655:) are found throughout the range and occasionally both dolphin species are sighted in mixed groups. Where both species overlap in prey selection, Heaviside's take larger prey items, potentially because they are outcompeted by the larger dusky dolphins for their preferred, smaller sized prey. In central Namibia (especially Walvis Bay) Heaviside's dolphins overlap with a small populations of fewer than 100 common bottlenose dolphins REF. The bottlenose dolphin population use only water less than 15m depth while in this area Heaviside's dolphins are almost always encounteres in water deeper than 20 m (66 ft), suggesting some form of competitive exclusion.
664:
outside of the detectable frequencies of killer whales. Although NBHF clicks are limited in acoustic range, they have a better resolution for small targets and are thought to provide a foraging advantage in the often cluttered, nearshore environment in which these species occur. Heaviside's also produce a second click type, of lower frequency and broader bandwidth, that is within the hearing range of killer whales. These calls are produced most frequently in groups engaging in social behaviour. It is likely that the dolphins use these calls when socialising away from predator threat and switch to high frequency clicks when foraging and travelling.
42:
60:
126:
565:
coastal waves. Iconic vertical leaps clear the water before re-entering headfirst with almost no splash. Heaviside's dolphins use echolocation to find and capture prey. Mating typically occurs in social groups of 3–7 individuals which remain in a small area exhibiting extensive rolling, touching and position changes with frequent leaps by one of pairs of animals which potentially serve a competitive function.
84:
507:
170 km into Angola but well south of the defined northern boundary of the
Benguela Ecosystem. The northern boundary of the Benguela current shifts north and south seasonally and as Heaviside's dolphins appear closely linked to its cool waters, their northern range limit may shift along with water conditions. Systematic surveys have dedicated effort to describing the distribution in southern
410:
498:, the real southern limit beyond which sightings are extremely rare is Hout Bay, some 40 km (25 mi) to the north (a considerable distance for a species which shows very high site fidelity to quite small spatial scales when nearshore). The species occurs more or less continuously for 2,500 km (1,600 mi) to the north of this along the South African coast, through
759:
739:
behaviour in Walvis Bay includes a reduction in resting behaviour and an increase in socialising behaviour in the presence of tour boats (indurkyhua). One
Heaviside's dolphin was documented in 2010 with evidence of a propeller strike along its flank. As individuals have small home ranges, they may be vulnerable to localised threats.
552:
most bays throughout South Africa and
Namibia, but they are rarely seen in the protected shallows of these bays. Outside of bays, they show relatively high densities along exposed sandy beaches, but these may be a secondary choice after a preference for areas where there is a high abundance of their main prey item; juvenile hake (
257:
751:(IUCN) listed the Heaviside's as 'Data Deficient' however, as of 2017 the status was changed to 'Near Threatened', owing to improved knowledge on the species from multiple studies. Despite this, the overall population trend remains unknown, and there are many aspects of the species biology that remain to be studied.
451:
Information on reproduction is limited for
Heaviside's dolphins, however they are thought to be comparable to Hector's and Commerson's dolphins. Females and males reach sexual maturity approximately between 5–9 years. Mating is thought to occur year-round, however individual females may only produce
692:
are 508 and 494 respectively. A visual and acoustic line-transect ship survey estimated an average of 1594 dolphins in the
Namibian Islands' Marine Protected Area (NIMPA), which spans 400 km (250 mi) of coastline along southern Namibia (REF Martin et al. 2020). Quantification of abundance
551:
Heaviside's dolphins typically remain nearshore in the mornings where they typically socialise and rest. When nearshore their distribution patterns are remarkably predictable within and between years, with the animals showing highly consistent use of aggregation sites at the exposed western tips of
564:
Heaviside's dolphins are energetic and social animals, especially when nearshore in the mornings. Behaviour when offshore tends to be less playful and may include a rest phase when moving offshore to feed. They are attracted to boats and frequently bow-ride. Individuals can also be seen surfing in
620:
move offshore in the afternoon to feed on prey rising vertically to the surface at night. Movement inshore to rest and socialise occurs in the morning. However, the pattern is different in
Luderitz and Walvis Bay, Namibia where the movement is less pronounced and dolphins appear to stay inshore
417:
Heaviside's are small and stocky with adults reaching a maximum length and weight of 1.7 m (5.6 ft) and 75 kg (165 lb) respectively. The dolphin has a distinct black, grey and white body pattern, and is not easily confused with any other species in its range. The head is cone
663:
As is the case with all species in the genus, Heaviside's dolphins produce narrow-band high-frequency (NBHF) echolocation clicks (centred around 125–130 kHz), and do not whistle. This adaptation is theorised to allow acoustic crypsis from eavesdropping predators, as the sounds produced are
738:
Heaviside's dolphins may be exposed to increase in the marine eco-tourism business in
Namibia, which has grown without regulation in Walvis Bay (Leeney, 2014). Negative effects have been demonstrated for other coastal cetacean species, for example the impact of tour boats on bottlenose dolphin
730:
warm-water currents at both its northern and southern limits. Consequently, small increases in water temperature could result in all shelf waters in this area becoming too warm for both species, and populations are expected to decline rather than shift poleward due to a lack of shelf habitat.
729:
Heaviside's dolphins are listed amongst the cetacean species most vulnerable to climate change as they are limited to a distribution range that includes both suitable shelf habitat and cool water temperatures (Best, 2007). The
Benguela Current is the only eastern-boundary current bordered by
506:
where the northern boundary for the species remains poorly defined. Several dolphins have been sighted or accidentally caught by fishing vessels north of the Angola–Namibia border, but no sightings were reported during a series of coastal scientific surveys at Tombua which is approximately
638:
There has been limited research into
Heaviside's diving behaviour, however a study of two dolphins fitted with satellite tags was undertaken in South Africa in 1997. The maximum dive depth recorded was 147 meters; however, the majority of dives were less than 50 meters. Dive duration were
534:
Recent genetic research has demonstrated evidence of population structure across the range, indicating two metapopulations (north and south) with limited genetic exchange. This pattern of fragmentation is a common feature amongst the other three species in the genus
710:
Heaviside's dolphins are exposed to several poorly quantified and rapidly changing human threats including fisheries bycatch related mortality and illegal directed catch (Elwen and Gopal, 2018; Alfaro-Shigueto et al., 2019). Recently developed mid water trawls for
1845:
Kyhn, L. A.; Jensen, F. H.; Beedholm, K.; Tougaard, J.; Hansen, M.; Madsen, P. T. (2010). "Echolocation in sympatric Peale's dolphins (Lagenorhynchus australis) and
Commerson's dolphins (Cephalorhynchus commersonii) producing narrow-band high-frequency clicks".
701:
Heaviside's dolphins are exposed to a variety of threats given their limited range in coastal shallow waters which are subject to a range of anthropogenic activities. Directed catch has occurred historically, with meat being used for human consumption.
629:
Heaviside's have small home ranges of 50–80 km (31–50 mi) as measured using satellite telemetry over 2–3 months and photographic resightings over up to 3 years. Some individuals have been resighted at the same location for up to 10 years.
464:
Typically occurs in small groups of 2–3, but numbers of 1–10 are frequent and large aggregations of ~100 individuals or more are known to form in high density areas. Nursery groups (exclusively females and calves) are not formed in this species.
1348:
Hamner, Rebecca M.; Pichler, Franz B.; Heimeier, Dorothea; Constantine, Rochelle; Baker, C. Scott (2012). "Genetic differentiation and limited gene flow among fragmented populations of New Zealand endemic Hector's and Maui's dolphins".
334:, who was known for his own biological collections at the time. "Heaviside's dolphin" is the recognised common name, though amongst others, "Haviside's dolphin" and "Benguela dolphin" are also used, the latter especially in Namibia.
1501:
Elwen, Simon H.; Best, Peter B.; Reeb, Desray; Thornton, Meredith (2009). "Diurnal Movements and Behaviour of Heaviside's Dolphins,Cephalorhynchus heavisidii, with some Comparative Data for Dusky Dolphins,Lagenothynchus obscutus".
1073:
Elwen, S.H.; Thornton, M.; Reeb, D.; Best, P.B. (2010). "Near-shore distribution of Heaviside's (Cephalorhynchus heavisidii) and dusky dolphins (Lagenorhynchus obscurus) at the southern limit of their range in South Africa".
4079:
755:
1646:
Davis, RW; David, JHM; Meÿer, MA; Sekiguchi, K; Best, PB; Dassis, M; Rodríguez, DH (2014). "Home range and diving behaviour of Heaviside's dolphins monitored by satellite off the west coast of South Africa".
442:
is minimal, however variation in the shape of the white patch covering the genital slit is distinct between genders. In males, the patch ends in a point, but in females widens out to cover the mammary slits.
