31:
341:
is correlated with both sex and size. The smallest crabs have green claws, followed by orange and pink, with the largest male crabs having purple claws; females with purple claws have small claws. The purple colour is the easiest of the four to distinguish against the
279:. Adults are around 25 mm (1 in) wide, with males being larger and having larger and more conspicuously coloured claws. The males wave their claws to communicate with other crabs, giving them their
651:
639:
943:
956:
1038:
411:
moves its carapace up and down; this capability allows the crab to breathe air without losing the water held in the gill chamber under the carapace.
917:
433:
that it uses to eat larger pieces of plants and animals. Both males and females use their claws alternately; while one is transferring food to the
969:
727:
1033:
1028:
337:; a male's claws thus become proportionally larger as the crab grows, becoming much larger than those of females. The colour of the
995:
668:
568:
891:
878:
576:
961:
517:
of the crabs in the family
Ocypodidae, and erected a new subfamily for the genus, which was later raised to the
846:
30:
896:
514:
183:
1023:
1000:
434:
430:
374:
607:
480:
139:
287:. They can breathe both in air and under water, and feed at low tide on detritus in the sediment.
826:
768:
744:
526:
452:
Males signal to other crabs by waving their brightly coloured claws, which gives the species its
205:
38:
948:
403:, where it is the most abundant crab species, usually living among the roots of the mangroves.
982:
904:
785:
694:
611:
569:"Systema Brachyurorum: Part I. An annotated checklist of extant Brachyuran crabs of the world"
502:
326:
160:
883:
987:
818:
777:
736:
660:
393:
366:
721:
David P. Maitland (1992). "Carapace movements aid air breathing in the semaphore crab,
518:
472:
457:
422:
781:
1017:
748:
909:
807:(H. Milne Edwards, 1837) (Decapoda, Ocypodidae) as revealed by larval morphology".
585:
497:
370:
343:
315:
85:
690:
930:
809:
664:
453:
280:
121:
869:
542:
484:
362:
73:
822:
330:
276:
61:
789:
863:
492:
426:
418:
396:
378:
358:
334:
311:
268:
97:
830:
762:
G. R. MacFarlane; D. J. Booth; K. R. Brown (2000). "The semaphore crab,
922:
740:
400:
272:
260:
935:
308:
49:
840:
803:
D. R. Fielder; J. G. Greenwood (1985). "The systematic position of
766:: bio-indication potential for heavy metals in estuarine systems".
974:
638:
Tanya Detto; Jochen Zeil; Robert D. Magrath; Sarah Hunt (2004).
446:
442:
338:
322:
264:
109:
844:
407:
can breathe both in air and in water. While out of the water,
525:
and other species described in the genus are all considered
307:
are around 25 mm (1 in) wide, with a dark purple,
437:, the other claw is collecting more material. Predators of
567:
Peter K. L. Ng; Danièle Guinot; Peter J. F. Davie (2008).
853:
417:is primarily a deposit feeder, sifting through the
652:Journal of Experimental Marine Biology and Ecology
640:"Sex, size and colour in a semi-terrestrial crab,
505:in 1851, alongside a newly described species,
8:
841:
716:
714:
712:
29:
20:
346:of the mudflats on which the crabs live.
685:
683:
681:
602:
600:
598:
501:). It was transferred to a new genus by
633:
631:
629:
554:
329:, with males' claws showing positive
7:
725:(Decapoda: Brachyura: Ocypodidae)".
562:
560:
558:
728:Journal of Comparative Physiology B
429:from mangroves, but also has large
521:of family, as Heloeciidae. Dana's
14:
1039:Taxa named by Henri Milne-Edwards
291:is the only species in the genus
377:), as well as eastern parts of
509:. A 1983 study concluded that
1:
782:10.1016/S0166-445X(00)00083-7
608:"Semaphore Crab Illustration"
333:, while females' claws grow
295:and the family Heloeciidae.
665:10.1016/j.jembe.2003.09.023
577:Raffles Bulletin of Zoology
219:H. Milne-Edwards, 1837
1055:
644:(H. Milne Edwards, 1837)"
212:
204:
189:
182:
39:Scientific classification
37:
28:
23:
1034:Monotypic decapod genera
1029:Crustaceans of Australia
275:along the east coast of
199:(H. Milne-Edwards, 1837)
823:10.1163/156854085X00954
584:: 1β286. Archived from
318:are on long eyestalks.
365:), along the coast of
884:Heloecius_cordiformis
855:Heloecius cordiformis
805:Heloecius cordiformis
764:Heloecius cordiformis
723:Heloecius cordiformis
642:Heloecius cordiformis
477:Gelasimus cordiformis
469:Heloecius cordiformis
441:include a variety of
385:Ecology and behaviour
256:Heloecius cordiformis
216:Gelasimus cordiformis
193:Heloecius cordiformis
344:reflectance spectrum
481:Henri Milne-Edwards
263:of semiterrestrial
234:Heloecius inornatus
225:Heloecius areolatus
175:H. cordiformis
769:Aquatic Toxicology
741:10.1007/BF00260766
527:taxonomic synonyms
327:sexually dimorphic
242:Heloecius signatus
1011:
1010:
983:Open Tree of Life
847:Taxon identifiers
695:Australian Museum
612:Australian Museum
503:James Dwight Dana
252:
251:
246:
238:
229:
220:
1046:
1004:
1003:
991:
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978:
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718:
707:
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697:. 5 January 2010
691:"Semaphore Crab"
687:
676:
675:
674:on 22 June 2007.
