3056:
3048:
3043:
3035:
3030:
3022:
3017:
3009:
3004:
2996:
2991:
2984:Graciliceratops
2982:
2981:
2976:
2967:
2966:
2961:
2954:Graciliceratops
2948:
2938:
2933:
2905:
2898:
2873:
2832:
2781:
2762:Furcatoceratops
2733:
2714:Albertaceratops
2699:
2673:Menefeeceratops
2636:
2609:Tatankaceratops
2566:
2512:Mercuriceratops
2463:Arrhinoceratops
2430:
2378:
2365:
2348:
2336:
2308:
2288:
2249:
2198:
2165:Montanoceratops
2094:
2089:Graciliceratops
2077:
2064:
2044:Archaeoceratops
2029:
1943:
1930:
1918:Xuanhuaceratops
1904:Hualianceratops
1882:
1863:Albalophosaurus
1847:
1834:
1805:
1785:
1758:
1752:Avemetatarsalia
1737:Avemetatarsalia
1727:
1721:Marginocephalia
1683:
1682:
1672:
1667:
1637:
1636:
1610:
1605:
1604:
1600:
1592:
1569:
1564:
1563:
1559:
1531:
1530:
1526:
1485:(2): e0118116.
1472:
1471:
1467:
1455:
1454:
1450:
1419:
1418:
1414:
1380:
1379:
1375:
1355:
1350:
1349:
1342:
1328:
1327:
1323:
1285:
1284:
1277:
1236:(12): e112055.
1223:
1222:
1218:
1207:. Random House.
1202:
1201:
1197:
1189:
1182:
1181:
1172:
1142:
1141:
1137:
1121:
1116:
1115:
1104:
1099:
1074:
1069:
1067:
1064:
902:Graciliceratops
899:
894:
885:
876:
867:
858:
849:
840:
790:
675:
610:Graciliceratops
584:Graciliceratops
570:
541:Graciliceratops
504:
466:Graciliceratops
333:Graciliceratops
322:
314:Late Cretaceous
294:Graciliceratops
286:
280:
257:
254:Graciliceratops
249:
236:
220:
204:
188:
128:
106:
104:
103:
98:
93:
88:
83:
78:
73:
68:
63:
58:
53:
48:
37:
36:
34:
31:Late Cretaceous
27:
25:Graciliceratops
17:
12:
11:
5:
3116:
3114:
3106:
3105:
3100:
3095:
3090:
3085:
3080:
3075:
3065:
3064:
3058:
3057:
3055:
3054:
3041:
3028:
3015:
3002:
2989:
2974:
2958:
2956:
2950:
2949:
2944:
2935:
2934:
2932:
2931:
2921:
2914:
2911:
2910:
2907:
2906:
2892:
2890:
2887:
2886:
2883:
2882:
2879:
2878:
2875:
2874:
2872:
2871:
2864:
2857:
2850:
2842:
2840:
2834:
2833:
2831:
2830:
2822:
2815:
2808:
2800:
2798:
2789:
2783:
2782:
2780:
2779:
2772:
2769:Nasutoceratops
2765:
2758:
2751:
2743:
2741:
2735:
2734:
2732:
2731:
2728:Medusaceratops
2724:
2717:
2709:
2707:
2701:
2700:
2698:
2697:
2690:
2683:
2676:
2669:
2662:
2659:Diabloceratops
2654:
2648:
2646:Centrosaurinae
2642:
2641:
2638:
2637:
2635:
2634:
2627:
2620:
2617:Titanoceratops
2613:
2605:
2602:Regaliceratops
2598:
2591:
2584:
2576:
2574:
2572:Triceratopsini
2568:
2567:
2565:
2564:
2557:
2550:
2543:
2536:
2533:Sierraceratops
2529:
2522:
2519:Navajoceratops
2515:
2508:
2501:
2494:
2487:
2480:
2473:
2466:
2459:
2452:
2444:
2438:
2436:Chasmosaurinae
2432:
2431:
2429:
2428:
2421:
2414:
2407:
2400:
2397:Brachyceratops
2393:
2385:
2379:
2374:
2367:
2366:
2361:
2354:
2353:
2350:
2349:
2335:
2333:
2330:
2329:
2326:
2325:
2322:
2321:
2318:
2317:
2314:
2313:
2310:
2309:
2307:
2306:
2305:
2304:
2298:
2296:
2290:
2289:
2287:
2286:
2279:
2276:Turanoceratops
2272:
2263:
2257:
2251:
2250:
2248:
2247:
2240:
2233:
2226:
2217:
2215:
2206:
2200:
2199:
2197:
2196:
2189:
2186:Unescoceratops
2182:
2175:
2168:
2161:
2154:
2151:Ischioceratops
2147:
2144:Gryphoceratops
2140:
2133:
2126:
2119:
2112:
2104:
2102:
2096:
2095:
2093:
2092:
2084:
2078:
2073:
2070:
2069:
2066:
2065:
2063:
2062:
2054:
2047:
2039:
2037:
2031:
2030:
2028:
2027:
2024:Sasayamagnomus
2020:
2013:
2006:
1999:
1991:
1984:
1977:
1974:Auroraceratops
1970:
1963:
1954:
1944:
1939:
1936:
1935:
1932:
1931:
1929:
1928:
1921:
1914:
1907:
1900:
1897:Chaoyangsaurus
1892:
1890:
1884:
1883:
1881:
1880:
1877:Psittacosaurus
1873:
1866:
1858:
1848:
1843:
1836:
1835:
1830:
1823:
1822:
1819:
1818:
1815:
1814:
1811:
1810:
1807:
1806:
1804:
1803:
1802:
1801:
1795:
1793:
1787:
1786:
1784:
1783:
1782:
1781:
1772:
1766:
1760:
1759:
1757:
1756:
1755:
1754:
1745:
1739:
1729:
1728:
1724:
1723:
1714:
1705:
1696:
1690:
1681:
1674:
1673:
1668:
1666:
1665:
1658:
1651:
1643:
1635:
1634:
1621:(2): 231–242.
1598:
1595:on 2019-03-07.
1580:(2): 462–485.
1557:
1544:(4): 601–614.
1524:
1465:
1448:
1429:(4): 246–252.
1412:
1393:(6): 829–850.
1373:
1362:(3508): 1–16.
1340:
1321:
1294:(4): 345–377.
1275:
1216:
1195:
1170:
1135:
1101:
1100:
1098:
1095:
1094:
1093:
1086:
1080:
1079:
1063:
1060:
1021:and the basal
949:tyrannosauroid
898:
895:
891:
890:
887:
886:
882:
881:
878:
877:
873:
872:
869:
868:
864:
863:
860:
859:
855:
854:
851:
850:
846:
845:
842:
841:
837:
836:
833:
832:
823:
820:
819:
816:
815:
811:Diabloceratops
806:
801:
799:
796:
795:
792:
791:
787:
786:
783:
782:
773:
770:
769:
766:
765:
756:
751:
749:
744:
742:
734:
733:
730:
729:
725:Turanoceratops
720:
715:
713:
710:
709:
706:
705:
696:
691:
689:
681:
680:
677:
676:
672:
671:
668:
667:
658:
655:
654:
651:
650:
641:
636:
634:
624:
622:
619:
618:
615:
614:
605:
600:
598:
588:
569:
568:Classification
566:
503:
500:
341:ZPAL MgD-I/156
339:The holotype,
321:
318:
308:that lived in
303:neoceratopsian
288:
287:
281:
269:
268:
262:
261:
247:
243:
242:
234:
227:
226:
218:
211:
210:
208:Neornithischia
202:
195:
194:
186:
179:
178:
173:
166:
165:
160:
156:
155:
150:
146:
145:
140:
136:
135:
122:
121:
117:
116:
108:
107:
99:
94:
89:
84:
79:
74:
69:
64:
59:
54:
49:
44:
43:
28:
15:
13:
10:
9:
6:
4:
3:
2:
3115:
3104:
3101:
3099:
3096:
3094:
3091:
3089:
3086:
3084:
3081:
3079:
3076:
3074:
3071:
3070:
3068:
3051:
3046:
3042:
3038:
3033:
3029:
3025:
3020:
3016:
3012:
3007:
3003:
2999:
2994:
2990:
2985:
2979:
2975:
2970:
2964:
2960:
2959:
2957:
2955:
2951:
2947:
2942:
2930:
2922:
2920:
2916:
2915:
2912:
2902:
2896:
2870:
2869:
2865:
2863:
2862:
2858:
2856:
2855:
2851:
2849:
2848:
2847:Achelousaurus
2844:
2843:
2841:
2839:
2835:
2828:
2827:
2826:Styracosaurus
2823:
2821:
2820:
2816:
2814:
2813:
2809:
2807:
2806:
2802:
2801:
2799:
2797:
2796:Centrosaurini
2793:
2790:
2788:
2787:Eucentrosaura
2784:
2778:
2777:
2773:
2771:
2770:
2766:
2764:
2763:
2759:
2757:
2756:
2752:
2750:
2749:
2745:
2744:
2742:
2740:
2736:
2730:
2729:
2725:
2723:
2722:
2718:
2716:
2715:
2711:
2710:
2708:
2706:
2702:
2696:
2695:
2691:
2689:
2688:
2687:Wendiceratops
2684:
2682:
2681:
2677:
2675:
2674:
2670:
2668:
2667:
2663:
2661:
2660:
2656:
2655:
2652:
2649:
2647:
2643:
2633:
2632:
2628:
2626:
2625:
2621:
2619:
2618:
2614:
2611:
2610:
2606:
2604:
2603:
2599:
2597:
2596:
2592:
2590:
2589:
2585:
2583:
2582:
2581:Eotriceratops
2578:
2577:
2575:
2573:
2569:
2563:
2562:
2558:
2556:
2555:
2551:
2549:
2548:
2544:
2542:
2541:
2537:
2535:
2534:
2530:
2528:
2527:
2526:Pentaceratops
2523:
2521:
2520:
2516:
2514:
2513:
2509:
2507:
2506:
2505:Kosmoceratops
2502:
2500:
2499:
2495:
2493:
2492:
2488:
2486:
2485:
2481:
2479:
2478:
2477:Bravoceratops
2474:
2472:
2471:
2470:Bisticeratops
2467:
2465:
2464:
2460:
2458:
2457:
2456:Anchiceratops
2453:
2451:
2450:
2449:Agujaceratops
2446:
2445:
2442:
2439:
2437:
2433:
2427:
2426:
2422:
2420:
2419:
2415:
2413:
2412:
2408:
2406:
2405:
2401:
2399:
2398:
2394:
2392:
2391:
2387:
2386:
2383:
2380:
2377:
2372:
2368:
2364:
2359:
2355:
2346:
2340:
2302:
2301:
2300:
2299:
2297:
2295:
2291:
2285:
2284:
2280:
2278:
2277:
2273:
2270:
2269:
2265:
2264:
2261:
2258:
2256:
2255:Ceratopsoidea
2252:
2246:
2245:
2244:Protoceratops
2241:
2239:
2238:
2237:Breviceratops
2234:
2232:
2231:
2227:
2224:
2223:
2219:
2218:
2216:
2214:
2210:
2207:
2205:
2201:
2195:
2194:
2190:
2188:
2187:
2183:
2181:
2180:
2179:Udanoceratops
2176:
2174:
2173:
2172:Prenoceratops
2169:
2167:
2166:
2162:
2160:
2159:
2158:Leptoceratops
2155:
2153:
2152:
2148:
2146:
2145:
2141:
2139:
2138:
2134:
2132:
2131:
2127:
2125:
2124:
2120:
2118:
2117:
2116:Bainoceratops
2113:
2111:
2110:
2106:
2105:
2103:
2101:
2097:
2091:
2090:
2086:
2085:
2082:
2079:
2076:
2071:
2060:
2059:
2055:
2053:
2052:
2051:Helioceratops
2048:
2046:
2045:
2041:
2040:
2038:
2036:
2032:
2026:
2025:
2021:
2019:
2018:
2017:Mosaiceratops
2014:
2012:
2011:
2007:
2005:
2004:
2000:
1997:
1996:
1992:
1990:
1989:
1988:Koreaceratops
1985:
1983:
1982:
1978:
1976:
1975:
1971:
1969:
1968:
1964:
1961:
1960:
1956:
1955:
1952:
1948:
1945:
1942:
1941:Neoceratopsia
1937:
1927:
1926:
1922:
1920:
1919:
1915:
1913:
1912:
1908:
1906:
1905:
1901:
1899:
1898:
1894:
1893:
1891:
1889:
1885:
1879:
1878:
1874:
1872:
1871:
1867:
1865:
1864:
1860:
1859:
1856:
1852:
1849:
1846:
1841:
1837:
1833:
1828:
1824:
1799:
1798:
1797:
1796:
1794:
1792:
1788:
1780:
1776:
1775:
1774:
1773:
1770:
1767:
1765:
1761:
1753:
1749:
1748:
1747:
1746:
1743:
1740:
1738:
1734:
1730:
1722:
1718:
1715:
1713:
1709:
1706:
1704:
1700:
1697:
1695:
1691:
1689:
1685:
1684:
1679:
1675:
1671:
1664:
1659:
1657:
1652:
1650:
1645:
1644:
1641:
1629:
1624:
1620:
1616:
1609:
1602:
1599:
1591:
1587:
1583:
1579:
1575:
1568:
1561:
1558:
1552:
1547:
1543:
1539:
1535:
1528:
1525:
1520:
1516:
1511:
1506:
1501:
1496:
1492:
1488:
1484:
1480:
1476:
1469:
1466:
1461:
1460:
1452:
1449:
1444:
1440:
1436:
1432:
1428:
1424:
1416:
1413:
1408:
1404:
1400:
1396:
1392:
1388:
1384:
1377:
1374:
1369:
1365:
1361:
1354:
1347:
1345:
1341:
1336:
1332:
1325:
1322:
1317:
1313:
1309:
1305:
1301:
1297:
1293:
1289:
1282:
1280:
1276:
1271:
1267:
1262:
1257:
1252:
1247:
1243:
1239:
1235:
1231:
1227:
1220:
1217:
1214:
1211:
1206:
1199:
1196:
1188:
1187:
1179:
1177:
1175:
1171:
1166:
1162:
1158:
1154:
1150:
1146:
1139:
1136:
1131:
1127:
1120:
1113:
1111:
1109:
1107:
1103:
1096:
1092:
1091:
1087:
1085:
1082:
1081:
1077:
1066:
1061:
1059:
1057:
1054:
1050:
1049:
1045:
1041:
1040:
1035:
1034:
1029:
1028:
1024:
1020:
1019:
1015:
1011:
1010:
1005:
1001:
997:
996:
992:
988:
984:
980:
979:
975:
971:
970:
965:
961:
960:
955:
954:
953:Alectrosaurus
950:
946:
945:
941:
940:dromaeosaurid
936:
934:
930:
926:
922:
919:
915:
911:
908:
903:
896:
889:
888:
880:
879:
871:
870:
862:
861:
853:
852:
844:
843:
835:
834:
831:
830:
829:
822:
821:
818:
817:
814:
813:
812:
805:
804:
798:
797:
794:
793:
785:
784:
781:
780:
779:
772:
771:
768:
767:
764:
763:
762:
755:
754:
748:
747:
741:
740:
736:
735:
732:
731:
728:
727:
726:
719:
718:
712:
711:
708:
707:
704:
703:
702:
695:
694:
688:
687:
686:Ceratopsoidea
683:
682:
679:
678:
670:
669:
666:
665:
664:
657:
656:
653:
652:
649:
648:
647:
646:Protoceratops
640:
639:
633:
632:
628:
627:
621:
620:
617:
616:
613:
612:
611:
604:
603:
597:
596:
592:
591:
587:
585:
581:
577:
576:
567:
565:
563:
559:
555:
551:
547:
542:
538:
537:Protoceratops
534:
533:
532:Protoceratops
527:
523:
519:
512:
508:
501:
499:
497:
493:
492:specific name
489:
485:
484:
480:
476:
475:
471:
467:
463:
459:
456:and isolated
455:
451:
447:
443:
439:
435:
431:
427:
423:
419:
415:
411:
407:
403:
399:
395:
390:
388:
384:
380:
379:
374:
373:Microceratops
370:
366:
365:Microceratops
362:
358:
354:
350:
346:
342:
334:
330:
326:
319:
317:
315:
311:
307:
304:
300:
296:
295:
284:
279:
278:
270:
267:
263:
256:
255:
248:
245:
244:
241:
240:Neoceratopsia
235:
232:
229:
228:
225:
219:
216:
213:
212:
209:
203:
200:
197:
196:
193:
187:
184:
181:
180:
177:
174:
171:
168:
167:
164:
161:
158:
157:
154:
151:
148:
147:
144:
141:
138:
137:
132:
127:
123:
118:
114:
109:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
41:
32:
26:
22:
19:
3078:Ceratopsians
2953:
2866:
2861:Stellasaurus
