4535:). They also estimate that the Cohen modal haplotype must be older than 4500 years old, and maybe as much as 8600 years old - well before the supposed origin of the Cohanim. Only the "Palestinian & Israeli Arab" modal had a strong correlation to an ethnic group, but it was also rare. In conclusion, the authors were negative about the usefulness of STR defined modals for any "forensic or genealogical purposes" because "they were found across ethnic groups with different cultural or geographic affiliation".
5010:
3039:
47:
4391:). More generally, since then this cluster has been found to be frequent among men in the Middle East and North Africa, but less frequent in areas of Ethiopia and Europe where J-M267 is nevertheless common. The genetical pattern is therefore similar to the pattern of J-P58 generally, described above, and may be caused by the same movements/migration of people (
4556:
Tofanelli et al. 2014 responded by saying: "In conclusion, while the observed distribution of sub-clades of haplotypes at mitochondrial and Y chromosome non-recombinant genomes might be compatible with founder events in recent times at the origin of Jewish groups as
Cohenite, Levite, Ashkenazite, the
4520:
rejected this for J-P58 as a whole, but accepted that "some of the populations with low diversity, such as
Bedouins from Israel, Qatar, Sudan and UAE, are tightly clustered near high-frequency haplotypes suggesting founder effects with star burst expansion in the Arabian Desert". They did not comment
257:
populations. In
Ethiopia there are signs of older movements of J-M267 into Africa across the Red Sea, not only in the J-P58 form. This also appears to be associated with Semitic languages. According to a study in 2011, in Tunisia, J-M267 is significantly more abundant in the urban (31.3%) than in the
203:
Since the discovery of haplogroup J-P209 it has generally been recognized that it shows signs of having evolvedâ~â20,000 years ago somewhere in northwestern Iran, the
Caucasus, the Armenian Highlands, and northern Mesopotamia. The frequency and diversity of both its major branches, J-M267 and J-M172,
5903:
Chiaroni, Jacques; King, Roy J.; Myres, Natalie M.; Henn, Brenna M.; Ducourneau, Axel; Mitchell, Michael J.; Boetsch, Gilles; Sheikha, Issa; Lin, Alice A.; Nik-Ahd, Mahnoosh; Ahmad, Jabeen; Lattanzi, Francesca; Herrera, Rene J.; Ibrahim, Muntaser E.; Brody, Aaron; Semino, Ornella; Kivisild, Toomas;
4495:
then pointed out that the GMH is also the most frequent type of J-P209 haplotype found in north-west
Africans and Yemenis, so it is not restricted to Israel and Palestine. However, this particular variant "is absent" from two particular "non-Arab Middle Eastern populations", namely "Jews and Muslim
6891:
Skourtanioti, Eirini; Erdal, Yilmaz S.; Frangipane, Marcella; Balossi
Restelli, Francesca; Yener, K. Aslıhan; Pinnock, Frances; Matthiae, Paolo; Ăzbal, Rana; Schoop, Ulf-Dietrich; Guliyev, Farhad; Akhundov, Tufan; Lyonnet, Bertille; Hammer, Emily L.; Nugent, Selin E.; Burri, Marta; Neumann, Gunnar
6745:
Skourtanioti, Eirini; Erdal, Yilmaz S.; Frangipane, Marcella; Balossi
Restelli, Francesca; Yener, K. Aslıhan; Pinnock, Frances; Matthiae, Paolo; Ăzbal, Rana; Schoop, Ulf-Dietrich; Guliyev, Farhad; Akhundov, Tufan; Lyonnet, Bertille; Hammer, Emily L.; Nugent, Selin E.; Burri, Marta; Neumann, Gunnar
6671:
Skourtanioti, Eirini; Erdal, Yilmaz S.; Frangipane, Marcella; Balossi
Restelli, Francesca; Yener, K. Aslıhan; Pinnock, Frances; Matthiae, Paolo; Ăzbal, Rana; Schoop, Ulf-Dietrich; Guliyev, Farhad; Akhundov, Tufan; Lyonnet, Bertille; Hammer, Emily L.; Nugent, Selin E.; Burri, Marta; Neumann, Gunnar
6597:
Skourtanioti, Eirini; Erdal, Yilmaz S.; Frangipane, Marcella; Balossi
Restelli, Francesca; Yener, K. Aslıhan; Pinnock, Frances; Matthiae, Paolo; Ăzbal, Rana; Schoop, Ulf-Dietrich; Guliyev, Farhad; Akhundov, Tufan; Lyonnet, Bertille; Hammer, Emily L.; Nugent, Selin E.; Burri, Marta; Neumann, Gunnar
3269:
J-M267 is uncommon in most of
Northern and Central Europe. It is, however, found in significant pockets at levels of 5â10% among many populations in southern Europe. A recent study with the extant variation concludes that the Caucasus is likely to be the source of the Greek and Italian haplogroup
4565:
The correspondence between P58 and high DYS388 values, and YCAII=22-22 is not perfect. For example the J-M267 subclade of J-P58 defined by SNP M368 has DYS388=13 and YCAII=19-22, like other types of J-M267 outside the "Arabic" type of J-M267, and it is therefore believed to be a relatively old
8689:
Shen, Peidong; Lavi, Tal; Kivisild, Toomas; Chou, Vivian; Sengun, Deniz; Gefel, Dov; Shpirer, Issac; Woolf, Eilon; et al. (2004). "Reconstruction of patrilineages and matrilineages of Samaritans and other Israeli populations from Y-chromosome and mitochondrial DNA sequence variation".
5669:
K-M2313*, which as yet has no phylogenetic name, has been documented in two living individuals, who have ethnic ties to India and South East Asia. In addition, K-Y28299, which appears to be a primary branch of K-M2313, has been found in three living individuals from India. See: Poznik
8081:
King, Roy J; Di Cristofaro, Julie; Kouvatsi, Anastasia; Triantaphyllidis, Costas; Scheidel, Walter; Myres, Natalie M; Lin, Alice A; Eissautier, Alexandre; Mitchell, Michael; Binder, Didier; Semino, Ornella; Novelletto, Andrea; Underhill, Peter A; Chiaroni, Jacques (December 2011).
9136:
Behar, Doron M.; Garrigan, Daniel; Kaplan, Matthew E.; Mobasher, Zahra; Rosengarten, Dror; Karafet, Tatiana M.; Quintana-Murci, Lluis; Ostrer, Harry; et al. (2004). "Contrasting patterns of Y chromosome variation in Ashkenazi Jewish and host non-Jewish European populations".
4586:
There are several confirmed and proposed phylogenetic trees available for haplogroup J-M267. The following phylogeny or family tree of J-M267 haplogroup subclades is based on the ISOGG (2012) tree, which is in turn based upon the YCC 2008 tree and subsequent published research.
5807:
Sahakyan, Hovhannes; Margaryan, Ashot; Saag, Lauri; Karmin, Monika; Flores, Rodrigo; Haber, Marc; Kushniarevich, Alena; Khachatryan, Zaruhi; Bahmanimehr, Ardeshir; Parik, JĂŒri; Karafet, Tatiana; Yunusbayev, Bayazit; Reisberg, Tuuli; Solnik, Anu; Metspalu, Ene (2021-03-23).
7788:
Gonçalves, Rita; Freitas, Ana; Branco, Marta; Rosa, Alexandra; Fernandes, Ana T.; Zhivotovsky, Lev A.; Underhill, Peter A.; Kivisild, Toomas; Brehm, Antonio (2005). "Y-chromosome lineages from Portugal, Madeira and Açores record elements of Sephardim and Berber ancestry".
4512:
later agreed that this seemed consistent with the evidence and generalized from this that distribution of the entire YCAII=22-22 cluster of J-M267 in the Arabic-speaking areas of the Middle East and North Africa might in fact mainly have an origin in historical times.
6393:
Triki-Fendri, Soumaya; SĂĄnchez-Diz, Paula; Rey-GonzĂĄlez, Danel; Alfadhli, Suad; Ayadi, Imen; Ben Marzoug, Riadh; Carracedo, Ăngel; Rebai, Ahmed (4 March 2016). "Genetic structure of the Kuwaiti population revealed by paternal lineages: Genetic Structure of Kuwait".
8438:
Ottoni, Claudio; Larmuseau, Maarten H.D.; Vanderheyden, Nancy; MartĂnez-Labarga, Cristina; Primativo, Giuseppina; Biondi, Gianfranco; Decorte, Ronny; Rickards, Olga (2011). "Deep into the roots of the Libyan Tuareg: A genetic survey of their paternal heritage".
4339:
On the other hand, the authors agree that later waves of dispersion in and around this area have also had complex effects upon the distributions of some types of J-P58 in some regions. They list three regions which are particularly important to their proposal:
4301:, who moved into arid areas during periods known to have had low rainfall. Thus, while other haplogroups including J-M267 moved out of the area with agriculturalists who followed the rainfall, populations carrying J-M267 remained with their flocks (
6325:
Grugni, Viola; Battaglia, Vincenza; Hooshiar Kashani, Baharak; Parolo, Silvia; Al-Zahery, Nadia; Achilli, Alessandro; Olivieri, Anna; Gandini, Francesca; Houshmand, Massoud; Sanati, Mohammad Hossein; Torroni, Antonio; Semino, Ornella (2012-07-18).
8509:
Repping, S; van Daalen, SK; Brown, LG; Korver, Cindy M; Lange, Julian; Marszalek, Janet D; Pyntikova, Tatyana; van der Veen, Fulco; et al. (2006). "High mutation rates have driven extensive structural polymorphism among human Y chromosomes".
4441:(despite Ethiopia being outside of Eurasia), and is much more diverse. The authors also say that "Omanis show a mix of Eurasian pool-like and typical Arabic haplotypes as expected, considering the role of corridor played at different times by the
4787:
livingâ~â8.3 kya. This branch is absent in other ancient European hunter-gatherers. Unfortunately, it is not possible to put this sample in the context of the current haplogroup J1-M267 variation because of the poor quality of the DNA sequence.
4371:
Studies show that J-P58 group is not only in itself very dominant in many areas where J-M267 or J1 are common, but it also contains a large cluster which had been recognized before the discovery of P58. It is still a subject of research though.
4545:. They said that it was necessary to look at a more detailed STR haplotype in order to define a new "Extended Cohen Modal Haplotype" which is extremely rare outside Jewish populations, and even within Jewish populations is mainly only found in
8249:
Nebel, A; Filon, D; Weiss, DA; Weale, M; Faerman, M; Oppenheim, A; Thomas, MG (2000). "High-resolution Y chromosome haplotypes of Israeli and Palestinian Arabs reveal geographic substructure and substantial overlap with haplotypes of Jews".
6892:
U.; Penske, Sandra; Ingman, Tara; Akar, Murat; Shafiq, Rula; Palumbi, Giulio; Eisenmann, Stefanie; d'Andrea, Marta; Rohrlach, Adam B.; Warinner, Christina; Jeong, Choongwon; Stockhammer, Philipp W.; Haak, Wolfgang; Krause, Johannes (2020).
6746:
U.; Penske, Sandra; Ingman, Tara; Akar, Murat; Shafiq, Rula; Palumbi, Giulio; Eisenmann, Stefanie; d'Andrea, Marta; Rohrlach, Adam B.; Warinner, Christina; Jeong, Choongwon; Stockhammer, Philipp W.; Haak, Wolfgang; Krause, Johannes (2020).
6672:
U.; Penske, Sandra; Ingman, Tara; Akar, Murat; Shafiq, Rula; Palumbi, Giulio; Eisenmann, Stefanie; d'Andrea, Marta; Rohrlach, Adam B.; Warinner, Christina; Jeong, Choongwon; Stockhammer, Philipp W.; Haak, Wolfgang; Krause, Johannes (2020).
6598:
U.; Penske, Sandra; Ingman, Tara; Akar, Murat; Shafiq, Rula; Palumbi, Giulio; Eisenmann, Stefanie; d'Andrea, Marta; Rohrlach, Adam B.; Warinner, Christina; Jeong, Choongwon; Stockhammer, Philipp W.; Haak, Wolfgang; Krause, Johannes (2020).
9252:
Semino, O.; Passarino, G; Oefner, PJ; Lin, AA; Arbuzova, S; Beckman, LE; De Benedictis, G; Francalacci, P; et al. (2000). "The Genetic Legacy of Paleolithic Homo sapiens sapiens in Extant Europeans: A Y Chromosome Perspective".
8414:
Onofri, Valerio; Alessandrini, Federica; Turchi, Chiara; Pesaresi, Mauro; Tagliabracci, Adriano (2008). "Y-chromosome markers distribution in Northern Africa: High-resolution SNP and STR analysis in Tunisia and Morocco populations".
7909:
Karachanak, Sena; Grugni, Viola; Fornarino, Simona; Nesheva, Desislava; Al-Zahery, Nadia; Battaglia, Vincenza; Carossa, Valeria; Yordanov, Yordan; Torroni, Antonio; Galabov, Angel S.; Toncheva, Draga; Semino, Ornella (6 March 2013).
6805:
Olivieri, A.; Sidore, C.; Achilli, A.; Angius, A.; Posth, C.; FurtwÀngler, A.; Brandini, S.; Rosario Capodiferro, M.; Gandini, F.; Zoledziewska, M.; Pitzalis, M.; Maschio, A.; Busonero, F.; Lai, L.; Skeates, R. (2017-05-01).
4285:
period, "from a geographical zone, including northeast Syria, northern Iraq and eastern Turkey toward Mediterranean Anatolia, Ismaili from southern Syria, Jordan, Palestine and northern Egypt." They further propose that the
2036:
The area including eastern Turkey and the Zagros and Taurus mountains, has been identified as a likely area of ancient J-M267 diversity. Both J-P58 and other types of J-M267 are present, sometimes with similar frequencies.
8559:
Robino, C.; Crobu, F.; Di Gaetano, C.; Bekada, A.; Benhamamouch, S.; Cerutti, N.; Piazza, A.; Inturri, S.; Torre, C. (2008). "Analysis of Y-chromosomal SNP haplogroups and STR haplotypes in an Algerian population sample".
8043:
King, R. J.; Ăzcan, S. S.; Carter, T.; KalfoÄlu, E.; Atasoy, S.; Triantaphyllidis, C.; Kouvatsi, A.; Lin, A. A.; et al. (2008). "Differential Y-chromosome Anatolian influences on the Greek and Cretan Neolithic".
7450:
CinnioÄlu, Cengiz; King, Roy; Kivisild, Toomas; KalfoÄlu, Ersi; Atasoy, Sevil; Cavalleri, Gianpiero L; Lillie, Anita S.; Roseman, Charles C; et al. (2004). "Excavating Y-chromosome haplotype strata in Anatolia".
7377:
Chiaroni, J; King, Roy J; Underhill, Peter A (2008). "Correlation of annual precipitation with human Y-chromosome diversity and the emergence of Neolithic agricultural and pastoral economies in the Fertile Crescent".
7609:
Fadhlaoui-Zid, Karima; Martinez-Cruz, Begoña; Khodjet-El-Khil, Houssein; Mendizabal, Isabel; Benammar-Elgaaïed, Amel; Comas, David (2011). "Genetic structure of Tunisian ethnic groups revealed by paternal lineages".
7748:
Di Giacomo, F.; Luca, F.; Popa, L. O.; Akar, N.; Anagnou, N.; Banyko, J.; Brdicka, R.; Barbujani, G.; et al. (2004). "Y chromosomal haplogroup J as a signature of the post-neolithic colonization of Europe".
7879:
Hassan, Hisham Y.; Underhill, Peter A.; Cavalli-Sforza, Luca L.; Ibrahim, Muntaser E. (2008). "Y-chromosome variation among Sudanese: restricted gene flow, concordance with language, geography, and history".
5953:
Alvarez, Luis; Ciria, Estela; Marques, Sofia L.; Santos, Cristina; Aluja, Maria Pilar (2014). "Y-chromosome analysis in a Northwest Iberian population: Unraveling the impact of Northern African lineages".
6421:
Finocchio, Andrea; Trombetta, Beniamino; Messina, Francesco; DâAtanasio, Eugenia; Akar, Nejat; Loutradis, Aphrodite; Michalodimitrakis, Emmanuel I.; Cruciani, Fulvio; Novelletto, Andrea (2018-05-10).
249:
is not known to have had any Semitic language. In North Africa J-M267 is dominated by J-P58, and dispersed in a very uneven manner according to studies so far, often but not always being lower among
4557:
overall substantial polyphyletism as well as their systematic occurrence in non-Jewish groups highlights the lack of support for using them either as markers of Jewish ancestry or Biblical tales."
8646:"Polarity and temporality of high-resolution y-chromosome distributions in India identify both indigenous and exogenous expansions and reveal minor genetic influence of Central Asian pastoralists"
6965:
Moran, CN; Scott RA; Adams SM; Warrington SJ; Jobling MA; Wilson RH; Goodwin WH; Georgiades E; Wolde B; Pitsiladis YP. (Nov 2004). "Y chromosome haplogroups of elite Ethiopian endurance runners".
5505:
4904:
4348:
note a "patchy distribution of J1c3 or J-P58 frequency" which is difficult to interpret, and which "may reflect the complex demographic dynamics of religion and ethnicity in the region".
237:
North Africa received Semitic migrations, according to some studies it may have been diffused in recent time by Arabs who, mainly from the 7th century A.D., expanded to northern Africa (
8127:
KujanovĂĄ, Martina; Pereira, LuĂsa; Fernandes, VerĂłnica; Pereira, Joana B.; ÄernĂœ, Viktor (2009). "Near eastern neolithic genetic input in a small oasis of the Egyptian Western Desert".
