46:
2086:
33:
249:
area. The majority are dark in color and range in size from approximately 1.5 to 5.0 mm. Most specimens have been collected in arid climates, but one species from
Madagascar is known to occur in a humid forest habitat. Although males have functional wings, heterogynaid females are typically
360:
Manuela Sann, Oliver
Niehuis, Ralph S. Peters, Christoph Mayer, Alexey Kozlov, Lars Podsiadlowski, Sarah Bank, Karen Meusemann, Bernhard Misof, Christoph Bleidorn and Michael Ohl (2018) Phylogenomic analysis of Apoidea sheds new light on the sister group of bees.
1959:
424:
Ohl, M. & Bleidorn, C. (2006) "The phylogenetic position of the enigmatic wasp family
Heterogynaidae based on molecular data, with a new description of a new, nocturnal species (Hymenoptera: Apoidea)".
265:
adapted to hunt in tight spaces, such as under tree bark. This is speculative and has not yet been confirmed by actual observations of behavior. It is also possible that modifications of the female
312:
Branstetter, Michael G.; Danforth, Bryan N.; Pitts, James P.; Faircloth, Brant C.; Ward, Philip S.; Buffington, Matthew L.; Gates, Michael W.; Kula, Robert R.; Brady, Seán G. (2017).
2032:
2058:
258:. Other aspects of their biology are completely unknown, but details of their morphology have prompted researchers to hypothesize that they may be non-
2006:
2045:
2127:
458:
254:, a trait which is unique among spheciform wasps. Wing venation is reduced in both sexes. All species are diurnal, with the exception of
2050:
421:
Goulet, H., Huber, J.T. (1993) Hymenoptera of the World. Agriculture Canada
Research Branch, publication 1894/E. 668pp.
1985:
45:
2120:
2063:
1998:
451:
2146:
155:
2151:
1910:
176:
169:
197:
148:
2113:
246:
162:
1202:
190:
183:
1247:
1157:
977:
920:
444:
394:
325:
242:
1212:
1107:
1275:
1237:
1152:
997:
1497:
1257:
1222:
1112:
410:
40:
1735:
272:
VI and gonostyli may represent a unique prey transport mechanism, but this is also unconfirmed.
1280:
2037:
1967:
1340:
1172:
855:
343:
2097:
314:"Phylogenomic Insights into the Evolution of Stinging Wasps and the Origins of Ants and Bees"
1593:
1525:
1325:
882:
633:
402:
366:
333:
1948:
1554:
1055:
958:
864:
566:
550:
283:, but recent studies utilizing molecular characters do not support this hypothesis. While
398:
329:
2085:
1993:
1980:
1750:
1502:
1375:
1290:
1252:
1142:
1132:
1127:
1063:
1002:
933:
892:
872:
832:
803:
598:
406:
2140:
1867:
1831:
1823:
1507:
1330:
1310:
1270:
1242:
902:
877:
840:
819:
625:
251:
414:
1893:
1854:
1755:
1717:
1700:
1466:
1429:
1421:
1315:
1305:
1295:
1207:
1177:
1073:
1045:
1025:
1007:
845:
688:
680:
638:
511:
276:
234:
222:
1972:
2019:
1942:
1740:
1722:
1627:
1606:
1580:
1492:
1479:
1411:
1403:
1393:
1385:
1227:
1217:
1197:
1147:
1137:
1097:
1040:
1017:
992:
982:
948:
897:
659:
582:
468:
288:
262:
107:
97:
1933:
436:
1862:
1846:
1838:
1827:
1819:
1798:
1745:
1622:
1614:
1572:
1546:
1474:
1354:
1320:
1232:
1192:
1182:
1167:
1030:
912:
887:
811:
775:
651:
574:
370:
338:
313:
292:
287:
of the genus is strongly supported, it is now considered to be related to the
226:
1815:
1811:
1765:
1730:
1656:
1648:
1635:
1585:
1559:
1517:
1487:
1367:
1362:
1335:
1300:
1285:
1187:
1162:
1117:
1089:
1035:
987:
943:
938:
925:
730:
722:
701:
499:
487:
284:
259:
117:
77:
57:
347:
32:
1927:
1760:
1533:
1443:
1262:
1102:
751:
709:
667:
617:
611:
590:
522:
505:
266:
230:
2093:
2011:
1690:
558:
280:
212:
2024:
1806:
1122:
493:
481:
269:
87:
67:
1904:
965:
1536:(paper wasps, potter wasps, pollen wasps, yellowjackets, hornets)
238:
1908:
440:
1782:
1677:
1669:
2101:
211:
is a minor and disputed lineage (only eight described
275:
1917:
1781:
1698:
1689:
1668:
1647:
1605:
1571:
1545:
1516:
1465:
1442:
1420:
1402:
1384:
1353:
1072:
1054:
1016:
957:
911:
863:
854:
831:
802:
773:
750:
721:
700:
679:
650:
624:
610:
549:
520:
385:Nagy (Hymenoptera: Sphecidae; Heterogyninae)".