1948:
Martin, Morgan J.; Elwen, Simon H.; Kassanjee, Reshma; Gridley, Tess (2019). "To buzz or burst-pulse? The functional role of Heaviside's dolphin, Cephalorhynchus heavisidii, rapidly pulsed signals".
1727:
Morisaka, Tadamichi; Karczmarski, Leszek; Akamatsu, Tomonari; Sakai, Mai; Dawson, Steve; Thornton, Meredith (2011). "Echolocation signals of Heaviside's dolphins (Cephalorhynchus heavisidii)".
923:"A conservation assessment of Cephalorhynchus heavisidii. In Child MF, Roxburgh L, Do Linh San E, Raimondo D, Davies-Mostert HT. The Red List of Mammals of South Africa, Swaziland and Lesotho"
1700:
Heinrich, Sonja; Elwen, Simon; Bräger, Stefan (2010). "Patterns of sympatry in Lagenorhynchus and Cephalorhynchus: dolphins in different habitats". In Würsig, Bernd; Wursig, Melany (eds.).
1295:
Gopal, K; Karczmarski, L; Tolley, K.A (2019). "Patterns of geographic variation between mitochondrial and nuclear markers in Heaviside's (Benguela) dolphins (Cephalorhynchus heavisidii)".
1397:
Elwen, Simon H.; Reeb, Desray; Thornton, Meredith; Best, Peter B. (2009). "A population estimate of Heaviside's dolphins,Cephalorhynchus heavisidii, at the southern end of their range".
2054:"Appendices I and II of the Convention on the Conservation of Migratory Species of Wild Animals (CMS)" (PDF). 5 March 2009. p. 6. Archived from the original (PDF) on 21 February 2012.
1545:
Leeney, R. H; Carslake, D; Elwen, S. H (2011). "Using static acoustic monitoring to describe echolocation behaviour of Heaviside's dolphins (Cephalorhynchus heavisidii) in Namibia".
1154:
Morias, Miguel (2012). "Marine mammal sightings off the Angolan coast recorded from the R/V Dr. Fridtjof Nansenin August 2004 and July 2005". In Van Waerebeek, Koen (ed.).
556:) in adjacent offshore waters. Most commonly sighted within sea surface temperatures of 9 to 15 °C (48 to 59 °F) and depths less than 100 metres (330 ft).
1010:
Pichler, F; D. Robineau, R; Goodall, M; Meyer, M; Olivarria, C; Baker, C (2001). "Origin and radiation of Southern Hemisphere coastal dolphins (genus Cephalorhynchus)".
4131:
4356:
748:
4209:
337:
The genus name "Cephalorhynchus" comes from the Greek kephale for 'head' and rhynchos for 'beak'. For the species name "heavisidii" see the above description.
1127:
Best, Peter B.; Abernethy, R. Blake (1994). "Heaviside's dolphin, Cephalorhynchus heavisidii (Gray, 1828)". In Ridgway, Sam H.; Harrison, Richard (eds.).
531:, South Africa. Sightings are common from land and there are several dolphin watching tour companies by which Heaviside's dolphins can be seen by boat.
4386:
4366:
4328:
4105:
4396:
4144:
4222:
1711:
1191:
961:
896:
922:
600:) are also consumed especially in Namibia. Foraging occurs mostly at the seabed, in shallow depths. Feeding nearshore is rarely observed.
494:
The species is strongly associated with the cool waters of the Benguela Ecosystem. Although the southern limit of the range is defined as
4391:
1138:
762:. The Memorandum of Understanding was established in 2008 and aims to protect these species at a national, regional and global level.
1444:"Using static acoustic monitoring to describe echolocation behaviour of heaviside's dolphins (Cephalorhynchus heavisidii) in Namibia"
2099:
991:
434:. The underbelly is white, with bands that extend onto the lower rear of the body. Small white patches are located just behind the
2016:"Global climate change, range changes and potential implications for the conservation of marine cetaceans: a review and synthesis"
1790:"Predation by killer whales (Orcinus orca) and the evolution of whistle loss and narrow-band high frequency clicks in odontocetes"
4381:
4287:
4170:
3139:
1443:
3121:
1587:"Range and Movements of Female Heaviside's Dolphins (Cephalorhynchus heavisidii), as Determined by Satellite-Linked Telemetry"
4188:
4149:
3845:
760:
Memorandum of Understanding Concerning the Conservation of the Manatee and Small Cetaceans of Western Africa and Macaronesia
4018:
125:
4196:
3262:
2987:
2978:
1174:
Weir, C. R (2019). "The Cetaceans (Whales and Dolphins) of Angola". In Huntley, B; Russo, V; Lages, F; Ferrand, N (eds.).
385:
3791:
2808:
2799:
2731:
41:
4376:
4183:
4053:
3818:
3029:
2996:
2758:
4045:
3890:
418:
shaped with a blunt beak. The dorsal fin is triangular in shape and centred in the middle of the back. The head and
4371:
4214:
3923:
3914:
2841:
2249:
323:
3809:
3764:
3732:
3723:
3708:
3699:
3549:
3148:
3005:
2258:
4361:
3271:
3056:
2641:
384:, from which Heaviside's dolphin are the basal species. Radiation around the southern hemisphere following the
3947:
3938:
3872:
3836:
3800:
3773:
3649:
3322:
1585:
Elwen, Simon; Meÿer, Michael A.; Best, Peter B.; Kotze, P. G H.; Thornton, Meredith; Swanson, Stephan (2006).
1053:
Best, P. B (1988). "The external appearance of Heaviside's dolphin, Cephalorhynchus heavisidii (Gray, 1828)".
980:
Dawson, S (26 February 2009). "Cephalorhynchus dolphins". In Würsig, B; Thewissen, J.G.M; Kovacs, K.M (eds.).
256:
3658:
3466:
2650:
684:, South Africa represents the southernmost populations in the species range. Local population estimates for
430:
and dorsal cape are a dark grey to almost black with a band that extends forward from the dorsal fin to the
3980:
3863:
3827:
3782:
3667:
3475:
2534:
2072:
1891:"Heaviside's dolphins (Cephalorhynchus heavisidii) relax acoustic crypsis to increase communication range"
354:
4058:
4248:
3089:
3080:
2312:
2092:
230:
1462:"Comparative analysis of the diets of smaller odontocete cetaceans along the coast of southern Africa"
4292:
4092:
4027:
3558:
3484:
2722:
2240:
331:
456:
time is unknown. Maximum known lifespan is based on the oldest recorded individual at 26 years old.
4032:
3899:
3854:
3567:
3295:
2782:
2698:
2689:
2561:
1155:
673:
540:
370:
362:
73:
811:
4266:
3881:
3104:
2913:
2904:
2889:
2880:
2609:
2600:
2303:
1983:
1827:
1770:
1682:
1527:
1374:
1330:
1197:
1109:
1035:
717:
431:
120:
3346:
3337:
2740:
2674:
2665:
519:, Namibia. These locations are also popular hotspots for watching these dolphins in addition to
4253:
4136:
639:
predominantly less than 2 minutes with most dives between 0 and 1 minutes (Davis et al. 2014).
330:, who described the species in his Spicilegia Zoologica, misidentified Haviside as the surgeon
4235:
4066:
3747:
3429:
3420:
2832:
2749:
2460:
2451:
2037:
1975:
1930:
1912:
1871:
1863:
1819:
1811:
1762:
1754:
1707:
1674:
1608:
1519:
1483:
1424:
1366:
1322:
1241:
1187:
1134:
1101:
1027:
987:
957:
892:
439:
318:
Early in the 19th century, a specimen was caught off the Cape of Good Hope and brought to the
2000:
Best, P; Ros, G.J.B (1977). Exploitation of small cetaceans of the coast of Southern Africa.
4240:
4084:
3682:
3604:
3595:
3585:
3304:
3245:
3130:
2961:
2357:
2339:
2330:
2085:
2027:
1965:
1957:
1920:
1902:
1855:
1801:
1744:
1736:
1664:
1656:
1598:
1562:
1554:
1511:
1473:
1414:
1406:
1358:
1312:
1304:
1277:
1231:
1179:
1091:
1083:
1019:
862:
806:
327:
246:
59:
1220:"The distribution of small odontocete cetaceans off the coasts of South Africa and Namibia"
4162:
3065:
3047:
3038:
2928:
2543:
2525:
2507:
1258:
Payne, A. I. L; Brink, K. H; Mann, K. H; Hillborn, R (eds.) Benguela Trophic Functioning.
381:
346:
285:
207:
197:
88:
2375:
930:
South African National Biodiversity Institute and Endangered Wildlife Trust, South Africa
676:
estimate currently exists, however a population estimate of 6,345 for the region between
17:
2946:
2713:
2585:
2576:
2423:
2414:
2146:
1925:
1890:
1163:(CMS Technical Series No. 26 ed.). Bonn, Germany: CMS Secretariat. pp. 26–30.
