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Gryposaurus

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5284: 5279: 1134: 4369: 870: 4383: 1604: 5266: 149: 4376: 1587:, like other cranial modifications in duckbills, may have been used for a variety of social functions, such as identification of sexes or species and social ranking. It could also have functioned as a tool for broadside pushing or butting in social contests, and there may have been inflatable air sacs flanking it for both visual and auditory signaling. The top of the arch is roughened in some specimens, suggesting that it was covered by thick, 5290: 556: 1051: 1122: 697: 125: 5273: 3322: 1090:-shaped scutes upwards of 3.8 centimeters long (1.5 inches) on the flank and tail; uniform polygonal scales on the neck and sides of the body; and pyramidal structures, flattened side-to-side, with fluted sides, longer than tall and found along the top of the back in a single midline row. In 2016, Gregory S. Paul estimated the size of 528:, and for many years the two were thought to be synonyms. It is known from numerous skulls, some skeletons, and even some skin impressions that show it to have had pyramidal scales projecting along the midline of the back. It is most easily distinguished from other duckbills by its narrow arching nasal hump, sometimes described as similar to a " 1461: 812:
had enlarged prongs along its upper margin, where the lower jaw's beak was based. This new species greatly expands the geographic range of this genus, and there may be a second, more lightly built species present as well. Multiple gryposaur species are known from the Kaiparowits Formation,
1959:; Hernández,-Rivera, René; Gates, Terry; Paul, Gregory S.; Nesbitt, Sterling; Serrano-Brañas, Claudia Inés; Garcia-de la Garza, Juan Pablo (2006). "Large hadrosaurine dinosaurs from the latest Campanian of Coahuila, Mexico". In Lucas, Spencer G.; Sullivan Robert M. (eds.). 3210:
Eaton, J.G., 2002. Multituberculate mammals from the Wahweap (Campanian, Aquilan) and Kaiparowits (Campanian, Judithian) formations, within and near Grand Staircase–Escalante National Monument, southern Utah. Miscellaneous Publication 02-4, Utah Geological Survey, 66
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swamps, ponds and lakes, and was bordered by highlands. The climate was wet and humid, and supported an abundant and diverse range of organisms. This formation contains one of the best and most continuous records of Late Cretaceous terrestrial life in the world.
1113:. In profile view, they rise into a rounded hump in front of the eyes, reaching a height as tall as the highest point of the back of the skull. The skeleton is known in great detail, making it a useful point of reference for other duckbill skeletons. 2404: 3296:
Eaton, Jeffrey G.; Cifelli, Richard L.; Hutchinson, J. Howard; Kirkland, James I.; Parrish, J. Michael (1999). "Cretaceous vertebrate faunas from the Kaiparowits Plateau, south-central Utah". In Gillete David D. (ed.).
3198: 2111:(1979). "Upper Cretaceous dinosaurs from the Bearpaw Shale (marine) of south-central Montana with a checklist of Upper Cretaceous dinosaur remains from marine sediments in North America". 5831: 3073: 3199:
40Ar/39Ar age of the Kaiparowits Formation, southern Utah, and correlation of contemporaneous Campanian strata and vertebrate faunas along the margin of the Western Interior Basin
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Lull, Richard Swann; Wright, Nelda E. (1942). Hadrosaurian Dinosaurs of North America. Geological Society of America Special Paper 40. Geological Society of America. pp. 164–172.
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that contained hundreds of teeth, only a relative handful of which were in use at any time. Plant material would have been cropped by its broad beak, and held in the jaws by a
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Titus, Alan L. and Mark A. Loewen (editors). At the Top of the Grand Staircase: The Late Cretaceous of Southern Utah. 2013. Indiana University Press. Hardbound: 634 pp.
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appears to have only existed for part of the duration of time that the rocks were being formed. As the formation was being laid down, it recorded a change to more
2023:(1916). "Contributions to the geology and paleontology of San Juan County, New Mexico. 2. Vertebrate faunas of the Ojo Alamo, Kirtland and Fruitland Formations". 2389:
The hadrosaurian dinosaurs (ornithischia: hadrosauria) of Big Bend National Park, Brewster County, Texas, with implications for late Cretaceous paleozoogeography
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was once thought to fall into this group as well, before it was described (then known under the nickname "Antonio"). A subfamily, Gryposaurinae, was coined by
5861: 5762: 5283: 3020: 777:, from slightly older rocks in Montana than the classic gryposaur localities of Alberta. Based on two parts of a skeleton collected in 1916 for the 2784:(1902). "On Vertebrata of the mid-Cretaceous of the Northwest Territory. 2. New genera and species from the Belly River Series (mid-Cretaceous)". 661:
type specimen is missing the front part of the skull, so the full shape of the nasal arch cannot be seen. The 1942 publication of the influential
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have been unearthed for decades, but none were considered to be identifiable as a determined genus of hadrosaur, but do resemble some species of
4368: 5876: 5278: 3081: 5826: 3355: 3306: 2959: 2897: 2657: 2626: 2590: 2288:(Ornithischia: Hadrosauridae) with descriptions of two new hadrosaurid species and an evaluation of hadrosaurid phylogenetic relationships". 2204: 2176: 2145: 2089: 1870: 1827: 1626:
indicates that the Kaiparowits Formation was deposited between 76.1 and 74.0 million years ago, during the Campanian stage of the Late
5866: 4382: 3261:
Zanno, Lindsay E.; Sampson, Scott D. (2005). "A new oviraptorosaur (Theropoda; Maniraptora) from the Late Cretaceous (Campanian) of Utah".
1511:-like organ. Its feeding range would have extended from the ground to about 4 m (13 ft) above. The paleontologists who unearthed 3103: 2573:; Weishampel, David B.; Forster, Catherine A (2004). "Hadrosauridae". In Weishampel, David B.; Dodson, Peter; OsmĂłlska Halszka (eds.). 778: 750:
of Montana (which have since fallen out of the literature), by 1990 he had changed his position, and was among the first to again use
3231: 5265: 532:," and which may have been used for species or sexual identification, and/or combat with individuals of the same species. A large 5851: 5841: 5644: 1956: 1896:, a new genus and species of trachodont dinosaur from the Belly River Formation of Alberta, with a description of the skull of 955:, Montana, USA, is known from partial skulls and skeletons from several individuals. Its nasal arch is prominent like that of 5856: 5667: 5654: 2921: 2042: 1791: 634: 136: 1630:
period. During the Late Cretaceous period, the site of the Kaiparowits Formation was located near the western shore of the
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to the east. The plateau where dinosaurs lived was an ancient floodplain dominated by large channels and abundant wetland
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as part of a larger revision that promoted Hadrosaurinae to family status, but is not now in use. A rough equivalent is
1158: 355: 2677: 1133: 607:. This skull was missing the snout, which had eroded into fragments; Brown restored it after the duckbill now known as 5289: 1109:
differ in details of the skull and lower jaw. The prominent nasal arch found in this genus is formed from the paired
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back in the 2000s brought the fact that this creature dined on tough, fibrous plant material which would imply that
613:, which was flat-headed, and believed that some unusual pieces were evidence of compression. Lambe's description of 578: 4375: 1094:
at 7.5 metres (25 ft) in length and 2.5 metric tons (2.8 short tons) in body mass and the other two species (
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based on the phylogenetic analysis conducted by Prieto-Márquez and Wagner in 2012, showing the relationships of
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The Illustrated Encyclopedia of Dinosaurs: An Original and Compelling Insight into Life in the Dinosaur Kingdom
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without hollow head crests. The general term "gryposaur" is sometimes used for duckbills with arched nasals.
2803: 2080:; Horner, Jack R. (1990). "Hadrosauridae". In Weishampel, David B.; Dodson, Peter; OsmĂłlska Halszka (eds.). 2020: 1631: 952: 907:
Dinosaur Park Formation of Alberta, Canada. It is now thought that another species from the same formation,
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Redescription of a remarkably large Gryposaurus notabilis (Dinosauria: Hadrosauridae) from Alberta, Canada
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was a hadrosaurid of typical size and shape; one of the best specimens of this genus, the nearly complete
948: 497: 5788: 5624: 4180: 3482: 1723: 1612: 861:, was unearthed in the Javelina Formation in Texas. Further research is needed to confirm its validity. 797: 505: 2984:, a new species of hadrosaurid dinosaur from the Late Cretaceous of the Pacific coast of North America" 722:
was most common in the late 1970s–early 1980s, and appears in some popular books; one well-known work,
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is incompletely known, lacking most of the bones in front of the eyes, it was very similar to that of
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This situation is made more confusing by old suggestions by some authors, including Jack Horner, that
5749: 5044: 3341: 3164: 2740: 2525: 2416: 1994: 1796: 1527: 821: 5272: 1555:
present instead. Other dinosaurs known from only the lower part of the formation include the horned
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Hopson, James A. (1975). "The evolution of cranial display structures in hadrosaurian dinosaurs".
1078:) came from an animal about 8.2 meters (27 feet) long. This specimen also has the best example of 919:. The two had been differentiated by the size of the nasal arch (larger and closer to the eyes in 813:
represented by cranial and postcranial remains, and were larger than their northern counterparts.
