113:
135:
348:
795:
1543:
790:
576:
Whiteside, D. I., Duffin, C. J., Gill, P. G., Marshall, J. E. A., & Benton, M. J. (2016). The Late
Triassic and Early Jurassic fissure faunas from Bristol and South Wales: Stratigraphy and setting.
1587:
372:. More specimens from the same species were later found at Slickstones Quarry, and were first described by Fraser (1988a) as belonging to an indeterminate
1621:
609:
1626:
376:. The species is referred to as the 'Cromhall procolophonid' by Fraser (1988c) and as 'Procolophonid B' by Fraser (1994) and Edwards (2000).
554:
Fraser, N. C. (1994). Assemblages of small tetrapods from
British Late Triassic fissure deposits. In N. C. Fraser & H.-D. Sues (Eds.),
1631:
1580:
567:
Edwards, B. (2000). The Upper
Triassic microvertebrate assemblage of Ruthin Quarry, South Wales. PhD, University College London, 390 pp.
360:
The holotype, specimen UMZC 2023.4.1, which is a partial skull, was discovered during the 1970s by Jeanne Evans and was likely found at
1606:
1611:
1573:
479:
Butler, R. J.; Meade, L. E.; Cleary, T. J.; McWhirter, K. T.; Brown, E. E.; Kemp, T. S.; Benito, J.; Fraser, N. C. (2023).
361:
602:
134:
1616:
794:
595:
369:
326:
1483:
1169:
1241:
1306:
1278:
1102:
964:
694:
112:
1454:
824:
485:
gen. et sp. nov., a new leptopleuronine procolophonid from the Late
Triassic of southwest England"
973:
789:
205:
129:
1462:
1439:
1314:
1425:
804:
502:
395:
251:
1557:
1177:
1297:
1227:
1076:
850:
814:
757:
492:
422:
313:
231:
1409:
1370:
1335:
1155:
1134:
541:
Fraser, N. C. (1988c). Latest
Triassic terrestrial vertebrates and their biostratigraphy.
418:
373:
218:
1363:
1270:
1206:
1185:
1042:
932:
839:
774:
347:
1600:
1447:
1432:
1249:
1056:
994:
987:
953:
939:
887:
859:
732:
706:
618:
323:
316:
70:
30:
1384:
1342:
1328:
1234:
1220:
1147:
1035:
880:
831:
449:
437:
432:
386:
353:
263:
192:
1505:
1476:
1349:
1263:
1256:
1213:
1123:
1086:
1049:
1014:
925:
911:
767:
524:
319:
45:
1512:
1491:
1418:
1391:
1192:
1116:
1095:
1005:
946:
918:
898:
873:
866:
679:
663:
306:
90:
55:
401:(2023) identified that the specimens belonged to a separate genus, naming it
1498:
1469:
1356:
1286:
1162:
1028:
980:
146:
95:
39:
506:
1542:
1550:
1398:
1377:
1199:
1109:
648:
642:
443:
365:
166:
85:
80:
65:
60:
50:
1553:
1021:
657:
330:
176:
100:
75:
636:
497:
480:
302:
298:
156:
346:
309:
121:
587:
629:
591:
381:
522:
Fraser, N. C. (1988a). The osteology and relationships of
530:
1561:
270:
1408:
1296:
1133:
1085:
1074:
1004:
963:
897:
849:
813:
802:
756:
705:
678:
421:. In 2023, it was classified into the subfamily
1581:
603:
8:
333:. The type, and currently only, species is
1588:
1574:
1139:
1082:
903:
810:
753:
749:
684:
675:
626:
610:
596:
588:
558:(pp. 214β226). Cambridge University Press.
111:
20:
496:
462:
7:
1539:
1537:
518:
516:
474:
472:
470:
468:
466:
379:The specimens were then referred to
351:Life reconstruction of the related
1560:. You can help Knowledge (XXG) by
14:
1622:Late Triassic reptiles of Europe
1541:
793:
788:
133:
556:In the shadow of the dinosaurs
1:
1627:Fossil taxa described in 2023
124:as seen from multiple angles
528:(Reptilia: Sphenodontida).
417:has been classified within
16:Fossil genus of parareptile
1648:
1632:Prehistoric reptile genera
1536:
413:Ever since its discovery,
1142:
906:
786:
752:
748:
729:
687:
674:
625:
269:
262:
130:Scientific classification
128:
119:
110:
23:
579:Palaeontologia Polonica
1607:Triassic reptile stubs
483:Hwiccewyrm trispiculum
403:Hwiccewyrm trispiculum
370:Magnesian Conglomerate
357:
327:Magnesian Conglomerate
275:Hwiccewyrm trispiculum
1612:Triassic parareptiles
1549:This article about a
489:The Anatomical Record
368:in an outcrop of the
350:
581:, 67, 257β287.