712:
319:
4350:
4157:
3539:
3364:
2767:
2492:
2441:
2208:
2199:
1806:
1789:
1410:
1201:
1113:
1023:
797:
648:
609:
528:
435:
393:
93:
2067:
1987:
1831:
1686:
1531:
1378:
1334:
1039:
788:
4319:
4071:
3622:
3185:
3176:
3166:
2294:
2174:
2158:
1774:
1087:
681:
613:
508:
474:
427:
1961:
1701:
1660:
1183:
1128:
981:
951:
886:
4279:
4261:
4201:
4175:
4118:
4012:
3383:
2624:
2223:
2152:
689:
516:
409:
389:
177:
50:
1274:
Spatial and temporal patterns of habitat use of Heaviside's dolphins in Namibia
2399:
2390:
2348:
2189:
1603:
1586:
1478:
1461:
1362:
1236:
1219:
685:
621:
during the night, which is likely associated with foraging on different prey.
581:
512:
495:
423:
4003:
2041:
1979:
1916:
1889:
Martin, Morgan J.; Gridley, Tess; Elwen, Simon H.; Jensen, Frants H. (2018).
1867:
1815:
1758:
1678:
1612:
1558:
1523:
1487:
1428:
1370:
1245:
1105:
481:) are known predators and there is evidence of shark attack from body scars.
3640:
3502:
2366:
2321:
1308:
677:
524:
520:
453:
137:
1934:
1907:
1875:
1823:
1766:
1326:
1031:
833:
4313:
3997:
3401:
3374:
3286:
3227:
3020:
2823:
2140:
2128:
2077:
1515:
574:
298:
290:
157:
1749:
1628:
Heavy metal analysis in Heaviside's dolphins (Cephalorhyncus heavisidii)
864:
Spicilegia Zoologica: Original Figures and Short Systematic Descriptions
396:, is thought to have led to the subsequent speciation within the genus.
4333:
4110:
3447:
3313:
3209:
3200:
2856:
2276:
2109:
1859:
617:
589:
499:
294:
280:
187:
1970:
1740:
1669:
1567:
1419:
1317:
1281:
1218:
Findlay, K. P.; Best, P. B.; Ross, G. J. B.; Cockcroft, V. G. (1992).
1096:
4123:
4040:
2937:
2134:
2122:
2032:
2015:
503:
419:
302:
167:
147:
3974:
756:
Convention on the Conservation of Migratory Species of Wild Animals
4274:
3521:
3512:
3411:
438:
and a single white patch extends between these fins on the chest.
408:
110:
104:
4097:
4227:
3457:
2865:
577:
422:
are coloured light grey with darker patches around the eye. The
3978:
2489:
2171:
2081:
1157:
Conserving cetaceans and manatees in the western African region
782:
780:
543:, which displays genetic isolation over very short distances.
345:
The three other species in the genus Cephalorhynchus are the
511:
and current research efforts focus on local populations in
1133:. Vol. 5 The First Book of Dolphins. Academic Press.
369:). All are located in cool temperate shelf waters in the
1276:(M.Sc). Pretoria, South Africa: University of Pretoria.
1895:
Proceedings of the Royal Society B: Biological Sciences
1630:(Thesis). Pokfulam, Hong Kong: University of Hong Kong.
1703:
The Dusky Dolphin: Master Acrobat Off Different Shores
867:. Vol. 1. Soho, London: Treüttel, Würtz & Co.
1460:
Sekiguchi, K.; Klages, N. T. W.; Best, P. B. (1992).
888:
Whales and Dolphins of the Southern African Subregion
380:
dolphins originated from a single common ancestor in
1068:
1066:
1064:
754:
Heaviside's dolphin is listed on Appendix II of the
4303:
3987:
3936:
3912:
3745:
3721:
3680:
3638:
3621:
3593:
3584:
3547:
3538:
3510:
3501:
3455:
3446:
3409:
3400:
3372:
3363:
3335:
3284:
3243:
3226:
3198:
3174:
3165:
3102:
3078:
3018:
2959:
2926:
2902:
2878:
2854:
2821:
2780:
2711:
2687:
2663:
2622:
2598:
2574:
2523:
2506:
2449:
2440:
2412:
2388:
2292:
2275:
2221:
2197:
2188:
1005:
1003:
975:
973:
388:(otherwise known as the West Wind Drift), first to
1213:
1211:
916:
914:
912:
910:
908:
1641:
1639:
1637:
1580:
1578:
1729:The Journal of the Acoustical Society of America
1392:
1390:
1388:
945:
943:
941:
939:
2002:Report to the International Whaling Commission
1055:Report of the International Whaling Commission
749:International Union for Conservation of Nature
2093:
812:10.2305/IUCN.UK.2018-2.RLTS.T4161A50352086.en
8:
413:Heaviside's dolphins off Walvis Bay, Namibia
956:. Bloomsbury Publishing. pp. 359–361.
891:. Cambridge University Press. p. 388.
647:Whilst typically found further from shore,
3975:
3635:
3590:
3544:
3507:
3452:
3406:
3369:
3240:
3171:
2520:
2503:
2486:
2446:
2289:
2194:
2185:
2168:
2100:
2086:
2078:
1504:South African Journal of Wildlife Research
953:Handbook of Whales, Dolphins and Porpoises
880:
878:
876:
874:
473:Levels of predation are unknown; however,
341:Closely related species and genetic origin
255:
82:
58:
40:
31:
2031:
1969:
1924:
1906:
1805:
1748:
1668:
1602:
1566:
1477:
1418:
1316:
1235:
1095:
810:
986:(3rd ed.). London: Academic Press.
693:throughout the range is still required.
1466:South African Journal of Marine Science
1260:South African Journal of Marine Science
1224:South African Journal of Marine Science
776:
301:ecosystem along the southwest coast of
856:
854:
584:species, predominantly juvenile hake (
4357:IUCN Red List near threatened species
721:) are considered an emerging threat.
7:
1788:Morisaka, T.; Connor, R. C. (2007).
1706:. Academic Press. pp. 313–332.
1178:. Cham: Springer. pp. 445–470.
921:Gopal, K; Elwen, S; Plön, S (2016).
798:IUCN Red List of Threatened Species
67:Size compared to an average human
25:
3122:Indo-Pacific bottlenose dolphin (
1649:African Journal of Marine Science
376:Genetic studies suggest that the
1807:10.1111/j.1420-9101.2007.01336.x
1411:10.1111/j.1748-7692.2008.00246.x
1024:10.1046/j.0962-1083.2001.01360.x
124:
4387:Marine fauna of Southern Africa
4367:Cetaceans of the Atlantic Ocean
3263:Indo-Pacific finless porpoise (
3140:Tamanend's bottlenose dolphin (
2988:Indian Ocean humpback dolphin (
2979:Indo-Pacific humpback dolphin (
1848:Journal of Experimental Biology
1794:Journal of Evolutionary Biology
612:movement pattern is present in
592:species such as juvenile goby (
4397:Taxa named by John Edward Gray
2809:Southern right whale dolphin (
2800:Northern right whale dolphin (
2732:Atlantic white-sided dolphin (
1088:10.1080/15627020.2010.11657256
983:Encyclopedia of Marine Mammals
643:Sympatry with other delphinids
1:
3846:Deraniyagala's beaked whale (
3819:Ginkgo-toothed beaked whale (
3030:Pantropical spotted dolphin (
2997:Australian humpback dolphin (
2759:Pacific white-sided dolphin (
1962:10.1016/j.anbehav.2019.01.007
787:Elwen, S.; Gopal, K. (2018).
573:Prey items consist of mostly
386:Antarctic Circumpolar Current
322:by a Captain Haviside of the
2842:Australian snubfin dolphin (
2250:North Atlantic right whale (
1661:10.2989/1814232X.2014.973903
1184:10.1007/978-3-030-03083-4_16
625:Home range and site fidelity
598:Trachurus trachurus capensis
3792:Blainville's beaked whale (
3733:Tropical bottlenose whale (
3709:Southern bottlenose whale (
3700:Northern bottlenose whale (
3149:Common bottlenose dolphin (
3006:Atlantic humpback dolphin (
2259:North Pacific right whale (
2020:Endangered Species Research
1057:(Special issue 9): 279–299.
4413:
4085:cephalorhynchus-heavisidii
4046:cephalorhynchus-heavisidii
4033:Cephalorhynchus_heavisidii
4019:Cephalorhynchus heavisidii
3989:Cephalorhynchus heavisidii
3891:Stejneger's beaked whale (
3272:Yangtze finless porpoise (
3057:Atlantic spotted dolphin (
2642:Short-finned pilot whale (
1130:Handbook of Marine Mammals
861:Gray, John Edward (1828).