4504:
but also that J-M267 in this region had very little diversity. They concluded that J-M267 in this region is a result of two distinct migration events: "early Neolithic dispersion" and "
1889:
1855:
9448:
This table shows the historic names for J-P209 (AKA J-12f2.1 or J-M304) in published peer reviewed literature. Note that in Semino 2000 Eu09 is a subclade of Eu10 and in Karafet 2001
4362:
recognize similar signs, but reject it as possibly a result of "either sampling variability and/or demographic complexity associated with multiple founders and multiple migrations."
4275:
and throughout the Arabian peninsula. It also makes up approximately 70% of the J-M267 among the Amhara of Ethiopia. Notably, it is not common among the J-M267 of the Caucasus.
7639:
Flores, Carlos; Maca-Meyer, Nicole; PeĆez, Jose A.; Cabrera, Vicente M. (September 2001). "The peopling of the Canary Islands: a CD4/Alu microsatellite haplotype perspective".
4894:
4425:, and it is a relatively homogeneous group, implying that it might have dispersed relatively recently compared to J-M267 generally. The more diverse "Eurasian" group includes
8601:"Origin, diffusion, and differentiation of Y-chromosome haplogroups E and J: inferences on the neolithization of Europe and later migratory events in the Mediterranean area"
7574:
Ennafaa, Hajer; Fregel, Rosa; Khodjet-El-Khil, Houssein; Gonzålez, Ana M; Mahmoudi, Hejer Abdallah El; Cabrera, Vicente M; Larruga, José M; Benammar-Elgaaïed, Amel (2011).
8644:
Sengupta, Sanghamitra; Zhivotovsky, Lev A.; King, Roy; Mehdi, S.Q; Edmonds, Christopher A.; Chow, Cheryl-Emiliane T.; Lin, Alice A.; Mitra, Mitashree; et al. (2006).
7836:
Hammer, Michael F.; Behar, Doron M.; Karafet, Tatiana M.; Mendez, Fernando L.; Hallmark, Brian; Erez, Tamar; Zhivotovsky, Lev A.; Rosset, Saharon; Skorecki, Karl (2009).
7334:
Capelli, Cristian; Onofri, Valerio; Brisighelli, Francesca; Boschi, Ilaria; Scarnicci, Francesca; Masullo, Mara; Ferri, Gianmarco; Tofanelli, Sergio; et al. (2009).
8599:
Semino, Ornella; Magri, Chiara; Benuzzi, Giorgia; Lin, Alice A.; Al-Zahery, Nadia; Battaglia, Vincenza; Maccioni, Liliana; Triantaphyllidis, Costas; et al. (2004).
4909:
4919:
4914:
7488:
Di Gaetano, Cornelia; Cerutti, Nicoletta; Crobu, Francesca; Robino, Carlo; Inturri, Serena; Gino, Sarah; Guarrera, Simonetta; Underhill, Peter A; et al. (2009).
4812:
circa 11,000 BC, specifically belonging to the rare J1-FT34521 subclade. The ancient individual from Satsurblia was male with black hair, brown eyes, and light skin.
8206:
Msaidie, Said; Ducourneau, Axel; Boetsch, Gilles; Longepied, Guy; Papa, Kassim; Allibert, Claude; Yahaya, Ali Ahmed; Chiaroni, Jacques; Mitchell, Michael J (2011).
9093:
Behar, Doron M.; Thomas, Mark G.; Skorecki, Karl; Hammer, Michael F.; Bulygina, Ekaterina; Rosengarten, Dror; Jones, Abigail L.; Held, Karen; et al. (2003).
7221:"High-Resolution Analysis of Human Y-Chromosome Variation Shows a Sharp Discontinuity and Limited Gene Flow between Northwestern Africa and the Iberian Peninsula"
5532:
Van Oven M, Van Geystelen A, Kayser M, Decorte R, Larmuseau HD (2014). "Seeing the wood for the trees: a minimal reference phylogeny for the human Y chromosome".
9217:
Flores, Carlos; Maca-Meyer, Nicole; GonzĂĄlez, Ana M; Oefner, Peter J; Shen, Peidong; PĂ©rez, Jose A; Rojas, Antonio; Larruga, Jose M; Underhill, Peter A (2004).
4849:
9571:
8729:
Shlush, Liran I.; Behar, Doron M.; Yudkovsky, Guennady; Templeton, Alan; Hadid, Yarin; Basis, Fuad; Hammer, Michael; Itzkovitz, Shalev; Skorecki, Karl (2008).
3863:
The Caucasus has areas of both high and low J-M267 frequency. The J-M267 in the Caucasus is also notable because most of it is not within the J-P58 subclade.
7407:
Chiaroni, Jacques; King, Roy J; Myres, Natalie M; Henn, Brenna M; Ducourneau, Axel; Mitchell, Michael J; Boetsch, Gilles; Sheikha, Issa; et al. (2010).
8784:
Tofanelli, Sergio; Ferri, Gianmarco; Bulayeva, Kazima; Caciagli, Laura; Onofri, Valerio; Taglioli, Luca; Bulayev, Oleg; Boschi, Ilaria; et al. (2009).
4527:
take a stronger position of rejecting any strong correlation between the Arab expansion and either the YCAII=22-22 STR-defined sub-cluster as discussed by
7531:
El-Sibai, Mirvat; Platt, Daniel E.; Haber, Marc; Xue, Yali; Youhanna, Sonia C.; Wells, R. Spencer; Izaabel, Hassan; Sanyoura, May F.; et al. (2009).
6112:
8874:
Zalloua, Pierre A.; Xue, Yali; Khalife, Jade; Makhoul, Nadine; Debiane, Labib; Platt, Daniel E.; Royyuru, Ajay K.; Herrera, Rene J.; et al. (2008).
7176:
Battaglia, Vincenza; Fornarino, Simona; Al-Zahery, Nadia; Olivieri, Anna; Pala, Maria; Myres, Natalie M; King, Roy J; Rootsi, Siiri; et al. (2009).
4899:
219:). Because of the dominance of J-P58 in J-M267 populations in many areas, discussion of J-M267's origins require a discussion of J-P58 at the same time.
4549:. They also said that by using more markers and a more restrictive definition, the estimated age of the Cohanim lineage is lower than the estimates of
8919:
Zalloua, Pierre A.; Platt, Daniel E.; El Sibai, Mirvat; Khalife, Jade; Makhoul, Nadine; Haber, Marc; Xue, Yali; Izaabel, Hassan; et al. (2008).
4516:
More recent studies have emphasized doubt that the Islamic expansions are old enough to completely explain the major patterns of J-M267 frequencies.
8470:
Regueiro, M.; Cadenas, A.M.; Gayden, T.; Underhill, P.A.; Herrera, R.J. (2006). "Iran: tricontinental nexus for Y-chromosome driven migration".
8208:"Genetic diversity on the Comoros Islands shows early seafaring as major determinant of human biocultural evolution in the Western Indian Ocean"
5510:
8197:
4553:, and it is consistent with a common ancestor at the approximate time of founding of the priesthood which is the source of Cohen surnames.
9303:
This table shows the historic names for J-M267 and its earlier discovered and named subclade J-M62 in published peer reviewed literature.
7047:
Alvarez, Luis; Santos, Cristina; Montiel, Rafael; Caeiro, Blazquez; Baali, Abdellatif; Dugoujona, Jean-Michel; Aluja, Maria Pilar (2009).
6160:
Bekada, Asmahan; Fregel, Rosa; Cabrera, Vicente M.; Larruga, José M.; Pestano, José; Benhamamouch, Soraya; Gonzålez, Ana M. (2013-02-19).
6423:"A finely resolved phylogeny of Y chromosome Hg J illuminates the processes of Phoenician and Greek colonizations in the Mediterranean"
4839:
4355:
recognize signs that J-M267 might have an older presence, and on balance they accept the evidence but note that it could be in error.
9174:
Firasat, Sadaf; Khaliq, Shagufta; Mohyuddin, Aisha; Papaioannou, Myrto; Tyler-Smith, Chris; Underhill, Peter A; Ayub, Qasim (2006).
4994:
4924:
4869:
8084:"The coming of the Greeks to Provence and Corsica: Y-chromosome models of archaic Greek colonization of the western Mediterranean"
2343:
J-M267 is very common throughout this region, dominated by J-P58, but some specific sub-populations have notably low frequencies.
9620:
5997:
Fadhlaoui-Zid, Karima; Haber, Marc; MartĂnez-Cruz, Begoña; Zalloua, Pierre; Benammar Elgaaied, Amel; Comas, David (2013-11-27).
9576:
4984:
4379:
haplotypes - specifically YCAII as 22-22, and DYS388 having unusual repeat values of 15 or higher, instead of more typical 13 (
9219:"Reduced genetic structure of the Iberian peninsula revealed by Y-chromosome analysis: Implications for population demography"
7090:
Arredi, B; Poloni, E; Paracchini, S; Zerjal, T; Fathallah, D; Makrelouf, M; Pascali, V; Novelletto, A; Tyler-Smith, C (2004).
211:
J-M267 has several recognized subclades, some of which were recognized before J-M267 itself was recognized, for example J-M62
7703:
Fregel, Rosa; Gomes, VerĂłnica; GusmĂŁo, Leonor; GonzĂĄlez, Ana M; Cabrera, Vicente M; Amorim, AntĂłnio; Larruga, Jose M (2009).
7133:
Balanovsky, O.; Dibirova, K.; Dybo, A.; Mudrak, O.; Frolova, S.; Pocheshkhova, E.; Haber, M.; Platt, D.; et al. (2011).
4460:
also noted the anomalously high apparent age of Omani J-M267 when looking more generally at J-P58 and J-M267 more generally.
144:
4328:. The authors propose that this involved a spread of some J-P58 from the direction of Syria towards Arab populations of the
8156:
Luis, J; Rowold, D; Regueiro, M; Caeiro, B; CinnioÄlu, C; Roseman, C; Underhill, P; Cavalli-Sforza, L; Herrera, R (2004).
7219:
Bosch, Elena; Calafell, Francesc; Comas, David; Oefner, Peter J.; Underhill, Peter A.; Bertranpetit, Jaume (April 2001).
5890:"Synthetic review on the genetic relatedness between North Africa and Arabia deduced from paternal lineage distributions"
5875:"Synthetic review on the genetic relatedness between North Africa and Arabia deduced from paternal lineage distributions"
4467:", found among Jewish populations, but especially in men with surnames related to Cohen. It also contains the "Galilee
7490:"Differential Greek and northern African migrations to Sicily are supported by genetic evidence from the Y chromosome"
2893:
7668:
Flores, Carlos; Maca-Meyer, Nicole; Larruga, Jose M.; Cabrera, Vicente M.; Karadsheh, Naif; Gonzalez, Ana M. (2005).
7002:
Abu-Amero, Khaled K; Hellani, Ali; GonzĂĄlez, Ana M; Larruga, Jose M; Cabrera, Vicente M; Underhill, Peter A (2009).
4463:
This cluster in turn contains three well-known related sub-clusters. First, it contains the majority of the Jewish "
2897:
5651:
Haplogroup K2b1 (P397/P399) is also known as Haplogroup MS, but has a broader and more complex internal structure.
4948:
4943:
4844:
118:
82:
5173:
4938:
4383:) This cluster was found to be relevant in some well-publicized studies of Jewish and Palestinian populations (
187:
surnames. It can also be found much less commonly, but still occasionally in significant amounts, in parts of
5736:"Dissecting the influence of Neolithic demic diffusion on Indian Y-chromosome pool through J2-M172 haplogroup"
6500:"Mitochondrial and Y chromosome haplotype motifs as diagnostic markers of Jewish ancestry: a reconsideration"
5113:
9095:"Multiple Origins of Ashkenazi Levites: Y Chromosome Evidence for Both Near Eastern and European Ancestries"
5697:
Haplogroup S, as of 2017, is also known as K2b1a. (Previously the name Haplogroup S was assigned to K2b1a4.)
5376:
5103:
4874:
4282:
7963:
Karafet, T. M.; Mendez, F. L.; Meilerman, M. B.; Underhill, Peter A.; Zegura, S. L.; Hammer, M. F. (2008).
7262:
Cadenas, Alicia M; Zhivotovsky, Lev A; Cavalli-Sforza, Luca L; Underhill, Peter A; Herrera, Rene J (2008).
6328:"Ancient Migratory Events in the Middle East: New Clues from the Y-Chromosome Variation of Modern Iranians"
5706:
Haplogroup M, as of 2017, is also known as K2b1b. (Previously the name Haplogroup M was assigned to K2b1d.)
5200:
5183:
4566:
offshoot of J-P58, that did not take part in the most recent waves of J-M267 expansion in the Middle East (
8519:
5365:
5357:
5097:
4962:
2541:
2507:
1599:
430:
6135:
5191:
7533:"Geographical structure of the Y-chromosomal genetic landscape of the Levant: a coastal-inland contrast"
5427:
5273:
4542:
4464:
4293:
may be ancestral. They also propose that this movement of people may also be linked to the dispersal of
6162:"Introducing the Algerian Mitochondrial DNA and Y-Chromosome Profiles into the North African Landscape"
5350:
5343:
5336:
4401:
refers to this overall cluster with YCAII=22-22 and high DYS388 values as an "Arabic" as opposed to a "
204:
in that region makes them candidates as genetic markers of the spread of farming technology during the
8012:"Congruent distribution of Neolithic painted pottery and ceramic figurines with Y-chromosome lineages"
7670:"Isolates in a corridor of migrations: a high-resolution analysis of Y-chromosome variation in Jordan"
9262:
8742:
7965:"New binary polymorphisms reshape and increase resolution of the human Y chromosomal haplogroup tree"
7923:
6434:
6339:
6173:
6010:
5821:
5747:
5466:
5385:
5320:
5264:
5249:
5235:
5156:
5128:
3153:
122:
31:
8524:
7336:"Moors and Saracens in Europe: estimating the medieval North African male legacy in southern Europe"
5734:
Singh S; Singh A; Rajkumar R; Sampath Kumar K; Kadarkarai Samy S; Nizamuddin S; et al. (2016).
5459:
5454:
4505:
4977:
4889:
4693:
Accounts for the majority of J1c3d in Saudi Arabia. An important element of J1c3d in North Africa.
4376:
4298:
7705:"Demographic history of Canary Islands male gene-pool: replacement of native lineages by European"
5889:
5874:
4800:. found a J1c3 haplotype in one of their ancient samples from Sardinia, dated to 6190â6000 calBP.
9162:
8921:"Identifying Genetic Traces of Historical Expansions: Phoenician Footprints in the Mediterranean"
8715:
8585:
8545:
8495:
8275:
8069:
8031:
7824:
7774:
7476:
7395:
7322:
7078:
6990:
6933:
6787:
6713:
6639:
6550:
6258:
6093:
5979:
5557:
5395:
5069:
5052:
4438:
3431:
2128:
8334:"Genetic evidence for the expansion of Arabian tribes into the Southern Levant and North Africa"
8158:"The Levant versus the Horn of Africa: evidence for bidirectional corridors of human migrations"
7838:"Extended Y chromosome haplotypes resolve multiple and unique lineages of the Jewish priesthood"
6486:
5999:"Genome-Wide and Paternal Diversity Reveal a Recent Origin of Human Populations in North Africa"
9278:
9240:
9205:
9176:"Y-chromosomal evidence for a limited Greek contribution to the Pathan population of Pakistan"
9154:
9124:
8950:
8905:
8860:
8815:
8770:
8707:
8675:
8630:
8577:
8537:
8487:
8456:
8400:
8363:
8318:
8267:
8237:
8187:
8144:
8115:
8061:
7994:
7951:
7897:
7867:
7816:
7766:
7736:
7691:
7656:
7627:
7597:
7562:
7519:
7468:
7438:
7365:
7314:
7285:
7250:
7207:
7164:
7121:
7070:
7035:
6982:
6925:
6845:
6827:
6779:
6705:
6631:
6531:
6468:
6450:
6375:
6357:
6307:
6250:
6209:
6191:
6127:
6085:
6046:
6028:
5971:
5935:
5855:
5837:
5773:
5626:
5607:
F-Y27277, sometimes known as F2'4, is both the parent clade of F2 and F4 and a child of F-M89.
