Knowledge (XXG)

Homalocephale

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1026: 140: 393: 1719: 118: 1742: 506: 465: 1303: 1127:
Evans, D. C.; Hayashi, S.; Chiba, K.; Watabe, M.; Ryan, M. J.; Lee, Y.-N.; Currie, P. J.; Tsogtbaatar, K.; Barsbold, R. (2018). "Morphology and histology of new cranial specimens of Pachycephalosauridae (Dinosauria: Ornithischia) from the Nemegt Formation, Mongolia".
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bone. Paleontologists concluded that the specimen was an adult, despite the fact that the sutures are discernible and that it had a flat skull (a juvenile trait in many pachycephalosaurid species).
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In 2010, a study by Nick Longrich and colleagues suggested that flat-headed pachycephalosaurs were just juvenile forms of dome-headed adults, a view also supported by the earlier analysis of
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from the Nemegt Formation, noting a relatively linear growth in this pachycephalosaur characterized by a rounded dome. This differs from the flat skull of
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Maryańska, Teresa; Chapman, Ralph E.; Weishampel, David B. (2004). "Pachycephalosauria". In Weishampel, David B.; Dodson, Peter; Osmólska Halszka (eds.).
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is also noted for having an unusually broad pelvis and some have suggested that the width served to protect vital organs from harm during flank-butting.
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was about 1.8 m (5.9 ft) long. Unlike other definitely adult pachycephalosaurs (though similar to probable juvenile specimens referred to
1904: 1735: 1059: 1321: 1889: 1287: 1266: 1316: 1341: 126: 420:, and a large, round eye socket. The forehead is notably rough, with multiple nodules on the lateral and posterior sides of the 1106:, a new genus of pachycephalosaurid (Dinosauria: Ornithischia) from the upper Campanian Aguja Formation, southern Texas, USA." 865: 1919: 1894: 428: 139: 1334: 1745: 1230:(1997). "Agonistic behavior in pachycephalosaurs (Ornithischia:Dinosauria): a new look at head-butting behavior". 898: 1439: 1180:"The oldest North American pachycephalosaurid and the hidden diversity of small-bodied ornithischian dinosaurs" 614: 544: 1718: 848: 1762: 1166: 915: 736: 664: 416:. The specimen has large openings on the top of the skull, a distinct frontoparietal suture, low and long 1899: 1861: 1466: 760: 597: 580: 296: 1025: 1809: 1613: 1561: 1322:
https://web.archive.org/web/20070518001124/http://www.leute.server.de/frankmuster/H/Homalocephale.htm
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mainly based on the lack of a dome and being discovered in the same locality (Nemegt) as the latter.
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https://web.archive.org/web/20061006225059/http://www.dino-nakasato.org/en/special97/Homa-e.html
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Dinosaurs: The Most Complete, Up-to-Date Encyclopedia for Dinosaur Lovers of All Ages
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possessed a rather rounded dome, both taxa should be regarded as separate.
1779: 1390: 1381: 479: 468: 409: 355: 343: 184: 171: 83: 78: 63: 58: 48: 1827: 1204: 1179: 98: 73: 1375: 1326: 380:, juvenile specimens of the latter indicate that they were distinct. 161: 1756: 1257:(2nd ed.). Berkeley: University of California Press. pp.  408:, was described from an incomplete skull and postcranial material ( 1024: 504: 463: 391: 358:, about 70 million years ago. The genus was described in 1974 by 1814: 1760: 1368: 1330: 1083:"Pachycephalosauria, a new suborder of ornithischian dinosaurs" 444:
David C. Evans and team in 2018 reported juvenile specimens of
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and Goodwin in 2009. Longrich and colleagues suggested that
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was 1.8 m (5.9 ft) long and possibly a omnivore.
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Evans DC, Schott RK, Larson DW, Brown CM, Ryan MJ (2013).
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also had rather long legs, indicating a fast-moving gait.