431:was found to be the
343:Discovery and naming
965:Pumiliopareiasauria
362:Slickstones Quarry
358:
206:Procolophonomorpha
1569:
1568:
1534:
1533:
1530:
1529:
1526:
1525:
1522:
1521:
1070:
1069:
1066:
1065:
805:Pareiasauromorpha
784:
783:
744:
743:
740:
739:
725:
724:
289:
288:
258:
1639:
1617:Leptopleuronines
1590:
1583:
1576:
1545:
1538:
1298:Leptopleuroninae
1228:Smilodonterpeton
1170:Gomphiosauridion
1140:
1083:
1077:Procolophonoidea
904:
851:Nycteroleteridae
815:Lanthanosuchidae
811:
797:
792:
758:Nyctiphruretidae
754:
750:
685:
676:
669:
668:
627:
612:
605:
598:
589:
582:
574:
568:
565:
559:
552:
546:
539:
533:
532:, B321, 125β178.
520:
511:
510:
500:
498:10.1002/ar.25316
476:
423:Leptopleuroninae
272:
250:
243:
232:Leptopleuroninae
230:
217:
204:
191:
138:
137:
115:
105:
42:
29:Temporal range:
21:
1647:
1646:
1642:
1641:
1640:
1638:
1637:
1636:
1597:
1596:
1595:
1594:
1535:
1518:
1410:Procolophoninae
1404:
1371:Pentaedrusaurus
1336:Koiloskiosaurus
1292:
1135:Procolophonidae
1129:
1079:
1062:
1000:
959:
893:
845:
807:
798:
780:
736:
721:
701:
670:
632:
631:
621:
616:
586:
585:
575:
571:
566:
562:
553:
549:
540:
536:
521:
514:
478:
477:
464:
459:
419:Procolophonidae
411:
345:
314:leptopleuronine
285:
278:
249:
241:
228:
219:Procolophonidae
215:
202:
189:
132:
106:
104:
103:
98:
93:
88:
83:
78:
73:
68:
63:
58:
53:
48:
37:
36:
34:
27:
17:
12:
11:
5:
1645:
1643:
1635:
1634:
1629:
1624:
1619:
1614:
1609:
1599:
1598:
1593:
1592:
1585:
1578:
1570:
1567:
1566:
1546:
1532:
1531:
1528:
1527:
1524:
1523:
1520:
1519:
1517:
1516:
1509:
1502:
1495:
1488:
1480:
1473:
1466:
1459:
1451:
1444:
1436:
1429:
1422:
1414:
1412:
1406:
1405:
1403:
1402:
1395:
1388:
1381:
1374:
1367:
1364:Neoprocolophon
1360:
1353:
1346:
1339:
1332:
1325:
1318:
1311:
1302:
1300:
1294:
1293:
1291:
1290:
1283:
1275:
1271:Tricuspisaurus
1267:
1260:
1253:
1246:
1242:Spondylolestes
1238:
1231:
1224:
1217:
1210:
1207:Phaanthosaurus
1203:
1196:
1189:
1186:Kitchingnathus
1182:
1174:
1166:
1159:
1152:
1143:
1137:
1131:
1130:
1128:
1127:
1120:
1113:
1106:
1099:
1091:
1089:
1080:
1075:
1072:
1071:
1068:
1067:
1064:
1063:
1061:
1060:
1053:
1046:
1043:Sanchuansaurus
1039:
1032:
1025:
1018:
1010:
1008:
1002:
1001:
999:
998:
991:
984:
977:
969:
967:
961:
960:
958:
957:
950:
943:
936:
933:Embrithosaurus
929:
922:
915:
907:
901:
895:
894:
892:
891:
884:
877:
870:
863:
855:
853:
847:
846:
844:
843:
840:Lanthanosuchus
836:
828:
820:
818:
808:
803:
800:
799:
787:
785:
782:
781:
779:
778:
775:Nyctiphruretus
771:
763:
761:
746:
745:
742:
741:
738:
737:
730:
727:
726:
723:
722:
720:
719:
718:
717:
711:
709:
703:
702:
700:
699:
698:
697:
688:
682:
672:
671:
667:
666:
660:
651:
645:
639:
630:
623:
622:
617:
615:
614:
607:
600:
592:
584:
583:
569:
560:
547:
545:, 13, 125β140.