5396: 5007: 3592: 3278: 3180: 3046: 3001: 2864: 2842: 2756: 2729:"Hadrosaurian dinosaurs from the Maastrichtian Javelina Formation, Big Bend National Park, Texas" 2541: 2458: 2440: 2405:"Hadrosaurian dinosaurs from the Maastrichtian Javelina Formation, Big Bend National Park, Texas" 2368: 2250: 1623: 992: 817: 513: 397: 370: 143: 2618: 2610: 1922:(1910). "The Cretaceous Ojo Alamo beds of New Mexico with description of the new dinosaur genus 5793: 2582: 5780: 5403: 5142: 5063: 5023: 3907: 3793: 3739: 3710: 3302: 2955: 2903: 2893: 2653: 2622: 2586: 2489: 2432: 2316:(Dinosauria: Hadrosauridae) from the late Campanian Kaiparowits Formation, southern Utah, USA" 2221: 2200: 2172: 2168: 2162: 2141: 2137: 2131: 2085: 1967:. Albuquerque, New Mexico: New Mexico Museum of Natural History and Science. pp. 299–315. 1866: 1823: 5740: 3232:"Presidential Proclamation: Establishment of the Grand Staircase–Escalante National Monument" 2649: 2643: 1086:, showing this dinosaur to have had several different types of scalation: pyramidal, ridged, 597:
shortly thereafter, Lambe drawing attention to its unusual nasal crest. A few years earlier,
5544: 5537: 5347: 5340: 4869: 4780: 4618: 4519: 4505: 4219: 4187: 3822: 3731: 3645: 3580: 3327: 3301:. Miscellaneous Publication 99-1. Salt Lake City: Utah Geological Survey. pp. 345–353. 3270: 3172: 2991: 2748: 2574: 2533: 2424: 2360: 2327: 2002: 1935: 1854: 1771: 1734: 1504: 1147:
was a saurolophine (hadrosaurine of older references) hadrosaurid, a member of the duckbill
1057: 904: 5629: 5424: 5375: 5238: 5169: 5072: 4892: 4611: 4597: 4576: 4194: 4067: 3834: 3602: 3440: 2226:, from the Campanian of the East Coast of North America, with a reevaluation of the genus" 2192: 1708: 1658: 555: 466: 37: 2954:. Vol. Supplement 2. Jefferson, North Carolina: McFarland & Co. pp. 79–80. 1855: 3168: 2744: 2529: 2420: 1998: 880:
As of 2016, there are currently three named species that are recognized as valid today:
5639: 5634: 5572: 5509: 5382: 5231: 5194: 5176: 5121: 4753: 4569: 4562: 4491: 4413: 4400: 4328: 4316: 4285: 4260: 4124: 4046: 4007: 3993: 3870: 3781: 3703: 3681: 3674: 3415: 2947: 2781: 1889: 1850: 1743: 1684: 1669: 1551: 1017: 594: 590: 319: 225: 2045:(1919). "Preliminary description of a new species of trachodont dinosaur of the genus 1102:) at 8 metres (26 ft) in length and 3 metric tons (3.3 short tons) in body mass. 1050: 5810: 5579: 5523: 5502: 5473: 5389: 5326: 5224: 5217: 5203: 5113: 5093: 5086: 4983: 4933: 4926: 4910: 4885: 4858: 4851: 4844: 4815: 4766: 4728: 4693: 4674: 4646: 4590: 4253: 4246: 4201: 4131: 4000: 3859: 3844: 3806: 3717: 3610: 3573: 3566: 3282: 3239: 2760: 2728: 2575: 2444: 2372: 2351:(Dinosauria: Hadrosauridae) from the Upper Campanian Kaiparowits Formation of Utah". 2332: 2311: 1729: 1695: 1689: 1674: 1563: 1378: 1067: 996: 747: 481: 433: 374: 267: 254: 75: 3184: 3005: 2545: 1818:
Creisler, Benjamin S. (2007). "Deciphering duckbills". In Carpenter, Kenneth (ed.).
5487: 5463: 5451: 5417: 5410: 5361: 5354: 5160: 5104: 5079: 5051: 4940: 4742: 4707: 4653: 4583: 4555: 4441: 4434: 4274: 4239: 4166: 4159: 4081: 4022: 3977: 3879: 3753: 3746: 3695: 3659: 3617: 3467: 3406: 2364: 1978: 1919: 1717: 1713: 1700: 1679: 1665: 1557: 1534: 1354: 1216: 1202: 1162: 1153: 1004: 960: 893: 833: 790: 767: 598: 477: 302: 280: 209: 3128: 3104:"New Dinosaur Species, Gryposaurus Monumentensis, Unearthed In Utah | Science 2.0" 5734: 5676: 5565: 5530: 5368: 5210: 5183: 5135: 4961: 4947: 4919: 4899: 4878: 4822: 4801: 4794: 4787: 4721: 4714: 4667: 4632: 4604: 4540: 4533: 4302: 4152: 4138: 4110: 4088: 4030: 3893: 3724: 3688: 3652: 3548: 3535: 3494: 3364: 2702: 1779: 1704: 1496: 1473: 1395: 1240: 1167: 1121: 1009: 968: 838: 706: 696: 681:. Reviews beginning in the 1990s, however, called into question the identity of 603: 537: 524: 412: 241: 50: 124: 5618: 5593: 5586: 5558: 5551: 5442: 5431: 5317: 5016: 4996: 4954: 4829: 4681: 4548: 4498: 4463: 4456: 4427: 4267: 4145: 4117: 3986: 3959: 3934: 3774: 3767: 3760: 3638: 3317: 3176: 2537: 2493: 1763: 1751: 1639: 1627: 1481: 1110: 1079: 964: 932: 809: 746:
for a partial skull and skeleton and a second less-complete skeleton from the
719: 662: 617:
provided evidence of a different type of skull configuration, and by 1916 the
533: 95: 60: 20: 5725: 2907: 2436: 1495:. Its skull had special joints that permitted a grinding motion analogous to 516:
in Texas may extend the temporal range of the genus to 66 million years ago.
5600: 5516: 5495: 5333: 5030: 4968: 4477: 4449: 4309: 4173: 4103: 4074: 3900: 3886: 3629: 2681: 2084:(1st ed.). Berkeley: University of California Press. pp. 534–561. 1822:. Bloomington and Indianapolis: Indiana University Press. pp. 185–210. 1775: 1738: 1635: 1592: 1488: 1485: 1460: 1187: 1148: 944: 900: 855: 666: 637:'s naming of a nearly complete skeleton from the Dinosaur Park Formation as 601:
had collected and described a partial skull from New Mexico, which he named
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material. Horner, who described the specimens, considered it to be a less
5719: 5480: 5149: 5037: 4837: 4808: 4735: 4660: 4625: 4526: 4512: 4470: 4294: 4095: 4060: 3966: 3952: 3914: 3424: 3397: 3388: 2816:
Canada Department of Mines Geological Survey Bulletin (Geological Series)
1661: 786: 574: 529: 462: 193: 180: 90: 85: 70: 65: 55: 2752: 2428: 2006: 1983:"Cretaceous Eocene correlation in New Mexico, Wyoming, Montana, Alberta" 685:, which has limited material for comparison with other duckbills. Thus, 5767: 4773: 4208: 2220:
Prieto-Márquez, Alberto; Weishampel, David B.; Horner, John R. (2006).
1588: 763: 501: 493: 444: 105: 80: 2727:
Lehman, Thomas M.; Wick, Steve L.; Wagner, Jonathan R. (1 July 2016).
2516:(Dinosauria, Hadrosauridae), with a taxonomic revision of the genus". 2483: 2403:
Lehman, Thomas M.; Wick, Steven L.; Wagner, Jonathan R. (March 2016).
935:, as well as with two skeletons with skulls that had been assigned to 758:, although known from fragmentary material, can be distinguished from 4639: 4232: 3382: 2887: 1939: 1767: 1759: 1755: 1087: 858: 851: 742:
in a photo caption. Although Horner in 1979 used the new combination
440: 170: 5696: 1982: 808:. Its skull was more robust than that of the other species, and its 2581:(2nd ed.). Berkeley: University of California Press. pp.  1634:, a large inland sea that split North America into two landmasses, 979:
was listed second on the top 10 list of new species in 2008 by the
927:). Ten complete skulls and twelve fragmentary skulls are known for 804:, includes a skull and partial skeleton that represent the species 4484: 4014: 2615:
Dinosaur Provincial Park: A Spectacular Ancient Ecosystem Revealed
1602: 1572: 1542: 1508: 1500: 1492: 1459: 1132: 1120: 1049: 868: 695: 554: 544: 455: 448: 2512:
Prieto–Marquez, Alberto (2010). "The braincase and skull roof of
1742:. Other paleofauna present in the Kaiparowits Formation included 1034:. However, this synonymy was rejected in the 2004 edition of the 5754: 3852: 1747: 1643: 773:
Further research has revealed the presence of a second species,
509: 5700: 5305: 4398: 3533: 3375: 3337: 1175:, but it is slightly younger. Additionally, while the skull of 3333: 1963:. New Mexico Museum of Natural History and Science Bulletin, 496:, Canada, and two formations in the United States: the Lower 3275:
10.1671/0272-4634(2005)025[0897:ANOTMF]2.0.CO;2
971:"Hadrosauravus" is an early, unused name for this species. 959:, but farther forward on the snout, and its teeth are less 923:) and the form of the upper arm (longer and more robust in 621:
skull had been redone with a nasal arch and both Brown and
1861:. Jefferson, North Carolina: McFarland & Co. pp.  689:
has once again been separated, at least temporarily, from
581:
2278, a skull and partial skeleton collected in 1913 by
2347:
Gates, Terry; Sampson, Scott (2006). "A new species of
975:
is known from a skull and partial skeleton from Utah.