791:Cephalorhynchus heavisidii
539:and most prevalent in the
324:British East India Company
276:Cephalorhynchus heavisidii
264:Heaviside's dolphin range
240:Cephalorhynchus heavisidii
4392:Mammals described in 1828
3924:Shepherd's beaked whale (
3467:Araguaian river dolphin (
2651:Long-finned pilot whale (
2502:
2485:
2184:
2167:
2117:
1604:10.1644/05-MAMM-A-307R2.1
1479:10.2989/02577619209504746
1363:10.1007/s10592-012-0347-9
1237:10.2989/02577619209504706
950:Carwardine, Mark (2020).
263:
254:
236:
229:
121:Scientific classification
119:
102:
80:
71:
66:
57:
48:
39:
34:
3810:Gervais's beaked whale (
3765:Sowerby's beaked whale (
3476:Bolivian river dolphin (
2068:Namibian Dolphin Project
1559:10.1578/AM.37.2.2011.151
452:calves every 2–4 years.
49:Heaviside's dolphin off
18:Heaviside's Dolphin
4382:Mammals of South Africa
3948:Cuvier's beaked whale (
3873:Perrin's beaked whale (
3837:Hector's beaked whale (
3801:Ramari's beaked whale (
3774:Andrews' beaked whale (
3650:Arnoux's beaked whale (
3323:Burmeister's porpoise (
3090:Rough-toothed dolphin (
2514:(Oceanic dolphins)
2313:Antarctic minke whale (
1309:10.1111/1749-4877.12380
885:Best, Peter B. (2007).
758:and is included in the
653:Lagenorhynchus obscurus
594:Sufflogobius bibarbatus
588:) and octopus, however
3659:Baird's beaked whale (
3559:Ganges river dolphin (
3485:Amazon river dolphin (
2866:Orca or killer whale (
2790:(Right whale dolphins)
2723:White-beaked dolphin (
2241:Southern right whale (
1908:10.1098/rspb.2018.1178
1176:Biodiversity of Angola
596:) and horse mackerel (
414:
309:Taxonomy and evolution
4249:Paleobiology Database
3900:Spade-toothed whale (
3864:True's beaked whale (
3855:Strap-toothed whale (
3828:Gray's beaked whale (
3783:Hubbs' beaked whale (
3668:Sato's beaked whale (
3568:Indus river dolphin (
3296:Spectacled porpoise (
3112:(Bottlenose dolphins)
2553:Heaviside's dolphin (
2535:Commerson's dolphin (
2340:Pygmy Bryde's whale (
1442:Leeney, Ruth (2011).
1399:Marine Mammal Science
1351:Conservation Genetics
412:
4305:Delphinus heavisidii
3882:Pygmy beaked whale (
3755:(Mesoplodont whales)
2914:False killer whale (
2890:Melon-headed whale (
2610:Pygmy killer whale (
2304:Common minke whale (
2014:MacLeod, CD (2009).
1626:Serot, J. L (2013).
1591:Journal of Mammalogy
1516:10.3957/056.039.0204
834:"Appendices | CITES"
805:: e.T4161A50352086.
35:Heaviside's dolphin
3690:(Bottlenose whales)
3430:Dwarf sperm whale (
3421:Pygmy sperm whale (
3253:(Finless porpoises)
2969:(Humpback dolphins)
2833:Irrawaddy dolphin (
2750:Hourglass dolphin (
2461:Pygmy right whale (
1297:Integrative Zoology
1272:Golaski, S (2015).
747:Prior to 2018, the
743:Conservation status
706:Bycatch and hunting
634:Dive time and depth
586:Merluccius capensis
554:Merluccius capensis
547:Habitat preferences
371:Southern Hemisphere
359:C. commersonii
355:Commerson's dolphin
271:Heaviside's dolphin
74:Conservation status
4377:Mammals of Namibia
3605:La Plata dolphin (
3305:Harbour porpoise (
3274:N. asiaeorientalis
3131:Burrunan dolphin (
2699:Fraser's dolphin (
2562:Hector's dolphin (
1901:(1883): 20181178.
1860:10.1242/jeb.042440
718:Trachurus capensis
502:and into southern
415:
222:C. heavisidii
4372:Mammals of Angola
4344:
4343:
4236:Open Tree of Life
3981:Taxon identifiers
3972:
3971:
3968:
3967:
3964:
3963:
3960:
3959:
3758:
3693:
3632:
3617:
3616:
3580:
3579:
3534:
3533:
3497:
3496:
3442:
3441:
3396:
3395:
3359:
3358:
3347:Dall's porpoise (
3256:
3237:
3222:
3221:
3161:
3160:
3115:
3066:Spinner dolphin (
3048:Striped dolphin (
3039:Clymene dolphin (
2972:
2793:
2741:Peale's dolphin (
2675:Risso's dolphin (
2635:
2544:Chilean dolphin (
2517:
2496:
2481:
2480:
2477:
2476:
2473:
2472:
2436:
2435:
2286:
2271:
2270:
2234:
2178:
1854:(11): 1940–1949.
1741:10.1121/1.3519401
1713:978-0-08-092035-1
1193:978-3-030-03082-7
1012:Molecular Ecology
963:978-1-4729-7715-1
898:978-0-521-89710-5
734:Boat interactions
668:Population status
604:Movement patterns
569:Diet and Foraging
440:Sexual dimorphism
279:) is one of four
268:
267:
114:
97:
27:Species of mammal
16:(Redirected from
4404:
4337:
4336:
4324:
4323:
4322:
4296:
4295:
4283:
4282:
4270:
4269:
4257:
4256:
4244:
4243:
4231:
4230:
4218:
4217:
4205:
4204:
4202:NHMSYS0000376516
4192:
4191:
4179:
4178:
4166:
4165:
4153:
4152:
4140:
4139:
4127:
4126:
4114:
4113:
4101:
4100:
4088:
4087:
4075:
4074:
4062:
4061:
4049:
4048:
4036:
4035:
4023:
4022:
4021:
4008:
4007:
4006:
3976:
3757:
3756:
3752:
3692:
3691:
3687:
3636:
3631:
3630:
3626:
3591:
3545:
3508:
3469:I. araguaiaensis
3453:
3407:
3386:P. macrocephalus
3370:
3255:
3254:
3250:
3241:
3236:
3235:
3231:
3172:
3114:
3113:
3109:
2971:
2970:
2966:
2947:Guiana dolphin (
2792:
2791:
2787:
2644:G. macrorhynchus
2634:
2633:
2629:
2586:Common dolphin (
2521:
2516:
2515:
2511:
2504:
2495:(Toothed whales)
2494:
2487:
2447:
2424:Humpback whale (
2306:B. acutorostrata
2290:
2285:
2284:
2280:
2233:
2232:
2228:
2195:
2186:
2176:
2169:
2102:
2095:
2088:
2079:
2055:
2052:
2046:
2045:
2035:
2033:10.3354/esr00197
2011:
2005:
1998:
1992:
1991:
1973:
1950:Animal Behaviour
1945:
1939:
1938:
1928:
1910:
1886:
1880:
1879:
1842:
1836:
1835:
1809:
1800:(4): 1439–1458.
1785:
1779:
1778:
1752:
1724:
1718:
1717:
1697:
1691:
1690:
1672:
1643:
1632:
1631:
1623:
1617:
1616:
1606:
1582:
1573:
1572:
1570:
1542:
1536:
1535:
1498:
1492:
1491:
1481:
1457:
1451:
1450:
1448:
1439:
1433:
1432:
1422:
1394:
1383:
1382:
1345:
1339:
1338:
1320:
1292:
1286:
1285:
1269:
1263:
1256:
1250:
1249:
1239:
1215:
1206:
1205:
1171:
1165:
1164:
1162:
1151:
1145:
1144:
1124:
1118:
1117:
1099:
1070:
1059:
1058:
1050:
1044:
1043:
1018:(9): 2215–2223.