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has been regarded as a synonym (and juvenile form) of
1769: 1612: 1560: 1528: 1477: 1450: 1423: 1102:Longrich, N.R., Sankey, J. and Tanke, D. (2010). " 1130:Palaeogeography, Palaeoclimatology, Palaeoecology 412:MPC-D 100/1201) from the Nemegt locality of the 435:is actually the juvenile or sub-adult stage of 1342: 8: 1757: 1566: 1534: 1525: 1456: 1429: 1420: 1365: 1349: 1335: 1327: 1076: 1074: 116: 20: 1203: 1070: 1060:Timeline of pachycephalosaur research 7: 1081:Maryańska, T.; Osmólska, H. (1974). 366:, and consists of a single species, 890: 840: 833: 783: 776: 752: 728: 704: 680: 656: 572: 565: 536: 524: 14: 1741: 1740: 1717: 1301: 396:Skeletal diagram of the holotype 138: 127:Mongolian Natural History Museum 1280:Fantastic Facts About Dinosaurs 866:Pachycephalosaurus wyomingensis 1915:Taxa named by Halszka Osmólska 1910:Taxa named by Teresa Maryańska 315:Maryanska & Osmolska, 1974 1: 1905:Fossil taxa described in 1974 1116:10.1016/j.cretres.2009.12.002 493:sported a flat, wedge-shaped 452:and given that even juvenile 1162:Holtz, Thomas R. Jr. (2008) 1150:10.1016/j.palaeo.2017.11.029 689:Homalocephale calathocercos 309:Homalocephale calathocercos 1936: 509:Cast of the holotype skull 350:period of what is now the 16:Extinct genus of dinosaurs 1890:Pachycephalosaurs of Asia 1730: 1715: 1569: 1537: 1524: 1505: 1459: 1432: 1419: 1364: 1167:Supplementary Information 912: 899:Sphaerotholus buchholtzae 895: 888: 862: 845: 838: 831: 805: 788: 781: 774: 757: 750: 733: 726: 709: 702: 685: 678: 661: 654: 628: 611: 594: 577: 570: 563: 541: 534: 302: 295: 135:Scientific classification 133: 124: 115: 23: 1232:Contributions to Geology 1090:Palaeontologica Polonica 615:Stegoceras novomexicanum 545:Wannanosaurus yansiensis 339:, "head") is a genus of 125:Mounted holotype in the 849:Alaskacephale gangloffi 418:infratemporal fenestrae 1040: 916:Sphaerotholus goodwini 737:Foraminacephale brevis 665:Goyocephale lattimorei 510: 471: 397: 346:that lived during the 335:, "even", and κεφαλή, 1862:Paleobiology Database 1184:Nature Communications 1104:Texacephale langstoni 1028: 761:Amtocephale gobiensis 598:Hanssuesia sternbergi 581:Colepiocephale lambei 508: 467: 395: 288:H. calathocercos 1920:Ornithischian genera 1895:Maastrichtian genera 1614:Pachycephalosaurinae 1562:Pachycephalosauridae 713:Tylocephale gilmorei 559:Pachycephalosauridae 246:Pachycephalosauridae 1196:2013NatCo...4.1828E 1142:2018PPP...494..121E 1108:Cretaceous Research 809:Prenocephale prenes 519:Evans et al. (2013) 1665:Pachycephalosaurus 1530:Pachycephalosauria 1509:Pachycephalosauria 1479:Pachycephalosauria 1358:Pachycephalosauria 1228:Carpenter, Kenneth 1205:10.1038/ncomms2749 1041: 632:Stegoceras validum 530:Pachycephalosauria 511: 472: 398: 341:pachycephalosaurid 233:Pachycephalosauria 1877: 1876: 1849:Open Tree of Life 1763:Taxon identifiers 1754: 1753: 1726: 1725: 1713: 1712: 1709: 1708: 1520: 1519: 1516: 1515: 1501: 1500: 1497: 1496: 1017: 1016: 1008: 1007: 999: 998: 990: 989: 981: 980: 972: 