842:. However, in 2016, a possibly forth valid species of 543:
around 8 meters (26 feet) long, it may have preferred
3129:"Toothy dinosaur newest to come out of southern Utah" 1961:
Late Cretaceous vertebrates from the Western Interior
1820:
Horns and Beaks: Ceratopsian and Ornithopod Dinosaurs
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is absent from the upper part of the formation, with
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that lived about 80 to 75 million years ago, in the
381: 362: 347: 335: 309: 5709: 5610: 5461: 5441: 5316: 5193: 5159: 5103: 5062: 5006: 4982: 4909: 4868: 4752: 4691: 4411: 4327: 4284: 4218: 4044: 3976: 3932: 3869: 3833: 3792: 3628: 3591: 3546: 3493: 3466: 3439: 3021:"Plant-eating dinosaur a "Cretaceous weed whacker"" 2703:"Scientists Announce Top 10 New Species; Issue SOS" 2924:(1920). "The osteology of the trachodont dinosaur 2875:. Geological Society of America. pp. 110–117. 2617:. Bloomington: Indiana University Press. pp.  2025:United States Geological Survey Professional Paper 1928:Bulletin of the American Museum of Natural History 633:were one and the same. This idea was reflected in 2892:. Princeton University Press. pp. 337–339. 2707:International Institute for Species Exploration 2613:. In Currie, Phillip J.; Koppelhus Eva (eds.). 981:International Institute for Species Exploration 2871:. Geological Society of America Special Paper 2849:. Geological Society of America Special Paper 2257:. Geological Society of America Special Paper 2199:. New York: Crescent Books. pp. 116–121. 3349: 2930:University of Toronto Studies, Geology Series 2853:. Geological Society of America. p. 226. 2810:, with a description of the type specimen of 824:, indeterminate hadrosaur remains resembling 649:in its own genus). Direct comparison between 8: 2261:. Geological Society of America. p. 21. 677:question for nearly fifty years in favor of 5832:Grand Staircase–Escalante National Monument 2310:Gates, Terry A.; Sampson, Scott D. (2007). 2055:Transactions of the Royal Society of Canada 1611:(left background) and other animals of the 1503:were continually replacing and packed into 802:Grand Staircase–Escalante National Monument 593:. This specimen was described and named by 5822:Late Cretaceous dinosaurs of North America 5697: 5313: 5302: 4988: 4758: 4699: 4419: 4408: 4395: 4224: 4052: 3944: 3940: 3798: 3558: 3554: 3543: 3530: 3472: 3445: 3436: 3372: 3356: 3342: 3334: 2642:Lambert, David; the Diagram Group (1990). 2609:Ryan, Michael J.; Evans, David C. (2005). 2161:Lambert, David; the Diagram Group (1983). 1951: 1949: 734:), but identifies the mounted skeleton of 123: 27: 2995: 2978:Prieto-Márquez, A.; Wagner, J.R. (2011). 2973: 2971: 2459:"Current Research | Big Bend Conservancy" 2331: 2320:Zoological Journal of the Linnean Society 983:. Recently, a possible fourth species of 724:The Illustrated Encyclopedia of Dinosaurs 512:. A possible additional species from the 3150: 3148: 2565: 2563: 2561: 2559: 2557: 2555: 2507: 2505: 2503: 2276: 2274: 2272: 2270: 2268: 443:"; sometimes incorrectly translated as " 5882:Multispecific non-avian dinosaur genera 2869:Hadrosaurian Dinosaurs of North America 2847:Hadrosaurian Dinosaurs of North America 2604: 2602: 2392:(Thesis thesis). Texas Tech University. 2305: 2303: 2255:Hadrosaurian Dinosaurs of North America 2136:. Secaucus, NJ: Citadel Press. p.  2103: 2101: 1845: 1843: 1841: 1839: 1807: 3047:"Huge New Dinosaur Had A Serious Bite" 2889:The Princeton Field Guide to Dinosaurs 2786:Contributions to Canadian Paleontology 2290:Museum of the Rockies Occasional Paper 2072: 2070: 2068: 1987:Geological Society of America Bulletin 1884: 1882: 1813: 1811: 3197:Roberts EM, Deino AL, Chan MA (2005) 1171:has been proposed to be a synonym of 1165:, as used by Alberto Prieto-Márquez. 1022:was considered a possible species of 7: 1125:Size comparison of three species of 1370: 1346: 1287: 1263: 1256: 1232: 1208: 1197: 999:, along with an unnamed species of 3263:Journal of Vertebrate Paleontology 3201:. Cretaceous Research 26: 307–318. 2518:Journal of Vertebrate Paleontology 2353:Journal of Vertebrate Paleontology 779:American Museum of Natural History 754:in print. Current thought is that 14: 5862:Campanian genus first appearances 854:, who was the last leader of the 657:is hindered by the fact that the 5288: 5282: 5277: 5271: 5264: 4381: 4374: 4367: 3320: 2648:. New York: Avon Books. p.  2386:Wagner, Jonathan R. (May 2001). 2333:10.1111/j.1096-3642.2007.00349.x 2167:. New York: Avon Books. p.  1561:and the hollow-crested duckbill 1537:of the Dinosaur Park Formation, 147: 5645:List of informal dinosaur names 3299:Vertebrate Paleontology in Utah 3019:Dunham, Will (3 October 2007). 2678:"Re: What are these dinosaurs?" 2284:(1992). "Cranial morphology of 1013:, but with a more solid crest. 5655:Timeline of hadrosaur research 3074:"New duck-dinosaur discovered" 2365:10.1080/02724634.2006.10010069 1792:Timeline of hadrosaur research 1709:Parasaurolophus cyrtocristatus 1657:with other dinosaurs, such as 1624:Argon-argon radiometric dating 1583:The distinctive nasal arch of 1194:among the other kritosaurins: 1074:(now regarded as a synonym of 816:In Texas, specifically at the 585:from what is now known as the 137:Natural History Museum of Utah 1: 5877:Fossil taxa described in 1914 2988:Acta Palaeontologica Polonica 2233:Acta Palaeontologica Polonica 5827:Taxa named by Lawrence Lambe 3072:Than, Ker (3 October 2007). 1042:being tabulated as dubious. 1026:, following the synonymy of 949:Lower Two Medicine Formation 836:or at least some species of 5867:Campanian genus extinctions 2952:Dinosaurs: The Encyclopedia 2867:; Wright, Nelda E. (1942). 2845:; Wright, Nelda E. (1942). 2253:; Wright, Nelda E. (1942). 2133:The New Dinosaur Dictionary 1857:Dinosaurs: The Encyclopedia 1105:The three named species of 967:-like characteristics. The 730:for the Canadian material ( 710:is also the same as either 700:Skull, Royal Tyrrell Museum 5898: 3080:. NBC News. Archived from 2463:fossildiscoveryexhibit.com 2195:(1985). "Hadrosaurids I". 2164:A Field Guide to Dinosaurs 1591:skin, or that there was a 995:, which dates to the late 389:Lehman, Wick, Wagner, 2016 18: 5312: 5301: 5262: 4991: 4761: 4702: 4422: 4407: 4394: 4365: 4227: 4055: 3947: 3943: 3801: 3561: 3557: 3542: 3529: 3475: 3448: 3435: 3371: 3177:10.1017/S0094837300002165 2886:Paul, Gregory S. (2016). 2611:"Ornithischian Dinosaurs" 2538:10.1080/02724631003762971 1730:Kosmoceratops richardsoni 1651:Gryposaurus monumentensis 1392: 1375: 1368: 1351: 1344: 1313:Gryposaurus monumentensis 1309: 1292: 1285: 1268: 1261: 1254: 1237: 1230: 1213: 1206: 1019:Stephanosaurus marginatus 669:on hadrosaurs sealed the 403: 396: 332: 327: 308: 301: 144:Scientific classification 142: 131: 122: 30: 16:Extinct genus of reptiles 2926:Kritosaurus incurvimanus 2130:Glut, Donald F. (1982). 2051:Kritosaurus incurvimanus 1545:-influenced conditions. 1072:Kritosaurus incurvimanus 1016:The dubious hadrosaurid 1007:which closely resembles 913:Gryposaurus incurvimanus 909:Kritosaurus incurvimanus 850:, which was named after 651:Kritosaurus incurvimanus 643:Gryposaurus incurvimanus 639:Kritosaurus incurvimanus 432:(meaning "hooked-nosed ( 19:Not to be confused with 5852:Paleontology in Alberta 5842:Paleontology in Montana 2733:Journal of Paleontology 2409:Journal of Paleontology 2113:Journal of Paleontology 1853:(1997). "Gryposaurus". 1770:were present including 1632:Western Interior Seaway 991:, was unearthed in the 610:Edmontosaurus annectens 587:Dinosaur Park Formation 490:Dinosaur Park Formation 2806:(1924). "On the genus 2709:. Newswise. 