Knowledge (XXG)

Gravitholus

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1109: 136: 967: 990: 428: 118: 346:). It was a pachycephalosaur, and like other pachycephalosaurids the skull roof formed a thick dome made of dense bone, which may have been used in head-butting contests over mates or territory. It lived in what is now 409:, thus decreasing the diversity of Dinosaur Park pachcephalosaurids to only two valid taxa. This conclusion was reached because new morphological data recovered from synchrotron scanning of the skull revealed 550: 508:
Wall, W. P. & Galton, P. M. (1979). "Notes on pachycephalosaurid dinosaurs (Reptilia: Ornithischia) from North America, with comments on their status as ornithopods".
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There is some debate amongst paleontologists as to whether the animal represents a distinct genus, or if it is synonymous with
1136: 467:"Problematic putative pachycephalosaurids: Synchrotron µCT imaging shines new light on the anatomy and taxonomic validity of 1194: 1199: 565: 393:. It is unclear whether all of these species would have lived in the area at the same time. A 2023 publication by Dyer 582: 135: 993: 1189: 687: 466: 966: 1010: 389: 1081: 714: 383: 289: 861: 809: 266: 527: 519: 515: 1108: 375:
was initially described as one of several pachycephalosaurids known from Dinosaur Park, including
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from the Belly River Group (Campanian) of Alberta, Canada"
1124: 350:, Canada, and was described in 1979 by W. P. Wall and 1017: 860: 808: 776: 725: 698: 671: 522:(2004). "Pachycephalosauria". In D. B. Weishampel, 1144: 590: 8: 475:Vertebrate Anatomy Morphology Palaeontology 1151: 1137: 1005: 814: 782: 773: 704: 677: 668: 613: 597: 583: 575: 116: 20: 486: 453: 442:Timeline of pachycephalosaur research 7: 1105: 1103: 413:to be virtually identical to mature 1123:. You can help Knowledge (XXG) by 510:Canadian Journal of Earth Sciences 14: 461:Dyer, Aaron D.; Powers, Mark J.; 1107: 989: 988: 965: 426: 134: 1: 1170:Fossil taxa described in 1979 320:(meaning 'heavy dome') was a 563:on the Dinosaur Mailing List 548:at the Paleobiology Database 1216: 1180:Taxa named by Peter Galton 1102: 16:Extinct genus of dinosaurs 1175:Pachycephalosaurs of Asia 978: 963: 817: 785: 772: 753: 707: 680: 667: 612: 295: 288: 131:Scientific classification 129: 124: 115: 23: 1115:This article related to 1185:Paleontology in Alberta 308:Wall & Galton, 1979 518:, Chapman, R. E., and 405:to be synonymous with 390:Foraminacephale brevis 1082:Paleobiology Database 384:Hanssuesia sternbergi 29:Temporal range: Late 1195:Ornithischian genera 862:Pachycephalosaurinae 810:Pachycephalosauridae 469:Gravitholus albertae 360:Gravitholus albertae 302:Gravitholus albertae 242:Pachycephalosauridae 1200:Ornithischian stubs 125:Skull TMP 72.27.01 913:Pachycephalosaurus 778:Pachycephalosauria 757:Pachycephalosauria 727:Pachycephalosauria 606:Pachycephalosauria 568:2016-03-03 at the 553:2012-02-06 at the 488:10.18435/vamp29388 378:Stegoceras validum 326:pachycephalosaurid 229:Pachycephalosauria 1132: 1131: 1097: 1096: 1069:Open Tree of Life 1011:Taxon identifiers 1002: 1001: 974: 973: 