971: 963: 962: 954: 953: 945: 944: 936: 935: 927: 926: 877: 876: 820: 819: 792:Acrotholus audeti 643: 642: 320: 319: 276: 1927: 1870: 1869: 1857: 1856: 1844: 1843: 1831: 1830: 1818: 1817: 1805: 1804: 1803: 1790: 1789: 1788: 1758: 1744: 1743: 1721: 1567: 1535: 1526: 1457: 1430: 1421: 1414: 1413: 1366: 1351: 1344: 1337: 1328: 1311: 1309:Dinosaurs portal 1306: 1305: 1304: 1273: 1272: 1256: 1246: 1240: 1239: 1224: 1218: 1217: 1207: 1175: 1169: 1160: 1154: 1153: 1124: 1118: 1100: 1094: 1093: 1087: 1078: 891: 841: 834: 784: 777: 753: 729: 705: 681: 657: 573: 566: 537: 525: 469:Life restoration 414:Nemegt Formation 406:H. calathocercos 368:H. calathocercos 364:Teresa Maryańska 360:Halszka Osmólska 352:Nemegt Formation 311: 307: 264: 257: 244: 231: 215: 199: 143: 142: 120: 110: 40: 29:Temporal range: 21: 1935: 1934: 1930: 1929: 1928: 1926: 1925: 1924: 1880: 1879: 1878: 1873: 1865: 1860: 1852: 1847: 1839: 1834: 1826: 1821: 1813: 1808: 1799: 1798: 1793: 1784: 1783: 1778: 1765: 1755: 1750: 1722: 1705: 1644:Foraminacephale 1608: 1556: 1512: 1493: 1473: 1446: 1440:Avemetatarsalia 1425:Avemetatarsalia 1415: 1409:Marginocephalia 1371: 1370: 1360: 1355: 1307: 1302: 1300: 1297: 1276: 1269: 1248: 1247: 1243: 1226: 1225: 1221: 1177: 1176: 1172: 1161: 1157: 1126: 1125: 1121: 1101: 1097: 1085: 1080: 1079: 1072: 1068: 1056: 1023: 1018: 1009: 1000: 991: 982: 973: 964: 955: 946: 937: 928: 878: 821: 644: 503: 462: 390: 348:Late Cretaceous 316: 313: 305: 304: 291: 263: 255: 242: 229: 213: 197: 137: 111: 109: 108: 107: 106: 101: 96: 91: 86: 81: 76: 71: 66: 61: 56: 51: 46: 35: 34: 31:Late Cretaceous 27: 17: 12: 11: 5: 1933: 1931: 1923: 1922: 1917: 1912: 1907: 1902: 1897: 1892: 1882: 1881: 1875: 1874: 1872: 1871: 1858: 1845: 1832: 1819: 1806: 1791: 1775: 1773: 1767: 1766: 1761: 1752: 1751: 1749: 1748: 1738: 1731: 1728: 1727: 1724: 1723: 1716: 1714: 1711: 1710: 1707: 1706: 1704: 1703: 1696: 1689: 1682: 1675: 1668: 1661: 1654: 1647: 1640: 1633: 1626: 1618: 1616: 1610: 1609: 1607: 1606: 1599: 1592: 1585: 1578: 1575:Colepiocephale 1570: 1564: 1558: 1557: 1555: 1554: 1547: 1543:Ferganocephale 1538: 1532: 1522: 1521: 1518: 1517: 1514: 1513: 1506: 1503: 1502: 1499: 1498: 1495: 1494: 1492: 1491: 1490: 1489: 1483: 1481: 1475: 1474: 1472: 1471: 1470: 1469: 1460: 1454: 1448: 1447: 1445: 1444: 1443: 1442: 1433: 1427: 1417: 1416: 1412: 1411: 1402: 1393: 1384: 1378: 1369: 1362: 1361: 1356: 1354: 1353: 1346: 1339: 1331: 1325: 1324: 1319: 1313: 1312: 1296: 1295:External links 1293: 1292: 1291: 1275: 1274: 1267: 1253:The Dinosauria 1241: 1219: 1170: 1155: 1119: 1095: 1069: 1067: 1064: 1063: 1062: 1055: 1052: 1022: 1019: 1015: 1014: 1011: 1010: 1006: 1005: 1002: 1001: 997: 996: 993: 992: 988: 987: 984: 983: 979: 978: 975: 974: 970: 969: 966: 965: 961: 960: 957: 956: 952: 951: 948: 947: 943: 942: 939: 938: 934: 933: 930: 929: 925: 924: 921: 920: 911: 908: 907: 904: 903: 894: 889: 887: 884: 883: 880: 879: 875: 874: 871: 870: 861: 858: 857: 854: 853: 844: 839: 837: 832: 830: 827: 826: 823: 822: 818: 817: 814: 813: 804: 801: 800: 797: 796: 787: 