2859:
2852:
2845:
2824:
2817:
2812:Coronosaurus
2810:
2805:Centrosaurus
2803:
2774:
2767:
2760:
2753:
2746:
2726:
2721:Lokiceratops
2719:
2712:
2694:Xenoceratops
2692:
2685:
2680:Sinoceratops
2678:
2671:
2664:
2657:
2629:
2622:
2615:
2607:
2600:
2593:
2588:Nedoceratops
2586:
2579:
2561:Vagaceratops
2559:
2554:Utahceratops
2552:
2547:Terminocavus
2545:
2538:
2531:
2524:
2517:
2510:
2503:
2498:Judiceratops
2496:
2489:
2484:Chasmosaurus
2482:
2475:
2468:
2461:
2454:
2447:
2423:
2416:
2409:
2402:
2395:
2388:
2376:Ceratopsidae
2363:Ceratopsidae
2294:Ceratopsidae
2283:Zuniceratops
2281:
2274:
2268:Ajkaceratops
2266:
2242:
2235:
2230:Bagaceratops
2228:
2222:Ajkaceratops
2220:
2204:Coronosauria
2191:
2184:
2177:
2170:
2163:
2156:
2149:
2142:
2135:
2128:
2121:
2114:
2109:Asiaceratops
2107:
2088:
2087:
2075:Euceratopsia
2058:Yamaceratops
2056:
2049:
2042:
2022:
2015:
2010:Microceratus
2008:
2003:Liaoceratops
2001:
1993:
1986:
1979:
1972:
1965:
1959:Ajkaceratops
1957:
1923:
1916:
1909:
1902:
1895:
1875:
1868:
1861:
1779:Ornithischia
1764:Ornithischia
1716:
1712:Ornithischia
1707:
1698:
1618:
1614:
1601:
1590:the original
1577:
1573:
1560:
1541:
1537:
1527:
1482:
1478:
1468:
1458:
1451:
1426:
1422:
1415:
1390:
1386:
1376:
1359:
1334:
1330:
1324:
1291:
1287:
1233:
1229:
1219:
1204:
1198:
1185:
1148:
1144:
1138:
1129:
1125:
1090:Microceratus
1088:
1046:
1039:Lindholmemys
1037:
1033:Paralligator
1031:
1025:
1023:hadrosauroid
1016:
1012:, the large
1007:
993:
983:ornithomimid
976:
967:
964:Enigmosaurus
959:Erlikosaurus
957:
951:
944:Achillobator
942:
937:
901:
900:
897:Paleoecology
828:Centrosaurus
826:
825:
809:
808:
776:
775:
761:Chasmosaurus
759:
758:
739:Ceratopsidae
737:
723:
722:
701:Zuniceratops
699:
698:
684:
663:Bagaceratops
661:
660:
644:
643:
629:
609:
608:
607:
595:Coronosauria
593:
583:
573:
571:
560:. The pedal
554:articulation
540:
536:
530:
515:
496:mongolienses
495:
482:
481:
473:
472:
465:
462:generic name
391:
386:
383:nomen dubium
378:Microceratus
376:
372:
364:
340:
338:
332:
293:
292:
291:
273:
272:
266:Type species
259:Sereno, 2000
253:
252:
230:
214:
198:
192:Ornithischia
182:
169:
24:
18:
2978:Wikispecies
2854:Einiosaurus
2748:Avaceratops
2631:Triceratops
2595:Ojoceratops
2418:Monoclonius
2130:Ferrisaurus
1995:Kulceratops
1053:azhdarchids
1009:Tsagantegia
991:ankylosaurs
987:Garudimimus
978:Amtocephale
969:Segnosaurus
778:Triceratops
502:Description
375:(now named
369:Paul Sereno
367:. However,
312:during the
38:96–89
3067:Categories
2917:See also:
2624:Torosaurus
2540:Spiclypeus
2390:Agathaumas
2303:see below↓
2123:Cerasinops
1911:Stenopelix
1845:Ceratopsia
1832:Ceratopsia
1800:see below↓
1791:Ceratopsia
1703:Dinosauria
1670:Ceratopsia
1423:Terra Nova
1337:: 113–182.