2121:
452:
8:
2128:
2114:
1905:
1778:
1695:
1462:
860:
799:
770:
546:
542:
459:
445:
437:
31:
20:
337:
291:or, most recently, a lineage within the
381:Day, M.C. (1984). "The enigmatic genus
304:
7:
2082:
2080:
1999:f76db9c5-afc0-42d2-8bb5-0ddf64856515
2100:. You can help Knowledge (XXG) by
407:10.1111/j.1365-3113.1984.tb00054.x
14:
2084:
44:
1638:(sapygid, or club-horned wasps)
1:
1857:(mason bees, leafcutter bees)
2168:
2079:
1888:
545:
476:
371:10.1186/s12862-018-1155-8
339:10.1016/j.cub.2017.03.027
144:
139:
41:Scientific classification
39:
30:
23:
363:BMC Evolutionary Biology
279:to the remainder of the
641:(web-spinning sawflies)
2096:-related article is a
1768:(thread-waisted wasps)
1596:(sierolomorphid wasps)
1528:(rhopalosomatid wasps)
733:(parasitic wood wasps)
156:Heterogyna fantsilotra
427:Systematic Entomology
387:Systematic Entomology
247:Eastern Mediterranean
1994:Fauna Europaea (new)
243:United Arab Emirates
177:Heterogyna nocticola
170:Heterogyna madecassa
577:(cimbicid sawflies)
399:1984SysEn...9..293D
330:2017CBio...27.1019B
225:wasps occurring in
215:in a single genus,
198:Heterogyna ravenala
149:Heterogyna botswana
662:(cedar wood wasps)
585:(conifer sawflies)
163:Heterogyna kugleri
2109:
2108:
2074:
2073:
1911:Taxon identifiers
1902:
1901:
1884:
1883:
1880:
1879:
1876:
1875:
1858:
1850:
1842:
1835:
1802:
1792:
1777:
1776:
1769:
1726:
1725:(cockroach wasps)
1711:
1681:
1660:
1639:
1631:
1618:
1597:
1594:Sierolomorphidae
1589:
1563:
1537:
1529:
1526:Rhopalosomatidae
1483:
1438:
1437:
1371:
1349:
1348:
1341:Trichogrammatidae
1266:
1203:Eutrichosomatidae
1173:Epichrysomallidae
1093:
1083:
971:
929:
856:Proctotrupomorpha
823:
822:(ichneumon wasps)
815:
746:
745:
742:
741:
734:
713:
692:
671:
663:
642:
602:
601:(common sawflies)
594:
593:(pergid sawflies)
586:
578:
570:
562:
206:
205:
191:Heterogyna protea
184:Heterogyna polita
135:
2159:
2130:
2123:
2116:
2088:
2081:
2067:
2066:
2054:
2053:
2041:
2040:
2028:
2027:
2015:
2014:
2002:
2001:
1989:
1988:
1976:
1975:
1963:
1962:
1953:
1952:
1951:
1938:
1937:
1936:
1906:
1856:
1848:
1841:(plasterer bees)
1840:
1809:
1800:
1791:
1790:
1786:
1779:
1767:
1724:
1710:
1709:
1708:(sphecoid wasps)
1705:
1696:
1679:
1658:
1637:
1629:
1616:
1595:
1587:
1561:
1535:
1527:
1481:
1463:
1369:
1326:Tanaostigmatidae
1264:
1248:Melanosomellidae
1091:
1082:
1081:
1077:
970:
969:
961:
927:
883:Neuroscelionidae
861:
821:
813:
800:
771:
732:
711:
690:
669:
661:
640:
634:Megalodontesidae
600:
592:
584:
576:
568:
567:Blasticotomidae
561:(argid sawflies)
560:
547:
543:
461:
454:
447:
438:
418:
373:
358:
352:
351:
341:
324:(7): 1019–1025.
