108:
413:
was formally described by
Sellards in 1916 from fossil material collected from the Upper Bony Valley, Hardee and Polk counties, Florida, a typical Latest Hemphillian. The main holotype was a mandible (USNM 8838) but additional material of this species has since been collected from this locality and
427:
was described by
Dalquest, 1986 on a holotype (MWSU 12147) found from Coffee Ranch in Texas. The holotype specimen is a partial mandible preserving some molar teeth. All specimens of the species were originally classified as those of
453:
is a new species described by
Jiangzuo et al. (2023). The holotype (HMV 2005) is a nearly complete skull from the Wangjiashan fossil locality. This species differs from
461:
in not being as hypercarnivorous with more developed anterior premolars and the first and second molars longer. The skull is also much narrower in this species.
661:
414:
other locations from the southern United States and parts of Mexico. Differs from other species in the genus of shorter and smaller premolars and molars.
1442:
1467:
1447:
1432:
1437:
654:
581:
1462:
1457:
525:"New fossil giant panda relatives (Ailuropodinae, Ursidae): a basal lineage of gigantic Mio-Pliocene cursorial carnivores"
647:
374:
which led the authors of the study to do a comprehensive analysis of agriotheriin specimens to assess their taxonomy.
608:
from North
America, and implications for understanding transformations in the metaconid-entoconid complex of bears".
1452:
1356:
1059:
107:
1186:
1305:
574:
Evolution of
Tertiary Mammals of North America: Terrestrial carnivores, ungulates, and ungulatelike mammals
1472:
1372:
1404:
1196:
1137:
1285:
1335:
847:
735:
1315:
747:
1049:
801:
777:
625:
550:
102:
1409:
767:
1039:
577:
569:
1265:
1235:
617:
542:
532:
576:. Evolution of Tertiary Mammals of North America. Vol. 1. Cambridge University Press.
1255:
1225:
639:
1325:
1295:
1245:
1208:
1426:
1275:
1014:
929:
920:
629:
554:
315:
307:
304:
182:
61:
1395:
1104:
1078:
982:
941:
723:
352:
319:
208:
1166:
1114:
1004:
859:
787:
757:
356:. Hunt (1998) had noted differences in the structure of the carnassials between
323:
36:
621:
1176:
1126:
1026:
994:
869:
833:
710:
670:
81:
46:
961:
887:
704:
366:
335:
159:
119:
86:
382:
When
Jiangzuo et al. (2023) computed the phylogenetic analysis, they found
1389:
1094:
899:
823:
813:
698:
692:
139:
76:
71:
56:
51:
41:
1150:
971:
909:
674:
311:
91:
66:
481:
as well based on overall similarities of their molars. In the case of
360:
and Old World species of said genus, but lots of similarities between
686:
546:
149:
129:
1366:
537:
524:
523:
Jiangzuo, Q.; Flynn, J. J.; Wang, S.; Hou, S.; Deng, T. (2023).
169:
1370:
643:
334:
was a hypercarnivorous genus of bear that had adaptations for
350:
were originally classified as species of the related genus
604:
Qigao
Jiangzuo; John J. Flynn (2020). "A new species of
370:. This was further supported in the 2019 publication of
572:. In Janis, C. M.; Scott, K. M.; Jacobs, L. L. (eds.).
440:
in having much larger and longer premolars and molars.
386:
to be paraphyletic to the clade containing the genera
1331:
1321:
1311:
1301:
1291:
1281:
1271:
1261:
1251:
1241:
1231:
1221:
1192:
1182:
1172:
1162:
1133:
1110:
1100:
1090:
1055:
1045:
1035:
1010:
1000:
990:
967:
957:
947:
937:
905:
895:
884:
865:
855:
844:
829:
819:
809:
798:
783:
773:
763:
753:
743:
732:
720:
215:
1379:
1206:
1149:
1124:
1077:
1024:
981:
928:
919:
883:
843:
797:
731:
719:
518:
516:
514:
512:
510:
508:
506:
655:
8:
1367:
1214:
1155:
1083:
1021:
925:
728:
662:
648:
640:
20:
536:
502:
7:
469:Two species of agriotheriin bears,
14:
1443:Cenozoic mammals of North America
330:), which is a bamboo specialist,
322:. Unlike its modern relative the
314:period. Fossils were found from
106:
610:Journal of Mammalian Evolution
1:
1468:Fossil taxa described in 2023
1448:Prehistoric carnivoran genera
278:(Morales & Aguirre, 1976)
570:"Ursidae, pp. 174–195 (180)"
568:Hunt, Robert M. Jr. (1998).