5549:
5487:
5482:
5473:
5416:
5407:
5327:
5298:
5258:
5216:
5168:
5086:
4854:
4329:
4313:
4294:
3324:
152:
129:
6894:"Genomic History of Neolithic to Bronze Age Anatolia, Northern Levant, and Southern Caucasus"
6748:"Genomic History of Neolithic to Bronze Age Anatolia, Northern Levant, and Southern Caucasus"
6674:"Genomic History of Neolithic to Bronze Age Anatolia, Northern Levant, and Southern Caucasus"
6600:"Genomic History of Neolithic to Bronze Age Anatolia, Northern Levant, and Southern Caucasus"
9270:
9230:
9195:
9187:
9146:
9114:
9106:
9032:
8940:
8932:
8895:
8887:
8850:
8842:
8805:
8797:
8760:
8750:
8699:
8665:
8657:
8620:
8612:
8569:
8529:
8479:
8448:
8424:
8390:
8353:
8345:
8308:
8300:
8259:
8227:
8219:
8177:
8169:
8136:
8105:
8095:
8053:
8023:
7984:
7976:
7941:
7931:
7889:
7857:
7849:
7806:
7798:
7758:
7726:
7716:
7681:
7648:
7619:
7587:
7552:
7544:
7509:
7501:
7460:
7428:
7420:
7387:
7355:
7347:
7306:
7275:
7240:
7232:
7197:
7189:
7154:
7146:
7111:
7103:
7060:
7025:
7015:
6974:
6915:
6905:
6835:
6819:
6769:
6759:
6695:
6685:
6621:
6611:
6565:
6521:
6511:
6458:
6442:
6403:
6365:
6347:
6297:
6289:
6240:
6199:
6181:
6077:
6036:
6018:
5963:
5925:
5917:
5845:
5829:
5763:
5755:
5541:
5449:
5207:
5161:
4675:
4290:
9591:
5221:
4809:
4768:
4679:
4468:
4434:
4325:
4324:(approximately 3000â5000 BCE), and this involved the branch of Semitic which leads to the
3889:
3320:
2603:
2310:
2253:
2188:
188:
151:). This haplogroup is found today in significant frequencies in many areas in or near the
9008:
8207:
9600:
9266:
8746:
7927:
6438:
6343:
6177:
6014:
5825:
5751:
5023:
Please help update this article to reflect recent events or newly available information.
3052:
Please help update this article to reflect recent events or newly available information.
9200:
9175:
9119:
9094:
8945:
8920:
8900:
8875:
8810:
8785:
8765:
8730:
8670:
8645:
8625:
8600:
8313:
8288:
8232:
8182:
8157:
8110:
8083:
7989:
7964:
7946:
7911:
7862:
7837:
7731:
7704:
7557:
7532:
7514:
7489:
7433:
7408:
7360:
7335:
7245:
7220:
7202:
7177:
7159:
7134:
7116:
7091:
7030:
7003:
6840:
6526:
6499:
6463:
6422:
6370:
6327:
6302:
6277:
6204:
6161:
6041:
5998:
5930:
5905:
5850:
5809:
5768:
5735:
5437:
5230:
5063:
4970:
3937:
2966:
2405:
1562:
1481:
1452:
1422:
1392:
1191:
246:
140:
8855:
8830:
8358:
8333:
7652:
7049:"Y-chromosome variation in South Iberia: insights into the North African contribution"
5574:
4668:
Accounts for the majority of J1, the predominant haplogroup in the Arabian peninsula.
4316:, which possibly reached as far as Yemen, a more recent dispersal occurred during the
4281:
proposed that J-P58 (that they refer to as J1e) might have first dispersed during the
9614:
9063:
8057:
8035:
7802:
7548:
7399:
7178:"Y-chromosomal evidence of the cultural diffusion of agriculture in southeast Europe"
6937:
6791:
6717:
6643:
5983:
5630:
5312:
5306:
5292:
5281:
5244:
5147:
5142:
5133:
5119:
5080:
4784:
4480:
4271:). It is very prevalent in many areas where J-M267 is common, especially in parts of
4081:
4033:
3961:
2637:
1673:
464:
250:
180:
133:
9166:
8589:
8549:
8073:
8011:
7778:
7480:
7326:
6994:
6262:
6097:
6065:
5561:
3069:
J-P58 is the most common Y-Chromosome haplogroup among men from all of this region.
1772:
9044:
8719:
8499:
8379:"Phylogeographic analysis of paternal lineages in NE Portuguese Jewish communities"
8289:"The Y chromosome pool of Jews as part of the genetic landscape of the Middle East"
8279:
7828:
7082:
4756:
4476:
4442:
4317:
4272:
3098:
2753:
2726:
2722:
1868:
1834:
1830:
1636:
860:
254:
192:
176:
156:
72:
9274:
8988:
8004:
7310:
7092:"A predominantly neolithic origin for Y-chromosomal DNA variation in North Africa"
6808:"Mitogenome diversity in Sardinians : a genetic window onto an island's past"
4611:
Found at low frequency in the Armenian Highlands, Iran, Qatar and parts of Europe.
8755:
8428:
7936:
6352:
6186:
6023:
8976:
8287:
Nebel, A; Filon, D; Brinkmann, B; Majumder, P; Faerman, M; Oppenheim, A (2001).
7409:"The emergence of Y-chromosome haplogroup J1e among Arabic-speaking populations"
6278:"Traces of sub-Saharan and Middle Eastern lineages in Indian Muslim populations"
5906:"The emergence of Y-chromosome haplogroup J1e among Arabic-speaking populations"
4821:
4450:
4057:
2439:
1974:
1710:
9605:
9075:
9020:
8996:
8936:
8891:
7004:"Saudi Arabian Y-Chromosome diversity and its relationship with nearby regions"
6910:
6893:
6764:
6747:
6690:
6673:
6616:
6599:
6446:
5833:
9150:
9069:
This is an ongoing research project by citizen scientists. Over 950 J members.
8876:"Y-chromosomal diversity in Lebanon is structured by recent historical events"
8831:"A Nomenclature System for the Tree of Human Y-Chromosomal Binary Haplogroups"
8573:
8027:
7853:
7762:
7686:
7669:
7464:
7391:
6978:
4859:
4764:
4633:
Found sporadically in Anatolia, East Africa, the Arabian Peninsula and Europe.
4496:
Kurds" (even though both of these populations do have high levels of J-P209).
4321:
4216:
4133:
1942:
1228:
1046:
762:
426:
9595:
9038:
This is an ongoing research project by citizen scientists. Over 1050 members.
9002:
This is an ongoing research project by citizen scientists. Over 2300 members.
7912:"Y-Chromosome Diversity in Modern Bulgarians: New Clues about Their Ancestry"
6920:
6831:
6774:
6700:
6626:
6516:
6454:
6361:
6195:
6032:
5841:
5682:
4375:
This relatively young cluster, compared to J-M267 overall, was identified by
17:
9235:
9218:
9191:
9081:
This is an ongoing research project by citizen scientists. Over 400 members.
9050:
This is an ongoing research project by citizen scientists. Over 450 members.
9026:
This is an ongoing research project by citizen scientists. Over 550 members.
9014:
This is an ongoing research project by citizen scientists. Over 150 members.
8100:
7721:
7280:
7263:
7150:
7020:
6823:
4879:
4864:
4779:
A member of haplogroup J1-M267 is found among eastern hunter-gatherers from
4472:
4426:
4312:
According to this scenario, after the initial neolithic expansion involving
4105:
828:
205:
9282:
9244:
9209:
9158:
9128:
8954:
8909:
8864:
8819:
8786:"J1-M267 Y lineage marks climate-driven pre-historical human displacements"
8774:
8711:
8679:
8634:
8581:
8541:
8491:
8460:
8404:
8367:
8322:
8271:
8241:
8191:
8148:
8119:
8065:
7998:
7955:
7901:
7871:
7820:
7770:
7740:
7695:
7660:
7631:
7601:
7566:
7523:
7472:
7442:
7369:
7318:
7289:
7254:
7211:
7168:
7125:
7074:
7039:
6986:
6929:
6849:
6783:
6709:
6635:
6535:
6472:
6379:
6311:
6254:
6213:
6131:
6089:
6050:
5975:
5939:
5859:
5777:
5553:
4682:), but missing from Quraysh including Sharif of Makkah of Banu Hashem clan.
46:
8263:
8223:
7505:
7424:
7351:
7193:
6293:
6113:"History and geography of human Y-chromosome in Europe: a SNP perspective"
5921:
1791:
8801:
8332:
Nebel, A; Landau-Tasseron, E; Filon, D; Oppenheim, A; Faerman, M (2002).
7811:
4884:
4571:
4191:
4187:
4162:
4158:
4129:
3985:
3828:
3799:
3774:
3749:
3724:
3699:
3641:
3612:
3584:
3480:
2901:
2866:
2634:
2572:
2538:
2504:
2473:
2470:
1864:
1740:
1706:
1669:
1632:
1521:
1231:
498:
172:
168:
136:). (All these haplogroups have had other historical names listed below.)
114:
7592:
7575:
6807:
6245:
6228:
5589:
Haplogroup A0-T is also known as A-L1085 (and previously as A0'1'2'3'4).
4755:
who lived between 2565-1896 BC, belonged to J1-P58. Ebla was an ancient
8703:
8452:
8395:
8378:
8140:
7980:
7893:
7623:
7065:
7048:
6407:
5967:
5545:
4780:
4728:
4599:
clusters are found in the Armenian Highlands and parts of the Caucasus.
4546:
4501:
4410:
4402:
4287:
4009:
3913:
3832:
3552:
3504:
3296:
2993:
2690:
2664:
1801:
1769:
1364:
1360:
1328:
1299:
1295:
1261:
1224:
1194:
1187:
1157:
1150:
1113:
1083:
1079:
1049:
1042:
1013:
976:
944:
917:
888:
856:
824:
791:
758:
725:
695:
691:
661:
657:
625:
593:
559:
527:
332:
294:
212:
208:, which is proposed to have had a major impact upon human populations.
9076:"J2-Middle East Project Ù
ێ۱ÙŰč ŰłÙۧÙŰ© ŰŹ2 ÙÙ Ű§ÙŰčۧÙÙ
ۧÙŰčŰ±ŰšÙ ÙۧÙŰŽŰ±Ù Ű§ÙŰŁÙ۳۷"
8846:
8483:
6863:
6569:
6066:"Y chromosome sequence variation and the history of human populations"
5759:
5675:
5629:
was known as "Haplogroup K2". That name has since been re-assigned to
4351:
The Armenian Highlands, northern Iraq and western Iran. In this area,
5686:, 2017, "Details of the Y-SNP markers included in the minimal Y tree"
4406:
3803:
3695:
3670:
3559:
3556:
3528:
3456:
3427:
3402:
3373:
3211:
2862:
2831:
2805:
2779:
2306:
2066:
1154:
1117:
948:
795:
9586:
9110:
8661:
8616:
8533:
8377:
Nogueiro, I.; Manco, L.; Gomes, V.; Amorim, A.; GusmĂŁo, L. (2010).
8349:
8304:
8173:
7236:
7107:
5642:
Haplogroup K2b (M1221/P331/PF5911) is also known as Haplogroup MPS.
215:. With one notable exception, J-P58, most of these are not common (
143:
ancestor, which is demonstrated and defined by the presence of the
8972:
7135:"Parallel evolution of genes and languages in the Caucasus region"
6657:
6081:
4760:
4740:
4570:). These DYS388=13 haplotypes are most common in the Caucasus and
4453:
4430:
4422:
4414:
4333:
3666:
3637:
3608:
3377:
3349:
3182:
3124:
2997:
2963:
2932:
2835:
2693:
2630:
2599:
2568:
2534:
2500:
2466:
2432:
2401:
2372:
2219:
2157:
2002:
1970:
1938:
1595:
1558:
1517:
1265:
1120:
980:
729:
629:
597:
563:
531:
494:
460:
422:
393:
361:
298:
184:
167:. It is also found at very high but lesser extent in parts of the
164:
126:
6877:
6276:
Eaaswarkhanth, M; Haque, I; Ravesh, Z; et al. (March 2010).
9581:
7297:
Cann, H. M. (2002). "A Human Genome Diversity Cell Line Panel".
6731:
6583:
4752:
4446:
4418:
4344:
The Levant (Syria, Jordan, Israel and Palestine). In this area,
3234:
2959:
2928:
2436:
2280:
2249:
2215:
2184:
2153:
2124:
2095:
1332:
302:
160:
5810:"Origin and diffusion of human Y chromosome haplogroup J1-M267"
4966:
4731:
who lived between 1930-1325 BC, belonged to haplogroup J1-P58.
4644:
Found at low frequency in the Levant and the Arabian Peninsula.
7576:"Mitochondrial DNA and Y-chromosome microstructure in Tunisia"
5003:
3032:
5790:
4358:
The southern area of Oman, Yemen and Ethiopia. In this area,
4743:
who lived c. 3491-3122 BC, belonged to haplogroup J1-Z1824.
147:(SNP) mutation referred to as M267, which was announced in (
8731:"The Druze: a population genetic refugium of the Near East"
4905:
Y-DNA haplogroups in populations of East and Southeast Asia
4674:
Accounts for the majority of J1 in Yemen, Cohen Jews (both
4471:
haplotype" (GMH) and "Palestinian & Israeli Arab modal
8417:
Forensic Science International: Genetics Supplement Series
5575:
International Society of Genetic Genealogy (ISOGG; 2015),
4531:
or the smaller "Galilee modal haplotype" as discussed by (
5616:
Haplogroup LT (L298/P326) is also known as Haplogroup K1.
8979:. International Society of Genetic Genealogists "ISOGG".
30:"Haplogroup J1" redirects here. Not to be confused with
9057:
Haplogroup-Specific Ethnic/Geographical Group Projects
7264:"Y-chromosome diversity characterizes the Gulf of Oman"
6229:"Sousse: extreme genetic heterogeneity in North Africa"
6558:
Karadeniz AraĆtırmaları (Journal of Black Sea Studies)
1909:
J*(xJ-M172) was found in India among Indian Muslims.
8982:
Ongoing Corrections/Additions by citizen scientists.
4895:
Y-chromosome haplogroups in populations of the world
8995:Schrack; Janzen; Rottensteiner; Ricci; Mas (2013).
5045:
4824:who lived circa 5706-5622 BC, belonged to J1-L620.
96:
88:
78:
68:
56:
39:
4910:Y-DNA haplogroups in populations of the Near East
4259:), but had been announced earlier under the name
51:Interpolated geographical frequency distribution.
4920:Y-DNA haplogroups in populations of the Caucasus
4915:Y-DNA haplogroups in populations of North Africa
5688:(Access date of these pages: 9 December 2017)
4500:noted not only the presence of the GMH in the
4978:
4850:Conversion table for Y chromosome haplogroups
2023:
1995:
1963:
1286:
1252:
1215:
8:
4405:" type of J-M267. This Arabic type includes
9606:Y-DNA Haplogroup J and its Subclades - 2011
5660:Haplogroup P (P295) is also klnown as K2b2.
179:and amongst most Levant peoples, including
8987:Haplotype/SNP research Projects. See also
4985:
4971:
4963:
4900:Y-DNA haplogroups in populations of Europe
4233:
4209:
4180:
4151:
4122:
4098:
4074:
4050:
4026:
4002:
3978:
3954:
3930:
3906:
3865:
3688:
3545:
3521:
3497:
3473:
3272:
3071:
2345:
2039:
1911:
264:
163:, it is found at very high frequencies in
61:
45:
9234:
9199:
9118:
8944:
8899:
8854:
8809:
8764:
8754:
8669:
8624:
8523:
8441:American Journal of Physical Anthropology
8394:
8383:American Journal of Physical Anthropology
8357:
8312:
8231:
8181:
8129:American Journal of Physical Anthropology
8109:
8099:
7988:
7945:
7935:
7882:American Journal of Physical Anthropology
7861:
7810:
7730:
7720:
7685:
7612:American Journal of Physical Anthropology
7591:
7556:
7513:
7432:
7359:
7279:
7244:
7201:
7158:
7115:
7064:
7029:
7019:
6919:
6909:
6839:
6773:
6763:
6699:
6689:
6625:
6615:
6525:
6515:
6462:
6369:
6351:
6301:
6244:
6203:
6185:
6040:
6022:
5929:
5849:
5767:
4575:
4550:
4524:
4398:
3850:
3821:
3792:
3767:
3742:
3630:
3601:
3577:
3342:
3115:
2242:
2088:
1547:
216:
148:
100:M267, L255, L321, L765, L814, L827, L1030
9457:
9304:
9045:"J-L24-Y-DNA Haplogroup Project (J2a1h)"
9043:Aburto; Katz; Al Gazzah; Janzen (2013).
5598:Haplogroup A1 is also known as A1'2'3'4.
4567:
4517:
4457:
4392:
4380:
4359:
4352:
4345:
4306:
4278:
4268:
3366:
3251:
3199:
3170:
3141:
3015:
2919:
2822:
2796:
2770:
2744:
2710:
2681:
2655:
2621:
2590:
2559:
2525:
2491:
2457:
2423:
2394:
2389:
2328:
2297:
2271:
2237:
2206:
2175:
2117:
2112:
2083:
1694:
1657:
1620:
1583:
1542:
451:
413:
381:
9296:
8562:International Journal of Legal Medicine
5726:
5524:
4662:Found at low frequency in South Arabia.
4367:The "YCAII=22-22 and DYS388â„15" cluster
4264:
4256:
3659:
3204:
3175:
3146:
1821:
1178:
1141:
1104:
1070:
259:
9099:The American Journal of Human Genetics
8989:Y-DNA haplogroup projects (ISOGG Wiki)
8977:"Y-DNA Haplogroup J and its Subclades"
8829:Y Chromosome Consortium "YCC" (2002).
8162:The American Journal of Human Genetics
7225:The American Journal of Human Genetics
6498:Tofanelli, Sergio (10 November 2014).
4605:Found at very low frequency in Europe.
4538:
4528:
4509:
4488:
4388:
4251:The P58 marker which defines subgroup
3449:
3313:
2984:
2950:
2884:
2853:
2715:
1699:
1662:
1625:
1588:
1551:
1510:
1501:
1472:
1443:
1413:
1033:
1006:
1001:
908:
716:
682:
584:
485:
352:
323:
242:
238:
36:
4808:An ancient sample of J1 was found at
4532:
4506:expansions from the Arabian peninsula
4497:
4492:
4484:
4475:", both of which are associated with
4384:
1760:
1731:
1506:
139:Men from this lineage share a common
7:
8010:King, R; Underhill, Peter A (2002).