961: 960: 957: 956: 768: 767: 764: 763: 749: 748: 745: 744: 520:Weishampel, D. B. 463:Currie, Philip J. 344:million years ago 313: 312: 269: 1207: 1190:Campanian genera 1153: 1146: 1139: 1117:pachycephalosaur 1111: 1104: 1090: 1089: 1077: 1076: 1064: 1063: 1051: 1050: 1038: 1037: 1036: 1006: 992: 991: 969: 815: 783: 774: 705: 678: 669: 662: 661: 614: 599: 592: 585: 576: 501: 500: 490: 458: 436: 434:Dinosaurs portal 431: 430: 429: 304: 300: 281:G. albertae 260: 253: 240: 227: 211: 195: 139: 138: 120: 110: 40: 21: 1215: 1214: 1210: 1209: 1208: 1206: 1205: 1204: 1160: 1159: 1158: 1157: 1119:dinosaurs is a 1100: 1098: 1093: 1085: 1080: 1072: 1067: 1059: 1054: 1046: 1041: 1032: 1031: 1026: 1013: 1003: 998: 970: 953: 892:Foraminacephale 856: 804: 760: 741: 721: 694: 688:Avemetatarsalia 673:Avemetatarsalia 663: 657:Marginocephalia 619: 618: 608: 603: 570:Wayback Machine 555:Wayback Machine 541: 505: 504: 460: 459: 455: 450: 432: 427: 425: 422: 309: 306: 298: 297: 284: 259: 251: 238: 225: 209: 193: 133: 111: 109: 108: 107: 106: 101: 96: 91: 86: 81: 76: 71: 66: 61: 56: 51: 46: 35: 34: 27: 17: 12: 11: 5: 1213: 1211: 1203: 1202: 1197: 1192: 1187: 1182: 1177: 1172: 1162: 1161: 1156: 1155: 1148: 1141: 1133: 1130: 1129: 1112: 1095: 1094: 1092: 1091: 1078: 1065: 1052: 1039: 1023: 1021: 1015: 1014: 1009: 1000: 999: 997: 996: 986: 979: 976: 975: 972: 971: 964: 962: 959: 958: 955: 954: 952: 951: 944: 937: 930: 923: 916: 909: 902: 895: 888: 881: 874: 866: 864: 858: 857: 855: 854: 847: 840: 833: 826: 823:Colepiocephale 818: 812: 806: 805: 803: 802: 795: 791:Ferganocephale 786: 780: 770: 769: 766: 765: 762: 761: 754: 751: 750: 747: 746: 743: 742: 740: 739: 738: 737: 731: 729: 723: 722: 720: 719: 718: 717: 708: 702: 696: 695: 693: 692: 691: 690: 681: 675: 665: 664: 660: 659: 650: 641: 632: 626: 617: 610: 609: 604: 602: 601: 594: 587: 579: 573: 572: 557: 540: 539:External links 537: 536: 535: 532:The Dinosauria 513: 512:. 16:1176-1186 503: 502: 452: 451: 449: 446: 445: 444: 438: 437: 421: 418: 331:from the late 311: 310: 307: 293: 292: 286: 285: 277: 275: 271: 270: 249: 245: 244: 236: 232: 231: 223: 216: 215: 213:Neornithischia 207: 200: 199: 191: 184: 183: 178: 171: 170: 165: 161: 160: 155: 151: 150: 145: 141: 140: 127: 126: 122: 121: 113: 112: 104: 103: 102: 97: 92: 87: 82: 77: 72: 67: 62: 57: 52: 47: 42: 41: 28: 15: 13: 10: 9: 6: 4: 3: 2: 1212: 1201: 1198: 1196: 1193: 1191: 1188: 1186: 1183: 1181: 1178: 1176: 1173: 1171: 1168: 1167: 1165: 1154: 1149: 1147: 1142: 1140: 1135: 1134: 1128: 1126: 1122: 1118: 1113: 1110: 1106: 1101: 1088: 1083: 1079: 1075: 1070: 1066: 1062: 1057: 1053: 1049: 1044: 1040: 1035: 1029: 1025: 1024: 1022: 1020: 1016: 1012: 1007: 995: 987: 985: 981: 980: 977: 968: 950: 949: 945: 943: 942: 941:Sphaerotholus 938: 936: 935: 931: 929: 928: 924: 922: 921: 917: 915: 914: 910: 908: 907: 906:Homalocephale 903: 901: 900: 896: 894: 893: 889: 887: 886: 882: 880: 879: 878:Alaskacephale 875: 873: 872: 868: 867: 865: 863: 859: 853: 852: 848: 846: 845: 841: 839: 838: 834: 832: 831: 827: 825: 824: 820: 819: 816: 813: 811: 807: 801: 800: 799:Wannanosaurus 796: 793: 792: 788: 787: 784: 781: 779: 775: 771: 759: 758: 752: 735: 734: 733: 732: 730: 728: 724: 716: 712: 711: 710: 709: 706: 703: 701: 697: 689: 685: 684: 683: 682: 679: 676: 674: 670: 666: 658: 654: 651: 649: 645: 642: 640: 636: 633: 631: 627: 625: 621: 620: 615: 611: 607: 600: 595: 593: 588: 586: 581: 580: 577: 571: 567: 564: 562: 558: 556: 552: 549: 547: 543: 542: 538: 533: 529: 525: 521: 517: 516:Maryańska, T. 514: 511: 507: 506: 498: 494: 489: 484: 481:(1): 65–110. 