1489:
1433:Cenozoic mammals of Europe
1357:Category:Prehistoric bears
622:10.1007/s10914-019-09480-x
1353:
1217:
1158:
1086:
681:
529:American Museum Novitates
235:
230:
214:
207:
103:Scientific classification
101:
23:
1438:Cenozoic mammals of Asia
1463:Pleistocene extinctions
1458:Pleistocene carnivorans
1187:Helarctos sinomalayanus
430:Agriotherium schneideri
358:Agriotherium schneideri
303:is an extinct genus of
220:Agriotherium schneideri
673:members of the family
465:Other possible species
409:The genotype species,
328:Ailuropoda melanoleuca
16:Extinct genus of bears
1405:Paleobiology Database
267:Jiangzuo et al., 2023
202:Jiangzuo et al., 2023
1306:U. sackdillingensis
32:Tortonian–Calabrian
1197:Melursus theobaldi
1060:A. wulingshanensis
404:Huracan schneideri
1453:Pleistocene bears
1420:
1419:
1373:Taxon identifiers
1364:
1363:
1349:
1348:
1345:
1344:
1145:
1144:
1073:
1072:
1069:
1068:
879:
878:
485:is sister to the
296:
295:
290:
279:
268:
257:
246:
203:
1480:
1413:
1412:
1400:
1399:
1398:
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1333:
1323:
1313:
1303:
1293:
1283:
1273:
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1233:
1223:
1215:
1194:
1184:
1174:
1164:
1156:
1135:
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1102:
1092:
1084:
1057:
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1022:
1012:
1002:
992:
969:
959:
949:
939:
926:
907:
897:
886:
867:
857:
846:
831:
821:
811:
800:
785:
775:
765:
755:
745:
734:
729:
722:
664:
657:
650:
641:
634:
633:
601:
595:
594:
592:
590:
565:
559:
558:
540:
520:
477:might belong to
289:(Lydekker, 1884)
288:
277:
266:
256:(Dalquest, 1986)
255:
245:(Sellards, 1916)
244:
217:
201:
194:
181:
111:
110:
96:
33:
29:Temporal range:
21:
1488:
1487:
1483:
1482:
1481:
1479:
1478:
1477:
1423:
1422:
1421:
1416:
1408:
1403:
1394:
1393:
1388:
1375:
1365:
1360:
1341:
1202:
1141:
1120:
1065:
1020:
977:
915:
875:
839:
793:
715:
677:
668:
638:
637:
603:
602:
598:
588:
586:
584:
567:
566:
562:
522:
521:
504:
499:
467:
448:
422:
407:
400:
380:
344:
285:H?. punjabensis
226:
223:
200:
192:
179:
105:
97:
95:
94:
89:
84:
79:
74:
69:
64:
59:
54:
49:
44:
39:
31:
30:
27:
17:
12:
11:
5:
1486:
1484:
1476:
1475:
1470:
1465:
1460:
1455:
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1445:
1440:
1435:
1425:
1424:
1418:
1417:
1415:
1414:
1401:
1385:
1383:
1377:
1376:
1371:
1362:
1361:
1354:
1351:
1350:
1347:
1346:
1343:
1342:
1340:
1339:
1329:
1319:
1309:
1299:
1289:
1279:
1269:
1259:
1249:
1239:
1229:
1218:
1212:
1204:
1203:
1201:
1200:
1190:
1180:
1170:
1159:
1153:
1147:
1146:
1143:
1142:
1132:
1130:
1122:
1121:
1119:
1118:
1108:
1098:
1087:
1081:
1075:
1074:
1071:
1070:
1067:
1066:
1064:
1063:
1053:
1043:
1032:
1030:
1019:
1018:
1008:
998:
987:
985:
979:
978:
976:
975:
965:
955:
945:
934:
932:
923:
917:
916:
914:
913:
903:
892:
890:
881:
880:
877:
876:
874:
873:
863:
852:
850:
848:Phoberocyonini
841:
840:
838:
837:
827:
817:
806:
804:
795:
794:
792:
791:
781:
771:
761:
751:
740:
738:
726:
717:
716:
714:
713:
707:
701:
695:
689:
682:
679:
678:
669:
667:
666:
659:
652:
644:
636:
635:
616:(4): 775–787.
596:
582:
560:
538:10.1206/3996.1
531:(3996): 1–71.