6064:Underhill, Peter A (December 2000).
5802:
5800:
5798:
5511:Y-DNA haplogroups of historic people
5486:
5481:
5472:
5470:
5465:
5463:
5458:
5453:
5448:
5442:
5435:
5425:
5423:
5413:
5405:
5393:
5391:
5382:
5374:
5332:
5325:
5319:
5317:
5311:
5305:
5303:
5297:
5291:
5279:
5269:
5263:
5257:
5248:
5243:
5234:
5229:
5220:
5215:
5179:
5165:
5048:
4739:One out of 18 male individuals from
4302:
3717:
3420:
3395:
3256:
3020:
1796:J-P58 might be 5% in upcoming study
386:
60:17,000â24,000 years before present (
6120:Journal of Anthropological Sciences
5153:
5093:
5076:
4820:One out of 4 male individuals from
4541:disagreed, at least concerning the
9223:European Journal of Human Genetics
9180:European Journal of Human Genetics
8925:American Journal of Human Genetics
8880:American Journal of Human Genetics
8790:European Journal of Human Genetics
8650:American Journal of Human Genetics
8605:American Journal of Human Genetics
8338:American Journal of Human Genetics
8293:American Journal of Human Genetics
8212:European Journal of Human Genetics
7494:European Journal of Human Genetics
7413:European Journal of Human Genetics
7340:European Journal of Human Genetics
7268:European Journal of Human Genetics
7182:European Journal of Human Genetics
7096:American Journal of Human Genetics
6551:"The Genetics of Crimean Karaites"
6282:European Journal of Human Genetics
5910:European Journal of Human Genetics
4840:Genetic history of the Middle East
4727:Five out 12 male individuals from
3093:Previous research on same samples
2367:Previous research on same samples
2061:Previous research on same samples
289:previous research on same samples
213:Y Chromosome Consortium "YCC" 2002
25:
9031:Cone; Al Gazzah; Sanders (2013).
7053:American Journal of Human Biology
6396:American Journal of Human Biology
5956:American Journal of Human Biology
5580:. (Access date: 1 February 2015.)
4925:Y-DNA haplogroups by ethnic group
4870:Human Y-chromosome DNA haplogroup
4735:Arslantepe archaeological complex
4622:Found at low frequency in Europe.
27:Human Y-chromosome DNA haplogroup
9007:Givargidze; Hrechdakian (2013).
8058:10.1111/j.1469-1809.2007.00414.x
7803:10.1111/j.1529-8817.2005.00161.x
7549:10.1111/j.1469-1809.2009.00538.x
5008:
4751:Three out of 6 individuals from
3037:
92:J-M62, J-M365.1, J-L136, J-Z1828
9582:Y-Full J-M267 Phylogenetic Tree
7139:Molecular Biology and Evolution
6812:Molecular Biology and Evolution
6549:Brook, Kevin A. (Summer 2014).
5633:, the sibling of Haplogroup LT.
258:rural total population (2.5%) (
233:North Africa and Horn of Africa
6227:Fadhlaoui-Zid, Karima (2014).
4759:-speaking city and kingdom in
4582:Phylogenetics and distribution
4574:, but also found in Ethiopia (
2339:Levant and Semitic populations
145:single nucleotide polymorphism
1:
9275:10.1126/science.290.5494.1155
7653:10.1016/S0198-8859(01)00311-1
7311:10.1126/science.296.5566.261b
9577:J-M267 Relevant Publications
9471:Jobling and Tyler-Smith 2000
9319:Jobling and Tyler-Smith 2000
8997:"Y-DNA J Haplogroup Project"
8756:10.1371/journal.pone.0002105
8429:10.1016/j.fsigss.2007.10.173
7937:10.1371/journal.pone.0056779
6353:10.1371/journal.pone.0041252
6187:10.1371/journal.pone.0056775
6024:10.1371/journal.pone.0080293
5904:Underhill, Peter A. (2010).
4995:Y-chromosome DNA haplogroups
9033:"J-M172 Y-DNA Project (J2)"
6111:Francalacci, Paolo (2008).
4993:Phylogenetic tree of human
9637:
9572:J-M267 Ancient Samples Map
9461:YCC 2002/2008 (Shorthand)
9308:YCC 2002/2008 (Shorthand)
8975:; Schrack, Janzen (2013).
8937:10.1016/j.ajhg.2008.10.012
8892:10.1016/j.ajhg.2008.01.020
6911:10.1016/j.cell.2020.04.044
6765:10.1016/j.cell.2020.04.044
6691:10.1016/j.cell.2020.04.044
6617:10.1016/j.cell.2020.04.044
6447:10.1038/s41598-018-25912-9
5834:10.1038/s41598-021-85883-2
5577:Y-DNA Haplogroup Tree 2015
5479:
5446:
5403:
5372:
5363:
5355:
5348:
5341:
5334:
5289:
5255:
5241:
5227:
5213:
5197:
5181:
5050:
4767:that was destroyed by the
4723:Alalakh Amorite city-state
29:
9466:(AKA J-12f2.1 or J-M304)
9151:10.1007/s00439-003-1073-7
8574:10.1007/s00414-007-0203-5
8028:10.1017/s0003598x00091158
7854:10.1007/s00439-009-0727-5
7763:10.1007/s00439-004-1168-9
7687:10.1007/s10038-005-0274-4
7674:Journal of Human Genetics
7580:Journal of Human Genetics
7465:10.1007/s00439-003-1031-4
7392:10.1017/s0003598x00096800
6979:10.1007/s00439-004-1202-y
6487:Percentage of haplogroups
6233:Journal of Human Genetics
5500:
5278:
5271:
5205:
5188:
5172:
5167:
5160:
5155:
5146:
5141:
5139:
5132:
5127:
5125:
5118:
5111:
5109:
5102:
5095:
5085:
5078:
5068:
5061:
5059:
5043:
5017:This article needs to be
5001:
4845:Genetic history of Europe
4656:Previously known as J1e2.
4650:Previously known as J1e1.
4508:" during the 7th century.
4267:and called that again in
3046:This article needs to be
937:Pestano J, et al. (2013)
125:) along with its sibling
109:, also commonly known as
44:
9587:J-M267 Phylogenetic Tree
8088:BMC Evolutionary Biology
8046:Annals of Human Genetics
7791:Annals of Human Genetics
7709:BMC Evolutionary Biology
7537:Annals of Human Genetics
6564:(42): 69â84 on page 83.
6517:10.3389/fgene.2014.00384
183:, especially those with
69:Possible place of origin
9621:Human Y-DNA haplogroups
9236:10.1038/sj.ejhg.5201225
9192:10.1038/sj.ejhg.5201726
8101:10.1186/1471-2148-11-69
8005:Supplementary Material.
7722:10.1186/1471-2148-9-181
7281:10.1038/sj.ejhg.5201934
7021:10.1186/1471-2156-10-59
5625:Between 2002 and 2008,
4875:Molecular phylogenetics
4283:Pre-Pottery Neolithic B
119:Y-DNA haplogroup J-P209
57:Possible time of origin
8967:Haplogroups/Phylogeny
6485:El-Sibai et al.,2009,
4702:J1c3d2a (L65.2/S159.2)
4648:J1c3a (M367.1, M368.1)
8264:10.1007/s004390000426
8224:10.1038/ejhg.2010.128
7506:10.1038/ejhg.2008.120
7425:10.1038/ejhg.2009.166
7352:10.1038/ejhg.2008.258
7194:10.1038/ejhg.2008.249
7151:10.1093/molbev/msr126
6824:10.1093/molbev/msx082
6504:Frontiers in Genetics
6294:10.1038/ejhg.2009.168
5922:10.1038/ejhg.2009.166
4591:J1 (L255, L321, M267)
4543:Cohen modal haplotype
4465:Cohen modal haplotype
4242:Subclade Distribution
9551:YCC 2010r (Longhand)
9429:YCC 2010r (Longhand)
8802:10.1038/ejhg.2009.58
6904:(5): 1158â1175.e28.
6758:(5): 1158â1175.e28.
6684:(5): 1158â1175.e28.
6610:(5): 1158â1175.e28.
4747:Ancient city of Ebla
4445:in the dispersal of
3270:J-M267 chromosomes.
3154:United Arab Emirates
159:. Out of its native
32:Haplogroup J (mtDNA)
9543:YCC 2008 (Longhand)
9535:YCC 2005 (Longhand)
9527:YCC 2002 (Longhand)
9418:YCC 2008 (Longhand)
9407:YCC 2005 (Longhand)
9396:YCC 2002 (Longhand)
9267:2000Sci...290.1155S
9009:"J1* Y-DNA Project"
8747:2008PLoSO...3.2105S
7928:2013PLoSO...856779K
7593:10.1038/jhg.2011.92
6439:2018NatSR...8.7465F
6344:2012PLoSO...741252G
6246:10.1038/jhg.2014.99
6178:2013PLoSO...856775B
6015:2013PLoSO...880293F
5826:2021NatSR..11.6659S
5752:2016NatSR...619157S
5506:Y-DNA by population
5421:
4959:Y-DNA Backbone Tree
4890:Y-chromosomal Aaron
969:Fadhlaoui-Zid 2015
191:and as far east as
132:(commonly known as
121:(commonly known as
9291:Phylogenetic Notes
9078:. Family Tree DNA.
9074:Al Gazzah (2013).
9066:. Family Tree DNA.
9047:. Family Tree DNA.
9035:. Family Tree DNA.
9023:. Family Tree DNA.
9019:Al Haddad (2013).
9011:. Family Tree DNA.
8999:. Family Tree DNA.
8704:10.1002/humu.20077
8453:10.1002/ajpa.21473
8396:10.1002/ajpa.21154
8141:10.1002/ajpa.21078
7981:10.1101/gr.7172008
7894:10.1002/ajpa.20876
7624:10.1002/ajpa.21581
7066:10.1002/ajhb.20888
6427:Scientific Reports
6408:10.1002/ajhb.22773
5968:10.1002/ajhb.22602
5814:Scientific Reports
5740:Scientific Reports
5546:10.1002/humu.22468
5272:
5053:Y-chromosomal Adam
4255:was announced in (
2024:Eaaswarkhanth 2009
1996:Eaaswarkhanth 2009
1964:Eaaswarkhanth 2009
1287:Fadhlaoui-Zid 2011
1253:Fadhlaoui-Zid 2011
1216:Fadhlaoui-Zid 2011
552:Fadhlaoui-Zid 2013
97:Defining mutations
9558:
9557:
9452:is a subclade of
9439:
9438:
8847:10.1101/gr.217602
8796:(11): 1520â1524.
8484:10.1159/000093774
8052:(Pt 2): 205â214.
7797:(Pt 4): 443â454.
7543:(Pt 6): 568â581.
7305:(5566): 261â262.
6921:20.500.12154/1254
6775:20.500.12154/1254
6701:20.500.12154/1254
6627:20.500.12154/1254
6570:10.12787/KARAM859
5760:10.1038/srep19157
5717:
5716:
5679:, 2017, "K-M2335"
5677:YFull YTree v5.08
5627:Haplogroup T-M184
5495:
5494:
5440:
5430:
5420:
5411:
5399:
5388:
5380:
5368:
5360:
5353:
5346:
5339:
5284:
5203:
5195:
5186:
5116:
5100:
5083:
5066:
5038:
5037:
4932:Y-DNA J Subclades
4855:Genetic Genealogy
4330:Arabian Peninsula
4314:Semitic languages
4295:Semitic languages
4239:
4238:
3856:
3855:
3325:Albanian speakers
3262:
3261:
3067:
3066:
3029:Arabian peninsula
3026:
3025:
2336:
2335:
2029:
2028:
1897:
1896:
650:Francalacci 2008
153:Arabian Peninsula
130:haplogroup J-M172
107:Haplogroup J-M267
104:
103:
40:Haplogroup J-M267
16:(Redirected from
9628:
9559:
9458:
9446:
9440:
9305:
9301:
9286:
9261:(5494): 1155â9.
9248:
9238:
9213:
9203:
9170:
9132:
9122:
9079:
9067:
9048:
9036:
9024:
9012:
9000:
8980:
8958:
8948:
8913:
8903:
8868:
8858:
8823:
8813:
8778:
8768:
8758:
8723:
8683:
8673:
8638:
8628:
8611:(5): 1023â1034.
8593:
8553:
8527:
8503:
8464:
8432:
8408:
8398:
8371:
8361:
8344:(6): 1594â1596.
8326:
8316:
8299:(5): 1095â1112.
8283:
8245:
8235:
8195:
8185:
8152:
8123:
8113:
8103:
8077:
8039:
8022:(293): 707â714.
8002:
7992:
7959:
7949:
7939:
7905:
7875:
7865:
7832:
7814:
7782:
7744:
7734:
7724:
7699:
7689:
7664:
7641:Human Immunology
7635:
7605:
7595:
7570:
7560:
7527:
7517:
7484:
7446:
7436:
7403:
7386:(316): 281â289.
7373:
7363:
7330:
7293:
7283:
7258:
7248:
7231:(4): 1019â1029.
7215:
7205:
7172:
7162:
7129:
7119:
7086:
7068:
7043:
7033:
7023:
6998:
6942:
6941:
6923:
6913:
6888:
6882:
6881:
6874:
6868:
6867:
6860:
6854:
6853:
6843:
6818:(5): 1230â1239.
6802:
6796:
6795:
6777:
6767:
6742:
6736:
6735:
6728:
6722:
6721:
6703:
6693:
6668:
6662:
6661:
6654:
6648:
6647:
6629:
6619:
6594:
6588:
6587:
6580:
6574:
6573:
6555:
6546:
6540:
6539:
6529:
6519:
6495:
6489:
6483:
6477:
6476:
6466:
6418:
6412:
6411:
6390:
6384:
6383:
6373:
6355:
6322:
6316:
6315:
6305:
6273:
6267:
6266:
6248:
6224:
6218:
6217:
6207:
6189:
6157:
6151:
6150:
6148:
6146:
6141:on 28 March 2012
6140:
6134:. Archived from
6117:
6108:
6102:
6101:
6061:
6055:
6054:
6044:
6026:
5994:
5988:
5987:
5950:
5944:
5943:
5933:
5900:
5894:
5893:
5885:
5879:
5878:
5870:
5864:
5863:
5853:
5804:
5793:
5788:
5782:
5781:
5771:
5731:
5707:
5704:
5698:
5695:
5689:
5667:
5661:
5658:
5652:
5649:
5643:
5640:
5634:
5623:
5617:
5614:
5608:
5605:
5599:
5596:
5590:
5587:
5581:
5572:
5566:
5565:
5529:
5436:
5426:
5414:
5406:
5394:
5384:
5375:
5364:
5356:
5349:
5342:
5335:
5280:
5199:
5190:
5182:
5112:
5096:
5079:
5062:
5046:
5033:
5030:
5024:
5012:
5011:
5004:
4987:
4980:
4973:
4964:
4785:Northeast Europe
4660:J1c3c (L92, L93)
4521:on the Maghreb.
4291:material culture
3866:
3273:
3072:
3062:
3059:
3053:
3041:
3040:
3033:
2346:
2040:
1912:
753:Aboukhalid 2010
265:
49:
37:
21:
9636:
9635:
9631:
9630:
9629:
9627:
9626:
9625:
9611:
9610:
9596:J1-M267 project
9568:
9563:
9562:
9465:
9447:
9443:
9302:
9298:
9293:
9251:
9216:
9173:
9135:
9092:
9089:
9087:Further reading
9073:
9062:Eddali (2013).
9061:
9042:
9030:
9021:"J1c3 (J-L147)"
9018:
9006:
8994:
8971:
8965:
8918:
8873:
8835:Genome Research
8828:
8783:
8728:
8688:
8643:
8598:
8558:
8525:10.1.1.537.1822
8508:
8469:
8437:
8413:
8376:
8331:
8286:
8248:
8205:
8155:
8126:
8080:
8042:
8009:
7969:Genome Research
7962:
7908:
7878:
7835:
7787:
7747:
7702:
7667:
7638:
7608:
7586:(10): 734â741.
7573:
7530:
7487:
7449:
7406:
7376:
7333:
7296:
7261:
7218:
7175:
7145:(10): 2905â20.