480: 476: 472: 470: 464: 457: 454: 447: 443: 440: 439: 435: 424: 419: 417: 416: 412: 408: 404: 400: 396: 392: 391: 386: 385: 380: 379: 374: 370: 369: 363: 361: 357: 353: 349: 345: 341: 338: 334: 330: 327: 323: 319: 318: 305: 303: 294: 291: 290:Binomial name 287: 283: 282: 276: 273: 272: 268: 264: 258: 257: 250: 247: 246: 243: 237: 234: 233: 230: 224: 221: 218: 217: 214: 208: 205: 202: 201: 198: 192: 189: 186: 185: 182: 179: 176: 173: 172: 169: 166: 163: 162: 159: 156: 153: 152: 149: 146: 143: 142: 137: 132: 128: 123: 119: 114: 100: 95: 90: 85: 80: 75: 70: 65: 60: 55: 50: 45: 39: 32: 26: 22: 19: 1125:expanding it 1114: 1099: 1018: 946: 939: 932: 927:Prenocephale 925: 918: 911: 904: 897: 890: 883: 876: 869: 849: 842: 835: 829: 828: 821: 797: 789: 755: 715:Ornithischia 700:Ornithischia 652: 648:Ornithischia 643: 634: 560: 545: 531: 509: 478: 474: 468: 456: 414: 410: 406: 402: 398: 394: 388: 382: 376: 372: 366: 364: 359: 356:type species 352:Peter Galton 342:, around 75 316: 315: 314: 301: 296: 280: 279: 255: 254: 219: 203: 197:Ornithischia 187: 174: 24: 18: 1019:Gravitholus 948:Tylocephale 934:Sinocephale 920:Platytholus 899:Goyocephale 885:Amtocephale 851:Texacephale 830:Gravitholus 561:Gravitholus 546:Gravitholus 528:H. Osmólska 415:Stegoceras. 411:Gravitholus 399:Gravitholus 373:Gravitholus 317:Gravitholus 261:Wall & 256:Gravitholus 25:Gravitholus 1164:Categories 982:See also: 871:Acrotholus 844:Stegoceras 837:Hanssuesia 736:see below↓ 639:Dinosauria 448:References 407:Stegoceras 403:Hanssuesia 368:Stegoceras 333:Cretaceous 181:Dinosauria 622:Kingdom: 524:P. Dodson 497:2292-1389 337:Campanian 274:Species: 154:Kingdom: 148:Eukaryota 31:Campanian 1028:Wikidata 994:Category 984:Timeline 630:Chordata 628:Phylum: 624:Animalia 566:Archived 551:Archived 530:(eds.), 465:(2023). 420:See also 397:. found 335:period ( 329:dinosaur 235:Family: 168:Chordata 164:Phylum: 158:Animalia 144:Domain: 36:75  1074:4128762 1061:1034182 1048:4823084 1034:Q134642 348:Alberta 299:† 278:† 248:Genus: 105:↓ 526:& 495:  354:. The 263:Galton 1087:38784 1056:IRMNG 653:Clade 644:Clade 635:Clade 395:et al 340:stage 322:genus 220:Clade 204:Clade 188:Clade 175:Clade 1121:stub 1043:GBIF 713:see 686:see 493:ISSN 401:and 387:and 267:1979 44:PreꞒ 483:doi 358:is 324:of 1166:: 1084:: 1071:: 1058:: 1045:: 1030:: 655:: 646:: 637:: 491:. 479:10 477:. 473:. 381:, 362:. 265:, 222:: 206:: 190:: 177:: 94:Pg 38:Ma 33:, 1152:e 1145:t 1138:v 1127:. 794:? 598:e 591:t 584:v 499:. 485:: 252:† 239:† 226:† 210:† 194:† 99:N 89:K 84:J 79:T 74:P 69:C 64:D 59:S 54:O 49:Ꞓ

Index

Campanian
Ma
PreꞒ

O
S
D
C
P
T
J
K
Pg
N

Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Dinosauria
Ornithischia
Neornithischia
Pachycephalosauria
Pachycephalosauridae
Gravitholus
Galton
1979
Binomial name
genus

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