673:
667:. Archived from
648:
635:
624:
623:
621:
619:
614:. 7 October 2009
604:
593:
592:
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483:, in the family
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140:H. Milne-Edwards
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21:
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839:
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627:
617:
615:
606:
605:
596:
591:on 6 June 2011.
588:
571:
566:
565:
556:
551:
539:
473:first described
466:
387:
371:Port Philip Bay
367:New South Wales
352:
301:
269:mangrove swamps
223:
200:
197:
191:
178:
164:
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157:
143:
137:
136:
124:
112:
100:
88:
76:
64:
52:
24:Semaphore crab
17:
12:
11:
5:
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1042:
1041:
1036:
1031:
1026:
1016:
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1009:
1008:
1006:
1005:
992:
979:
966:
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940:
927:
914:
901:
888:
875:
859:
857:
851:
850:
845:
837:
836:
817:(3): 244β248.
795:
776:(3): 153β166.
754:
735:(4): 375β382.
708:
677:
625:
594:
553:
552:
550:
547:
546:
545:
538:
535:
531:H. cordiformis
465:
462:
439:H. cordiformis
423:organic matter
415:H. cordiformis
409:H. cordiformis
405:H. cordiformis
390:H. cordiformis
386:
383:
355:H. cordiformis
351:
348:
305:H. cordiformis
300:
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289:H. cordiformis
285:semaphore crab
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35:
34:
26:
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16:Genus of crabs
15:
13:
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4:
3:
2:
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1040:
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828:
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548:
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541:
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536:
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524:
520:
516:
513:was the most
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384:
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356:
349:
347:
345:
340:
336:
335:isometrically
332:
328:
324:
319:
317:
316:compound eyes
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217:
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196:
194:
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185:
184:Binomial name
181:
177:
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167:
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156:
155:
150:
147:
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141:
135:
131:
128:
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119:
117:Superfamily:
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27:
22:
19:
854:
814:
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773:
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732:
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722:
699:. Retrieved
669:the original
656:
650:
641:
616:. Retrieved
586:the original
581:
575:
530:
523:H. inornatus
522:
510:
507:H. inornatus
506:
496:
488:
476:
468:
467:
451:
438:
414:
413:
408:
404:
389:
388:
354:
353:
350:Distribution
320:
304:
302:
292:
288:
284:
255:
254:
253:
241:
233:
228:Heller, 1862
224:
215:
192:
190:
174:
173:
153:
152:
133:
105:Infraorder:
86:Malacostraca
18:
1024:Ocypodoidea
931:iNaturalist
810:Crustaceana
464:Systematics
454:common name
357:lives from
299:Description
281:common name
134:Heloeciidae
122:Ocypodoidea
69:Subphylum:
1018:Categories
549:References
543:Ghost crab
485:Ocypodidae
435:mouthparts
394:intertidal
363:Queensland
303:Adults of
245:Hess, 1865
237:Dana, 1851
62:Arthropoda
515:primitive
511:Heloecius
491:is now a
489:Gelasimus
458:semaphore
431:mandibles
401:estuaries
399:, and in
397:mangroves
392:lives in
331:allometry
293:Heloecius
277:Australia
267:found in
169:Species:
154:Heloecius
110:Brachyura
74:Crustacea
45:Kingdom:
949:10793541
870:Q2523176
864:Wikidata
831:20104041
790:10958951
749:27520948
701:21 April
659:: 1β15.
618:21 April
537:See also
493:subgenus
427:detritus
425:and the
419:sediment
379:Tasmania
375:Victoria
359:Brisbane
312:carapace
273:mudflats
206:Synonyms
129:Family:
98:Decapoda
57:Phylum:
50:Animalia
923:4647276
460:crab".
309:mottled
261:species
208:
148:Genus:
93:Order:
81:Class:
1001:444926
988:813111
975:444926
936:418176
829:
788:
747:
314:. The
163:, 1851
142:, 1852
996:WoRMS
962:53299
944:IRMNG
910:3KH2Z
897:69699
827:JSTOR
745:S2CID
672:(PDF)
647:(PDF)
589:(PDF)
572:(PDF)
443:birds
339:claws
323:claws
259:is a
970:OBIS
957:NCBI
918:GBIF
892:BOLD
786:PMID
703:2010
620:2010
519:rank
471:was
456:of "
447:fish
445:and
421:for
325:are
321:The
271:and
265:crab
161:Dana
905:CoL
879:AFD
819:doi
778:doi
737:doi
733:162
661:doi
657:302
529:of
498:Uca
495:of
479:by
475:as
369:to
283:of
1020::
998::
985::
972::
959::
946::
933::
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894::
881::
866::
825:.
815:48
813:.
784:.
774:50
772:.
743:.
731:.
711:^
693:.
680:^
655:.
649:.
628:^
610:.
597:^
582:17
580:.
574:.
557:^
533:.
449:.
381:.
833:.
821::
792:.
780::
751:.
739::
705:.
663::
622:.
487:(
373:(
361:(
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