7132:
7089:
7046:
7001:
6964:
6961:
6956:
6951:
6946:
6945:
6890:
6889:
6885:
6876:
6875:
6871:
6864:"J-Y6313 YTree"
6862:
6861:
6857:
6804:
6803:
6799:
6744:
6743:
6739:
6730:
6729:
6725:
6670:
6669:
6665:
6658:"J-Z1842 YTree"
6656:
6655:
6651:
6596:
6595:
6591:
6582:
6581:
6577:
6553:
6548:
6547:
6543:
6497:
6496:
6492:
6484:
6480:
6420:
6419:
6415:
6392:
6391:
6387:
6324:
6323:
6319:
6275:
6274:
6270:
6226:
6225:
6221:
6159:
6158:
6154:
6144:
6142:
6138:
6115:
6110:
6109:
6105:
6070:Nature Genetics
6063:
6062:
6058:
5996:
5995:
5991:
5952:
5951:
5947:
5902:
5901:
5897:
5887:
5886:
5882:
5872:
5871:
5867:
5806:
5805:
5796:
5789:
5785:
5733:
5732:
5728:
5723:
5718:
5713:
5712:
5711:
5710:
5705:
5701:
5696:
5692:
5668:
5664:
5659:
5655:
5650:
5646:
5641:
5637:
5624:
5620:
5615:
5611:
5606:
5602:
5597:
5593:
5588:
5584:
5573:
5569:
5531:
5530:
5526:
5521:
5515:
5496:
5039:
5034:
5028:
5025:
5022:
5013:
5009:
4997:
4991:
4961:
4956:
4934:
4929:
4835:
4830:
4818:
4810:Satsurblia Cave
4806:
4794:
4777:
4749:
4737:
4725:
4720:
4691:J1c3d2 (L222.2)
4686:J1c3d1 (L174.1)
4584:
4563:
4369:
4326:Arabic language
4249:
4244:
4234:Di Giacomo 2004
4210:Balanovsky 2011
4181:Balanovsky 2011
4152:Balanovsky 2011
4123:Balanovsky 2011
4099:Balanovsky 2011
4075:Balanovsky 2011
4051:Balanovsky 2011
4027:Balanovsky 2011
4003:Balanovsky 2011
3979:Balanovsky 2011
3955:Balanovsky 2011
3931:Balanovsky 2011
3907:Balanovsky 2011
3861:
3689:Di Gaetano 2008
3546:Di Giacomo 2004
3522:Di Giacomo 2004
3498:Karachanak 2013
3474:Di Giacomo 2004
3321:North Macedonia
3267:
3063:
3057:
3054:
3051:
3042:
3038:
3031:
2542:Syriac Catholic
2508:Syriac Catholic
2341:
2034:
1907:
1902:
618:Underhill 2000
245:). However the
235:
230:
225:
201:
189:southern Europe
161:Asian Continent
62:Di Giacomo 2004
52:
35:
28:
23:
22:
15:
12:
11:
5:
9634:
9632:
9624:
9623:
9613:
9612:
9609:
9608:
9603:
9598:
9589:
9584:
9579:
9574:
9567:
9566:External links
9564:
9561:
9560:
9556:
9555:
9552:
9548:
9547:
9544:
9540:
9539:
9536:
9532:
9531:
9528:
9524:
9523:
9520:
9516:
9515:
9512:
9508:
9507:
9504:
9500:
9499:
9496:
9492:
9491:
9488:
9484:
9483:
9480:
9479:Underhill 2000
9476:
9475:
9472:
9468:
9467:
9462:
9441:
9437:
9436:
9433:
9430:
9426:
9425:
9422:
9419:
9415:
9414:
9411:
9408:
9404:
9403:
9400:
9397:
9393:
9392:
9389:
9386:
9382:
9381:
9378:
9375:
9371:
9370:
9367:
9364:
9360:
9359:
9356:
9353:
9349:
9348:
9345:
9342:
9338:
9337:
9334:
9331:
9330:Underhill 2000
9327:
9326:
9323:
9320:
9316:
9315:
9312:
9309:
9295:
9294:
9292:
9289:
9288:
9287:
9249:
9229:(10): 855â63.
9214:
9171:
9139:Human Genetics
9133:
9111:10.1086/378506
9088:
9085:
9084:
9083:
9071:
9055:
9054:
9053:
9052:
9040:
9028:
9016:
8985:
8984:
8964:
8961:
8960:
8959:
8931:(5): 633â642.
8915:
8914:
8886:(4): 873â882.
8870:
8869:
8825:
8824:
8780:
8779:
8725:
8724:
8698:(3): 248â260.
8692:Human Mutation
8685:
8684:
8662:10.1086/499411
8656:(2): 202â221.
8640:
8639:
8617:10.1086/386295
8595:
8594:
8568:(3): 251â255.
8555:
8554:
8534:10.1038/ng1754
8518:(4): 463â467.
8505:
8504:
8478:(3): 132â143.
8472:Human Heredity
8466:
8465:
8447:(1): 118â124.
8434:
8433:
8410:
8409:
8389:(3): 373â381.
8373:
8372:
8350:10.1086/340669
8328:
8327:
8305:10.1086/324070
8284:
8258:(6): 630â641.
8246:
8202:
8201:
8174:10.1086/382286
8168:(3): 532â544.
8153:
8124:
8078:
8040:
8007:
7975:(5): 830â838.
7960:
7906:
7888:(3): 316â323.
7876:
7848:(5): 707â717.
7842:Human Genetics
7833:
7784:
7783:
7757:(5): 357â371.
7751:Human Genetics
7745:
7700:
7680:(9): 435â441.
7665:
7647:(9): 949â953.
7636:
7618:(2): 271â280.
7606:
7571:
7528:
7485:
7459:(2): 127â148.
7453:Human Genetics
7447:
7419:(3): 348â353.
7404:
7374:
7331:
7294:
7259:
7237:10.1086/319521
7216:
7173:
7130:
7108:10.1086/423147
7087:
7044:
6999:
6967:Human Genetics
6960:
6957:
6955:
6952:
6950:
6947:
6944:
6943:
6883:
6878:"J-L620 YTree"
6869:
6855:
6797:
6737:
6723:
6663:
6649:
6589:
6575:
6541:
6490:
6478:
6413:
6402:(2): 203â212.
6385:
6317:
6268:
6219:
6152:
6103:
6056:
6009:(11): e80293.
5989:
5962:(6): 740â746.
5945:
5916:(3): 348â353.
5895:
5888:Rebai, Ahmed.
5880:
5873:Rebai, Ahmed.
5865:
5794:
5783:
5725:
5724:
5722:
5719:
5715:
5714:
5709:
5708:
5699:
5690:
5662:
5653:
5644:
5635:
5618:
5609:
5600:
5591:
5582:
5567:
5534:Human Mutation
5523:
5522:
5517:
5516:
5514:
5513:
5508:
5502:
5501:
5498:
5497:
5493:
5492:
5490:
5485:
5480:
5477:
5476:
5471:
5469:
5464:
5462:
5457:
5452:
5447:
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5443:
5441:
5434:
5432:
5424:
5422:
5412:
5404:
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5400:
5392:
5390:
5381:
5373:
5370:
5369:
5362:
5354:
5347:
5340:
5333:
5331:
5324:
5318:
5316:
5310:
5304:
5302:
5296:
5290:
5287:
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5270:
5268:
5262:
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5242:
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5233:
5228:
5225:
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5211:
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5204:
5196:
5187:
5180:
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5159:
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5131:
5126:
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5117:
5110:
5107:
5106:
5101:
5094:
5091:
5090:
5084:
5077:
5074:
5073:
5067:
5060:
5057:
5056:
5049:
5044:
5041:
5040:
5036:
5035:
5016:
5014:
5007:
5002:
4999:
4998:
4992:
4990:
4989:
4982:
4975:
4967:
4960:
4957:
4955:
4954:
4951:
4946:
4941:
4935:
4933:
4930:
4928:
4927:
4922:
4917:
4912:
4907:
4902:
4897:
4892:
4887:
4882:
4877:
4872:
4867:
4862:
4857:
4852:
4847:
4842:
4836:
4834:
4831:
4829:
4826:
4817:
4814:
4805:
4802:
4793:
4790:
4776:
4773:
4748:
4745:
4736:
4733:
4724:
4721:
4719:
4716:
4715:
4714:
4713:
4712:
4711:
4710:
4709:
4708:
4707:
4706:
4705:
4704:
4688:
4683:
4666:J1c3d (L147.1)
4663:
4657:
4651:
4645:
4634:
4628:
4623:
4612:
4606:
4600:
4583:
4580:
4576:Tofanelli 2009
4562:
4559:
4551:Tofanelli 2009
4525:Tofanelli 2009
4409:speakers from
4399:Tofanelli 2009
4368:
4365:
4364:
4363:
4356:
4349:
4299:hunter-herders
4248:
4245:
4243:
4240:
4237:
4236:
4231:
4228:
4225:
4222:
4219:
4213:
4212:
4207:
4204:
4201:
4198:
4195:
4184:
4183:
4178:
4175:
4172:
4169:
4166:
4155:
4154:
4149:
4146:
4143:
4140:
4137:
4126:
4125:
4120:
4117:
4114:
4111:
4108:
4102:
4101:
4096:
4093:
4090:
4087:
4084:
4078:
4077:
4072:
4069:
4066:
4063:
4060:
4054:
4053:
4048:
4045:
4042:
4039:
4036:
4030:
4029:
4024:
4021:
4018:
4015:
4012:
4006:
4005:
4000:
3997:
3994:
3991:
3988:
3982:
3981:
3976:
3973:
3970:
3967:
3964:
3958:
3957:
3952:
3949:
3946:
3943:
3940:
3934:
3933:
3928:
3925:
3922:
3919:
3916:
3910:
3909:
3904:
3901:
3898:
3895:
3892:
3886:
3885:
3882:
3879:
3876:
3873:
3870:
3860:
3857:
3854:
3853:
3851:Gonçalves 2005
3848:
3845:
3842:
3839:
3836:
3825:
3824:
3822:Gonçalves 2005
3819:
3816:
3813:
3810:
3807:
3796:
3795:
3793:Gonçalves 2005
3790:
3787:
3784:
3781:
3778:
3771:
3770:
3768:Gonçalves 2005
3765:
3762:
3759:
3756:
3753:
3746:
3745:
3743:Gonçalves 2005
3740:
3737:
3734:
3731:
3728:
3721:
3720:
3715:
3712:
3709:
3706:
3703:
3692:
3691:
3686:
3683:
3680:
3677:
3674:
3663:
3662:
3657:
3654:
3651:
3648:
3645:
3634:
3633:
3631:Battaglia 2008
3628:
3625:
3622:
3619:
3616:
3605:
3604:
3602:Battaglia 2008
3599:
3596:
3593:
3590:
3587:
3581:
3580:
3578:Battaglia 2008
3575:
3572:
3569:
3566:
3563:
3549:
3548:
3543:
3540:
3537:
3534:
3531:
3525:
3524:
3519:
3516:
3513:
3510:
3507:
3501:
3500:
3495:
3492:
3489:
3486:
3483:
3477:
3476:
3471:
3468:
3465:
3462:
3459:
3453:
3452:
3447:
3444:
3441:
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3435:
3424:
3423:
3418:
3415:
3412:
3409:
3406:
3399:
3398:
3393:
3390:
3387:
3384:
3381:
3370:
3369:
3364:
3361:
3358:
3355:
3352:
3346:
3345:
3343:Battaglia 2008
3340:
3337:
3334:
3331:
3328:
3317:
3316:
3311:
3308:
3305:
3302:
3299:
3293:
3292:
3289:
3286:
3283:
3280:
3277:
3266:
3263:
3260:
3259:
3254:
3249:
3246:
3243:
3240:
3237:
3231:
3230:
3228:
3226:
3223:
3220:
3217:
3214:
3208:
3207:
3202:
3197:
3194:
3191:
3188:
3185:
3179:
3178:
3173:
3168:
3165:
3162:
3159:
3156:
3150:
3149:
3144:
3139:
3136:
3133:
3130:
3127:
3121:
3120:
3118:
3116:Abu-Amero 2009
3113:
3110:
3107:
3104:
3101:
3095:
3094:
3091:
3088:
3085:
3082:
3079:
3076:
3065:
3064:
3045:
3043:
3036:
3030:
3027:
3024:
3023:
3018:
3013:
3010:
3007:
3004:
3001:
2990:
2989:
2987:
2982:
2979:
2976:
2973:
2970:
2956:
2955:
2953:
2948:
2945:
2942:
2939:
2936:
2925:
2924:
2922:
2917:
2914:
2911:
2908:
2905:
2898:TrĂĄs-os-Montes
2890:
2889:
2887:
2882:
2879:
2876:
2873:
2870:
2859:
2858:
2856:
2851:
2848:
2845:
2842:
2839:
2828:
2827:
2825:
2820:
2817:
2814:
2811:
2808:
2802:
2801:
2799:
2794:
2791:
2788:
2785:
2782:
2776:
2775:
2773:
2768:
2765:
2762:
2759:
2756:
2750:
2749:
2747:
2742:
2739:
2736:
2733:
2730:
2719:
2718:
2713:
2708:
2705:
2702:
2699:
2696:
2687:
2686:
2684:
2679:
2676:
2673:
2670:
2667:
2661:
2660:
2658:
2653:
2650:
2647:
2644:
2641:
2627:
2626:
2624:
2619:
2616:
2613:
2610:
2607:
2596:
2595:
2593:
2588:
2585:
2582:
2579:
2576:
2565:
2564:
2562:
2557:
2554:
2551:
2548:
2545:
2531:
2530:
2528:
2523:
2520:
2517:
2514:
2511:
2497:
2496:
2494:
2489:
2486:
2483:
2480:
2477:
2463:
2462:
2460:
2455:
2452:
2449:
2446:
2443:
2429:
2428:
2426:
2421:
2418:
2415:
2412:
2409:
2398:
2397:
2392:
2387:
2384:
2381:
2378:
2375:
2369:
2368:
2365:
2362:
2359:
2356:
2353:
2350:
2340:
2337:
2334:
2333:
2331:
2326:
2323:
2320:
2317:
2314:
2303:
2302:
2300:
2295:
2292:
2289:
2286:
2283:
2277:
2276:
2274:
2269:
2266:
2263:
2260:
2257:
2246:
2245:
2243:Tofanelli 2009
2240:
2235:
2232:
2229:
2226:
2223:
2212:
2211:
2209:
2204:
2201:
2198:
2195:
2192:
2181:
2180:
2178:
2173:
2170:
2167:
2164:
2161:
2150:
2149:
2147:
2146:Kivisild 2012
2144:
2141:
2138:
2135:
2132:
2121:
2120:
2115:
2110:
2107:
2104:
2101:
2098:
2092:
2091:
2089:CinnioÄlu 2004
2086:
2081:
2078:
2075:
2072:
2069:
2063:
2062:
2059:
2056:
2053:
2050:
2047:
2044:
2033:
2030:
2027:
2026:
2021:
2018:
2015:
2012:
2009:
2006:
1999:
1998:
1993:
1990:
1987:
1984:
1981:
1978:
1967:
1966:
1961:
1958:
1955:
1952:
1949:
1946:
1935:
1934:
1931:
1928:
1925:
1922:
1919:
1916:
1906:
1903:
1901:
1898:
1895:
1894:
1892:
1887:
1884:
1881:
1878:
1875:
1872:
1861:
1860:
1858:
1853:
1850:
1847:
1844:
1841:
1838:
1827:
1826:
1824:
1819:
1816:
1813:
1810:
1807:
1804:
1798:
1797:
1794:
1789:
1786:
1783:
1780:
1777:
1774:
1766:
1765:
1763:
1758:
1755:
1752:
1749:
1746:
1743:
1737:
1736:
1734:
1729:
1726:
1723:
1720:
1717:
1714:
1703:
1702:
1697:
1692:
1689:
1686:
1683:
1680:
1677:
1666:
1665:
1660:
1655:
1652:
1649:
1646:
1643:
1640:
1629:
1628:
1623:
1618:
1615:
1612:
1609:
1606:
1603:
1592:
1591:
1586:
1581:
1578:
1575:
1572:
1569:
1566:
1563:Sudanese Arabs
1555:
1554:
1548:Tofanelli 2009
1545:
1540:
1537:
1534:
1531:
1528:
1525:
1514:
1513:
1504:
1499:
1496:
1493:
1490:
1487:
1484:
1478:
1477:
1475:
1470:
1467:
1464:
1461:
1458:
1455:
1453:Canary Islands
1449:
1448:
1446:
1441:
1438:
1435:
1432:
1429:
1426:
1423:Canary Islands
1419:
1418:
1416:
1411:
1408:
1405:
1402:
1399:
1396:
1395:(pre-Hispanic)
1393:Canary Islands
1389:
1388:
1386:
1383:
1380:
1377:
1374:
1371:
1368:
1357:
1356:
1354:
1351:
1348:
1345:
1342:
1339:
1336:
1325:
1324:
1321:
1318:
1315:
1312:
1309:
1306:
1303:
1292:
1291:
1289:
1284:
1281:
1278:
1275:
1272:
1269:
1264:(Cosmopolitan
1258:
1257:
1255:
1250:
1247:
1244:
1241:
1238:
1235:
1221:
1220:
1218:
1213:
1210:
1207:
1204:
1201:
1198:
1184:
1183:
1181:
1176:
1173:
1170:
1167:
1164:
1161:
1147:
1146:
1144:
1139:
1136:
1133:
1130:
1127:
1124:
1110:
1109:
1107:
1102:
1099:
1096:
1093:
1090:
1087:
1076:
1075:
1073:
1068:
1065:
1062:
1059:
1056:
1053:
1039:
1038:
1036:
1031:
1028:
1025:
1022:
1019:
1016:
1010:
1009:
1004:
999:
996:
993:
990:
987:
984:
973:
972:
970:
967:
964:
961:
958:
955:
952:
941:
940:
938:
935:
932:
929:
926:
923:
920:
914:
913:
911:
906:
903:
900:
897:
894:
891:
885:
884:
882:
879:
876:
873:
870:
867:
864:
853:
852:
850:
847:
844:
841:
838:
835:
832:
821:
820:
817:
814:
811:
808:
805:
802:
799:
788:
787:
784:
781:
778:
775:
772:
769:
766:
755:
754:
751:
748:
745:
742:
739:
736:
733:
722:
721:
719:
714:
711:
708:
705:
702:
699:
688:
687:
685:
680:
677:
674:
671:
668:
665:
654:
653:
651:
648:
645:
642:
639:
636:
633:
622:
621:
619:
616:
613:
610:
607:
604:
601:
590:
589:
587:
582:
579:
576:
573:
570:
567:
556:
555:
553:
550:
547:
544:
541:
538:
535:
524:
523:
522:Elmrghni 2012
520:
517:
514:
511:
508:
505:
502:
491:
490:
488:
483:
480:
477:
474:
471:
468:
457:
456:
454:
449:
446:
443:
440:
437:
434:
431:Western Desert
419:
418:
416:
411:
408:
405:
402:
399:
396:
390:
389:
384:
379:
376:
373:
370:
367:
364:
358:
357:
355:
350:
347:
344:
341:
338:
335:
329:
328:
326:
321:
318:
315:
312:
309:
306:
291:
290:
287:
284:
281:
278:
275:
272:
269:
247:Canary Islands
234:
231:
229:
226:
224:
221:
217:Tofanelli 2009
200:
197:
149:CinnioÄlu 2004
102:
101:
98:
94:
93:
90:
86:
85:
80:
76:
75:
70:
66:
65:
58:
54:
53:
50:
42:
41:
26:
24:
14:
13:
10:
9:
6:
4:
3:
2:
9633:
9622:
9619:
9618:
9616:
9607:
9604:
9602:
9599:
9597:
9593:
9590:
9588:
9585:
9583:
9580:
9578:
9575:
9573:
9570:
9569:
9565:
9553:
9550:
9549:
9545:
9542:
9541:
9537:
9534:
9533:
9529:
9526:
9525:
9521:
9518:
9517:
9513:
9510:
9509:
9505:
9502:
9501:
9497:
9494:
9493:
9489:
9486:
9485:
9481:
9478:
9477:
9473:
9470:
9469:
9463:
9460:
9459:
9455:
9451:
9445:
9442:
9434:
9431:
9428:
9427:
9423:
9420:
9417:
9416:
9412:
9409:
9406:
9405:
9401:
9398:
9395:
9394:
9390:
9387:
9384:
9383:
9379:
9376:
9373:
9372:
9368:
9365:
9362:
9361:
9357:
9354:
9351:
9350:
9346:
9343:
9340:
9339:
9335:
9332:
9329:
9328:
9324:
9321:
9318:
9317:
9313:
9310:
9307:
9306:
9300:
9297:
9290:
9284:
9280:
9276:
9272:
9268:
9264:
9260:
9256:
9250:
9246:
9242:
9237:
9232:
9228:
9224:
9220:
9215:
9211:
9207:
9202:
9197:
9193:
9189:
9185:
9181:
9177:
9172:
9168:
9164:
9160:
9156:
9152:
9148:
9145:(4): 354â65.