1007:
998:
997:
977:
968:
967:
947:
934:
933:
927:
918:
903:
902:
882:
869:
868:
858:
849:
848:
846:
844:
830:
824:
823:
821:
819:
814:
784:
541:Hector's dolphin
490:Geographic range
363:Hector's dolphin
351:C. eutropia
328:John Edward Gray
259:
242:
129:
128:
108:
91:
86:
85:
62:
44:
32:
21:
4412:
4411:
4407:
4406:
4405:
4403:
4402:
4401:
4362:Cephalorhynchus
4347:
4346:
4345:
4340:
4332:
4327:
4318:
4317:
4312:
4299:
4291:
4286:
4278:
4273:
4265:
4260:
4252:
4247:
4239:
4234:
4226:
4221:
4213:
4208:
4200:
4195:
4187:
4182:
4174:
4169:
4161:
4156:
4148:
4143:
4135:
4130:
4122:
4117:
4109:
4104:
4096:
4091:
4083:
4078:
4070:
4065:
4057:
4052:
4044:
4039:
4031:
4026:
4017:
4016:
4011:
4002:
4001:
3996:
3983:
3973:
3956:
3932:
3908:
3794:M. densirostris
3754:
3753:
3751:
3741:
3717:
3689:
3688:
3686:
3676:
3629:(Beaked whales)
3628:
3627:
3625:
3613:
3576:
3530:
3493:
3438:
3392:
3355:
3331:
3280:
3265:N. phocaenoides
3252:
3251:
3249:
3233:
3232:
3230:
3218:
3194:
3157:
3111:
3110:
3108:
3098:
3074:
3068:S. longirostris
3050:S. coeruleoalba
3014:
2968:
2967:
2965:
2955:
2922:
2898:
2874:
2850:
2835:O. brevirostris
2817:
2789:
2788:
2786:
2776:
2768:Dusky dolphin (
2707:
2683:
2659:
2631:
2630:
2628:
2618:
2594:
2570:
2526:Cephalorhynchus
2513:
2512:
2510:
2498:
2469:
2432:
2426:M. novaeangliae
2408:
2384:
2358:Omura's whale (
2331:Bryde's whale (
2282:
2281:
2279:
2277:Balaenopteridae
2267:
2230:
2229:
2227:
2217:
2209:Bowhead whale (
2180:
2177:(Baleen whales)
2163:
2113:
2106:
2064:
2059:
2058:
2053:
2049:
2013:
2012:
2008:
1999:
1995:
1947:
1946:
1942:
1888:
1887:
1883:
1844:
1843:
1839:
1787:
1786:
1782:
1726:
1725:
1721:
1714:
1699:
1698:
1694:
1645:
1644:
1635:
1625:
1624:
1620:
1584:
1583:
1576:
1547:Aquatic Mammals
1544:
1543:
1539:
1500:
1499:
1495:
1459:
1458:
1454:
1446:
1441:
1440:
1436:
1396:
1395:
1386:
1357:(4): 987–1002.
1347:
1346:
1342:
1294:
1293:
1289:
1271:
1270:
1266:
1257:
1253:
1217:
1216:
1209:
1194:
1173:
1172:
1168:
1160:
1153:
1152:
1148:
1141:
1126:
1125:
1121:
1076:African Zoology
1072:
1071:
1062:
1052:
1051:
1047:
1009:
1008:
1001:
994:
979:
978:
971:
964:
949:
948:
937:
925:
920:
919:
906:
899:
884:
883:
872:
860:
859:
852:
842:
840:
832:
831:
827:
817:
815:
786:
785:
778:
773:
768:
745:
736:
727:
708:
699:
670:
661:
645:
636:
627:
606:
571:
562:
549:
537:Cephalorhynchus
492:
487:
471:
462:
449:
407:
402:
382:Southern Africa
378:Cephalorhynchus
367:C. hectori
347:Chilean dolphin
343:
316:
311:
286:Cephalorhynchus
250:
244:
238:
225:
209:Cephalorhynchus
123:
115:
98:
89:Near Threatened
87:
83:
76:
28:
23:
22:
15:
12:
11:
5:
4410:
4408:
4400:
4399:
4394:
4389:
4384:
4379:
4374:
4369:
4364:
4359:
4349:
4348:
4342:
4341:
4339:
4338:
4325:
4309:
4307:
4301:
4300:
4298:
4297:
4284:
4271:
4258:
4245:
4232:
4219:
4206:
4193:
4180:
4167:
4154:
4141:
4128:
4115:
4102:
4089:
4076:
4063:
4050:
4037:
4024:
4009:
3993:
3991:
3985:
3984:
3979:
3970:
3969:
3966:
3965:
3962:
3961:
3958:
3957:
3955:
3954:
3950:Z. cavirostris
3944:
3942:
3934:
3933:
3931:
3930:
3920:
3918:
3910:
3909:
3907:
3906:
3897:
3888:
3879:
3870:
3861:
3852:
3843:
3834:
3825:
3816:
3807:
3798:
3789:
3780:
3771:
3761:
3759:
3743:
3742:
3740:
3739:
3729:
3727:
3719:
3718:
3716:
3715:
3706:
3696:
3694:
3678:
3677:
3675:
3674:
3665:
3656:
3646:
3644:
3633:
3619:
3618:
3615:
3614:
3612:
3611:
3607:P. blainvillei
3601:
3599:
3588:
3582:
3581:
3578:
3577:
3575:
3574:
3565:
3555:
3553:
3542:
3536:
3535:
3532:
3531:
3529:
3528:
3518:
3516:
3505:
3499:
3498:
3495:
3494:
3492:
3491:
3487:I. geoffrensis
3482:
3478:I. boliviensis
3473:
3463:
3461:
3450:
3444:
3443:
3440:
3439:
3437:
3436:
3427:
3417:
3415:
3404:
3398:
3397:
3394:
3393:
3391:
3390:
3380:
3378:
3367:
3361:
3360:
3357:
3356:
3354:
3353:
3343:
3341:
3333:
3332:
3330:
3329:
3325:P. spinipinnis
3320:
3311:
3302:
3292:
3290:
3282:
3281:
3279:
3278:
3269:
3259:
3257:
3238:
3224:
3223:
3220:
3219:
3217:
3216:
3206:
3204:
3196:
3195:
3193:
3192:
3186:Beluga whale (
3182:
3180:
3177:Delphinapterus
3169:
3163:
3162:
3159:
3158:
3156:
3155:
3146:
3137:
3128:
3118:
3116:
3100:
3099:
3097:
3096:
3092:S. bredanensis
3086:
3084:
3076:
3075:
3073:
3072:
3063:
3054:
3045:
3036:
3026:
3024:
3016:
3015:
3013:
3012:
3003:
2994:
2985:
2975:
2973:
2957:
2956:
2954:
2953:
2944:
2940:S. fluviatilis
2934:
2932:
2924:
2923:
2921:
2920:
2910:
2908:
2900:
2899:
2897:
2896:
2886:
2884:
2876:
2875:
2873:
2872:
2862:
2860:
2852:
2851:
2849:
2848:
2839:
2829:
2827:
2819:
2818:
2816:
2815:
2806:
2796:
2794:
2778:
2777:
2775:
2774:
2765:
2761:L. obliquidens
2756:
2747:
2738:
2729:
2725:L. albirostris
2719:
2717:
2714:Lagenorhynchus
2709:
2708:
2706:
2705:
2695:
2693:
2685:
2684:
2682:
2681:
2671:
2669:
2661:
2660:
2658:
2657:
2648:
2638:
2636:
2632:(Pilot whales)
2620:
2619:
2617:
2616:
2606:
2604:
2596:
2595:
2593:
2592:
2582:
2580:
2572:
2571:
2569:
2568:
2559:
2550:
2541:
2537:C. commersonii
2531:
2529:
2518:
2500:
2499:
2490:
2483:
2482:
2479:
2478:
2475:
2474:
2471:
2470:
2468:
2467:
2457:
2455:
2444:
2438:
2437:
2434:
2433:
2431:
2430:
2420:
2418:
2410:
2409:
2407:
2406:
2396:
2394:
2386:
2385:
2383:
2382:
2376:Rice's whale (
2373:
2364:
2355:
2346:
2337:
2328:
2319:
2315:B. bonaerensis
2310:
2300:
2298:
2287:
2273:
2272:
2269:
2268:
2266:
2265:
2256:
2247:
2237:
2235:
2231:(Right whales)
2219:
2218:
2216:
2215:
2205:
2203:
2192:
2182:
2181:
2172:
2165:
2164:
2162:
2161:
2155:
2149:
2147:Laurasiatheria
2143:
2137:
2131:
2125:
2118:
2115:
2114:
2107:
2105:
2104:
2097:
2090:
2082:
2076:
2075:
2070:
2063:
2062:External links
2060:
2057:
2056:
2047:
2006:
1993:
1940:
1881:
1837:
1780:
1735:(1): 449–457.
1719:
1712:
1692:
1655:(4): 455–466.
1633:
1618:
1597:(5): 866–877.
1574:
1553:(2): 151–160.
1537:
1510:(2): 143–154.
1493:
1472:(1): 843–861.
1452:
1434:
1405:(1): 107–124.
1384:
1340:
1303:(5): 506–526.
1287:
1264:
1251:
1230:(1): 237–270.