309:
133:
49:
48:
35:
21:
2167:
2166:
2162:
2161:
2160:
2158:
2157:
2156:
2137:
2136:
2135:
2134:
2077:
2075:
2070:
2062:
2057:
2049:
2044:
2036:
2031:
2023:
2018:
2010:
2005:
1997:
1992:
1984:
1979:
1971:
1966:
1958:
1956:
1947:
1946:
1941:
1932:
1931:
1926:
1913:
1903:
1898:
1872:
1788:
1787:
1785:
1773:
1707:
1706:
1704:
1685:
1664:
1659:(scoliid wasps)
1643:
1601:
1567:
1562:(tiphiid wasps)
1555:Bradynobaenidae
1541:
1512:
1457:
1455:
1453:
1451:
1449:
1447:
1445:
1434:
1416:
1398:
1380:
1345:
1158:Cynipencyrtidae
1080:(chalcid wasps)
1079:
1078:
1076:
1068:
1056:Mymarommatoidea
1050:
1012:
978:Proctorenyxidae
963:
962:
959:Proctotrupoidea
953:
921:Austrocynipidae
907:
865:Platygastroidea
850:
827:
793:
791:
789:
787:
785:
783:
781:
779:
777:
765:
763:
761:
759:
757:
755:
753:
738:
717:
712:(stem sawflies)
696:
675:
646:
620:
606:
599:Tenthredinidae
569:(fern sawflies)
551:Tenthredinoidea
536:
534:
532:
530:
528:
526:
524:
516:
472:
465:
380:
377:
376:
359:
355:
318:Current Biology
311:
310:
306:
301:
132:
43:
17:
12:
11:
5:
2165:
2163:
2155:
2154:
2149:
2147:Apoidea genera
2139:
2138:
2133:
2132:
2125:
2118:
2110:
2107:
2106:
2089:
2072:
2071:
2069:
2068:
2055:
2042:
2029:
2016:
2003:
1990:
1981:Fauna Europaea
1977:
1964:
1954:
1939:
1923:
1921:
1915:
1914:
1909:
1900:
1899:
1897:
1896:
1889:
1886:
1885:
1882:
1881:
1878:
1877:
1874:
1873:
1871:
1870:
1865:
1860:
1852:
1844:
1836:
1832:stingless bees
1824:carpenter bees
1804:
1795:
1793:
1775:
1774:
1772:
1771:
1763:
1758:
1753:
1751:Pemphredonidae
1748:
1743:
1738:
1733:
1728:
1720:
1714:
1712:
1693:
1687:
1686:
1684:
1683:
1674:
1672:
1666:
1665:
1663:
1662:
1653:
1651:
1645:
1644:
1642:
1641:
1633:
1630:(spider wasps)
1625:
1620:
1611:
1609:
1603:
1602:
1600:
1599:
1591:
1588:(flower wasps)
1583:
1577:
1575:
1569:
1568:
1566:
1565:
1557:
1551:
1549:
1543:
1542:
1540:
1539:
1531:
1522:
1520:
1514:
1513:
1511:
1510:
1505:
1503:Sclerogibbidae
1500:
1495:
1490:
1485:
1482:(cuckoo wasps)
1477:
1471:
1469:
1460:
1440:
1439:
1436:
1435:
1433:
1432:
1426:
1424:
1418:
1417:
1415:
1414:
1408:
1406:
1400:
1399:
1397:
1396:
1390:
1388:
1382:
1381:
1379:
1378:
1376:Gasteruptiidae
1373:
1370:(ensign wasps)
1365:
1359:
1357:
1351:
1350:
1347:
1346:
1344:
1343:
1338:
1333:
1328:
1323:
1318:
1313:
1308:
1303:
1298:
1293:
1291:Pelecinellidae
1288:
1283:
1278:
1273:
1268:
1260:
1255:
1253:Metapelmatidae
1250:
1245:
1240:
1235:
1230:
1225:
1220:
1215:
1213:Hetreulophidae
1210:
1205:
1200:
1195:
1190:
1185:
1180:
1175:
1170:
1165:
1160:
1155:
1150:
1145:
1143:Chrysolampidae
1140:
1135:
1133:Chalcedectidae
1130:
1128:Cerocephalidae
1125:
1120:
1115:
1110:
1105:
1100:
1095:
1086:
1084:
1070:
1069:
1067:
1066:
1064:Mymarommatidae
1060:
1058:
1052:
1051:
1049:
1048:
1043:
1038:
1033:
1028:
1022:
1020:
1014:
1013:
1011:
1010:
1005:
1003:Proctotrupidae
1000:
995:
990:
985:
980:
974:
972:
955:
954:
952:
951:
946:
941:
936:
934:Diplolepididae
931:
923:
917:
915:
909:
908:
906:
905:
900:
895:
893:Platygastridae
890:
885:
880:
875:
873:Geoscelionidae
869:
867:
858:
852:
851:
849:
848:
843:
837:
835:
833:Ceraphronoidea
829:
828:
826:
825:
820:Ichneumonidae
817:
808:
806:
804:Ichneumonoidea
797:
768:
748:
747:
744:
743:
740:
739:
737:
736:
727:
725:
719:
718:
716:
715:
706:
704:
698:
697:
695:
694:
685:
683:
677:
676:
674:
673:
665:
656:
654:
648:
647:
645:
644:
636:
630:
628:
622:
621:
616:
614:
608:
607:
605:
604:
596:
588:
580:
572:
564:
555:
553:
540:
518:
517:
515:
514:
508:
502:
496:
490:
484:
477:
474:
473:
466:
464:
463:
456:
449:
441:
435:
434:
422:
419:
393:(3): 293–307.
375:
374:
353:
303:
302:
300:
297:
209:Heterogynaidae
204:
203:
202:
201:
194:
187:
180:
173:
166:
159:
152:
142:
141:
137:
136:
125:
121:
120:
115:
111:
110:
105:
101:
100:
95:
91:
90:
85:
81:
80:
75:
71:
70:
65:
61:
60:
55:
51:
50:
37:
36:
28:
27:
16:Genus of wasps
15:
13:
10:
9:
6:
4:
3:
2:
2164:
2153:
2152:Apoidea stubs
2150:
2148:
2145:
2144:
2142:
2131:
2126:
2124:
2119:
2117:
2112:
2111:
2105:
2103:
2099:
2095:
2090:
2087:
2083:
2078:
2065:
2060:
2056:
2052:
2047:
2043:
2039:
2034:
2030:
2026:
2021:
2017:
2013:
2008:
2004:
2000:
1995:
1991:
1987:
1982:
1978:
1974:
1969:
1965:
1961:
1955:
1950:
1944:
1940:
1935:
1929:
1925:
1924:
1922:
1920:
1916:
1912:
1907:
1895:
1891:
1890:
1887:
1869:
1868:Stenotritidae
1866:
1864:
1861:
1859:
1855:Megachilidae
1853:
1851:
1845:
1843:
1837:
1833:
1829:
1825:
1821:
1817:
1813:
1808:
1805:
1803:
1797:
1796:
1794:
1784:
1780:
1770:
1764:
1762:
1759:
1757:
1754:
1752:
1749:
1747:
1744:
1742:
1739:
1737:
1734:
1732:
1729:
1727:
1721:
1719:
1716:
1715:
1713:
1703:
1702:
1697:
1694:
1692:
1688:
1682:
1676:
1675:
1673:
1671:
1667:
1661:
1655:
1654:
1652:
1650:
1646:
1640:
1634:
1632:
1626:
1624:
1621:
1619:
1617:(velvet ants)
1613:
1612:
1610:
1608:
1604:
1598:
1592:
1590:
1584:
1582:
1579:
1578:
1576:
1574:
1570:
1564:
1558:
1556:
1553:
1552:
1550:
1548:
1544:
1538:
1532:
1530:
1524:
1523:
1521:
1519:
1515:
1509:
1508:Scolebythidae
1506:
1504:
1501:
1499:
1496:
1494:
1491:
1489:
1486:
1484:
1478:
1476:
1473:
1472:
1470:
1468:
1464:
1461:
1459:
1441:
1431:
1428:
1427:
1425:
1423:
1422:Trigonaloidea
1419:
1413:
1410:
1409:
1407:
1405:
1401:
1395:
1392:
1391:
1389:
1387:
1383:
1377:
1374:
1372:
1366:
1364:
1361:
1360:
1358:
1356:
1352:
1342:
1339:
1337:
1334:
1332:
1331:Tetracampidae
1329:
1327:
1324:
1322:
1319:
1317:
1314:
1312:
1311:Signiphoridae
1309:
1307:
1304:
1302:
1299:
1297:
1294:
1292:
1289:
1287:
1284:
1282:
1279:
1277:
1274:
1272:
1271:Neanastatidae
1269:
1267:
1261:
1259:
1256:
1254:
1251:
1249:
1246:
1244:
1243:Megastigmidae
1241:
1239:
1236:
1234:
1231:
1229:
1226:
1224:
1221:
1219:
1216:
1214:
1211:
1209:
1206:
1204:
1201:
1199:
1196:
1194:
1191:
1189:
1186:
1184:
1181:
1179:
1176:
1174:
1171:
1169:
1166:
1164:
1161:
1159:
1156:
1154:
1151:
1149:
1146:
1144:
1141:
1139:
1136:
1134:
1131:
1129:
1126:
1124:
1121:
1119:
1116:
1114:
1111:
1109:
1108:Baeomorphidae
1106:
1104:
1101:
1099:
1096:
1094:
1088:
1087:
1085:
1075:
1071:
1065:
1062:
1061:
1059:
1057:
1053:
1047:
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1039:
1037:
1034:
1032:
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1027:
1024:
1023:
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1019:
1015:
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989:
986:
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947:
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937:
935:
932:
930:
924:
922:
919:
918:
916:
914:
910:
904:
903:Sparasionidae
901:
899:
896:
894:
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878:Janzenellidae
876:
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870:
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859:
857:
853:
847:
844:
842:
841:Ceraphronidae
839:
838:
836:
834:
830:
824:
818:
816:
810:
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807:
805:
801:
798:
796:
795:
772:
769:
767:
749:
735:
729:
728:
726:
724:
720:
714:
708:
707:
705:
703:
699:
693:
689:Xiphydriidae
687:
686:
684:
682:
681:Xiphydrioidea
678:
672:
666:
664:
658:
657:
655:
653:
649:
643:
639:Pamphiliidae
637:
635:
632:
631:
629:
627:
626:Pamphilioidea
623:
619:
615:
613:
609:
603:
597:
595:
589:
587:
581:
579:
573:
571:
565:
563:
557:
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548:
544:
541:
539:
538:
519:
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364:
357:
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308:
305:
298:
296:
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290:
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278:
273:
271:
268:
264:
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257:
253:
252:brachypterous
248:
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236:
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228:
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63:
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59:
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53:
52:
47:
42:
38:
34:
29:
26:
22:
19:
2102:expanding it
2091:
2076:
1918:
1894:Unicalcarida
1849:(sweat bees)
1801:(mason bees)
1756:Philanthidae
1723:Ampulicidae
1718:Ammoplanidae
1701:Spheciformes
1699:
1480:Chrysididae
1467:Chrysidoidea
1430:Trigonalidae
1404:Megalyroidea
1386:Stephanoidea
1316:Spalangiidae
1306:Pteromalidae
1296:Perilampidae
1276:Neodiparidae
1265:(fairyflies)
1238:Macromesidae
1208:Herbertiidae
1178:Eucharitidae
1153:Coelocybidae
1074:Chalcidoidea
1046:Monomachidae
1026:Austroniidae
1008:Vanhorniidae
998:Peradeniidae
928:(gall wasps)
846:Megaspilidae
774:
691:(wood wasps)
660:Anaxyelidae
583:Diprionidae
521:
512:Holometabola
510:Superorder:
504:Infraclass:
469:Hymenopteran
430:
426:
390:
386:
382:
362:
356:
321:
317:
307:
277:sister taxon
274:
256:H. nocticola
255:
235:Turkmenistan
217:
216:
208:
207:
196:
189:
182:
175:
168:
161:
154:
147:
128:
127:
24:
18:
2020:iNaturalist
1943:Wikispecies
1847:Halictidae
1839:Colletidae
1828:orchid bees
1820:cuckoo bees
1799:Andrenidae
1741:Crabronidae
1678:Formicidae
1670:Formicoidea
1628:Pompilidae