782: 780: 775: 773: 770: 769: 766: 765: 756: 751: 749: 746: 745: 742: 741: 732: 727: 725: 722: 721: 718: 717: 708: 703: 701: 698: 697: 694: 693: 684: 679: 677: 674: 673: 670: 669: 660: 655: 653: 650: 649: 646: 645: 641: 640: 637: 636: 627: 624: 623: 620: 619: 610: 607: 606: 603: 602: 593: 590: 589: 586: 585: 576: 571: 569: 564: 562: 554: 553: 550: 549: 540: 535: 533: 523: 502: 501:Classification 499: 461: 458: 389: 386: 318: 317: 314: 300: 299: 293: 292: 284: 282: 278: 277: 253: 249: 248: 240: 236: 235: 227: 220: 219: 217:Neornithischia 211: 204: 203: 195: 188: 187: 182: 175: 174: 169: 165: 164: 159: 155: 154: 149: 145: 144: 131: 130: 122: 121: 113: 112: 104: 103: 102: 97: 92: 87: 82: 77: 72: 67: 62: 57: 52: 47: 42: 41: 28: 15: 13: 10: 9: 6: 4: 3: 2: 1932: 1921: 1918: 1916: 1913: 1911: 1908: 1906: 1903: 1901: 1898: 1896: 1893: 1891: 1888: 1887: 1885: 1868: 1863: 1859: 1855: 1850: 1846: 1842: 1837: 1833: 1829: 1824: 1820: 1816: 1811: 1807: 1802: 1801:Homalocephale 1796: 1792: 1787: 1781: 1777: 1776: 1774: 1772: 1771:Homalocephale 1768: 1764: 1759: 1747: 1739: 1737: 1733: 1732: 1729: 1720: 1702: 1701: 1697: 1695: 1694: 1693:Sphaerotholus 1690: 1688: 1687: 1683: 1681: 1680: 1676: 1674: 1673: 1669: 1667: 1666: 1662: 1660: 1659: 1658:Homalocephale 1655: 1653: 1652: 1648: 1646: 1645: 1641: 1639: 1638: 1634: 1632: 1631: 1630:Alaskacephale 1627: 1625: 1624: 1620: 1619: 1617: 1615: 1611: 1605: 1604: 1600: 1598: 1597: 1593: 1591: 1590: 1586: 1584: 1583: 1579: 1577: 1576: 1572: 1571: 1568: 1565: 1563: 1559: 1553: 1552: 1551:Wannanosaurus 1548: 1545: 1544: 1540: 1539: 1536: 1533: 1531: 1527: 1523: 1511: 1510: 1504: 1487: 1486: 1485: 1484: 1482: 1480: 1476: 1468: 1464: 1463: 1462: 1461: 1458: 1455: 1453: 1449: 1441: 1437: 1436: 1435: 1434: 1431: 1428: 1426: 1422: 1418: 1410: 1406: 1403: 1401: 1397: 1394: 1392: 1388: 1385: 1383: 1379: 1377: 1373: 1372: 1367: 1363: 1359: 1352: 1347: 1345: 1340: 1338: 1333: 1332: 1329: 1323: 1320: 1318: 1315: 1314: 1310: 1299: 1294: 1289: 1288:0-7525-3166-2 1285: 1281: 1278: 1277: 1270: 1268:0-520-24209-2 1264: 1260: 1255: 1254: 1245: 1242: 1237: 1233: 1229: 1223: 1220: 1215: 1211: 1206: 1201: 1197: 1193: 1189: 1185: 1181: 1174: 1171: 1168: 1165: 1159: 1156: 1151: 1147: 1143: 1139: 1135: 1131: 1123: 1120: 1117: 1113: 1109: 1105: 1099: 1096: 1092:(30): 45−102. 1091: 1084: 1077: 1075: 1071: 1065: 1061: 1058: 1057: 1053: 1051: 1049: 1048:Homalocephale 1045: 1044:Homalocephale 1038: 1037: 1032: 1031:Homalocephale 1027: 1020: 1013: 1012: 1004: 1003: 995: 994: 986: 985: 977: 976: 968: 967: 959: 958: 950: 949: 941: 940: 932: 931: 923: 922: 919: 918: 917: 910: 909: 906: 905: 902: 901: 900: 893: 892: 886: 885: 882: 881: 873: 872: 869: 868: 867: 860: 859: 856: 855: 852: 851: 850: 843: 842: 836: 835: 829: 828: 825: 824: 816: 815: 812: 811: 810: 803: 802: 799: 798: 795: 794: 793: 786: 785: 779: 778: 772: 771: 768: 767: 764: 763: 762: 755: 754: 748: 747: 744: 743: 740: 739: 738: 731: 730: 724: 723: 720: 719: 716: 715: 714: 707: 706: 700: 699: 696: 695: 692: 691: 690: 683: 682: 676: 675: 672: 671: 