1207:
1192:
1166:
1146:
1140:978-0125885058
1139:
1119:
1060:
1045:
999:
992:
969:
962:
935:
904:
897:
870:
850:
825:
775:
774:
772:
769:
767:
764:
744:
741:
735:
732:
726:
725:Climate change
723:
715:(specifically
713:horse mackerel
707:
704:
698:
695:
669:
666:
660:
657:
649:dusky dolphins
644:
641:
635:
632:
626:
623:
616:, whereby the
605:
602:
570:
567:
561:
558:
548:
545:
491:
488:
486:
483:
470:
467:
461:
458:
448:
445:
406:
403:
401:
398:
342:
339:
332:John Heaviside
320:United Kingdom
315:
312:
310:
307:
266:
265:
261:
260:
252:
251:
245:
234:
233:
227:
226:
219:
217:
213:
212:
205:
201:
200:
195:
191:
190:
185:
181:
180:
175:
171:
170:
165:
161:
160:
155:
151:
150:
145:
141:
140:
135:
131:
130:
117:
116:
103:
100:
99:
81:
78:
77:
72:
69:
68:
64:
63:
55:
54:
46:
45:
37:
36:
26:
24:
14:
13:
10:
9:
6:
4:
3:
2:
4409:
4398:
4395:
4393:
4390:
4388:
4385:
4383:
4380:
4378:
4375:
4373:
4370:
4368:
4365:
4363:
4360:
4358:
4355:
4354:
4352:
4335:
4330:
4326:
4321:
4315:
4311:
4310:
4308:
4306:
4302:
4294:
4289:
4285:
4281:
4276:
4272:
4268:
4263:
4259:
4255:
4250:
4246:
4242:
4237:
4233:
4229:
4224:
4220:
4216:
4211:
4207:
4203:
4198:
4194:
4190:
4185:
4181:
4177:
4172:
4168:
4164:
4159:
4155:
4151:
4146:
4142:
4138:
4133:
4129:
4125:
4120:
4116:
4112:
4107:
4103:
4099:
4094:
4090:
4086:
4081:
4077:
4073:
4068:
4064:
4060:
4055:
4051:
4047:
4042:
4038:
4034:
4029:
4025:
4020:
4014:
4010:
4005:
3999:
3995:
3994:
3992:
3990:
3986:
3982:
3977:
3953:
3951:
3946:
3945:
3943:
3941:
3940:
3935:
3929:
3927:
3922:
3921:
3919:
3917:
3916:
3911:
3905:
3903:
3898:
3896:
3894:
3893:M. stejnegeri
3889:
3887:
3885:
3884:M. peruvianus
3880:
3878:
3876:
3871:
3869:
3867:
3862:
3860:
3858:
3853:
3851:
3849:
3844:
3842:
3840:
3835:
3833:
3831:
3826:
3824:
3822:
3821:M. ginkgodens
3817:
3815:
3813:
3808:
3806:
3804:
3799:
3797:
3795:
3790:
3788:
3786:
3785:M. carlhubbsi
3781:
3779:
3777:
3772:
3770:
3768:
3763:
3762:
3760:
3750:
3749:
3744:
3738:
3736:
3731:
3730:
3728:
3726:
3725:
3720:
3714:
3712:
3711:H. planifrons
3707:
3705:
3703:
3702:H. ampullatus
3698:
3697:
3695:
3685:
3684:
3679:
3673:
3671:
3666:
3664:
3662:
3657:
3655:
3653:
3648:
3647:
3645:
3643:
3642:
3637:
3634:
3624:
3620:
3610:
3608:
3603:
3602:
3600:
3598:
3597:
3592:
3589:
3587:
3586:Pontoporiidae
3583:
3573:
3571:
3566:
3564:
3562:
3557:
3556:
3554:
3552:
3551:
3546:
3543:
3541:
3540:Platanistidae
3537:
3527:
3525:
3524:L. vexillifer
3520:
3519:
3517:
3515:
3514:
3509:
3506:
3504:
3500:
3490:
3488:
3483:
3481:
3479:
3474:
3472:
3470:
3465:
3464:
3462:
3460:
3459:
3454:
3451:
3449:
3445:
3435:
3433:
3428:
3426:
3424:
3419:
3418:
3416:
3414:
3413:
3408:
3405:
3403:
3399:
3389:
3387:
3384:Sperm whale (
3382:
3381:
3379:
3377:
3376:
3371:
3368:
3366:
3362:
3352:
3350:
3345:
3344:
3342:
3340:
3339:
3334:
3328:
3326:
3321:
3319:
3317:
3312:
3310:
3308:
3303:
3301:
3299:
3294:
3293:
3291:
3289:
3288:
3283:
3277:
3275:
3270:
3268:
3266:
3261:
3260:
3258:
3248:
3247:
3242:
3239:
3229:
3225:
3215:
3213:
3208:
3207:
3205:
3203:
3202:
3197:
3191:
3189:
3184:
3183:
3181:
3179:
3178:
3173:
3170:
3168:
3164:
3154:
3152:
3147:
3145:
3143:
3138:
3136:
3134:
3129:
3127:
3125:
3120:
3119:
3117:
3107:
3106:
3101:
3095:
3093:
3088:
3087:
3085:
3083:
3082:
3077:
3071:
3069:
3064:
3062:
3060:
3055:
3053:
3051:
3046:
3044:
3042:
3037:
3035:
3033:
3028:
3027:
3025:
3023:
3022:
3017:
3011:
3009:
3004:
3002:
3000:
2999:S. sahulensis
2995:
2993:
2991:
2986:
2984:
2982:
2977:
2976:
2974:
2964:
2963:
2958:
2952:
2950:
2949:S. guianensis
2945:
2943:
2941:
2936:
2935:
2933:
2931:
2930:
2925:
2919:
2917:
2916:P. crassidens
2912:
2911:
2909:
2907:
2906:
2901:
2895:
2893:
2888:
2887:
2885:
2883:
2882:
2881:Peponocephala
2877:
2871:
2869:
2864:
2863:
2861:
2859:
2858:
2853:
2847:
2845:
2840:
2838:
2836:
2831:
2830:
2828:
2826:
2825:
2820:
2814:
2812:
2807:
2805:
2803:
2798:
2797:
2795:
2785:
2784:
2779:
2773:
2771:
2766:
2764:
2762:
2757:
2755:
2753:
2748:
2746:
2744:
2739:
2737:
2735:
2730:
2728:
2726:
2721:
2720:
2718:
2716:
2715:
2710:
2704:
2702:
2697:
2696:
2694:
2692:
2691:
2690:Lagenodelphis
2686:
2680:
2678:
2673:
2672:
2670:
2668:
2667:
2662:
2656:
2654:
2649:
2647:
2645:
2640:
2639:
2637:
2627:
2626:
2621:
2615:
2613:
2608:
2607:
2605:
2603:
2602:
2597:
2591:
2589:
2584:
2583:
2581:
2579:
2578:
2573:
2567:
2565:
2560:
2558:
2556:
2555:C. heavisidii
2551:
2549:
2547:
2542:
2540:
2538:
2533:
2532:
2530:
2528:
2527:
2522:
2519:
2509:
2505:
2501:
2497:
2488:
2484:
2466:
2464:
2459:
2458:
2456:
2454:
2453:
2448:
2445:
2443:
2442:Cetotheriidae
2439:
2429:
2427:
2422:
2421:
2419:
2417:
2416:
2411:
2405:
2403:
2398:
2397:
2395:
2393:
2392:
2387:
2381:
2379:
2374:
2372:
2370:
2365:
2363:
2361:
2356:
2354:
2352:
2347:
2345:
2343:
2338:
2336:
2334:
2329:
2327:
2325:
2320:
2318:
2316:
2311:
2309:
2307:
2302:
2301:
2299:
2297:
2296:
2291:
2288:
2278:
2274:
2264:
2262:
2257:
2255:
2253:
2248:
2246:
2244:
2239:
2238:
2236:
2226:
2225:
2220:
2214:
2212:
2211:B. mysticetus
2207:
2206:
2204:
2202:
2201:
2196:
2193:
2191:
2187:
2183:
2179:
2170:
2166:
2160:
2156:
2154:
2150:
2148:
2144:
2142:
2138:
2136:
2132:
2130:
2126:
2124:
2120:
2119:
2116:
2111:
2103:
2098:
2096:
2091:
2089:
2084:
2083:
2080:
2074:
2073:uk.whales.org
2071:
2069:
2066:
2065:
2061:
2051:
2048:
2043:
2039:
2034:
2029:
2025:
2021:
2017:
2010:
2007:
2003:
1997:
1994:
1989:
1985:
1981:
1977:
1972:
1967:
1963:
1959:
1955:
1951:
1944:
1941:
1936:
1932:
1927:
1922:
1918:
1914:
1909:
1904:
1900:
1896:
1892:
1885:
1882:
1877:
1873:
1869:
1865:
1861:
1857:
1853:
1849:
1841:
1838:
1833:
1829:
1825:
1821:
1817:
1813:
1808:
1803:
1799:
1795:
1791:
1784:
1781:
1776:
1772:
1768:
1764:
1760:
1756:
1751:
1746:
1742:
1738:
1734:
1730:
1723:
1720:
1715:
1709:
1705:
1704:
1696:
1693:
1688:
1684:
1680:
1676:
1671:
1666:
1662:
1658:
1654:
1650:
1642:
1640:
1638:
1634:
1629:
1622:
1619:
1614:
1610:
1605:
1600:
1596:
1592:
1588:
1581:
1579:
1575:
1569:
1564:
1560:
1556:
1552:
1548:
1541:
1538:
1533:
1529:
1525:
1521:
1517:
1513:
1509:
1505:
1497:
1494:
1489:
1485:
1480:
1475:
1471:
1467:
1463:
1456:
1453:
1445:
1438:
1435:
1430:
1426:
1421:
1416:
1412:
1408:
1404:
1400:
1393:
1391:
1389:
1385:
1380:
1376:
1372:
1368:
1364:
1360:
1356:
1352:
1344:
1341:
1336:
1332:
1328:
1324:
1319:
1314:
1310:
1306:
1302:
1298:
1291:
1288:
1283:
1279:
1275:
1268:
1265:
1261:
1255:
1252:
1247:
1243:
1238:
1233:
1229:
1225:
1221:
1214:
1212:
1208:
1203:
1199:
1195:
1189:
1185:
1181:
1177:
1170:
1167:
1159:
1158:
1150:
1147:
1142:
1136:
1132:
1131:
1123:
1120:
1115:
1111:
1107:
1103:
1098:
1093:
1089:
1085:
1081:
1077:
1069:
1067:
1065:
1061:
1056:
1049:
1046:
1041:
1037:
1033:
1029:
1025:
1021:
1017:
1013:
1006:
1004:
1000:
995:
993:9780080919935
989:
985:
984:
976:
974:
970:
965:
959:
955:
954:
946:
944:
942:
940:
936:
931:
924:
917:
915:
913:
911:
909:
905:
900:
894:
890:
889:
881:
879:
877:
875:
871:
866:
865:
857:
855:
851:
839:
835:
829:
826:
813:
808:
804:
800:
799:
794:
792:
783:
781:
777:
770:
765:
763:
761:
757:
752:
750:
742:
740:
733:
731:
724:
722:
720:
719:
714:
705:
703:
696:
694:
691:
687:
683:
679:
675:
667:
665:
659:Vocalisations
658:
656:
654:
650:
642:
640:
633:
631:
624:
622:
619:
615:
611:
603:
601:
599:
595:
591:
587:
583:
579:
576:
568:
566:
559:
557:
555:
546:
544:
542:
538:
532:
530:
529:Britannia Bay
526:
522:
518:
514:
510:
505:
501:
497:
489:
484:
482:
480:
476:
475:killer whales
468:
466:
459:
457:
455:
446:
444:
441:
437:
436:pectoral fins
433:
429:
425:
421:
411:
404:
399:
397:
395:
394:South America
391:
387:
383:
379:
374:
372:
368:
364:
360:
356:
352:
348:
340:
338:
335:
333:
329:
325:
321:
313:
308:
306:
304:
300:
296:
292:
288:
287:
283:in the genus
282:
278:
277:
272:
262:
258:
253:
248:
243:
241:
235:
232:
231:Binomial name
228:
224:
223:
218:
215:
214:
211:
210:
206:
203:
202:
199:
196:
193:
192:
189:
186:
183:
182:
179:
176:
173:
172:
169:
166:
163:
162:
159:
156:
153:
152:
149:
146:
143:
142:
139:
136:
133:
132:
127:
122:
118:
112:
106:
101:
95:
90:
79:
75:
70:
65:
61:
56:
52:
47:
43:
38:
33:
30:
19:
4304:
3988:
3949:
3937:
3926:T. shepherdi
3925:
3913:
3902:M. traversii
3901:
3892:
3883:
3874:
3865:
3856:
3847:
3838:
3829:
3820:
3812:M. europaeus
3811:
3802:
3793:
3784:
3775:
3766:
3746:
3735:I. pacificus
3734:
3722:
3710:
3701:
3681:
3669:
3660:
3651:
3639:
3606:
3594:
3569:
3561:P. gangetica
3560:
3548:
3523:
3511:
3486:
3477:
3468:
3456:
3431:
3423:K. breviceps
3422:
3410:
3385:
3373:
3365:Physeteridae
3348:
3338:Phocoenoides
3336:
3324:
3315:
3306:
3298:P. dioptrica
3297:
3285:
3273:
3264:
3244:
3212:M. monoceros
3211:
3199:
3187:
3175:
3167:Monodontidae
3151:T. truncatus
3150:
3142:T. erebennus
3141:
3133:T. australis
3132:
3123:
3103:
3091:
3079:
3067:
3059:S. frontalis
3058:
3049:
3040:
3032:S. attenuata
3031:
3019:
3007:
2998:
2989:
2981:S. chinensis
2980:
2960:
2948:
2939:
2927:
2915:
2903:
2891:
2879:
2867:
2855:
2844:O. heinsohni
2843:
2834:
2822:
2810:
2801:
2783:Lissodelphis
2781:
2769:
2760:
2751:
2743:L. australis
2742:
2733:
2724:
2712:
2700:
2688:
2676:
2664:
2652:
2643:
2625:Globicephala
2623:
2612:F. attenuata
2611:
2599:
2587:
2575:
2563:
2554:
2552:
2545:
2536:
2524:
2463:C. marginata
2462:
2450:
2425:
2413:
2401:
2400:Gray whale (
2391:Eschrichtius
2389:
2377:
2368:
2359:
2350:
2349:Blue whale (
2341:
2332:
2323:
2314:
2305:
2295:Balaenoptera
2293:
2260:
2252:E. glacialis
2251:
2243:E. australis
2242:
2222:
2210:
2198:
2159:Whippomorpha
2153:Artiodactyla
2050:
2023:
2019:
2009:
2001:
1996:
1953:
1949:
1943:
1898:
1894:
1884:
1851:
1847:
1840:
1797:
1793:
1783:
1750:10722/140937
1732:
1728:
1722:
1702:
1695:
1652:
1648:
1627:
1621:
1594:
1590:
1550:
1546:
1540:
1507:
1503:
1496:
1469:
1465:
1455:
1437:
1402:
1398:
1354:
1350:
1343:
1300:
1296:
1290:
1273:
1267:
1259:
1254:
1227:
1223:
1175:
1169:
1156:
1149:
1129:
1122:
1082:(1): 78–91.
1079:
1075:
1054:
1048:
1015:
1011:
982:
952:
929:
887:
863:
841:. Retrieved
837:
828:
816:. Retrieved
802:
796:
790:
753:
746:
737:
728:
716:
709:
700:
682:Lamberts Bay
671:
662:
652:
646:
637:
628:
614:South Africa
607:
597:
593:
585:
572:
563:
553:
550:
536:
533:
509:South Africa
493:
485:Distribution
479:Orcinus orca
478:
472:
463:
450:
447:Life history
416:
392:and then to
377:
375:
366:
358:
350:
344:
336:
326:. Zoologist
317:
314:Nomenclature
289:. The small
284:
275:
274:
270:
269:
239:
237:
221:
220:
208:
184:Infraorder:
178:Artiodactyla
29:
4262:SeaLifeBase
4119:iNaturalist
4013:Wikispecies
3857:M. layardii
3776:M. bowdoini
3724:Indopacetus
3307:P. phocoena
3246:Neophocoena
3234:(Porpoises)
3228:Phocoenidae
2802:L. borealis
2770:L. obscurus
2752:L. cruciger
2546:C. eutropia
2508:Delphinidae
2493:Odontoceti
2402:E. robustus
2369:B. physalus
2367:Fin whale (
2351:B. musculus
2324:B. borealis
2322:Sei whale (
2261:E. japonica
2145:Superorder
2139:Infraclass
2026:: 125–136.
1956:: 273–284.
1262:12:237–270.