9144:
9140:
9134:
9130:
9126:
9121:
9116:
9112:
9108:
9105:(4): 768â79.
9104:
9100:
9096:
9091:
9090:
9086:
9082:
9077:
9072:
9070:
9065:
9064:"Arab Tribes"
9060:
9059:
9058:
9051:
9046:
9041:
9039:
9034:
9029:
9027:
9022:
9017:
9015:
9010:
9005:
9004:
9003:
8998:
8993:
8992:
8991:
8990:
8983:
8978:
8974:
8970:
8969:
8968:
8962:
8956:
8952:
8947:
8942:
8938:
8934:
8930:
8926:
8922:
8917:
8916:
8911:
8907:
8902:
8897:
8893:
8889:
8885:
8881:
8877:
8872:
8871:
8866:
8862:
8857:
8852:
8848:
8844:
8841:(2): 339â48.
8840:
8836:
8832:
8827:
8826:
8821:
8817:
8812:
8807:
8803:
8799:
8795:
8791:
8787:
8782:
8781:
8776:
8772:
8767:
8762:
8757:
8752:
8748:
8744:
8740:
8736:
8732:
8727:
8726:
8721:
8717:
8713:
8709:
8705:
8701:
8697:
8693:
8687:
8686:
8681:
8677:
8672:
8667:
8663:
8659:
8655:
8651:
8647:
8642:
8641:
8636:
8632:
8627:
8622:
8618:
8614:
8610:
8606:
8602:
8597:
8596:
8591:
8587:
8583:
8579:
8575:
8571:
8567:
8563:
8557:
8556:
8551:
8547:
8543:
8539:
8535:
8531:
8526:
8521:
8517:
8513:
8507:
8506:
8501:
8497:
8493:
8489:
8485:
8481:
8477:
8473:
8468:
8467:
8462:
8458:
8454:
8450:
8446:
8442:
8436:
8435:
8430:
8426:
8422:
8418:
8412:
8411:
8406:
8402:
8397:
8392:
8388:
8384:
8380:
8375:
8374:
8369:
8365:
8360:
8355:
8351:
8347:
8343:
8339:
8335:
8330:
8329:
8324:
8320:
8315:
8310:
8306:
8302:
8298:
8294:
8290:
8285:
8281:
8277:
8273:
8269:
8265:
8261:
8257:
8253:
8247:
8243:
8239:
8234:
8229:
8225:
8221:
8217:
8213:
8209:
8204:
8203:
8199:
8193:
8189:
8184:
8179:
8175:
8171:
8167:
8163:
8159:
8154:
8150:
8146:
8142:
8138:
8135:(2): 336â46.
8134:
8130:
8125:
8121:
8117:
8112:
8107:
8102:
8097:
8093:
8089:
8085:
8079:
8075:
8071:
8067:
8063:
8059:
8055:
8051:
8047:
8041:
8037:
8033:
8029:
8025:
8021:
8017:
8013:
8008:
8006:
8000:
7996:
7991:
7986:
7982:
7978:
7974:
7970:
7966:
7961:
7957:
7953:
7948:
7943:
7938:
7933:
7929:
7925:
7922:(3): e56779.
7921:
7917:
7913:
7907:
7903:
7899:
7895:
7891:
7887:
7883:
7877:
7873:
7869:
7864:
7859:
7855:
7851:
7847:
7843:
7839:
7834:
7830:
7826:
7822:
7818:
7813:
7812:10400.13/3018
7808:
7804:
7800:
7796:
7792:
7786:
7785:
7780:
7776:
7772:
7768:
7764:
7760:
7756:
7752:
7746:
7742:
7738:
7733:
7728:
7723:
7718:
7714:
7710:
7706:
7701:
7697:
7693:
7688:
7683:
7679:
7675:
7671:
7666:
7662:
7658:
7654:
7650:
7646:
7642:
7637:
7633:
7629:
7625:
7621:
7617:
7613:
7607:
7603:
7599:
7594:
7589:
7585:
7581:
7577:
7572:
7568:
7564:
7559:
7554:
7550:
7546:
7542:
7538:
7534:
7529:
7525:
7521:
7516:
7511:
7507:
7503:
7499:
7495:
7491:
7486:
7482:
7478:
7474:
7470:
7466:
7462:
7458:
7454:
7448:
7444:
7440:
7435:
7430:
7426:
7422:
7418:
7414:
7410:
7405:
7401:
7397:
7393:
7389:
7385:
7381:
7375:
7371:
7367:
7362:
7357:
7353:
7349:
7346:(6): 848â52.
7345:
7341:
7337:
7332:
7328:
7324:
7320:
7316:
7312:
7308:
7304:
7300:
7295:
7291:
7287:
7282:
7277:
7274:(3): 374â86.
7273:
7269:
7265:
7260:
7256:
7252:
7247:
7242:
7238:
7234:
7230:
7226:
7222:
7217:
7213:
7209:
7204:
7199:
7195:
7191:
7188:(6): 820â30.
7187:
7183:
7179:
7174:
7170:
7166:
7161:
7156:
7152:
7148:
7144:
7140:
7136:
7131:
7127:
7123:
7118:
7113:
7109:
7105:
7102:(2): 338â45.
7101:
7097:
7093:
7088:
7084:
7080:
7076:
7072:
7067:
7062:
7058:
7054:
7050:
7045:
7041:
7037:
7032:
7027:
7022:
7017:
7013:
7009:
7005:
7000:
6996:
6992:
6988:
6984:
6980:
6976:
6972:
6968:
6963:
6962:
6958:
6953:
6948:
6939:
6935:
6931:
6927:
6922:
6917:
6912:
6907:
6903:
6899:
6895:
6887:
6884:
6879:
6873:
6870:
6865:
6859:
6856:
6851:
6847:
6842:
6837:
6833:
6829:
6825:
6821:
6817:
6813:
6809:
6801:
6798:
6793:
6789:
6785:
6781:
6776:
6771:
6766:
6761:
6757:
6753:
6749:
6741:
6738:
6733:
6732:"J-P58 YTree"
6727:
6724:
6719:
6715:
6711:
6707:
6702:
6697:
6692:
6687:
6683:
6679:
6675:
6667:
6664:
6659:
6653:
6650:
6645:
6641:
6637:
6633:
6628:
6623:
6618:
6613:
6609:
6605:
6601:
6593:
6590:
6585:
6584:"J-P58 YTree"
6579:
6576:
6571:
6567:
6563:
6559:
6552:
6545:
6542:
6537:
6533:
6528:
6523:
6518:
6513:
6509:
6505:
6501:
6494:
6491:
6488:
6482:
6479:
6474:
6470:
6465:
6460:
6456:
6452:
6448:
6444:
6440:
6436:
6432:
6428:
6424:
6417:
6414:
6409:
6405:
6401:
6397:
6389:
6386:
6381:
6377:
6372:
6367:
6363:
6359:
6354:
6349:
6345:
6341:
6338:(7): e41252.
6337:
6333:
6329:
6321:
6318:
6313:
6309:
6304:
6299:
6295:
6291:
6288:(3): 354â63.
6287:
6283:
6279:
6272:
6269:
6264:
6260:
6256:
6252:
6247:
6242:
6238:
6234:
6230:
6223:
6220:
6215:
6211:
6206:
6201:
6197:
6193:
6188:
6183:
6179:
6175:
6172:(2): e56775.
6171:
6167:
6163:
6156:
6153:
6137:
6133:
6129:
6125:
6121:
6114:
6107:
6104:
6099:
6095:
6091:
6087:
6083:
6082:10.1038/81685
6079:
6075:
6071:
6067:
6060:
6057:
6052:
6048:
6043:
6038:
6034:
6030:
6025:
6020:
6016:
6012:
6008:
6004:
6000:
5993:
5990:
5985:
5981:
5977:
5973:
5969:
5965:
5961:
5957:
5949:
5946:
5941:
5937:
5932:
5927:
5923:
5919:
5915:
5911:
5907:
5899:
5896:
5891:
5884:
5881:
5876:
5869:
5866:
5861:
5857:
5852:
5847:
5843:
5839:
5835:
5831:
5827:
5823:
5819:
5815:
5811:
5803:
5801:
5799:
5795:
5792:
5787:
5784:
5779:
5775:
5770:
5765:
5761:
5757:
5753:
5749:
5745:
5741:
5737:
5730:
5727:
5720:
5703:
5700:
5694:
5691:
5687:
5685:
5680:
5678:
5673:
5666:
5663:
5657:
5654:
5648:
5645:
5639:
5636:
5632:
5628:
5622:
5619:
5613:
5610:
5604:
5601:
5595:
5592:
5586:
5583:
5579:
5578:
5571:
5568:
5563:
5559:
5555:
5551:
5547:
5543:
5540:(2): 187â91.
5539:
5535:
5528:
5525:
5520:
5512:
5509:
5507:
5504:
5503:
5499:
5491:
5489:
5484:
5478:
5475:
5468:
5461:
5456:
5451:
5445:
5439:
5433:
5431:
5429:
5418:
5409:
5402:
5397:
5387:
5378:
5371:
5367:
5361:
5359:
5352:
5345:
5338:
5329:
5323:
5322:
5314:
5309:
5308:
5300:
5295:
5294:
5288:
5283:
5275:
5266:
5261:
5260:
5254:
5251:
5246:
5240:
5237:
5232:
5226:
5223:
5218:
5212:
5209:
5202:
5193:
5185:
5178:
5175:
5170:
5163:
5158:
5152:
5149:
5144:
5138:
5135:
5130:
5124:
5121:
5115:
5108:
5105:
5099:
5092:
5088:
5082:
5075:
5071:
5065:
5058:
5054:
5047:
5042:
5032:
5029:February 2021
5020:
5015:
5006:
5005:
5000:
4996:
4988:
4983:
4981:
4976:
4974:
4969:
4968:
4965:
4958:
4952:
4950:
4947:
4945:
4942:
4940:
4937:
4936:
4931:
4926:
4923:
4921:
4918:
4916:
4913:
4911:
4908:
4906:
4903:
4901:
4898:
4896:
4893:
4891:
4888:
4886:
4883:
4881:
4878:
4876:
4873:
4871:
4868:
4866:
4863:
4861:
4858:
4856:
4853:
4851:
4848:
4846:
4843:
4841:
4838:
4837:
4832:
4827:
4825:
4823:
4815:
4813:
4811:
4803:
4801:
4799:
4791:
4789:
4786:
4782:
4774:
4772:
4770:
4766:
4763:in the early
4762:
4758:
4754:
4746:
4744:
4742:
4734:
4732:
4730:
4722:
4717:
4703:
4700:
4699:
4698:
4695:
4694:
4692:
4689:
4687:
4684:
4681:
4677:
4673:
4670:
4669:
4667:
4664:
4661:
4658:
4655:
4652:
4649:
4646:
4643:
4640:
4639:
4638:
4635:
4632:
4629:
4627:
4624:
4621:
4618:
4617:
4616:
4613:
4610:
4607:
4604:
4601:
4598:
4595:
4594:
4593:
4592:
4588:
4581:
4579:
4577:
4573:
4569:
4568:Chiaroni 2009
4560:
4558:
4554:
4552:
4548:
4544:
4540:
4536:
4534:
4530:
4526:
4522:
4519:
4518:Chiaroni 2009
4514:
4511:
4507:
4503:
4499:
4494:
4490:
4486:
4482:
4481:Israeli Arabs
4478:
4474:
4470:
4466:
4461:
4459:
4458:Chiaroni 2009
4455:
4452:
4448:
4444:
4440:
4436:
4432:
4428:
4424:
4420:
4416:
4412:
4408:
4404:
4400:
4396:
4394:
4393:Chiaroni 2009
4390:
4386:
4382:
4381:Chiaroni 2009
4378:
4373:
4366:
4361:
4360:Chiaroni 2009
4357:
4354:
4353:Chiaroni 2009
4350:
4347:
4346:Chiaroni 2009
4343:
4342:
4341:
4337:
4335:
4331:
4327:
4323:
4319:
4315:
4310:
4308:
4307:Chiaroni 2008
4304:
4300:
4296:
4292:
4289:
4284:
4280:
4279:Chiaroni 2009
4276:
4274:
4270:
4269:Chiaroni 2009
4266:
4262:
4258:
4254:
4246:
4241:
4235:
4232:
4229:
4226:
4223:
4220:
4218:
4215:
4214:
4211:
4208:
4205:
4202:
4199:
4196:
4193:
4189:
4186:
4185:
4182:
4179:
4176:
4173:
4170:
4167:
4164:
4160:
4157:
4156:
4153:
4150:
4147:
4144:
4141:
4138:
4135:
4131:
4128:
4127:
4124:
4121:
4118:
4115:
4112:
4109:
4107:
4104:
4103:
4100:
4097:
4094:
4091:
4088:
4085:
4083:
4080:
4079:
4076:
4073:
4070:
4067:
4064:
4061:
4059:
4056:
4055:
4052:
4049:
4046:
4043:
4040:
4037:
4035:
4032:
4031:
4028:
4025:
4022:
4019:
4016:
4013:
4011:
4008:
4007:
4004:
4001:
3998:
3995:
3992:
3989:
3987:
3984:
3983:
3980:
3977:
3974:
3971:
3968:
3965:
3963:
3960:
3959:
3956:
3953:
3950:
3947:
3944:
3941:
3939:
3936:
3935:
3932:
3929:
3926:
3923:
3920:
3917:
3915:
3912:
3911:
3908:
3905:
3902:
3899:
3896:
3893:
3891:
3888:
3887:
3883:
3880:
3878:J-M267(xP58)
3877:
3875:Total J-M267
3874:
3871:
3868:
3867:
3864:
3858:
3852:
3849:
3846:
3843:
3840:
3837:
3834:
3830:
3827:
3826:
3823:
3820:
3817:
3814:
3811:
3808:
3805:
3801:
3798:
3797:
3794:
3791:
3788:
3785:
3782:
3779:
3776:
3773:
3772:
3769:
3766:
3763:
3760:
3757:
3754:
3751:
3748:
3747:
3744:
3741:
3738:
3735:
3732:
3729:
3726:
3723:
3722:
3719:
3716:
3713:
3710:
3707:
3704:
3701:
3697:
3694:
3693:
3690:
3687:
3684:
3681:
3678:
3675:
3672:
3668:
3665:
3664:
3661:
3658:
3655:
3652:
3649:
3646:
3643:
3639:
3636:
3635:
3632:
3629:
3626:
3623:
3620:
3617:
3614:
3610:
3607:
3606:
3603:
3600:
3597:
3594:
3591:
3588:
3586:
3583:
3582:
3579:
3576:
3573:
3570:
3567:
3564:
3561:
3558:
3554:
3551:
3550:
3547:
3544:
3541:
3538:
3535:
3532:
3530:
3527:
3526:
3523:
3520:
3517:
3514:
3511:
3508:
3506:
3503:
3502:
3499:
3496:
3493:
3490:
3487:
3484:
3482:
3479:
3478:
3475:
3472:
3469:
3466:
3463:
3460:
3458:
3455:
3454:
3451:
3448:
3445:
3442:
3439:
3436:
3433:
3429:
3426:
3425:
3422:
3419:
3416:
3413:
3410:
3407:
3404:
3401:
3400:
3397:
3394:
3391:
3388:
3385:
3382:
3379:
3375:
3372:
3371:
3368:
3367:El-Sibai 2009
3365:
3362:
3359:
3356:
3353:
3351:
3348:
3347:
3344:
3341:
3338:
3335:
3332:
3329:
3326:
3322:
3319:
3318:
3315:
3312:
3309:
3306:
3303:
3300:
3298:
3295:
3294:
3290:
3287:
3285:J-M267(xP58)
3284:
3282:Total J-M267
3281:
3278:
3275:
3274:
3271:
3264:
3258:
3255:
3253:
3252:Chiaroni 2009
3250:
3247:
3244:
3241:
3238:
3236:
3233:
3232:
3229:
3227:
3224:
3221:
3218:
3215:
3213:
3210:
3209:
3206:
3203:
3201:
3200:Chiaroni 2009
3198:
3195:
3192:
3189:
3186:
3184:
3181:
3180:
3177:
3174:
3172:
3171:Chiaroni 2009
3169:
3166:
3163:
3160:
3157:
3155:
3152:
3151:
3148:
3145:
3143:
3142:Chiaroni 2009
3140:
3137:
3134:
3131:
3128:
3126:
3123:
3122:
3119:
3117:
3114:
3111:
3108:
3105:
3102:
3100:
3097:
3096:
3092:
3089:
3086:
3084:J-M267(xP58)
3083:
3081:Total J-M267
3080:
3077:
3074:
3073:
3070:
3061:
3049:
3044:
3035:
3034:
3028:
3022:
3019:
3017:
3016:Chiaroni 2009
3014:
3011:
3008:
3005:
3002:
2999:
2995:
2992:
2991:
2988:
2986:
2983:
2980:
2977:
2974:
2971:
2968:
2965:
2961:
2958:
2957:
2954:
2952:
2949:
2946:
2943:
2940:
2937:
2934:
2930:
2927:
2926:
2923:
2921:
2920:Nogueiro 2009
2918:
2915:
2912:
2909:
2906:
2903:
2899:
2895:
2892:
2891:
2888:
2886:
2883:
2880:
2877:
2874:
2871:
2868:
2864:
2861:
2860:
2857:
2855:
2852:
2849:
2846:
2843:
2840:
2837:
2833:
2830:
2829:
2826:
2824:
2823:El-Sibai 2009
2821:
2818:
2815:
2812:
2809:
2807:
2804:
2803:
2800:
2798:
2797:Chiaroni 2009
2795:
2792:
2789:
2786:
2783:
2781:
2778:
2777:
2774:
2772:
2771:Chiaroni 2009
2769:
2766:
2763:
2760:
2757:
2755:
2752:
2751:
2748:
2746:
2745:Zalloua 2008b
2743:
2740:
2737:
2734:
2731:
2728:
2724:
2721:
2720:
2717:
2714:
2712:
2711:Chiaroni 2009
2709:
2706:
2703:
2700:
2697:
2695:
2692:
2689:
2688:
2685:
2683:
2682:Zalloua 2008a
2680:
2677:
2674:
2671:
2668:
2666:
2663:
2662:
2659:
2657:
2656:Chiaroni 2009
2654:
2651:
2648:
2645:
2642:
2639:
2636:
2632:
2629:
2628:
2625:
2623:
2622:Chiaroni 2009
2620:
2617:
2614:
2611:
2608:
2605:
2601:
2598:
2597:
2594:
2592:
2591:Chiaroni 2009
2589:
2586:
2583:
2580:
2577:
2574:
2570:
2567:
2566:
2563:
2561:
2560:Chiaroni 2009
2558:
2555:
2552:
2549:
2546:
2543:
2540:
2536:
2533:
2532:
2529:
2527:
2526:Chiaroni 2009
2524:
2521:
2518:
2515:
2512:
2509:
2506:
2502:
2499:
2498:
2495:
2493:
2492:Chiaroni 2009
2490:
2487:
2484:
2481:
2478:
2475:
2472:
2468:
2465:
2464:
2461:
2459:
2458:Chiaroni 2009
2456:
2453:
2450:
2447:
2444:
2441:
2438:
2434:
2431:
2430:
2427:
2425:
2424:Chiaroni 2009
2422:
2419:
2416:
2413:
2410:
2407:
2403:
2400:
2399:
2396:
2395:Zalloua 2008a
2393:
2391:
2390:El-Sibai 2009
2388:
2385:
2382:
2379:
2376:
2374:
2371:
2370:
2366:
2363:
2360:
2358:J-M267(xP58)
2357:
2355:Total J-M267
2354:
2351:
2348:
2347:
2344:
2338:
2332:
2330:
2329:Chiaroni 2009
2327:
2324:
2321:
2318:
2315:
2312:
2308:
2305:
2304:
2301:
2299:
2298:El-Sibai 2009
2296:
2293:
2290:
2287:
2284:
2282:
2279:
2278:
2275:
2273:
2272:Chiaroni 2009
2270:
2267:
2264:
2261:
2258:
2255:
2251:
2248:
2247:
2244:
2241:
2239:
2238:Chiaroni 2009
2236:
2233:
2230:
2227:
2224:
2221:
2217:
2214:
2213:
2210:
2208:
2207:Chiaroni 2009
2205:
2202:
2199:
2196:
2193:
2190:
2186:
2183:
2182:
2179:
2177:
2176:Chiaroni 2009
2174:
2171:
2168:
2165:
2162:
2159:
2155:
2152:
2151:
2148:
2145:
2142:
2139:
2136:
2133:
2130:
2126:
2123:
2122:
2119:
2118:Regueiro 2006
2116:
2114:
2113:Chiaroni 2009
2111:
2108:
2105:
2102:
2099:
2097:
2094:
2093:
2090:
2087:
2085:
2084:Chiaroni 2009
2082:
2079:
2076:
2073:
2070:
2068:
2065:
2064:
2060:
2057:
2054:
2052:J-M267(xP58)
2051:
2049:Total J-M267
2048:
2045:
2042:
2041:
2038:
2031:
2025:
2022:
2019:
2016:
2013:
2010:
2007:
2004:
2001:
2000:
1997:
1994:
1991:
1988:
1985:
1982:
1979:
1976:
1972:
1969:
1968:
1965:
1962:
1959:
1956:
1953:
1950:
1947:
1944:
1940:
1937:
1936:
1932:
1929:
1927:J-M267(xP58)
1926:
1924:total J-M267
1923:
1920:
1917:
1914:
1913:
1910:
1904:
1899:
1893:
1891:
1890:Soodyall 2011
1888:
1885:
1882:
1879:
1876:
1873:
1870:
1866:
1863:
1862:
1859:
1857:
1856:Soodyall 2011
1854:
1851:
1848:
1845:
1842:
1839:
1836:
1832:
1829:
1828:
1825:
1823:
1820:
1817:
1814:
1811:
1808:
1805:
1803:
1800:
1799:
1795:
1793:
1790:
1787:
1784:
1781:
1778:
1775:
1773:
1771:
1768:
1767:
1764:
1762:
1759:
1756:
1753:
1750:
1747:
1744:
1742:
1739:
1738:
1735:
1733:
1730:
1727:
1724:
1721:
1718:
1715:
1712:
1708:
1705:
1704:
1701:
1698:
1696:
1695:Chiaroni 2009
1693:
1690:
1687:
1684:
1681:
1678:
1675:
1671:
1668:
1667:
1664:
1661:
1659:
1658:Chiaroni 2009
1656:
1653:
1650:
1647:
1644:
1641:
1638:
1634:
1631:
1630:
1627:
1624:
1622:
1621:Chiaroni 2009
1619:
1616:
1613:
1610:
1607:
1604:
1601:
1600:Nilo-Saharans
1597:
1594:
1593:
1590:
1587:
1585:
1584:Chiaroni 2009
1582:
1579:
1576:
1573:
1570:
1567:
1564:
1560:
1557:
1556:
1553:
1549:
1546:
1544:
1543:Chiaroni 2009
1541:
1538:
1535:
1532:
1529:
1526:
1523:
1519:
1516:
1515:
1512:
1508:
1505:
1503:
1500:
1497:
1494:
1491:
1488:
1485:
1483:
1480:
1479:
1476:
1474:
1471:
1468:
1465:
1462:
1459:
1456:
1454:
1451:
1450:
1447:
1445:
1442:
1439:
1436:
1433:
1430:
1427:
1425:(17th-18th c)
1424:
1421:
1420:
1417:
1415:
1412:
1409:
1406:
1403:
1400:
1397:
1394:
1391:
1390:
1387:
1385:Regueiro 2015
1384:
1381:
1378:
1375:
1372:
1369:
1366:
1362:
1359:
1358:
1355:
1353:Regueiro 2015
1352:
1349:
1346:
1343:
1340:
1337:
1334:
1330:
1327:
1326:
1322:
1319:
1316:
1313:
1310:
1307:
1304:
1301:
1297:
1294:
1293:
1290:
1288:
1285:
1282:
1279:
1276:
1273:
1270:
1267:
1263:
1260:
1259:
1256:
1254:
1251:
1248:
1245:
1242:
1239:
1236:
1233:
1230:
1226:
1223:
1222:
1219:
1217:
1214:
1211:
1208:
1205:
1202:
1199:
1196:
1193:
1189:
1186:
1185:
1182:
1180:
1177:
1174:
1171:
1168:
1165:
1162:
1159:
1156:
1152:
1149:
1148:
1145:
1143:
1140:
1137:
1134:
1131:
1128:
1125:
1122:
1119:
1115:
1112:
1111:
1108:
1106:
1103:
1100:
1097:
1094:
1091:
1088:
1085:
1081:
1078:
1077:
1074:
1072:
1069:
1066:
1063:
1060:
1057:
1054:
1051:
1048:
1044:
1041:
1040:
1037:
1035:
1032:
1029:
1026:
1023:
1020:
1017:
1015:
1012:
1011:
1008:
1005:
1003:
1000:
997:
994:
991:
988:
985:
982:
978:
975:
974:
971:
968:
965:
962:
959:
956:
953:
950:
946:
943:
942:
939:
936:
933:
930:
927:
924:
921:
919:
916:
915:
912:
910:
907:
904:
901:
898:
895:
892:
890:
887:
886:
883:
881:Regueiro 2015
880:
877:
874:
871:
868:
865:
862:
858:
855:
854:
851:
848:
845:
842:
839:
836:
833:
830:
826:
823:
822:
818:
815:
812:
809:
806:
803:
800:
797:
793:
790:
789:
786:Laouina 2011
785:
782:
779:
776:
773:
770:
767:
764:
760:
757:
756:
752:
749:
746:
743:
740:
737:
734:
731:
727:
724:
723:
720:
718:
715:
712:
709:
706:
703:
700:
697:
693:
690:
689:
686:
684:
681:
678:
675:
672:
669:
666:
663:
659:
656:
655:
652:
649:
646:
643:
640:
637:
634:
631:
627:
624:
623:
620:
617:
614:
611:
608:
605:
602:
599:
595:
592:
591:
588:
586:
583:
580:
577:
574:
571:
568:
565:
561:
558:
557:
554:
551:
548:
545:
542:
539:
536:
533:
529:
526:
525:
521:
518:
515:
512:
509:
506:
503:
500:
496:
493:
492:
489:
487:
484:
481:
478:
475:
472:
469:
466:
462:
459:
458:
455:
453:
452:KujanovĂĄ 2009
450:
447:
444:
441:
438:
435:
432:
428:
424:
421:
420:
417:
415:
414:El-Sibai 2009
412:
409:
406:
403:
400:
397:
395:
392:
391:
388:
385:
383:
382:Chiaroni 2009
380:
377:
374:
371:
368:
365:
363:
360:
359:
356:
354:
351:
348:
345:
342:
339:
336:
334:
331:
330:
327:
325:
322:
319:
316:
313:
310:
307:
304:
300:
296:
293:
292:
288:
285:
282:
280:J-M267(xP58)
279:
277:total J-M267
276:
273:
270:
267:
266:
263:
261:
256:
252:
248:
244:
240:
232:
227:
222:
220:
218:
214:
209:
207:
198:
196:
194:
190:
186:
182:
181:Jewish groups
178:
175:and parts of
174:
170:
166:
162:
158:
154:
150:
146:
142:
137:
135:
134:haplogroup J2
131:
128:
124:
120:
116:
112:
111:Haplogroup J1
108:
99:
95:
91:
87:
84:
81:
77:
74:
71:
67:
63:
59:
55:
48:
43:
38:
33:
19:
18:Haplogroup J1
9519:Capelli 2001
9495:Karafet 2001
9453:
9449:
9444:
9385:Capelli 2001
9352:Karafet 2001
9299:
9258:
9254:
9226:
9222:
9186:(1): 121â6.
9183:
9179:
9142:
9138:
9102:
9098:
9080:
9068:
9056:
9049:
9037:
9025:
9013:
9001:
8986:
8981:
8966:
8928:
8924:
8883:
8879:
8838:
8834:
8793:
8789:
8741:(5): e2105.
8738:
8734:
8695:
8691:
8653:
8649:
8608:
8604:
8565:
8561:
8515:
8511:
8475:
8471:
8444:
8440:
8420:
8416:
8386:
8382:
8341:
8337:
8296:
8292:
8255:
8251:
8218:(1): 89â94.
8215:
8211:
8196:. (Also see
8165:
8161:
8132:
8128:
8091:
8087:
8049:
8045:
8019:
8015:
7972:
7968:
7919:
7915:
7885:
7881:
7845:
7841:
7794:
7790:
7754:
7750:
7712:
7708:
7677:
7673:
7644:
7640:
7615:
7611:
7583:
7579:
7540:
7536:
7500:(1): 91â99.
7497:
7493:
7456:
7452:
7416:
7412:
7383:
7379:
7343:
7339:
7302:
7298:
7271:
7267:
7228:
7224:
7185:
7181:
7142:
7138:
7099:
7095:
7059:(3): 407â9.
7056:
7052:
7011:
7008:BMC Genetics
7007:
6973:(6): 492â7.
6970:
6966:
6901:
6897:
6886:
6872:
6858:
6815:
6811:
6800:
6755:
6751:
6740:
6726:
6681:
6677:
6666:
6652:
6607:
6603:
6592:
6578:
6561:
6557:
6544:
6507:
6503:
6493:
6481:
6430:
6426:
6416:
6399:
6395:
6388:
6335:
6331:
6320:
6285:
6281:
6271:
6239:(1): 41â49.
6236:
6232:
6222:
6169:
6165:
6155:
6143:. Retrieved
6136:the original
6123:
6119:
6106:
6073:
6069:
6059:
6006:
6002:
5992:
5959:
5955:
5948:
5913:
5909:
5898:
5883:
5868:
5817:
5813:
5786:
5743:
5739:
5729:
5702:
5693:
5683:
5676:
5671:
5665:
5656:
5647:
5638:
5621:
5612:
5603:
5594:
5585:
5576:
5570:
5537:
5533:
5527:
5518:
5383:
5326:
5026:
5018:
4819:
4807:
4797:
4795:
4778:
4757:East Semitic
4750:
4738:
4726:
4701:
4696:
4690:
4685:
4671:
4665:
4659:
4654:J1c3b (M369)
4653:
4647:
4641:
4636:
4630:
4625:
4619:
4614:
4609:J1b (M365.1)
4608:
4602:
4596:
4590:
4589:
4585:
4564:
4555:
4537:
4523:
4515:
4462:
4451:East African
4443:Gulf of Oman
4397:
4374:
4370:
4338:
4318:Chalcolithic
4311:
4277:
4273:North Africa
4265:Repping 2006
4260:
4257:Karafet 2008
4252:
4250:
3884:Publication
3872:Sample size
3862:
3660:Capelli 2009
3291:publication
3279:Sample size
3268:
3205:Cadenas 2008
3176:Cadenas 2008
3147:Cadenas 2008
3099:Saudi Arabia
3090:Publication
3078:Sample size
3068:
3058:January 2018
3055:
3047:
2754:Palestinians
2723:Palestinians
2602:(North-east
2364:Publication
2352:Sample size
2342:
2058:Publication
2046:Sample size
2035:
1933:Publication
1921:J*(xJ-M172)
1918:Sample size
1908:
1831:South Africa
1822:Msaidie 2011
1792:Sanchez 2005
1320:Alvarez 2014
1179:Ennafaa 2011
1142:Ennafaa 2011
1105:Ennafaa 2011
1071:Ennafaa 2011
849:Alvarez 2014
816:Alvarez 2014
783:Alvarez 2014
750:Alvarez 2014
519:Alvarez 2014
286:publication
274:J*(xJ-M172)
271:Sample size
260:Ennafaa 2011
236:
223:Distribution
210:
202:
193:Central Asia
177:North Africa
157:Western Asia
138:
123:haplogroup J
117:(branch) of
110:
106:
105:
73:Western Asia
9601:J1b Project
9503:Semino 2000
9487:Hammer 2001
9363:Semino 2000
9341:Hammer 2001
8423:: 235â236.