1615:Mutillidae
1607:Pompiloidea
1581:Chyphotidae
1498:Plumariidae
1493:Embolemidae
1412:Megalyridae
1394:Stephanidae
1258:Moranilidae
1228:Leucospidae
1223:Idioporidae
1218:Heydeniidae
1198:Eurytomidae
1148:Cleonymidae
1138:Chalcididae
1113:Boucekiidae
1098:Aphelinidae
1092:(fig wasps)
1041:Maamingidae
1018:Diaprioidea
993:Pelecinidae
983:Roproniidae
949:Liopteridae
898:Scelionidae
814:(braconids)
812:Braconidae
670:(horntails)
575:Cimbicidae
289:Crabronidae
263:parasitoids
221:) of small
108:Crabronidae
98:Hymenoptera
2141:Categories
1949:Heterogyna
1919:Heterogyna
1892:See also:
1863:Melittidae
1816:bumblebees
1812:honey bees
1783:Anthophila
1766:Sphecidae
1746:Mellinidae
1736:Bembicidae
1657:Scoliidae
1649:Scolioidea
1636:Sapygidae
1623:Myrmosidae
1586:Thynnidae
1573:Thynnoidea
1560:Tiphiidae
1547:Tiphioidea
1475:Bethylidae
1368:Evaniidae
1355:Evanioidea
1321:Systasidae
1263:Mymaridae
1233:Lyciscidae
1193:Eupelmidae
1183:Eulophidae
1168:Encyrtidae
1090:Agaonidae
1031:Diapriidae
926:Cynipidae
913:Cynipoidea
888:Nixoniidae
731:Orussidae
723:Orussoidea
668:Siricidae
652:Siricoidea
498:Subclass:
488:Arthropoda
433:, 321–337.
383:Heterogyna
299:References
293:Bembicidae
245:, and the
227:Madagascar
223:spheciform
218:Heterogyna
134:Nagy, 1969
129:Heterogyna
78:Arthropoda
25:Heterogyna
1957:ButMoth:
1731:Astatidae
1534:Vespidae
1518:Vespoidea
1488:Dryinidae
1363:Aulacidae
1336:Torymidae
1301:Pirenidae
1286:Ormyridae
1281:Ooderidae
1188:Eunotidae
1163:Diparidae
1118:Calesidae
1036:Ismaridae
988:Heloridae
944:Ibaliidae
939:Figitidae
710:Cephidae
702:Cephoidea
612:Xyeloidea
591:Pergidae
500:Pterygota
480:Kingdom:
285:monophyly
267:metasomal
260:fossorial
118:Nyssonini
64:Kingdom:
58:Eukaryota
1934:Q1933488
1928:Wikidata
1761:Psenidae
1103:Azotidae
618:Xyelidae
559:Argidae
506:Neoptera
486:Phylum:
482:Animalia
471:families
415:84501607
348:28376325
231:Botswana
140:Species
104:Family:
74:Phylum:
68:Animalia
54:Domain:
2094:Apoidea
2038:1374534
2012:1345964
1960:13470.0
1691:Apoidea
494:Insecta
492:Class:
467:Extant
395:Bibcode
365:18:71.
326:Bibcode
281:Apoidea
213:species
124:Genus:
114:Tribe:
94:Order:
88:Insecta
84:Class:
2064:330849
2051:768228
2025:575116
1986:196117
1807:Apidae
1789:(bees)
1680:(ants)
1123:Ceidae
966:s.str.
413:
346:
270:tergum
241:, the
2092:This
2033:IRMNG
411:S2CID
2098:stub
2059:NCBI
2046:ITIS
2007:GBIF
1973:4WG7
344:PMID
239:Oman
1968:CoL
403:doi
367:doi
334:doi
2143::
2061::
2048::
2035::
2022::
2009::
1996::
1983::
1970::
1945::
1930::
1830:,
1826:,
1822:,
1818:,
1814:,
431:31
429:,
409:.
401:.
389:.
342:.
332:.
322:27
320:.
316:.
295:.
237:,
233:,
229:,
2129:e
2122:t
2115:v
2104:.
1834:)
1810:(
1458:a
1456:t
1454:a
1452:e
1450:l
1448:u
1446:c
1444:A
968:)
964:(
794:a
792:c
790:i
788:t
786:i
784:s
782:a
780:r
778:a
776:P
766:a
764:t
762:i
760:r
758:c
756:o
754:p
752:A
537:a
535:t
533:y
531:h
529:p
527:m
525:y
523:S
460:e
453:t
446:v
417:.
405::
397::
391:9
369::
350:.
336::
328::
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