501:
500:
498:
495:
483:I. punjabensis
475:I. punjabensis
466:
463:
447:
442:
421:
416:
406:
401:
399:
396:
379:
376:
343:
340:
294:
293:
292:
291:
280:
269:
258:
247:
233:
232:
228:
227:
225:Sellards, 1916
224:
212:
211:
205:
204:
190:
186:
185:
177:
173:
172:
167:
163:
162:
157:
153:
152:
147:
143:
142:
137:
133:
132:
127:
123:
122:
117:
113:
112:
99:
98:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
40:
35:
34:
28:
15:
13:
10:
9:
6:
4:
3:
2:
1485:
1474:
1473:Ailuropodinae
1471:
1469:
1466:
1464:
1461:
1459:
1456:
1454:
1451:
1449:
1446:
1444:
1441:
1439:
1436:
1434:
1431:
1430:
1428:
1411:
1406:
1402:
1397:
1391:
1387:
1386:
1384:
1382:
1378:
1374:
1369:
1359:
1358:
1352:
1338:
1337:
1330:
1328:
1327:
1320:
1318:
1317:
1310:
1308:
1307:
1300:
1298:
1297:
1290:
1288:
1287:
1286:U. pyrenaicus
1280:
1278:
1277:
1270:
1268:
1267:
1266:U. kudarensis
1260:
1258:
1257:
1250:
1248:
1247:
1240:
1238:
1237:
1236:U. dolinensis
1230:
1228:
1227:
1220:
1219:
1216:
1213:
1211:
1210:
1205:
1199:
1198:
1191:
1189:
1188:
1181:
1179:
1178:
1171:
1169:
1168:
1161:
1160:
1157:
1154:
1152:
1148:
1140:
1139:
1138:T. floridanus
1131:
1129:
1128:
1123:
1117:
1116:
1109:
1107:
1106:
1099:
1097:
1096:
1089:
1088:
1085:
1082:
1080:
1076:
1062:
1061:
1054:
1052:
1051:
1044:
1042:
1041:
1034:
1033:
1031:
1029:
1028:
1023:
1017:
1016:
1015:Kretzoiarctos
1009:
1007:
1006:
999:
997:
996:
989:
988:
986:
984:
980:
974:
973:
966:
964:
963:
956:
954:
953:
946:
944:
943:
936:
935:
933:
931:
930:Agriotheriini
927:
924:
922:
921:Ailuropodinae
918:
912:
911:
904:
902:
901:
894:
893:
891:
889:
882:
872:
871:
864:
862:
861:
854:
853:
851:
849:
842:
836:
835:
828:
826:
825:
818:
816:
815:
808:
807:
805:
803:
796:
790:
789:
782:
780:
779:
772:
770:
769:
762:
760:
759:
752:
750:
749:
742:
741:
739:
737:
736:Cephalogalini
730:
727:
725:
718:
712:
708:
706:
702:
700:
696:
694:
690:
688:
684:
683:
680:
676:
672:
665:
660:
658:
653:
651:
646:
645:
642:
631:
627:
623:
619:
615:
611:
607:
600:
597:
585:
583:9780521355193
579:
575:
571:
564:
561:
556:
552:
548:
544:
539:
534:
530:
526:
519:
517:
515:
513:
511:
509:
507:
503:
496:
494:
492:
488:
484:
480:
476:
472:
464:
462:
460:
459:H. schneideri
456:
452:
446:
443:
441:
439:
438:H. schneideri
436:differs from
435:
431:
426:
420:
417:
415:
412:
411:H. schneideri
405:
402:
397:
395:
393:
389:
385:
377:
375:
373:
369:
368:
363:
362:A. schneideri
359:
355:
354:
349:
346:Specimens of
341:
339:
337:
333:
329:
325:
321:
317:
316:North America
313:
309:
306:
302:
301:
287:
286:
281:
276:
275:
270:
265:
264:
259:
254:
253:
248:
243:
242:
241:H. schneideri
237:
236:
234:
229:
222:
221:
213:
210:
206:
199:
198:
191:
188:
187:
184:
183:Agriotheriini
178:
175:
174:
171:
168:
165:
164:
161:
158:
155:
154:
151:
148:
145:
144:
141:
138:
135:
134:
131:
128:
125:
124:
121:
118:
115:
114:
109:
104:
100:
93:
88:
83:
78:
73:
68:
63:
58:
53:
48:
43:
38:
26:
22:
19:
1380:
1355:
1336:U. vitabilis
1334:
1324:
1314:
1304:
1294:
1284:
1274:
1264:
1256:U. ingressus
1254:
1244:
1234:
1226:U. deningeri
1224:
1207:
1195:
1185:
1175:
1165:
1136:
1125:
1113:
1105:Arctotherium
1103:
1093:
1079:Tremarctinae
1058:
1048:
1038:
1025:
1013:
1003:
993:
983:Ailuropodini
970:
960:
951:
950:
942:Agriotherium
940:
908:
898:
868:
858:
832:
822:
812:
786:
776:
766:
756:
748:Adelpharctos
746:
724:Hemicyoninae
613:
609:
606:Agriotherium
605:
599:
587:. Retrieved
573:
563:
528:
490:
487:Agriotherium
486:
482:
478:
474:
470:
468:
458:
454:
450:
449:
444:
437:
433:
429:
424:
423:
418:
410:
408:
403:
392:Agriotherium
391:
387:
383:
381:
371:
365:
361:
357:
353:Agriotherium
351:
347:
345:
336:cursoriality
331:
327:
320:Eastern Asia
308:ailuropodine
305:agriotheriin
299:
298:
297:
284:
283:
273:
272:
262:
261:
251:
250:
240:
239:
219:
218:
209:Type species
196:
195:
24:
18:
1326:U. spelaeus
1296:U. rossicus
1246:U. etruscus
1167:Aurorarctos
1115:Plionarctos
1005:Ailurarctos
860:Phoberocyon
802:Hemicyonini
788:Phoberogale
778:Filholictis
758:Cephalogale
378:Systematics
324:giant panda
274:H?. roblesi
1427:Categories
1396:Q120024195
1276:U. minimus
1177:Protarctos
1127:Tremarctos
1050:A. microta
1027:Ailuropoda
995:Agriarctos
870:Plithocyon
834:Zaragocyon
768:Cyonarctos
711:Caniformia
709:Suborder:
497:References
471:A. roblesi
455:H. coffeyi
434:H. coffeyi
425:H. coffeyi
419:H. coffeyi
372:A. hendeyi
364:and genus
252:H. coffeyi
1316:U. savini
1040:A. baconi
962:Indarctos
888:Ursavinae
705:Carnivora
685:Kingdom:
630:201064589
555:257508340
547:2246/7315
432:, though
384:Indarctos
367:Indarctos
310:from the
160:Carnivora
126:Kingdom:
120:Eukaryota
1390:Wikidata
1095:Arctodus
900:Ballusia
824:Hemicyon
814:Dinocyon
699:Mammalia
693:Chordata
691:Phylum:
687:Animalia
231:Species
166:Family:
150:Mammalia
140:Chordata
136:Phylum:
130:Animalia
116:Domain:
1381:Huracan
1151:Ursinae
972:Miomaci
952:Huracan
910:Ursavus
703:Order:
697:Class:
675:Ursidae
671:Extinct
589:13 June
493:clade.
491:Huracan
479:Huracan
451:H. qiui
445:H. qiui
398:Species
388:Huracan
348:Huracan
342:History
332:Huracan
312:Neogene
300:Huracan
263:H. qiui
197:Huracan
189:Genus:
176:Tribe:
170:Ursidae
156:Order:
146:Class:
25:Huracan
1410:470226
628:
580:
553:
1209:Ursus
626:S2CID
551:S2CID
591:2023
578:ISBN
473:and
457:and
390:and
318:and
37:PreꞒ
618:doi
543:hdl
533:doi
1429::
1407::
1392::
624:.
614:27
612:.
549:.
541:.
527:.
505:^
394:.
338:.
87:Pg
1332:†
1322:†
1312:†
1302:†
1292:†
1282:†
1272:†
1262:†
1252:†
1242:†
1232:†
1222:†
1193:†
1183:†
1173:†
1163:†
1134:†
1111:†
1101:†
1091:†
1056:†
1046:†
1036:†
1011:†
1001:†
991:†
968:†
958:†
948:†
938:†
906:†
896:†
885:†
866:†
856:†
845:†
830:†
820:†
810:†
799:†
784:†
774:†
764:†
754:†
744:†
733:†
721:†
663:e
656:t
649:v
632:.
620::
593:.
557:.
545::
535::
489:–
326:(
282:†
271:†
260:†
249:†
238:†
216:†
193:†
180:†
92:N
82:K
77:J
72:T
67:P
62:C
57:D
52:S
47:O
42:Ꞓ
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