2008-05-23 2645:The Dinosaur Data Book 1615: 1491:, eating a variety of 1469: 1141: 1130: 1060: 877: 796:New material from the 762:by differences in the 744:Hadrosaurus notabilis 701: 683:Kritosaurus navajovius 589:of Alberta, along the 567: 498:Two Medicine Formation 5857:Kaiparowits Formation 5789:Paleobiology Database 5625:Basturs Poble bonebed 2997:10.4202/app.2011.0049 2514:Gryposaurus notabilis 1902:The Ottawa Naturalist 1894:Gryposaurus notabilis 1766:. A variety of early 1724:Nasutoceratops titusi 1701:armored ankylosaurids 1613:Kaiparowits Formation 1606: 1535:bird-hipped dinosaurs 1463: 1296:Gryposaurus notabilis 1136: 1124: 1076:Gryposaurus notabilis 1053: 872: 798:Kaiparowits Formation 699: 655:Gryposaurus notabilis 647:Gryposaurus notabilis 558: 506:Kaiparowits Formation 314:Gryposaurus notabilis 5872:Ornithischian genera 5847:Paleontology in Utah 5647:(contains hadrosaur 5045:Probrachylophosaurus 2078:Weishampel, David B. 1797:2016 in paleontology 1705:duckbilled hadrosaur 1272:Gryposaurus latidens 822:El Picacho Formation 551:History of discovery 5837:Dinosaur Park fauna 3169:1975Pbio....1...21H 3084:on October 15, 2013 2982:Saurolophus morrisi 2865:Lull, Richard Swann 2843:Lull, Richard Swann 2812:Lambeosaurus lambei 2804:Gilmore, Charles W. 2753:10.1017/jpa.2016.48 2745:2016JPal...90..333L 2530:2010JVPal..30..838P 2482:Bertozzo, Filippo. 2429:10.1017/jpa.2016.48 2421:2016JPal...90..333L 2251:Lull, Richard Swann 2224:Hadrosaurus foulkii 2021:Gilmore, Charles W. 2007:10.1130/gsab-25-355 1999:1914GSAB...25..355B 1746:(sharks and rays), 1739:Hagryphus giganteus 1718:Utahceratops gettyi 1575:-related settings. 1571:may have preferred 1186:The following is a 1003:and an undescribed 989:Gryposaurus alsatei 915:), is a synonym of 785:is also known from 583:George F. Sternberg 488:are known from the 484:. Named species of 5397:Ornithopodichnites 5008:Brachylophosaurini 3593:Hypsilophodontidae 3236:September 18, 1996 3230:Clinton, William. 2782:Lambe, Lawrence M. 2312:"A new species of 1957:Kirkland, James I. 1898:Chasmosaurus belli 1890:Lambe, Lawrence M. 1680:Ornithomimus velox 1616: 1480:would have been a 1470: 1142: 1131: 1061: 993:Javelina Formation 878: 818:Javelina Formation 702: 645:(although he left 625:had proposed that 568: 514:Javelina Formation 5804: 5803: 5703:Taxon identifiers 5694: 5693: 5690: 5689: 5686: 5685: 5404:Shiraminesauropus 5297: 5296: 5260: 5259: 5256: 5255: 5252: 5251: 5248: 5247: 5143:Naashoibitosaurus 5064:Austrokritosauria 5024:Brachylophosaurus 4978: 4977: 4390: 4389: 4363: 4362: 4359: 4358: 4355: 4354: 4351: 4350: 4347: 4346: 4040: 4039: 3928: 3927: 3924: 3923: 3908:Transylvanosaurus 3794:Rhabdodontomorpha 3740:Notohypsilophodon 3711:Macrogryphosaurus 3525: 3524: 3521: 3520: 3517: 3516: 3513: 3512: 3308:978-1-55791-634-1 3242:on 28 August 2013 3108:www.science20.com 2961:978-0-89950-917-4 2922:Parks, William A. 2899:978-1-78684-190-2 2674:Olshevsky, George 2659:978-0-380-75896-8 2628:978-0-253-34595-0 2592:978-0-520-24209-8 2359:(3, Suppl): 65A. 2206:978-0-517-46890-6 2193:Norman, David. B. 2178:978-0-380-83519-5 2147:978-0-8065-0782-8 2091:978-0-520-06727-1 2043:Parks, William A. 1872:978-0-89950-917-4 1829:978-0-253-34817-3 1772:multituberculates 1735:oviraptorosaurian 1464:Restored head of 1452: 1451: 1443: 1442: 1434: 1433: 1425: 1424: 1416: 1415: 1407: 1406: 1333: 1332: 1324: 1323: 947:-early Campanian 566:), collected 1918 454:) lizard") was a 425: 424: 419: 390: 378: 359: 344: 5889: 5797: 5796: 5784: 5783: 5771: 5770: 5758: 5757: 5745: 5744: 5743: 5730: 5729: 5728: 5698: 5671:mummy S.M. R4036 5545:Microhadrosaurus 5538:Mandschurosaurus 5348:Camptosaurichnus 5341:Bonaparteichnium 5314: 5307:Related articles 5303: 5292: 5286: 5281: 5276: 5275: 5268: 4989: 4870:Parasaurolophini 4781:Angulomastacator 4759: 4700: 4619:Shuangmiaosaurus 4520:Huehuecanauhtlus 4506:Gongpoquansaurus 4420: 4409: 4396: 4385: 4379: 4378: 4371: 4225: 4220:Hadrosauriformes 4188:Ratchasimasaurus 4053: 3945: 3941: 3823:Muttaburrasaurus 3799: 3732:Muttaburrasaurus 3646:Atlascopcosaurus 3581:Thescelosauridae 3559: 3555: 3544: 3531: 3473: 3446: 3437: 3430: 3429: 3373: 3358: 3351: 3344: 3335: 3330: 3328:Dinosaurs portal 3325: 3324: 3323: 3313: 3312: 3293: 3287: 3286: 3258: 3252: 3251: 3249: 3247: 3238:. Archived from 3227: 3221: 3218: 3212: 3208: 3202: 3195: 3189: 3188: 3152: 3143: 3142: 3140: 3139: 3125: 3119: 3118: 3116: 3115: 3100: 3094: 3093: 3091: 3089: 3069: 3063: 3062: 3060: 3058: 3051:ScienceDaily.com 3043: 3037: 3036: 3034: 3032: 3016: 3010: 3009: 2999: 2975: 2966: 2965: 2944: 2938: 2937: 2918: 2912: 2911: 2883: 2877: 2876: 2861: 2855: 2854: 2839: 2833: 2830: 2824: 2823: 2800: 2794: 2793: 2778: 2772: 2771: 2769: 2767: 2724: 2718: 2717: 2715: 2714: 2699: 2693: 2692: 2690: 2689: 2680:. Archived from 2670: 2664: 2663: 2639: 2633: 2632: 2606: 2597: 2596: 2580: 2567: 2550: 2549: 2509: 2498: 2497: 2479: 2473: 2472: 2470: 2469: 2455: 2449: 2448: 2400: 2394: 2393: 2383: 2377: 2376: 2344: 2338: 2337: 2335: 2307: 2298: 2297: 2278: 2263: 2262: 2247: 2241: 2240: 2230: 2217: 2211: 2210: 2189: 2183: 2182: 2158: 2152: 2151: 2127: 2121: 2120: 2105: 2096: 2095: 2074: 2063: 2062: 2039: 2033: 2032: 2017: 2011: 2010: 1975: 1969: 1968: 1953: 1944: 1943: 1916: 1910: 1909: 1886: 1877: 1876: 1860: 1847: 1834: 1833: 1815: 1655:paleoenvironment 1638:to the west and 1609:G. monumentensis 1505:dental batteries 1371: 1347: 1288: 1264: 1257: 1233: 1209: 1198: 1100:G. monumentensis 1082:impressions for 1058:life restoration 977:G. monumentensis 973:G. monumentensis 905:Upper Cretaceous 890:G. monumentensis 874:G. monumentensis 806:G. monumentensis 718:, or both. This 409: 388: 383: 369: 366:G. monumentensis 364: 354: 349: 342: 337: 311: 292: 279: 266: 253: 240: 224: 208: 152: 151: 135:skeleton in the 133:G. monumentensis 127: 117: 47: 36:Temporal range: 28: 5897: 5896: 5892: 5891: 5890: 5888: 5887: 5886: 5807: 5806: 5805: 5800: 5792: 5787: 5779: 5774: 5766: 5761: 5753: 5748: 5739: 5738: 5733: 5724: 5723: 5718: 5705: 5695: 5682: 5663:mummy AMNH 5060 5630:Dakota (fossil) 5606: 5457: 5437: 5425:Tetrapodosaurus 5376:Hadrosaurichnus 5308: 5293: 5270: 5244: 5239:Shantungosaurus 5189: 5170:Augustynolophus 5155: 5099: 5073:Bonapartesaurus 5058: 5002: 4974: 4905: 4893:Parasaurolophus 4864: 4748: 4696: 4687: 4612:Qianjiangsaurus 4598:Probactrosaurus 4577:Penelopognathus 4416: 4403: 4386: 4373: 4343: 4323: 4280: 4214: 4195:Riabininohadros 4068:Bayannurosaurus 4049: 4036: 3972: 3937: 3920: 3865: 3835:Tenontosauridae 3829: 3788: 3624: 3603:Gideonmantellia 3587: 3551: 3538: 3509: 3489: 3462: 3456:Avemetatarsalia 3441:Avemetatarsalia 3431: 3378: 3377: 3367: 3362: 3326: 3321: 3319: 3316: 3309: 3295: 3294: 3290: 3260: 3259: 3255: 3245: 3243: 3229: 3228: 3224: 3219: 3215: 3209: 3205: 3196: 3192: 3154: 3153: 3146: 3137: 3135: 3127: 3126: 3122: 3113: 3111: 3102: 3101: 3097: 3087: 3085: 3071: 3070: 3066: 3056: 3054: 3053:. Science Daily 3045: 3044: 3040: 3030: 3028: 3018: 3017: 3013: 2977: 2976: 2969: 2962: 2948:Glut, Donald F. 2946: 2945: 2941: 2920: 2919: 2915: 2900: 2885: 2884: 2880: 2863: 2862: 2858: 2841: 2840: 2836: 2831: 2827: 2802: 2801: 2797: 2780: 2779: 2775: 2765: 2763: 2726: 2725: 2721: 2712: 2710: 2701: 2700: 2696: 2687: 2685: 2672: 2671: 2667: 2660: 2641: 2640: 2636: 2629: 2608: 2607: 2600: 2593: 2571:Horner, John R. 2569: 2568: 2553: 2511: 2510: 2501: 2481: 2480: 2476: 2467: 2465: 2457: 2456: 2452: 2402: 2401: 2397: 2385: 2384: 2380: 2346: 2345: 2341: 2309: 2308: 2301: 2282:Horner, John R. 2280: 2279: 2266: 2249: 2248: 2244: 2228: 2219: 2218: 2214: 2207: 2191: 2190: 2186: 2179: 2160: 2159: 2155: 2148: 2129: 2128: 2124: 2109:Horner, John R. 2107: 2106: 2099: 2092: 2076: 2075: 2066: 2041: 2040: 2036: 2019: 2018: 2014: 1977: 1976: 1972: 1955: 1954: 1947: 1934:(24): 267–274. 