6954:Works cited
6433:(1): 7465.
5820:(1): 6659.
4718:Ancient DNA
4626:J1c1 (M390)
4539:Hammer 2009
4529:Semino 2004
4510:Semino 2004
4489:Hammer 2009
4477:Palestinian
4389:Hammer 2009
4377:STR markers
4058:Circassians
3869:Population
3611:(northeast
3450:Semino 2004
3314:Semino 2004
3276:Population
3075:Population
2985:Hammer 2009
2951:Hammer 2009
2885:Flores 2005
2854:Flores 2005
2716:Shlush 2008
2406:Jabel Druze
2349:Population
2043:Population
1915:Population
1700:Semino 2004
1663:Semino 2004
1626:Hassan 2008
1589:Hassan 2008
1552:Hassan 2008
1511:Flores 2001
1502:Fregel 2009
1473:Fregel 2009
1444:Fregel 2009
1414:Fregel 2009
1323:Frigi 2011
1034:Onofri 2008
1007:Arredi 2004
1002:Grugni 2012
909:Semino 2004
819:Palet 2010
717:Semino 2004
683:Semino 2004
585:Semino 2004
486:Ottoni 2011
353:Semino 2004
324:Robino 2007
268:Population
253:and/or non-
243:Semino 2004
239:Arredi 2004
89:Descendants
8252:Hum. Genet
7715:(1): 181.
6076:(3): 360.
5721:References
4860:Haplogroup
4822:Tell Kurdu
4816:Tell Kurdu
4804:Satsurblia
4765:Bronze age
4741:Arslantepe
4676:Rabbinical
4631:J1c2 (P56)
4615:J1c (L136)
4533:Nebel 2002
4498:Nebel 2002
4493:Nebel 2002
4485:Nebel 2000
4439:Ethiopians
4385:Nebel 2000
4322:Bronze Age
4217:Azerbaijan
4134:Ingushetia
3405:(mainland)
1905:South Asia
1761:Moran 2004
1732:Moran 2004
1507:Bosch 2001
1082:(Bou Saad
763:Casablanca
9592:J project
8520:CiteSeerX
8512:Nat Genet
8094:(1): 69.
8036:160359661
8016:Antiquity
8003:See also
7400:163297133
7380:Antiquity
7014:(1): 59.
6949:Footnotes
6938:219105572
6832:0737-4038
6792:219105572
6718:219105572
6644:219105572
6455:2045-2322
6362:1932-6203
6196:1932-6203
6145:9 October
6126:: 59â89.
6033:1932-6203
5984:205303372
5842:2045-2322
5746:: 19157.
5684:PhyloTree
5519:Footnotes
4880:Paragroup
4865:Haplotype
4796:Olivieri
4769:Akkadians
4603:J1a (M62)
4473:haplotype
4427:Europeans
4320:or Early
4303:King 2002
4106:Ossetians
3718:King 2011
3432:Macedonia
3421:King 2008
3396:King 2008
3257:Luis 2004
3021:Cann 2002
2967:Ashkenazi
2604:Assyrians
2474:Aramaeans
2311:Assyrians
2254:Assyrians
2189:Assyrians
2129:Khuzestan
2032:West Asia
1047:Bou Omran
829:El Jadida
387:Luis 2004
206:Neolithic
9615:Category
9283:11073453
9245:15280900
9210:17047675
9167:10310338
9159:14740294
9129:13680527
8963:Websites
8955:18976729
8910:18374297
8865:11827954
8820:19367321
8775:18461126
8735:PLOS ONE
8712:15300852
8680:16400607
8635:15069642
8590:11556974
8582:17909833
8550:17083896
8542:16501575
8492:16770078
8461:21312181
8405:19918998
8368:11992266
8323:11573163
8272:11153918
8242:20700146
8192:14973781
8149:19425100
8120:21401952
8074:22406638
8066:18269686
7999:18385274
7956:23483890
7916:PLOS ONE
7902:18618658
7872:19669163
7821:15996172
7779:18482536
7771:15322918
7741:19650893
7696:16142507
7661:11543897
7632:21915847
7602:21833004
7567:19686289
7524:18685561
7481:10763736
7473:14586639
7443:19826455
7370:19156170
7327:41595131
7319:11954565
7290:17928816
7255:11254456
7212:19107149
7169:21571925
7126:15202071
7075:19213004
7040:19772609
6995:13960753
6987:15503146
6959:Journals
6930:32470401
6850:28177087
6784:32470401
6710:32470401
6636:32470401
6536:25431579
6473:29748665
6380:22815981
6332:PLOS ONE
6312:19809480
6263:25186140
6255:25471516
6214:23431392
6166:PLOS ONE
6132:19934469
6098:12893406
6090:11062480
6051:24312208
6003:PLOS ONE
5976:25123837
5940:19826455
5860:33758277
5778:26754573
5681:, and;
5672:op. cit.
5562:23291764
5554:24166809
5192:F-Y27277
4885:Subclade
4833:Genetics
4828:See also
4792:Sardinia
4680:Karaitic
4572:Anatolia
4435:Iranians
4403:Eurasian
4192:Dagestan
4188:Chechens
4163:Chechnya
4159:Chechens
4130:Chechens
3986:Lezghins
3859:Caucasus
3829:Portugal
3800:Portugal
3775:Portugal
3752:(Centre)
3750:Portugal
3725:Portugal
3700:Provence
3642:Italians
3613:Italians
3585:Slovenia
3481:Bulgaria
2902:Portugal
2867:Dead Sea
2638:Ismailis
2635:Damascus
2573:Alawites
2539:Damascus
2471:Ma'loula
2005:(Mappla)
1865:Zimbabwe
1741:Ethiopia
1707:Ethiopia
1670:Ethiopia
1633:Ethiopia
1522:Khartoum
1482:Sahrawis
1300:Sejenane
1232:Zaghouan
1229:Andalusi
499:Benghazi
173:Ethiopia
169:Caucasus
141:paternal
115:subclade
79:Ancestor
9511:Su 1999
9374:Su 1999
9311:J-M267
9263:Bibcode
9255:Science
9201:2588664
9120:1180600
8946:2668035
8901:2427286
8811:2986692
8766:2324201
8743:Bibcode
8720:1571356
8671:1380230
8626:1181965
8500:7017701
8314:1274378
8280:8136092
8233:3039498
8183:1182266
8111:3068964
7990:2336805
7947:3590186
7924:Bibcode
7863:2771134
7829:3229760
7732:2728732
7558:3312577
7515:2985948
7434:2987219
7361:2947089
7299:Science
7246:1275654
7203:2947100
7160:3355373
7117:1216069
7083:7041905
7031:2759955
6841:5400395
6527:4229899
6510:: 384.
6464:5945646
6435:Bibcode
6371:3399854
6340:Bibcode
6303:2859343
6205:3576335
6174:Bibcode
6042:3842387
6011:Bibcode
5931:2987219
5851:7987999
5822:Bibcode
5769:4709632
5748:Bibcode
5396:K-M2313
5389:
5330:
5315:
5301:
5285:
5276:
5267:
5089:
5072:
5019:updated
4781:Karelia
4775:Karelia
4729:Alalakh
4697:J1c3d2*
4597:J1* J1*
4547:Cohanim
4502:Maghreb
4411:Maghreb
4288:Zarzian
4010:Shapsug
3938:Kubachi
3914:Dargins
3833:Madeira
3777:(South)
3727:(North)
3553:Croatia
3505:Romania
3297:Albania
3048:updated
2994:Bedouin
2964:Cohanim
2933:Cohanim
2691:Galilee
2665:Lebanon
2505:Sednaya
1802:Comoros
1770:Somalis
1361:Tunisia
1329:Tunisia
1296:Tunisia
1262:Tunisia
1225:Tunisia
1195:Berbers
1188:Tunisia
1158:Berbers
1155:Jerbian
1151:Tunisia
1118:Jerbian
1114:Tunisia
1084:Berbers
1080:Tunisia
1050:Berbers
1043:Tunisia
1014:Tunisia
977:Tunisia
945:Tunisia
928:16.64%
918:Tunisia
889:Tunisia
857:Morocco
825:Morocco
792:Morocco
759:Morocco
726:Morocco
696:Berbers
692:Morocco
662:Berbers
658:Morocco
626:Morocco
594:Morocco
560:Morocco
528:Morocco
333:Algeria
295:Algeria
199:Origins
113:, is a
9464:J-P209
9314:J-M62
9281:
9243:
9208:
9198:
9165:
9157:
9127:
9117:
8953:
8943:
8908:
8898:
8863:
8856:155271
8853:
8818:
8808:
8773:
8763:
8718:
8710:
8678:
8668:
8633:
8623:
8588:
8580:
8548:
8540:
8522:
8498:
8490:
8459:
8403:
8366:
8359:379148
8356:
8321:
8311:
8278:
8270:
8240:
8230:
8198:Errata
8190:
8180:
8147:
8118:
8108:
8072:
8064:
8034:
7997:
7987:
7954:
7944:
7900:
7870:
7860:
7827:
7819:
7777:
7769:
7739:
7729:
7694:
7659:
7630:
7600:
7565:
7555:
7522:
7512:
7479:
7471:
7441:
7431:
7398:
7368:
7358:
7325:
7317:
7288:
7253:
7243:
7210:
7200:
7167:
7157:
7124:
7114:
7081:
7073:
7038:
7028:
6993:
6985:
6936:
6928:
6848:
6838:
6830:
6790:
6782:
6716:
6708:
6642:
6634:
6534:
6524:
6471:
6461:
6453:
6378:
6368:
6360:
6310:
6300:
6261:
6253:
6212:
6202:
6194:
6130:
6096:
6088:
6049:
6039:
6031:
5982:
5974:
5938:
5928:
5858:
5848:
5840:
5776:
5766:
5631:K-M526
5560:
5552:
5419:
5415:
5410:
5398:
5379:
5206:
5198:
5194:
5189:
4953:J-M267
4949:J-M172
4944:J-P209
4672:J1c3d*
4561:J-M368
4407:Arabic
4261:Page08
4082:Ingush
4034:Abkhaz
3962:Kaitak
3881:J-P58
3804:Açores
3696:France
3671:Sicily
3560:Croats
3557:Osijek
3529:Russia
3457:Greece
3428:Greece
3403:Greece
3374:Greece
3288:J-P58
3265:Europe
3212:Kuwait
3087:J-P58
2863:Jordan
2832:Jordan
2806:Jordan
2780:Jordan
2440:Sunnis
2361:J-P58
2307:Turkey
2137:33.4%
2067:Turkey
2055:J-P58
1930:J-P58
1852:13.2%
1849:26.3%
1846:39.5%
1674:Amhara
949:Sousse
796:Figuig
641:31.5%
609:60.7%
465:Tuareg
283:J-P58
251:Berber
228:Africa
83:J-P209
9506:Eu10
9369:Eu10
9163:S2CID
8973:ISOGG
8716:S2CID
8586:S2CID
8546:S2CID
8496:S2CID
8276:S2CID
8070:S2CID
8032:S2CID
7825:S2CID
7775:S2CID
7477:S2CID
7396:S2CID
7323:S2CID
7079:S2CID
6991:S2CID
6934:S2CID
6788:S2CID
6714:S2CID
6640:S2CID
6554:(PDF)
6259:S2CID
6139:(PDF)
6116:(PDF)
6094:S2CID
5980:S2CID
5558:S2CID
5208:GHIJK
4939:J-P58
4798:et al
4761:Syria
4642:J1c3*
4469:modal
4454:genes
4447:Asian
4431:Kurds
4423:Qatar
4415:Sudan
4334:Negev
4247:J-P58
4224:15.2%
4065:11.9%
3996:44.4%
3993:44.4%
3890:Avars
3667:Italy
3638:Italy
3609:Italy
3378:Crete
3350:Malta
3248:37.2%
3242:38.0%
3219:45.2%
3196:67.7%
3190:72.6%
3183:Yemen
3167:34.8%
3161:34.8%
3138:56.9%
3132:58.3%
3125:Qatar
3106:40.1%
3012:64.3%
3006:67.9%
2998:Negev
2981:14.0%
2975:14.9%
2972:1,360
2962:(non-
2947:46.0%
2941:46.0%
2910:12.3%
2844:40.6%
2836:Amman
2813:35.5%
2793:48.7%
2787:48.7%
2767:32.7%
2761:32.7%
2735:39.2%
2707:12.2%
2701:13.4%
2694:Druze
2652:58.8%
2646:58.8%
2631:Syria
2600:Syria
2587:26.7%
2581:26.7%
2569:Syria
2535:Syria
2522:14.3%
2516:14.3%
2501:Syria
2467:Syria
2454:44.4%
2448:47.2%
2433:Syria
2420:11.8%
2414:14.7%
2402:Syria
2380:33.6%
2373:Syria
2322:16.0%
2319:20.0%
2262:16.1%
2234:62.3%
2228:64.1%
2220:Arabs
2203:10.7%
2200:17.9%
2197:28.6%
2166:11.8%
2158:Kurds
2103:11.3%
2003:India
1975:Sunni
1971:India
1951:10.6%
1939:India
1883:9.3%
1880:9.3%
1869:Lemba
1835:Lemba
1788:2.5%
1782:2.5%
1779:0.5%
1691:20.8%
1685:29.2%
1637:Oromo
1596:Sudan
1580:17.1%
1574:17.1%
1559:Sudan
1539:74.3%
1533:74.3%
1518:Sudan
1492:20.2%
1434:11.9%
1404:16.7%
1382:15.3%
1376:15.3%
1311:34.0%
1283:24.2%
1266:Tunis
1249:43.8%
1243:43.8%
1212:31.4%
1206:31.4%
1192:Sened
1121:Arabs
1024:34.6%
998:31.1%
992:32.4%
981:Tunis
966:25.9%
960:25.9%
899:34.2%
878:16.7%
872:16.7%
807:29.2%
774:15.7%
741:21.3%
730:Rabat
707:10.7%
630:Arabs
598:Arabs
575:20.4%
564:Arabs
543:26.4%
532:Arabs
510:39.5%
495:Libya
461:Libya
442:31.4%
423:Egypt
404:19.8%
394:Egypt
378:19.7%
372:21.1%
362:Egypt
314:22.5%
301:from
299:Arabs
255:urban
185:Cohen
165:Sudan
127:clade
9594:and
9490:Med
9435:J1a
9424:J1a
9413:J1a
9347:Med
9279:PMID
9241:PMID
9206:PMID
9155:PMID
9125:PMID
8951:PMID
8906:PMID
8861:PMID
8816:PMID
8771:PMID
8708:PMID
8676:PMID
8631:PMID
8578:PMID
8538:PMID
8488:PMID
8457:PMID
8401:PMID
8364:PMID
8319:PMID
8268:PMID
8238:PMID
8188:PMID
8145:PMID
8116:PMID
8062:PMID
7995:PMID
7952:PMID
7898:PMID
7868:PMID
7817:PMID
7767:PMID
7737:PMID
7692:PMID
7657:PMID
7628:PMID
7598:PMID
7563:PMID
7520:PMID
7469:PMID
7439:PMID
7366:PMID
7315:PMID
7286:PMID
7251:PMID
7208:PMID
7165:PMID
7122:PMID
7071:PMID
7036:PMID
6983:PMID
6926:PMID
6898:Cell
6846:PMID
6828:ISSN
6780:PMID
6752:Cell
6706:PMID
6678:Cell
6632:PMID
6604:Cell
6532:PMID
6469:PMID
6451:ISSN
6376:PMID
6358:ISSN
6308:PMID
6251:PMID
6210:PMID
6192:ISSN
6147:2022
6128:PMID
6086:PMID
6047:PMID
6029:ISSN
5972:PMID
5936:PMID
5856:PMID
5838:ISSN
5774:PMID
5550:PMID
5377:K2b1
5222:HIJK
5114:A1b1
5070:A0-T
4753:Ebla
4678:and
4637:J1c3
4620:J1c*
4487:and
4449:and
4437:and
4421:and
4419:Iraq
4387:and
4332:and
4305:and
4263:in (
4253:J1c3
4119:0.0%
4116:1.3%
4113:1.3%
4092:2.8%
4089:2.8%
4068:4.9%
3812:2.5%
3758:4.9%
3679:3.8%
3650:0.7%
3592:1.3%
3536:0.4%
3512:1.5%
3488:3.4%
3464:1.6%
3440:1.8%
3411:4.7%
3386:8.3%
3357:7.8%
3333:6.3%
3304:3.6%
3245:0.8%
3235:Oman
3193:4.8%
3164:0.0%
3135:1.4%
3009:3.6%
2978:0.9%
2960:Jews
2944:0.0%
2929:Jews
2894:Jews
2875:8.9%
2790:0.0%
2764:0.0%
2727:Akka
2704:1.2%
2649:0.0%
2618:3.3%
2615:0.0%
2612:3.3%
2584:0.0%
2556:9.5%
2553:0.0%
2550:9.5%
2519:0.0%
2488:2.3%
2485:4.5%
2482:6.8%
2451:2.8%
2437:Hama
2417:2.9%
2325:4.0%
2288:3.2%
2281:Iran
2268:6.5%
2265:9.7%
2250:Iran
2231:1.8%
2216:Iraq
2185:Iraq
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