1151:(2): 423.
1132:: 134–143.
1097:References
1056:pterosaurs
1027:Gobihadros
1000:Amtosaurus
907:Angiosperm
580:Ceratopsia
558:metatarsal
404:and seven
329:Cretaceous
224:Ceratopsia
176:Dinosauria
1686:Kingdom:
1316:0195-6671
995:Talarurus
989:’’, the
929:Santonian
400:, twelve
398:cervicals
353:Sainshand
306:dinosaurs
149:Kingdom:
143:Eukaryota
2963:Wikidata
2929:Category
2919:Timeline
2425:Polyonax
2411:Dysganus
2404:Ceratops
1967:Aquilops
1694:Chordata
1692:Phylum:
1688:Animalia
1586:54210424
1519:25714338
1479:PLOS ONE
1270:25494182
1230:PLOS ONE
1165:86021954
1062:See also
1042:and the
1014:sauropod
1006:’’, and
1004:Maleevus
966:’’, and
575:Aquilops
474:gracilis
440:; right
416:; right
414:coracoid
408:; right
349:Mongolia
316:period.
163:Chordata
159:Phylum:
153:Animalia
139:Domain:
3037:4946432
3024:1217488
3011:4823257
2998:4531827
2969:Q133140
2819:Spinops
2137:Gremlin
1925:Yinlong
1510:4340866
1487:Bibcode
1431:Bibcode
1395:Bibcode
1296:Bibcode
1261:4262212
1238:Bibcode
1048:Hybodus
918:calcite
562:unguals
546:glenoid
524:of the
454:tarsals
438:ischium
418:humerus
410:scapula
406:sacrals
402:dorsals
246:Genus:
1584:
1517:
1507:
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1268:
1258:
1163:
1018:Erketu
1002:’’, ‘’
981:, the
972:, the
947:, the
910:plants
522:centra
518:arches
460:. The
422:radius
285:, 2000
283:Sereno
3050:67673
3019:IRMNG
1717:Clade
1708:Clade
1699:Clade
1611:(PDF)
1593:(PDF)
1582:S2CID
1570:(PDF)
1356:(PDF)
1190:(PDF)
1161:S2CID
1122:(PDF)
1044:shark
483:κέρας
479:Greek
470:Latin
446:tibia
442:femur
434:ilium
430:pubis
394:skull
299:genus
231:Clade
215:Clade
199:Clade
183:Clade
170:Clade
3006:GBIF
1777:see
1750:see
1515:PMID
1312:ISSN
1266:PMID
998:, ‘’
962:, ‘’
921:U–Pb
916:and
520:and
488:horn
458:ribs
426:ulna
359:and
310:Asia
46:PreꞒ
2993:EoL
1981:Beg
1623:doi
1546:doi
1505:PMC
1495:doi
1439:doi
1403:doi
1364:doi
1304:doi
1256:PMC
1246:doi
1153:doi
450:pes
347:in
301:of
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2271:?
2225:?
2061:?
1998:?
1962:?
1662:e
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1648:v
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275:†
250:†
237:†
221:†
205:†
189:†
101:N
91:K
86:J
81:T
76:P
71:C
66:D
61:S
56:O
51:Ꞓ
35:~
33:,
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