1918: 1917: 1913: 1888: 1887: 1880: 1873: 1851:Glut, Donald F. 1849: 1848: 1837: 1830: 1817: 1816: 1809: 1805: 1788: 1744:chondrichthyans 1621: 1607:Restoration of 1601: 1581: 1458: 1453: 1444: 1435: 1426: 1417: 1408: 1334: 1325: 1119: 1048: 937:K. incurvimanus 925:K. incurvimanus 911:(also known as 867: 736:K. incurvimanus 623:Charles Gilmore 564:G. incurvimanus 553: 467:Late Cretaceous 417: 387: 368: 353: 341: 323: 317: 290: 277: 264: 251: 238: 222: 206: 146: 118: 116: 115: 114: 113: 108: 103: 98: 93: 88: 83: 78: 73: 68: 63: 58: 53: 42: 41: 38:Late Cretaceous 34: 24: 17: 12: 11: 5: 5895: 5893: 5885: 5884: 5879: 5874: 5869: 5864: 5859: 5854: 5849: 5844: 5839: 5834: 5829: 5824: 5819: 5809: 5808: 5802: 5801: 5799: 5798: 5785: 5772: 5759: 5746: 5731: 5715: 5713: 5707: 5706: 5701: 5692: 5691: 5688: 5687: 5684: 5683: 5681: 5680: 5673: 5665: 5657: 5652: 5642: 5640:Hypsilophodont 5637: 5635:Hadrosaur diet 5632: 5627: 5622: 5614: 5612: 5611:Other articles 5608: 5607: 5605: 5604: 5597: 5590: 5583: 5576: 5573:Stephanosaurus 5569: 5562: 5555: 5548: 5541: 5534: 5527: 5520: 5513: 5510:Fulgurotherium 5506: 5499: 5492: 5484: 5477: 5469: 5467: 5459: 5458: 5456: 5455: 5447: 5445: 5439: 5438: 5436: 5435: 5428: 5421: 5414: 5407: 5400: 5393: 5386: 5383:Iguanodontipus 5379: 5372: 5365: 5358: 5351: 5344: 5337: 5330: 5322: 5320: 5310: 5309: 5306: 5299: 5298: 5295: 5294: 5263: 5261: 5258: 5257: 5254: 5253: 5250: 5249: 5246: 5245: 5243: 5242: 5235: 5232:Laiyangosaurus 5228: 5221: 5214: 5207: 5199: 5197: 5195:Edmontosaurini 5191: 5190: 5188: 5187: 5180: 5177:Prosaurolophus 5173: 5165: 5163: 5157: 5156: 5154: 5153: 5146: 5139: 5132: 5125: 5122:Coahuilasaurus 5118: 5109: 5107: 5101: 5100: 5098: 5097: 5090: 5083: 5076: 5068: 5066: 5060: 5059: 5057: 5056: 5048: 5041: 5034: 5027: 5020: 5012: 5010: 5004: 5003: 5001: 5000: 4992: 4986: 4980: 4979: 4976: 4975: 4973: 4972: 4965: 4958: 4951: 4944: 4937: 4930: 4923: 4915: 4913: 4907: 4906: 4904: 4903: 4896: 4889: 4882: 4874: 4872: 4866: 4865: 4863: 4862: 4855: 4848: 4841: 4834: 4826: 4819: 4812: 4805: 4798: 4791: 4784: 4777: 4770: 4762: 4756: 4754:Lambeosaurinae 4750: 4749: 4747: 4746: 4739: 4732: 4725: 4718: 4711: 4703: 4697: 4692: 4689: 4688: 4686: 4685: 4678: 4671: 4664: 4657: 4650: 4643: 4636: 4629: 4622: 4615: 4608: 4601: 4594: 4587: 4580: 4573: 4570:Nanyangosaurus 4566: 4563:Nanningosaurus 4559: 4552: 4545: 4537: 4530: 4523: 4516: 4509: 4502: 4495: 4492:Gilmoreosaurus 4488: 4481: 4474: 4467: 4460: 4453: 4446: 4438: 4431: 4423: 4417: 4414:Hadrosauroidea 4412: 4405: 4404: 4401:Hadrosauroidea 4399: 4392: 4391: 4388: 4387: 4366: 4364: 4361: 4360: 4357: 4356: 4353: 4352: 4349: 4348: 4345: 4344: 4342: 4341: 4340: 4339: 4333: 4331: 4329:Hadrosauroidea 4325: 4324: 4322: 4321: 4317:Mantellisaurus 4313: 4306: 4299: 4290: 4288: 4286:Iguanodontidae 4282: 4281: 4279: 4278: 4271: 4264: 4261:Mantellisaurus 4257: 4250: 4243: 4236: 4228: 4222: 4216: 4215: 4213: 4212: 4205: 4198: 4191: 4184: 4177: 4170: 4163: 4156: 4149: 4142: 4135: 4128: 4125:Iguanacolossus 4121: 4114: 4107: 4100: 4092: 4085: 4078: 4071: 4064: 4056: 4050: 4047:Ankylopollexia 4045: 4042: 4041: 4038: 4037: 4035: 4034: 4027: 4019: 4011: 4008:Eousdryosaurus 4004: 3997: 3994:Dysalotosaurus 3990: 3982: 3980: 3974: 3973: 3971: 3970: 3963: 3956: 3948: 3938: 3933: 3930: 3929: 3926: 3925: 3922: 3921: 3919: 3918: 3911: 3904: 3897: 3890: 3883: 3875: 3873: 3871:Rhabdodontidae 3867: 3866: 3864: 3863: 3856: 3849: 3840: 3838: 3831: 3830: 3828: 3827: 3819: 3811: 3802: 3796: 3790: 3789: 3787: 3786: 3782:Weewarrasaurus 3778: 3771: 3764: 3757: 3750: 3743: 3736: 3728: 3721: 3714: 3707: 3704:Leaellynasaura 3700: 3692: 3685: 3682:Gasparinisaura 3678: 3675:Galleonosaurus 3671: 3663: 3656: 3649: 3642: 3634: 3632: 3626: 3625: 3623: 3622: 3614: 3607: 3598: 3596: 3589: 3588: 3586: 3585: 3577: 3570: 3562: 3552: 3547: 3540: 3539: 3534: 3527: 3526: 3523: 3522: 3519: 3518: 3515: 3514: 3511: 3510: 3508: 3507: 3506: 3505: 3499: 3497: 3491: 3490: 3488: 3487: 3486: 3485: 3476: 3470: 3464: 3463: 3461: 3460: 3459: 3458: 3449: 3443: 3433: 3432: 3428: 3427: 3418: 3416:Neornithischia 3409: 3400: 3391: 3385: 3376: 3369: 3368: 3363: 3361: 3360: 3353: 3346: 3338: 3332: 3331: 3315: 3314: 3307: 3288: 3269:(4): 897–904. 3253: 3222: 3213: 3203: 3190: 3144: 3120: 3095: 3064: 3038: 3011: 2967: 2960: 2939: 2913: 2898: 2878: 2856: 2834: 2825: 2808:Stephanosaurus 2795: 2773: 2739:(2): 333–356. 2719: 2694: 2676:(1999-11-16). 2665: 2658: 2634: 2627: 2598: 2591: 2577:The Dinosauria 2551: 2524:(3): 838–854. 2499: 2474: 2450: 2415:(2): 333–356. 2395: 2378: 2339: 2326:(2): 351–376. 2299: 2286:Prosaurolophus 2264: 2242: 2222:"The dinosaur 2212: 2205: 2184: 2177: 2153: 2146: 2122: 2097: 2090: 2082:The Dinosauria 2064: 2034: 2012: 1993:(1): 355–380. 1970: 1945: 1911: 1908:(11): 145–155. 1878: 1871: 1835: 1828: 1806: 1804: 1801: 1800: 1799: 1794: 1787: 1784: 1685:tyrannosaurids 1670:Talos sampsoni 1620: 1617: 1600: 1597: 1580: 1577: 1552:Prosaurolophus 1457: 1454: 1450: 1449: 1446: 1445: 1441: 1440: 1437: 1436: 1432: 1431: 1428: 1427: 1423: 1422: 1419: 1418: 1414: 1413: 1410: 1409: 1405: 1404: 1401: 1400: 1391: 1388: 1387: 1384: 1383: 1374: 1369: 1367: 1364: 1363: 1360: 1359: 1350: 1345: 1343: 1340: 1339: 1336: 1335: 1331: 1330: 1327: 1326: 1322: 1321: 1318: 1317: 1308: 1305: 1304: 1301: 1300: 1291: 1286: 1284: 1281: 1280: 1277: 1276: 1267: 1262: 1260: 1255: 1253: 1250: 1249: 1246: 1245: 1236: 1231: 1229: 1226: 1225: 1222: 1221: 1212: 1207: 1205: 1196: 1118: 1117:Classification 1115: 1047: 1044: 1040:Stephanosaurus 953:Pondera County 945:late Santonian 901:late Campanian 866: 863: 595:Lawrence Lambe 591:Red Deer River 552: 549: 522:is similar to 423: 422: 421: 420: 401: 400: 394: 393: 392: 391: 379: 360: 345: 330: 329: 325: 324: 318: 306: 305: 299: 298: 288: 284: 283: 275: 271: 270: 262: 258: 257: 249: 245: 244: 236: 229: 228: 226:Neornithischia 220: 213: 212: 204: 197: 196: 191: 184: 183: 178: 174: 173: 168: 164: 163: 158: 154: 153: 140: 139: 129: 128: 120: 119: 111: 110: 109: 104: 99: 94: 89: 84: 79: 74: 69: 64: 59: 54: 49: 48: 35: 15: 13: 10: 9: 6: 4: 3: 2: 5894: 5883: 5880: 5878: 5875: 5873: 5870: 5868: 5865: 5863: 5860: 5858: 5855: 5853: 5850: 5848: 5845: 5843: 5840: 5838: 5835: 5833: 5830: 5828: 5825: 5823: 5820: 5818: 5817:Saurolophines 5815: 5814: 5812: 5795: 5790: 5786: 5782: 5777: 5773: 5769: 5764: 5760: 5756: 5751: 5747: 5742: 5736: 5732: 5727: 5721: 5717: 5716: 5714: 5712: 5708: 5704: 5699: 5679: 5678: 5674: 5672: 5670: 5669:Edmontosaurus 5666: 5664: 5662: 5661:Edmontosaurus 5658: 5656: 5653: 5650: 5646: 5643: 5641: 5638: 5636: 5633: 5631: 5628: 5626: 5623: 5621: 5620: 5616: 5615: 5613: 5609: 5603: 5602: 5598: 5596: 5595: 5591: 5589: 5588: 5584: 5582: 5581: 5580:Syngonosaurus 5577: 5575: 5574: 5570: 5568: 5567: 5563: 5561: 5560: 5556: 5554: 5553: 5549: 5547: 5546: 5542: 5540: 5539: 5535: 5533: 5532: 5528: 5526: 5525: 5524:Koutalisaurus 5521: 5519: 5518: 5514: 5512: 5511: 5507: 5505: 5504: 5503:Eucercosaurus 5500: 5498: 5497: 5493: 5490: 5489: 5485: 5483: 5482: 5478: 5476: 5475: 5474:Arstanosaurus 5471: 5470: 5468: 5466: 5465: 5460: 5454: 5453: 5449: 5448: 5446: 5444: 5440: 5434: 5433: 5429: 5427: 5426: 5422: 5420: 5419: 5415: 5413: 5412: 5408: 5406: 5405: 5401: 5399: 5398: 5394: 5392: 5391: 5390:Orcauichnites 5387: 5385: 5384: 5380: 5378: 5377: 5373: 5371: 5370: 5366: 5364: 5363: 5359: 5357: 5356: 5352: 5350: 