818:19 November
400:Description
390:New Zealand
198:Delphinidae
107:Appendix II
4351:Categories
4320:Q125572400
3915:Tasmacetus
3875:M. perrini
3848:M. hotaula
3839:M. hectori
3748:Mesoplodon
3683:Hyperoodon
3670:B. minimus
3661:B. bairdii
3652:B. arnuxii
3596:Pontoporia
3550:Platanista
3124:T. aduncus
3041:S. clymene
3008:S. teuszii
2990:S. plumbea
2892:P. electra
2811:L. peronii
2677:G. griseus
2588:D. delphis
2564:C. hectori
2491:Parvorder
2283:(Rorquals)
2190:Balaenidae
2175:Mysticeti
2173:Parvorder
2004:27:494-497
1971:2263/71067
1670:2263/43900
1568:2263/17051
1420:2263/10207
1318:2263/74315
1282:2263/57243
1097:2263/14283
843:14 January
766:References
686:Walvis bay
582:cephalopod
513:Walvis Bay
496:Cape Point
460:Group size
424:dorsal fin
405:Morphology
361:) and the
53:, Namibia
3767:M. bidens
3641:Berardius
3623:Ziphiidae
3503:Lipotidae
3314:Vaquita (
3210:Narwhal (
3188:D. leucas
2905:Pseudorca
2734:L. acutus
2577:Delphinus
2415:Megaptera
2360:B. omurai
2333:B. brydei
2224:Eubalaena
2157:Suborder
2042:1863-5407
1980:0003-3472
1917:0962-8452
1868:0022-0949
1816:1010-061X
1759:0001-4966
1679:1814-232X
1613:0022-2372
1524:0379-4369
1488:0257-7615
1429:0824-0469
1371:1566-0621
1246:0257-7615
1202:134352239
1114:219289679
1106:1562-7020
838:cites.org
771:Citations
678:Table Bay
674:abundance
672:No total
560:Behaviour
525:Cape Town
521:Table Bay
469:Predation
454:Gestation
216:Species:
144:Kingdom:
138:Eukaryota
4314:Wikidata
4275:Species+
4189:14300039
4137:11241419
4098:46559320
3998:Wikidata
3939:Ziphius
3866:M. mirus
3830:M. grayi
3570:P. minor
3432:K. simus
3402:Kogiidae
3375:Physeter
3349:P. dalli
3316:P. sinus
3287:Phocoena
3105:Tursiops
3021:Stenella
2938:Tucuxi (
2824:Orcaella
2701:L. hosei
2653:G. melas
2378:B. ricei
2342:B. edeni
2141:Eutheria
2135:Mammalia
2129:Chordata
2123:Animalia
2121:Kingdom
1988:72334526
1935:30051842
1876:20472781
1832:23902777
1824:17584238
1767:21303024
1687:56296143
1532:85572457
1379:17218356
1335:59306988
1327:30688009
1040:24368161
1032:11555263
690:Lüderitz
618:dolphins
575:demersal
517:Lüderitz
432:blowhole
299:Benguela
291:cetacean
281:dolphins
194:Family:
168:Mammalia
158:Chordata
154:Phylum:
148:Animalia
134:Domain:
94:IUCN 3.1
51:Lüderitz
4334:2440521
4176:1006409
4111:5220041
4004:Q301048
3803:M. eueu
3522:Baiji (
3513:Lipotes
3448:Iniidae
3201:Monodon
2929:Sotalia
2868:O. orca
2857:Orcinus
2666:Grampus
2452:Caperea
2200:Balaena
2127:Phylum
2112:species
2110:Cetacea
2108:Extant
1926:6083265
1775:8219776
697:Threats
610:diurnal
590:pelagic
500:Namibia
353:), the
297:to the
295:endemic
204:Genus:
188:Cetacea
174:Order:
164:Class:
109: (
92: (
4293:254978
4241:860906
4228:254978
4215:103583
4150:180451
4059:462567
4041:ARKive
2601:Feresa
2151:Order
2133:Class
2040:
1986:
1978:
1933:
1923:
1915:
1874:
1866:
1830:
1822:
1814:
1773:
1765:
1757:
1710:
1685:
1677:
1611:
1530:
1522:
1486:
1427:
1377:
1369:
1333:
1325:
1244:
1200:
1190:
1137:
1112:
1104:
1038:
1030:
990:
960:
895:
527:) and
504:Angola
420:thorax
303:Africa
249:, 1828
4288:WoRMS
4267:69391
4254:82045
4132:IRMNG
4124:41517
4072:69J6N
3412:Kogia
3081:Steno
2962:Sousa
1984:S2CID
1828:S2CID
1771:S2CID
1683:S2CID
1528:S2CID
1447:(PDF)
1375:S2CID
1331:S2CID
1198:S2CID
1161:(PDF)
1110:S2CID
1036:S2CID
926:(PDF)
428:fluke
111:CITES
105:CITES
4329:GBIF
4280:4438
4223:OBIS
4210:NCBI
4163:4161
4158:IUCN
4145:ITIS
4106:GBIF
4054:BOLD
3458:Inia
2038:ISSN
1976:ISSN
1931:PMID
1913:ISSN
1872:PMID
1864:ISSN
1820:PMID
1812:ISSN
1763:PMID
1755:ISSN
1708:ISBN
1675:ISSN
1609:ISSN
1520:ISSN
1484:ISSN
1425:ISSN
1367:ISSN
1323:PMID
1242:ISSN
1188:ISBN
1135:ISBN
1102:ISSN
1028:PMID
988:ISBN
958:ISBN
893:ISBN
845:2022
820:2021
803:2018
688:and
680:and
580:and
578:fish
515:and
247:Gray
4197:NBN
4184:MSW
4171:MDD
4093:EoL
4080:CMS
4067:CoL
4028:ADW
2028:doi
1966:hdl
1958:doi
1954:150
1921:PMC
1903:doi
1899:285
1856:doi
1852:213
1802:doi
1745:hdl
1737:doi
1733:129
1665:hdl
1657:doi
1599:doi
1563:hdl
1555:doi
1512:doi
1474:doi
1415:hdl
1407:doi
1359:doi
1313:hdl
1305:doi
1278:hdl
1232:doi
1180:doi
1092:hdl
1084:doi
1020:doi
807:doi
293:is
4353::
4331::
4316::
4290::
4277::
4264::
4251::
4238::
4225::
4212::
4199::
4186::
4173::
4160::
4147::
4134::
4121::
4108::
4095::
4082::
4069::
4056::
4043::
4030::
4015::
4000::
2036:.
2022:.
2018:.
1982:.
1974:.
1964:.
1952:.
1929:.
1919:.
1911:.
1897:.
1893:.
1870:.
1862:.
1850:.
1826:.
1818:.
1810:.
1798:20
1796:.
1792:.
1769:.
1761:.
1753:.
1743:.
1731:.
1681:.
1673:.
1663:.
1653:36
1651:.
1636:^
1607:.
1595:87
1593:.
1589:.
1577:^
1561:.
1551:37
1549:.
1526:.
1518:.
1508:39
1506:.
1482:.
1470:12
1468:.
1464:.
1423:.
1413:.
1403:25
1401:.
1387:^
1373:.
1365:.
1355:13
1353:.
1329:.
1321:.
1311:.
1301:14
1299:.
1240:.
1228:12
1226:.
1222:.
1210:^
1196:.
1186:.
1108:.
1100:.
1090:.
1080:45
1078:.
1063:^
1034:.
1026:.
1016:10
1014:.
1002:^
972:^
938:^
928:.
907:^
873:^
853:^
836:.
801:.
795:.
779:^
608:A
426:,
373:.
305:.
3952:)
3928:)
3904:)
3895:)
3886:)
3877:)
3868:)
3859:)
3850:)
3841:)
3832:)
3823:)
3814:)
3805:)
3796:)
3787:)
3778:)
3769:)
3737:)
3713:)
3704:)
3672:)
3663:)
3654:)
3609:)
3572:)
3563:)
3526:)
3489:)
3480:)
3471:)
3434:)
3425:)
3388:)
3351:)
3327:)
3318:)
3309:)
3300:)
3276:)
3267:)
3214:)
3190:)
3153:)
3144:)
3135:)
3126:)
3094:)
3070:)
3061:)
3052:)
3043:)
3034:)
3010:)
3001:)
2992:)
2983:)
2951:)
2942:)
2918:)
2894:)
2870:)
2846:)
2837:)
2813:)
2804:)
2772:)
2763:)
2754:)
2745:)
2736:)
2727:)
2703:)
2679:)
2655:)
2646:)
2614:)
2590:)
2566:)
2557:)
2548:)
2539:)
2465:)
2428:)
2404:)
2380:)
2371:)
2362:)
2353:)
2344:)
2335:)
2326:)
2317:)
2308:)
2263:)
2254:)
2245:)
2213:)
2101:e
2094:t
2087:v
2044:.
2030::
2024:7
1990:.
1968::
1960::
1937:.
1905::
1878:.
1858::
1834:.
1804::
1777:.
1747::
1739::
1716:.
1689:.
1667::
1659::
1615:.
1601::
1571:.
1565::
1557::
1534:.
1514::
1490:.
1476::
1449:.
1431:.
1417::
1409::
1381:.
1361::
1337:.
1315::
1307::
1284:.
1280::
1248:.
1234::
1204:.
1182::
1143:.
1116:.
1094::
1086::
1042:.
1022::
996:.
966:.
932:.
901:.
847:.
822:.
809::
793:"
789:"
651:(
523:(
477:(
365:(
357:(
349:(
273:(
113:)
96:)
20:)
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