543:Modern Geology
534:
512:
461:
460:
458:
455:
410:
409:Classification
407:
344:
341:
336:H. trispiculum
287:
286:
279:
267:
266:
260:
259:
239:
235:
234:
226:
222:
221:
213:
209:
208:
200:
196:
195:
187:
180:
179:
174:
170:
169:
164:
160:
159:
154:
150:
149:
144:
140:
139:
126:
125:
117:
116:
108:
107:
99:
94:
89:
84:
79:
74:
69:
64:
59:
54:
49:
44:
43:
28:
15:
13:
10:
9:
6:
4:
3:
2:
1644:
1633:
1630:
1628:
1625:
1623:
1620:
1618:
1615:
1613:
1610:
1608:
1605:
1604:
1602:
1591:
1586:
1584:
1579:
1577:
1572:
1571:
1565:
1563:
1559:
1555:
1552:
1547:
1544:
1540:
1515:
1514:
1510:
1508:
1507:
1503:
1501:
1500:
1496:
1494:
1493:
1489:
1486:
1485:
1481:
1479:
1478:
1474:
1472:
1471:
1467:
1465:
1464:
1460:
1457:
1456:
1452:
1450:
1449:
1445:
1442:
1441:
1437:
1435:
1434:
1433:Eumetabolodon
1430:
1428:
1427:
1423:
1421:
1420:
1416:
1415:
1413:
1411:
1407:
1401:
1400:
1396:
1394:
1393:
1389:
1387:
1386:
1382:
1380:
1379:
1375:
1373:
1372:
1368:
1366:
1365:
1361:
1359:
1358:
1354:
1352:
1351:
1347:
1345:
1344:
1340:
1338:
1337:
1333:
1331:
1330:
1326:
1324:
1323:
1319:
1317:
1316:
1312:
1309:
1308:
1307:Basileosaurus
1304:
1303:
1301:
1299:
1295:
1289:
1288:
1284:
1281:
1280:
1279:Xenodiphyodon
1276:
1273:
1272:
1268:
1266:
1265:
1261:
1259:
1258:
1254:
1252:
1251:
1250:Suchonosaurus
1247:
1244:
1243:
1239:
1237:
1236:
1232:
1230:
1229:
1225:
1223:
1222:
1218:
1216:
1215:
1211:
1209:
1208:
1204:
1202:
1201:
1197:
1195:
1194:
1190:
1188:
1187:
1183:
1180:
1179:
1175:
1172:
1171:
1167:
1165:
1164:
1160:
1158:
1157:
1153:
1150:
1149:
1145:
1144:
1141:
1138:
1136:
1132:
1126:
1125:
1121:
1119:
1118:
1114:
1112:
1111:
1107:
1105:
1104:
1100:
1098:
1097:
1093:
1092:
1090:
1088:
1084:
1081:
1078:
1073:
1059:
1058:
1057:Shihtienfenia
1054:
1052:
1051:
1047:
1045:
1044:
1040:
1038:
1037:
1033:
1031:
1030:
1026:
1024:
1023:
1019:
1017:
1016:
1012:
1011:
1009:
1007:
1003:
997:
996:
995:Pumiliopareia
992:
990:
989:
988:Provelosaurus
985:
983:
982:
978:
976:
975:
971:
970:
968:
966:
962:
956:
955:
954:Senectosaurus
951:
949:
948:
944:
942:
941:
940:Nochelesaurus
937:
935:
934:
930:
928:
927:
923:
921:
920:
916:
914:
913:
909:
908:
905:
902:
900:
896:
890:
889:
888:Rhipaeosaurus
885:
883:
882:
878:
876:
875:
871:
869:
868:
864:
862:
861:
860:Bashkyroleter
857:
856:
854:
852:
848:
842:
841:
837:
834:
833:
829:
827:
826:
822:
821:
819:
816:
812:
809:
806:
801:
796:
791:
777:
776:
772:
770:
769:
765:
764:
762:
759:
755:
751:
747:
735:
734:
733:Procolophonia
728:
715:
714:
713:
712:
710:
708:
707:Procolophonia
704:
696:
692:
691:
690:
689:
686:
683:
681:
677:
673:
665:
661:
659:
655:
652:
650:
646:
644:
640:
638:
634:
633:
628:
624:
620:
619:Procolophonia
613:
608:
606:
601:
599:
594:
593:
590:
580:
573:
570:
564:
561:
557:
551:
548:
544:
538:
535:
531:
527:
526:
519:
517:
513:
508:
504:
499:
494:
490:
486:
484:
475:
473:
471:
469:
467:
463:
456:
454:
452:
451:
446:
445:
440:
439:
434:
430:
426:
424:
420:
416:
408:
406:
404:
400:
397:
393:
390:by Whiteside
389:
388:
384:
383:
377:
375:
374:procolophonid
371:
367:
363:
356:
355:
349:
342:
340:
338:
337:
332:
328:
325:
324:Late Triassic
321:
318:
317:procolophonid
315:
311:
308:
304:
301:
300:
295:
294:
283:
277:
276:
268:
265:
261:
256:
253:
248:
247:
240:
237:
236:
233:
227:
224:
223:
220:
214:
211:
210:
207:
201:
198:
197:
194:
188:
185:
182:
181:
178:
175:
172:
171:
168:
165:
162:
161:
158:
155:
152:
151:
148:
145:
142:
141:
136:
131:
127:
123:
118:
114:
109:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
41:
38:208β202
32:
31:Late Triassic
26:
22:
19:
1562:expanding it
1548:
1511:
1504:
1497:
1490:
1482:
1475:
1468:
1461:
1455:Lestanshoria
1453:
1446:
1438:
1431:
1424:
1417:
1397:
1390:
1385:Sclerosaurus
1383:
1376:
1369:
1362:
1355:
1348:
1343:Leptopleuron
1341:
1334:
1329:Hypsognathus
1327:
1321:
1320:
1313:
1305:
1285:
1277:
1269:
1262:
1255:
1248:
1240:
1235:Sphenosaurus
1233:
1226:
1221:Sauropareion
1219:
1212:
1205:
1198:
1191:
1184:
1176:
1168:
1161:
1154:
1148:Coartaredens
1146:
1122:
1115:
1108:
1101:
1094:
1055:
1048:
1041:
1036:Pareiasaurus
1034:
1027:
1020:
1013:
993:
986:
979:
972:
952:
945:
938:
931:
924:
917:
910:
899:Pareiasauria
886:
881:Nycteroleter
879:
872:
865:
858:
838:
832:Lanthaniscus
830:
825:Chalcosaurus
823:
773:
766:
731:
653:
647:Superclass:
578:
572:
563:
555:
550:
542:
537:
529:
523:
488:
482:
450:Leptopleuron
448:
442:
438:Hypsognathus
436:
433:sister-taxon
428:
427:
414:
412:
402:
398:
394:(2016), and
391:
387:Hypsognathus
385:
380:
378:
359:
354:Hypsognathus
352:
335:
334:
297:
292:
291:
290:
281:
274:
273:
264:Type species
254:
245:
244:
193:Parareptilia
183:
24:
18:
1506:Thelerpeton
1477:Procolophon
1350:Libognathus
1264:Tichvinskia
1257:Theledectes
1214:Pintosaurus
1124:Saurodektes
1087:Owenettidae
1050:Scutosaurus
1015:Arganaceras
926:Deltavjatia
912:Bradysaurus
768:Abyssomedon
525:Clevosaurus
320:parareptile
225:Subfamily:
1601:Categories
1513:Timanophon
1492:Teratophon
1463:Orenburgia
1440:Insulophon
1419:Anomoiodon
1392:Scoloparia
1322:Hwiccewyrm
1315:Cornualbus
1193:Lasasaurus
1117:Ruhuhuaria
1103:Candelaria
1096:Barasaurus
1006:Therischia
947:Parasaurus
919:Bunostegos
874:Macroleter
867:Emeroleter
716:see belowβ
695:Sauropsida
680:Sauropsida
664:Sauropsida
457:References
429:Hwiccewyrm
415:Hwiccewyrm
296:(meaning "
293:Hwiccewyrm
246:Hwiccewyrm
25:Hwiccewyrm
1499:Thelephon
1470:Procolina
1426:Burtensia
1357:Mandaphon
1287:Youngetta
1163:Eomurruna
1029:Obirkovia
981:Nanoparia
649:Tetrapoda
635:Kingdom:
322:from the
305:") is an
153:Kingdom:
147:Eukaryota
120:Holotype
1551:Triassic
1399:Soturnia
1378:Phonodus
1200:Oryporan
1110:Owenetta
974:Anthodon
643:Chordata
641:Phylum:
637:Animalia
507:37735997
444:Soturnia
366:Cromhall
212:Family:
177:Reptilia
167:Chordata
163:Phylum:
157:Animalia
143:Domain:
1554:reptile
1484:Samaria
1178:Kinelia
1156:Coletta
1022:Elginia
662:Class:
658:Amniota
331:England
307:extinct
280:Butler
238:Genus:
199:Order:
173:Class:
505:
399:et al.
396:Butler
392:et al.
303:dragon
299:Hwicce
284:, 2023
282:et al.
257:, 2023
255:et al.
252:Butler
1556:is a
1448:Kapes
654:Clade
310:genus
184:Clade
122:skull
1558:stub
693:see
503:PMID
447:and
46:Preκ
493:doi
435:to
425:.
382:cf.
329:of
312:of
1603::
656::
515:^
501:.
491:.
487:.
465:^
453:.
441:,
405:.
364:,
339:.
186::
96:Pg
40:Ma
33:,
1589:e
1582:t
1575:v
1564:.
1487:?
1458:?
1443:?
1310:?
1282:?
1274:?
1245:?
1181:?
1173:?
1151:?
835:?
817:?
760:?
611:e
604:t
597:v
509:.
495::
481:"
271:β
242:β
229:β
216:β
203:β
190:β
101:N
91:K
86:J
81:T
76:P
71:C
66:D
61:S
56:O
51:κ
35:~
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