5349: 5345: 5343: 5342: 5338: 5336: 5335: 5331: 5329: 5328: 5327:Amblydactylus 5324: 5323: 5321: 5319: 5315: 5311: 5304: 5300: 5291: 5287: 5285: 5280: 5274: 5267: 5241: 5240: 5236: 5234: 5233: 5229: 5227: 5226: 5225:Kundurosaurus 5222: 5220: 5219: 5218:Kerberosaurus 5215: 5213: 5212: 5208: 5206: 5205: 5204:Edmontosaurus 5201: 5200: 5198: 5196: 5192: 5186: 5185: 5181: 5179: 5178: 5174: 5172: 5171: 5167: 5166: 5164: 5162: 5158: 5152: 5151: 5147: 5145: 5144: 5140: 5138: 5137: 5133: 5131: 5130: 5126: 5124: 5123: 5119: 5116: 5115: 5114:Anasazisaurus 5111: 5110: 5108: 5106: 5102: 5096: 5095: 5094:Secernosaurus 5091: 5089: 5088: 5087:Kelumapusaura 5084: 5082: 5081: 5077: 5075: 5074: 5070: 5069: 5067: 5065: 5061: 5054: 5053: 5049: 5047: 5046: 5042: 5040: 5039: 5035: 5033: 5032: 5028: 5026: 5025: 5021: 5019: 5018: 5014: 5013: 5011: 5009: 5005: 4999: 4998: 4994: 4993: 4990: 4987: 4985: 4984:Saurolophinae 4981: 4971: 4970: 4966: 4964: 4963: 4959: 4957: 4956: 4952: 4950: 4949: 4945: 4943: 4942: 4938: 4936: 4935: 4934:Hypacrosaurus 4931: 4929: 4928: 4927:Corythosaurus 4924: 4922: 4921: 4917: 4916: 4914: 4912: 4911:Lambeosaurini 4908: 4902: 4901: 4897: 4895: 4894: 4890: 4888: 4887: 4886:Charonosaurus 4883: 4881: 4880: 4876: 4875: 4873: 4871: 4867: 4861: 4860: 4859:Tsintaosaurus 4856: 4854: 4853: 4852:Pararhabdodon 4849: 4847: 4846: 4845:Nipponosaurus 4842: 4840: 4839: 4835: 4832: 4831: 4827: 4825: 4824: 4820: 4818: 4817: 4816:Jaxartosaurus 4813: 4811: 4810: 4806: 4804: 4803: 4799: 4797: 4796: 4792: 4790: 4789: 4785: 4783: 4782: 4778: 4776: 4775: 4771: 4769: 4768: 4767:Adynomosaurus 4764: 4763: 4760: 4757: 4755: 4751: 4745: 4744: 4740: 4738: 4737: 4733: 4731: 4730: 4729:Lapampasaurus 4726: 4724: 4723: 4719: 4717: 4716: 4712: 4710: 4709: 4705: 4704: 4701: 4698: 4695: 4694:Hadrosauridae 4690: 4684: 4683: 4679: 4677: 4676: 4675:Zhanghenglong 4672: 4670: 4669: 4665: 4663: 4662: 4658: 4656: 4655: 4651: 4649: 4648: 4647:Telmatosaurus 4644: 4642: 4641: 4637: 4635: 4634: 4630: 4628: 4627: 4623: 4621: 4620: 4616: 4614: 4613: 4609: 4607: 4606: 4602: 4600: 4599: 4595: 4593: 4592: 4591:Portellsaurus 4588: 4586: 4585: 4581: 4579: 4578: 4574: 4572: 4571: 4567: 4565: 4564: 4560: 4558: 4557: 4553: 4551: 4550: 4546: 4543: 4542: 4538: 4536: 4535: 4531: 4529: 4528: 4524: 4522: 4521: 4517: 4515: 4514: 4510: 4508: 4507: 4503: 4501: 4500: 4496: 4494: 4493: 4489: 4487: 4486: 4482: 4480: 4479: 4475: 4473: 4472: 4468: 4466: 4465: 4461: 4459: 4458: 4454: 4452: 4451: 4447: 4444: 4443: 4439: 4437: 4436: 4432: 4430: 4429: 4425: 4424: 4421: 4418: 4415: 4410: 4406: 4402: 4397: 4393: 4384: 4380: 4377: 4370: 4337: 4336: 4335: 4334: 4332: 4330: 4326: 4319: 4318: 4314: 4312: 4311: 4307: 4305: 4304: 4300: 4297: 4296: 4292: 4291: 4289: 4287: 4283: 4277: 4276: 4272: 4270: 4269: 4265: 4263: 4262: 4258: 4256: 4255: 4254:Jinzhousaurus 4251: 4249: 4248: 4247:Hypselospinus 4244: 4242: 4241: 4237: 4235: 4234: 4230: 4229: 4226: 4223: 4221: 4217: 4211: 4210: 4206: 4204: 4203: 4202:Theiophytalia 4199: 4197: 4196: 4192: 4190: 4189: 4185: 4183: 4182: 4178: 4176: 4175: 4171: 4169: 4168: 4164: 4162: 4161: 4157: 4155: 4154: 4150: 4148: 4147: 4143: 4141: 4140: 4136: 4134: 4133: 4132:Lanzhousaurus 4129: 4127: 4126: 4122: 4120: 4119: 4115: 4113: 4112: 4108: 4106: 4105: 4101: 4098: 4097: 4093: 4091: 4090: 4086: 4084: 4083: 4079: 4077: 4076: 4072: 4070: 4069: 4065: 4063: 4062: 4058: 4057: 4054: 4051: 4048: 4043: 4033: 4032: 4028: 4025: 4024: 4020: 4017: 4016: 4012: 4010: 4009: 4005: 4003: 4002: 4001:Elrhazosaurus 3998: 3996: 3995: 3991: 3989: 3988: 3984: 3983: 3981: 3979: 3975: 3969: 3968: 3964: 3962: 3961: 3957: 3955: 3954: 3950: 3949: 3946: 3942: 3939: 3936: 3931: 3917: 3916: 3912: 3910: 3909: 3905: 3903: 3902: 3898: 3896: 3895: 3891: 3889: 3888: 3884: 3882: 3881: 3877: 3876: 3874: 3872: 3868: 3862: 3861: 3860:Tenontosaurus 3857: 3855: 3854: 3850: 3847: 3846: 3845:Convolosaurus 3842: 3841: 3839: 3836: 3832: 3825: 3824: 3820: 3817: 3816: 3812: 3809: 3808: 3807:Ampelognathus 3804: 3803: 3800: 3797: 3795: 3791: 3784: 3783: 3779: 3777: 3776: 3772: 3770: 3769: 3765: 3763: 3762: 3758: 3756: 3755: 3751: 3749: 3748: 3744: 3742: 3741: 3737: 3734: 3733: 3729: 3727: 3726: 3722: 3720: 3719: 3718:Mahuidacursor 3715: 3713: 3712: 3708: 3706: 3705: 3701: 3698: 3697: 3693: 3691: 3690: 3686: 3684: 3683: 3679: 3677: 3676: 3672: 3669: 3668: 3664: 3662: 3661: 3657: 3655: 3654: 3650: 3648: 3647: 3643: 3641: 3640: 3636: 3635: 3633: 3631: 3627: 3620: 3619: 3615: 3613: 3612: 3611:Hypsilophodon 3608: 3605: 3604: 3600: 3599: 3597: 3594: 3590: 3583: 3582: 3578: 3576: 3575: 3574:Callovosaurus 3571: 3569: 3568: 3567:Burianosaurus 3564: 3563: 3560: 3556: 3553: 3550: 3545: 3541: 3537: 3532: 3528: 3503: 3502: 3501: 3500: 3498: 3496: 3492: 3484: 3480: 3479: 3478: 3477: 3474: 3471: 3469: 3465: 3457: 3453: 3452: 3451: 3450: 3447: 3444: 3442: 3438: 3434: 3426: 3422: 3419: 3417: 3413: 3410: 3408: 3404: 3401: 3399: 3395: 3392: 3390: 3386: 3384: 3380: 3379: 3374: 3370: 3366: 3359: 3354: 3352: 3347: 3345: 3340: 3339: 3336: 3329: 3318: 3310: 3304: 3300: 3292: 3289: 3284: 3280: 3276: 3272: 3268: 3264: 3257: 3254: 3241: 3237: 3233: 3226: 3223: 3217: 3214: 3207: 3204: 3200: 3194: 3191: 3186: 3182: 3178: 3174: 3170: 3166: 3162: 3158: 3151: 3149: 3145: 3134: 3130: 3124: 3121: 3109: 3105: 3099: 3096: 3083: 3079: 3075: 3068: 3065: 3052: 3048: 3042: 3039: 3026: 3022: 3015: 3012: 3007: 3003: 2998: 2993: 2989: 2985: 2983: 2974: 2972: 2968: 2963: 2957: 2953: 2949: 2943: 2940: 2935: 2931: 2927: 2923: 2917: 2914: 2909: 2905: 2901: 2895: 2891: 2890: 2882: 2879: 2874: 2870: 2866: 2860: 2857: 2852: 2848: 2844: 2838: 2835: 2829: 2826: 2821: 2817: 2813: 2809: 2805: 2799: 2796: 2791: 2787: 2783: 2777: 2774: 2762: 2758: 2754: 2750: 2746: 2742: 2738: 2734: 2730: 2723: 2720: 2708: 2704: 2698: 2695: 2684:on 2008-06-26 2683: 2679: 2675: 2669: 2666: 2661: 2655: 2651: 2647: 2646: 2638: 2635: 2630: 2624: 2620: 2616: 2612: 2605: 2603: 2599: 2594: 2588: 2584: 2579: 2578: 2572: 2566: 2564: 2562: 2560: 2558: 2556: 2552: 2547: 2543: 2539: 2535: 2531: 2527: 2523: 2519: 2515: 2508: 2506: 2504: 2500: 2495: 2491: 2487: 2486: 2478: 2475: 2464: 2460: 2454: 2451: 2446: 2442: 2438: 2434: 2430: 2426: 2422: 2418: 2414: 2410: 2406: 2399: 2396: 2391: 2390: 2382: 2379: 2374: 2370: 2366: 2362: 2358: 2354: 2350: 2343: 2340: 2334: 2329: 2325: 2321: 2317: 2315: 2306: 2304: 2300: 2295: 2291: 2287: 2283: 2277: 2275: 2273: 2271: 2269: 2265: 2260: 2256: 2252: 2246: 2243: 2238: 2234: 2227: 2225: 2216: 2213: 2208: 2202: 2198: 2194: 2188: 2185: 2180: 2174: 2170: 2166: 2165: 2157: 2154: 2149: 2143: 2139: 2135: 2134: 2126: 2123: 2119:(3): 566–577. 2118: 2114: 2110: 2104: 2102: 2098: 2093: 2087: 2083: 2079: 2073: 2071: 2069: 2065: 2060: 2056: 2052: 2048: 2044: 2038: 2035: 2030: 2026: 2022: 2016: 2013: 2008: 2004: 2000: 1996: 1992: 1988: 1984: 1980: 1979:Brown, Barnum 1974: 1971: 1966: 1962: 1958: 1952: 1950: 1946: 1941: 1937: 1933: 1929: 1925: 1921: 1920:Brown, Barnum 1915: 1912: 1907: 1903: 1899: 1895: 1891: 1885: 1883: 1879: 1874: 1868: 1864: 1859: 1858: 1852: 1846: 1844: 1842: 1840: 1836: 1831: 1825: 1821: 1814: 1812: 1808: 1802: 1798: 1795: 1793: 1790: 1789: 1785: 1783: 1781: 1780:insectivorans 1777: 1773: 1769: 1765: 1761: 1757: 1753: 1749: 1745: 1741: 1740: 1736: 1732: 1731: 1726: 1725: 1720: 1719: 1715: 1711: 1710: 1706: 1702: 1698: 1697: 1696:Teratophoneus 1692: 1691: 1690:Albertosaurus 1686: 1682: 1681: 1676: 1675:ornithomimids 1672: 1671: 1667: 1663: 1660: 1659:dromaeosaurid 1656: 1652: 1648: 1645: 1641: 1637: 1633: 1629: 1625: 1618: 1614: 1610: 1605: 1598: 1596: 1594: 1593:cartilaginous 1590: 1586: 1578: 1576: 1574: 1570: 1566: 1565: 1564:Corythosaurus 1560: 1559: 1554: 1553: 1548: 1544: 1540: 1536: 1531: 1529: 1525: 1521: 1517: 1516:monumentensis 1514: 1510: 1506: 1502: 1498: 1494: 1490: 1487: 1483: 1479: 1475: 1467: 1462: 1455: 1448: 1447: 1439: 1438: 1430: 1429: 1421: 1420: 1412: 1411: 1403: 1402: 1399: 1398: 1397: 1390: 1389: 1386: 1385: 1382: 1381: 1380: 1379:Secernosaurus 1373: 1372: 1366: 1365: 1362: 1361: 1358: 1357: 1356: 1349: 1348: 1342: 1341: 1338: 1337: 1329: 1328: 1320: 1319: 1316: 1315: 1314: 1307: 1306: 1303: 1302: 1299: 1298: 1297: 1290: 1289: 1283: 1282: 1279: 1278: 1275: 1274: 1273: 1266: 1265: 1259: 1258: 1252: 1251: 1248: 1247: 1244: 1243: 1242: 1235: 1234: 1228: 1227: 1224: 1223: 1220: 1219: 1218: 1211: 1210: 1204: 1200: 1199: 1195: 1193: 1189: 1184: 1182: 1178: 1174: 1170: 1169: 1164: 1160: 1156: 1155: 1150: 1146: 1139: 1135: 1128: 1123: 1116: 1114: 1112: 1108: 1103: 1101: 1097: 1093: 1089: 1085: 1081: 1077: 1073: 1069: 1068:type specimen 1065: 1059: 1056: 1052: 1045: 1043: 1041: 1037: 1033: 1029: 1025: 1021: 1020: 1014: 1012: 1011: 1006: 1002: 998: 997:Maastrichtian 994: 990: 986: 982: 978: 974: 970: 969:informal name 966: 963:, reflecting 962: 958: 954: 950: 946: 942: 938: 934: 930: 926: 922: 918: 914: 910: 906: 902: 898: 895: 891: 887: 883: 875: 871: 864: 862: 860: 857: 853: 849: 845: 841: 840: 835: 831: 827: 823: 819: 814: 811: 807: 803: 799: 794: 792: 788: 784: 780: 776: 771: 769: 765: 761: 757: 753: 749: 748:Bearpaw Shale 745: 741: 737: 733: 729: 725: 721: 717: 713: 709: 708: 698: 694: 692: 688: 684: 680: 676: 672: 668: 664: 660: 656: 652: 648: 644: 640: 636: 635:William Parks 632: 628: 624: 620: 616: 612: 611: 606: 605: 600: 596: 592: 588: 584: 580: 576: 572: 565: 561: 557: 550: 548: 546: 542: 539: 535: 531: 527: 526: 521: 517: 515: 511: 507: 503: 499: 495: 491: 487: 483: 482:North America 479: 476: 472: 468: 464: 461: 457: 453: 450: 446: 442: 438: 435: 431: 430: 418:Lambert, 1990 415: 414: 408: 407:Hadrosauravus 405: 404: 402: 399: 395: 386: 380: 376: 372: 367: 361: 357: 352: 346: 340: 334: 333: 331: 326: 321: 316: 315: 307: 304: 300: 297: 296: 289: 286: 285: 282: 276: 273: 272: 269: 268:Saurolophinae 263: 260: 259: 256: 255:Hadrosauridae 250: 247: 246: 243: 237: 234: 231: 230: 227: 221: 218: 215: 214: 211: 205: 202: 199: 198: 195: 192: 189: 186: 185: 182: 179: 176: 175: 172: 169: 166: 165: 162: 159: 156: 155: 150: 145: 141: 138: 134: 130: 126: 121: 107: 102: 97: 92: 87: 82: 77: 72: 67: 62: 57: 52: 46: 39: 33: 29: 26: 22: 5710: 5675: 5668: 5660: 5648: 5617: 5599: 5592: 5585: 5578: 5571: 5564: 5557: 5550: 5543: 5536: 5529: 5522: 5515: 5508: 5501: 5494: 5488:Claorhynchus 5486: 5479: 5472: 5464:Nomina dubia 5462: 5452:Guegoolithus 5450: 5430: 5423: 5418:Staurichnium 5416: 5411:Sousaichnium 5409: 5402: 5395: 5388: 5381: 5374: 5367: 5362:Dakotasaurus 5360: 5355:Caririchnium 5353: 5346: 5339: 5332: 5325: 5269: 5237: 5230: 5223: 5216: 5209: 5202: 5182: 5175: 5168: 5161:Saurolophini 5148: 5141: 5134: 5128: 5127: 5120: 5112: 5105:Kritosaurini 5092: 5085: 5080:Huallasaurus 5078: 5071: 5052:Wulagasaurus 5050: 5043: 5036: 5029: 5022: 5015: 4995: 4967: 4960: 4953: 4946: 4941:Lambeosaurus 4939: 4932: 4925: 4918: 4898: 4891: 4884: 4877: 4857: 4850: 4843: 4836: 4828: 4821: 4814: 4807: 4800: 4793: 4786: 4779: 4772: 4765: 4743:Yamatosaurus 4741: 4734: 4727: 4720: 4713: 4708:Aquilarhinus 4706: 4680: 4673: 4666: 4659: 4654:Tethyshadros 4652: 4645: 4638: 4631: 4624: 4617: 4610: 4603: 4596: 4589: 4584:Plesiohadros 4582: 4575: 4568: 4561: 4556:Lophorhothon 4554: 4547: 4539: 4532: 4525: 4518: 4511: 4504: 4497: 4490: 4483: 4476: 4469: 4462: 4455: 4448: 4442:Batyrosaurus 4440: 4435:Bactrosaurus 4433: 4426: 4372: 4315: 4308: 4301: 4293: 4275:Ouranosaurus 4273: 4266: 4259: 4252: 4245: 4240:Brighstoneus 4238: 4231: 4207: 4200: 4193: 4186: 4179: 4172: 4167:Osmakasaurus 4165: 4160:Oblitosaurus 4158: 4151: 4144: 4137: 4130: 4123: 4116: 4109: 4102: 4094: 4087: 4082:Camptosaurus 4080: 4073: 4066: 4059: 4029: 4023:Kangnasaurus 4021: 4013: 4006: 3999: 3992: 3985: 3978:Dryosauridae 3965: 3958: 3951: 3913: 3906: 3899: 3892: 3885: 3880:Matheronodon 3878: 3858: 3851: 3843: 3821: 3813: 3805: 3780: 3773: 3766: 3759: 3754:Sektensaurus 3752: 3747:Qantassaurus 3745: 3738: 3730: 3723: 3716: 3709: 3702: 3696:Kangnasaurus 3694: 3687: 3680: 3673: 3665: 3660:Diluvicursor 3658: 3651: 3644: 3637: 3618:Vectidromeus 3616: 3609: 3601: 3579: 3572: 3565: 3483:Ornithischia 3468:Ornithischia 3420: 3411: 3407:Ornithischia 3402: 3393: 3298: 3291: 3266: 3262: 3256: 3244:. Retrieved 3240:the original 3235: 3225: 3216: 3206: 3193: 3163:(1): 21–43. 3160: 3157:Paleobiology 3156: 3136:. Retrieved 3132: 3123: 3112:. Retrieved 3110:. 2014-08-27 3107: 3098: 3086:. Retrieved 3082:the original 3078:NBC News.com 3077: 3067: 3055:. Retrieved 3050: 3041: 3029:. Retrieved 3024: 3014: 2990:. in press. 2987: 2981: 2951: 2942: 2933: 2929: 2925: 2916: 2888: 2881: 2872: 2868: 2859: 2850: 2846: 2837: 2828: 2822:(43): 29–48. 2819: 2815: 2811: 2807: 2798: 2789: 2785: 2776: 2766:21 September 2764:. Retrieved 2736: 2732: 2722: 2711:. Retrieved 2706: 2697: 2686:. Retrieved 2682:the original 2668: 2644: 2637: 2614: 2576: 2521: 2517: 2513: 2484: 2477: 2466:. Retrieved 2462: 2453: 2412: 2408: 2398: 2388: 2381: 2356: 2352: 2348: 2342: 2323: 2319: 2313: 2293: 2289: 2285: 2258: 2254: 2245: 2236: 2232: 2223: 2215: 2196: 2187: 2163: 2156: 2132: 2125: 2116: 2112: 2081: 2058: 2057:. Series 3. 2054: 2050: 2046: 2037: 2028: 2024: 2015: 1990: 1986: 1973: 1964: 1960: 1931: 1927: 1923: 1914: 1905: 1901: 1897: 1893: 1892:(1914). "On 1856: 1819: 1764:crocodilians 1737: 1728: 1722: 1716: 1714:ceratopsians 1707: 1694: 1688: 1678: 1668: 1650: 1649: 1622: 1608: 1599:Paleoecology 1584: 1582: 1568: 1562: 1558:Centrosaurus 1556: 1550: 1546: 1538: 1532: 1519: 1515: 1512: 1477: 1471: 1466:G. notabilis 1465: 1456:Paleobiology 1394: 1393: 1377: 1376: 1355:Aquilarhinus 1353: 1352: 1312: 1311: 1310: 1295: 1294: 1293: 1271: 1270: 1269: 1239: 1238: 1217:Wulagasaurus 1215: 1214: 1203:Kritosaurini 1191: 1185: 1180: 1176: 1172: 1166: 1163:Kritosaurini 1154:Tethyshadros 1152: 1144: 1143: 1137: 1126: 1106: 1104: 1099: 1096:G. notabilis 1095: 1091: 1083: 1075: 1071: 1063: 1062: 1055:G. notabilis 1054: 1039: 1035: 1031: 1027: 1023: 1018: 1015: 1008: 1005:saurolophine 1000: 988: 984: 976: 972: 957:G. notabilis 956: 940: 936: 929:G. notabilis 928: 924: 921:G. notabilis 920: 917:G. notabilis 916: 912: 908: 899:is from the 897:G. notabilis 896: 894:type species 889: 885: 882:G. notabilis 881: 879: 873: 847: 843: 837: 834:Kritosaurini 829: 825: 815: 805: 800:of Utah, in 795: 782: 774: 772: 759: 755: 751: 743: 739: 735: 731: 727: 723: 715: 711: 705: 703: 690: 686: 682: 678: 674: 670: 659:incurvimanus 658: 654: 650: 646: 642: 638: 630: 626: 618: 614: 608: 602: 599:Barnum Brown 570: 569: 563: 560:G. notabilis 559: 523: 519: 518: 485: 451: 436: 428: 427: 426: 411: 406: 384: 365: 350: 339:G. notabilis 338: 313: 312: 303:Type species 294: 293: 281:Kritosaurini 232: 216: 210:Ornithischia 200: 187: 132: 31: 25: 5741:Gryposaurus 5735:Wikispecies 5711:Gryposaurus 5677:Willinakaqe 5649:nomina nuda 5566:Siluosaurus 5531:Loncosaurus 5369:Dinehichnus 5211:Kamuysaurus 5184:Saurolophus 5136:Kritosaurus 5129:Gryposaurus 4962:Sahaliyania 4948:Magnapaulia 4920:Amurosaurus 4900:Tlatolophus 4879:Adelolophus 4823:Kazaklambia 4802:Blasisaurus 4795:Arenysaurus 4788:Aralosaurus 4722:Hadrosaurus 4715:Eotrachodon 4668:Yunganglong 4633:Sirindhorna 4605:Protohadros 4541:Koshisaurus 4534:Jintasaurus 4303:Comptonatus 4153:Napaisaurus 4139:Lurdusaurus 4111:Fukuisaurus 4089:Cedrorestes 4031:Valdosaurus 3894:Pareisactus 3725:Morrosaurus 3689:Isasicursor 3653:Chakisaurus 3549:Ornithopoda 3536:Ornithopoda 3495:Ornithopoda 3365:Ornithopoda 3133:EurekAlert! 3025:Reuters.com 2349:Gryposaurus 2314:Gryposaurus 2239:(1): 77–98. 2061:(4): 51–59. 2047:Kritosaurus 1924:Kritosaurus 1752:salamanders 1653:shared its 1595:extension. 1589:keratinized 1585:Gryposaurus 1569:Gryposaurus 1547:Gryposaurus 1539:Gryposaurus 1533:Like other 1522:was both a 1486:quadrupedal 1478:Gryposaurus 1474:hadrosaurid 1396:Willinakaqe 1241:Kritosaurus 1192:Gryposaurus 1181:Gryposaurus 1177:Kritosaurus 1173:Gryposaurus 1168:Kritosaurus 1159:Jack Horner 1145:Gryposaurus 1140:restoration 1138:G. alstasei 1127:Gryposaurus 1111:nasal bones 1107:Gryposaurus 1092:G. latidens 1084:Gryposaurus 1064:Gryposaurus 1046:Description 1032:Kritosaurus 1028:Gryposaurus 1024:Kritosaurus 1010:Saurolophus 1001:Kritosaurus 985:Gryposaurus 943:, from the 941:G. latidens 931:along with 886:G. latidens 844:Gryposaurus 839:Kritosaurus 830:Gryposaurus 826:Kritosaurus 783:G. latidens 775:G. latidens 760:Gryposaurus 756:Hadrosaurus 752:Gryposaurus 740:Hadrosaurus 732:Gryposaurus 728:Kritosaurus 716:Kritosaurus 712:Gryposaurus 707:Hadrosaurus 691:Kritosaurus 687:Gryposaurus 679:Kritosaurus 675:Gryposaurus 671:Kritosaurus 665:and Wright 631:Kritosaurus 627:Gryposaurus 619:Kritosaurus 615:Gryposaurus 604:Kritosaurus 571:Gryposaurus 538:quadrupedal 525:Kritosaurus 520:Gryposaurus 486:Gryposaurus 429:Gryposaurus 413:nomen nudum 385:G. alsatei? 351:G. latidens 343:Lambe, 1914 295:Gryposaurus 261:Subfamily: 242:Ornithopoda 43:80–75  32:Gryposaurus 5811:Categories 5619:Arkharavia 5594:Tichosteus 5587:Thespesius 5559:Pteropelyx 5552:Orthomerus 5432:Wintonopus 5017:Acristavus 4997:Barsboldia 4955:Olorotitan 4830:Latirhinus 4682:Zuoyunlong 4549:Levnesovia 4499:Gobihadros 4464:Datonglong 4457:Claosaurus 4428:Altirhinus 4338:see below↓ 4268:Morelladon 4146:Magnamanus 4118:Hippodraco 3987:Dryosaurus 3960:Hesperonyx 3935:Dryomorpha 3775:Trinisaura 3768:Tietasaura 3761:Talenkauen 3639:Anabisetia 3504:see below↓ 3398:Dinosauria 3246:9 November 3138:2021-08-22 3114:2021-08-22 2814:, Parks". 2713:2009-06-16 2688:2007-05-22 2494:1026402631 2468:2021-06-20 2031:: 279–302. 1803:References 1776:marsupials 1666:troodontid 1640:Appalachia 1628:Cretaceous 1579:Nasal arch 1520:Gryposaurs 1499:, and its 1036:Dinosauria 965:iguanodont 933:postcrania 848:G. alsatei 810:predentary 720:hypothesis 562:(formerly 547:settings. 530:Roman nose 460:duckbilled 194:Dinosauria 21:Gyposaurus 5601:Trachodon 5517:Glishades 5496:Diclonius 5334:Anomoepus 5318:Ichnotaxa 5031:Maiasaura 4969:Velafrons 4478:Equijubus 4450:Choyrodon 4310:Iguanodon 4174:Planicoxa 4104:Dakotadon 4075:Calvarius 3901:Rhabdodon 3887:Mochlodon 3630:Elasmaria 3381:Kingdom: 3283:131302174 3088:3 October 3057:3 October 3031:3 October 3027:. Reuters 2908:985402380 2761:133329640 2445:133329640 2437:0022-3360 2373:220413406 1940:2246/1398 1662:theropods 1636:Laramidia 1489:herbivore 1188:cladogram 1149:subfamily 856:Mescalero 793:species. 764:upper arm 667:monograph 577:specimen 541:herbivore 475:Campanian 471:Santonian 167:Kingdom: 161:Eukaryota 5720:Wikidata 5481:Cionodon 5150:Rhinorex 5038:Ornatops 4838:Minqaria 4809:Canardia 4736:Malefica 4661:Xuwulong 4626:Siamodon 4527:Jeyawati 4513:Gonkoken 4471:Eolambia 4295:Barilium 4096:Cumnoria 4061:Barilium 3967:Owenodon 3953:Draconyx 3915:Zalmoxes 3815:Fostoria 3667:Fostoria 3425:Cerapoda 3389:Chordata 3387:Phylum: 3383:Animalia 3185:88689241 3006:55969908 2950:(2002). 2792:: 25–81. 2546:83539808 2296:: 1–119. 1981:(1914). 1786:See also 1733:and the 820:and the 575:based on 504:and the 473:to late 463:dinosaur 398:Synonyms 328:Species 248:Family: 181:Chordata 177:Phylum: 171:Animalia 157:Domain: 5781:1095993 5768:4823293 5755:4531707 5726:Q385949 4774:Ajnabia 4209:Uteodon 3165:Bibcode 2936:: 1–76. 2741:Bibcode 2619:312–348 2526:Bibcode 2417:Bibcode 1995:Bibcode 1768:mammals 1760:lizards 1756:turtles 1528:browser 1497:chewing 1482:bipedal 1038:, with 961:derived 865:Species 859:Apaches 791:derived 787:bonebed 726:, uses 534:bipedal 502:Montana 494:Alberta 452:gryphus 445:griffin 375:Sampson 287:Genus: 274:Tribe: 112:↓ 5443:Ootaxa 4640:Tanius 4233:Bolong 3305:  3281:  3183:  3004:  2958:  2906:  2896:  2759:  2656:  2625:  2589:  2585:–463. 2544:  2492:  2443:  2435:  2371:  2203:  2175:  2144:  2088:  1869:  1865:–448. 1826:  1778:, and 1712:, the 1703:, the 1664:, the 1543:marine 1526:and a 1524:grazer 1493:plants 1201:  1088:limpet 892:. The 888:, and 852:Alsate 846:named 641:, not 478:stages 469:(late 441:lizard 437:grypos 377:, 2007 373:& 358:, 1992 356:Horner 322:, 1914 5794:53014 5776:IRMNG 4485:Fylax 4015:Iyuku 3421:Clade 3412:Clade 3403:Clade 3394:Clade 3279:S2CID 3181:S2CID 3002:S2CID 2757:S2CID 2542:S2CID 2441:S2CID 2369:S2CID 2229:(PDF) 1748:frogs 1687:like 1677:like 1573:river 1509:cheek 1501:teeth 1472:As a 1030:with 903:-age 876:skull 768:ilium 545:river 480:) of 456:genus 449:Latin 434:Greek 371:Gates 320:Lambe 233:Clade 217:Clade 201:Clade 188:Clade 5763:GBIF 4181:Proa 3853:Iani 3481:see 3454:see 3303:ISBN 3248:2013 3090:2007 3059:2007 3033:2007 2956:ISBN 2904:OCLC 2894:ISBN 2768:2016 2654:ISBN 2623:ISBN 2587:ISBN 2490:OCLC 2433:ISSN 2201:ISBN 2173:ISBN 2142:ISBN 2086:ISBN 2029:98-Q 1867:ISBN 1824:ISBN 1762:and 1727:and 1693:and 1644:peat 1619:Utah 1098:and 1080:skin 828:and 766:and 663:Lull 653:and 629:and 510:Utah 51:Preęž’ 5750:EoL 3271:doi 3211:pp. 3173:doi 2992:doi 2928:". 2749:doi 2583:438 2534:doi 2425:doi 2361:doi 2328:doi 2324:151 2169:161 2138:158 2053:". 2003:doi 1936:hdl 1926:". 1900:". 1863:445 1070:of 951:of 738:as 579:NMC 573:is 508:of 500:in 492:in 458:of 5813:: 5791:: 5778:: 5765:: 5752:: 5737:: 5722:: 3423:: 3414:: 3405:: 3396:: 3277:. 3267:25 3265:. 3234:. 3179:. 3171:. 3159:. 3147:^ 3131:. 3106:. 3076:. 3049:. 3023:. 3000:. 2986:. 2970:^ 2934:11 2932:. 2902:. 2873:40 2851:40 2820:38 2818:. 2788:. 2755:. 2747:. 2735:. 2731:. 2705:. 2652:. 2650:64 2621:. 2601:^ 2554:^ 2540:. 2532:. 2522:30 2520:. 2502:^ 2488:. 2461:. 2439:. 2431:. 2423:. 2413:90 2411:. 2407:. 2367:. 2357:26 2355:. 2322:. 2318:. 2302:^ 2292:. 2267:^ 2259:40 2237:51 2235:. 2231:. 2171:. 2140:. 2117:53 2115:. 2100:^ 2067:^ 2059:13 2049:, 2027:. 2001:. 1991:25 1989:. 1985:. 1965:35 1948:^ 1932:28 1930:. 1906:27 1904:. 1881:^ 1838:^ 1810:^ 1782:. 1774:, 1758:, 1754:, 1750:, 1721:, 1699:, 1683:, 1673:, 1567:. 1530:. 1513:G. 1476:, 1183:. 987:, 939:. 884:, 781:, 770:. 714:, 693:. 439:) 235:: 219:: 203:: 190:: 101:Pg 45:Ma 40:, 5651:) 5491:? 5117:? 5055:? 4833:? 4544:? 4445:? 4320:? 4298:? 4099:? 4026:? 4018:? 3848:? 3837:? 3826:? 3818:? 3810:? 3785:? 3735:? 3699:? 3670:? 3621:? 3606:? 3595:? 3584:? 3357:e 3350:t 3343:v 3311:. 3285:. 3273:: 3250:. 3187:. 3175:: 3167:: 3161:1 3141:. 3117:. 3092:. 3061:. 3035:. 3008:. 2994:: 2980:" 2964:. 2910:. 2790:3 2770:. 2751:: 2743:: 2737:1 2716:. 2691:. 2662:. 2631:. 2595:. 2548:. 2536:: 2528:: 2496:. 2471:. 2447:. 2427:: 2419:: 2375:. 2363:: 2336:. 2330:: 2294:2 2209:. 2181:. 2150:. 2094:. 2009:. 2005:: 1997:: 1942:. 1938:: 1875:. 1832:. 1484:/ 1468:. 1129:. 673:/ 536:/ 447:( 416:) 410:( 382:† 363:† 348:† 336:† 310:† 291:† 278:† 265:† 252:† 239:† 223:† 207:† 106:N 96:K 91:J 86:T 81:P 76:C 71:D 66:S 61:O 56:ęž’ 23:.

Index

Gyposaurus
Late Cretaceous
Ma
Preęž’
ęž’
O
S
D
C
P
T
J
K
Pg
N

Natural History Museum of Utah
Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Dinosauria
Ornithischia
Neornithischia
Ornithopoda
Hadrosauridae
Saurolophinae
Kritosaurini
Gryposaurus

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