668: 667: 666: 659: 658: 652: 651: 648: 647: 639: 638: 635: 634: 633: 626: 625: 622: 621: 618: 617: 616: 609: 608: 605: 604: 601: 600: 599: 592: 591: 588: 587: 584: 583: 582: 575: 574: 568: 567: 560: 556: 555: 552: 551: 548: 547: 546: 539: 538: 531: 527: 526: 522: 520: 516: 507: 500: 498: 496: 492: 491:Homalocephale 488: 487: 482: 481: 476: 475:Homalocephale 470: 466: 459: 457: 455: 451: 450:Homalocephale 447: 442: 440: 439: 434: 433:Homalocephale 430: 425: 423: 419: 415: 411: 407: 403: 394: 387: 385: 383: 382:Homalocephale 379: 378: 373: 372:Homalocephale 369: 365: 361: 357: 353: 349: 345: 342: 338: 334: 330: 326: 325: 324:Homalocephale 312: 310: 301: 298: 297:Binomial name 294: 290: 289: 283: 280: 279: 275: 271: 267: 262: 261: 260:Homalocephale 254: 251: 250: 247: 241: 238: 237: 234: 228: 225: 222: 221: 218: 212: 209: 206: 205: 202: 196: 193: 190: 189: 186: 183: 180: 177: 176: 173: 170: 167: 166: 163: 160: 157: 156: 153: 150: 147: 146: 141: 136: 132: 128: 123: 119: 114: 100: 95: 90: 85: 80: 75: 70: 65: 60: 55: 50: 45: 39: 32: 26: 25:Homalocephale 22: 19: 1900:Nemegt fauna 1770: 1698: 1691: 1684: 1679:Prenocephale 1677: 1670: 1663: 1657: 1656: 1649: 1642: 1635: 1628: 1621: 1601: 1594: 1587: 1580: 1573: 1549: 1541: 1507: 1467:Ornithischia 1452:Ornithischia 1404: 1400:Ornithischia 1395: 1386: 1279: 1252: 1244: 1235: 1231: 1222: 1187: 1183: 1173: 1163: 1158: 1133: 1129: 1122: 1107: 1103: 1098: 1089: 1047: 1043: 1042: 1034: 1030: 1021:Paleobiology 914: 913: 897: 896: 864: 863: 847: 846: 807: 806: 790: 789: 759: 758: 735: 734: 711: 710: 688: 687: 686: 663: 662: 630: 629: 613: 612: 596: 595: 579: 578: 543: 542: 512: 490: 484: 478: 474: 473: 454:Prenocephale 453: 449: 446:Prenocephale 445: 443: 438:Prenocephale 436: 432: 426: 405: 402:type species 399: 381: 377:Prenocephale 375: 371: 367: 336: 332: 323: 322: 321: 308: 303: 287: 286: 259: 258: 223: 207: 201:Ornithischia 191: 178: 24: 18: 1795:Wikispecies 1700:Tylocephale 1686:Sinocephale 1672:Platytholus 1651:Goyocephale 1637:Amtocephale 1603:Texacephale 1582:Gravitholus 1238:(1): 19–25. 1136:: 121−134. 1039:herd behind 1036:Saurolophus 513:Below is a 486:Goyocephale 460:Description 1884:Categories 1734:See also: 1623:Acrotholus 1596:Stegoceras 1589:Hanssuesia 1488:see below↓ 1391:Dinosauria 1066:References 495:skull roof 185:Dinosauria 1374:Kingdom: 515:cladogram 422:squamosal 388:Discovery 370:. Though 281:Species: 266:Maryanska 158:Kingdom: 152:Eukaryota 1780:Wikidata 1746:Category 1736:Timeline 1382:Chordata 1380:Phylum: 1376:Animalia 1214:23652016 1190:. 1828. 1054:See also 480:Dracorex 410:holotype 356:Mongolia 344:dinosaur 331:ὁμαλός, 270:Osmolska 239:Family: 172:Chordata 168:Phylum: 162:Animalia 148:Domain: 36:70  1854:4127650 1841:1048056 1828:4823180 1815:4530957 1786:Q134134 1192:Bibcode 1138:Bibcode 561:  532:  337:kephalē 333:homalos 306:† 285:† 252:Genus: 129:, 2002 105:↓ 1286:  1265:  1261:–477. 1212:  557:  528:  429:Horner 327:(from 268:& 1867:38795 1836:IRMNG 1405:Clade 1396:Clade 1387:Clade 1086:(PDF) 1033:with 517:from 329:Greek 224:Clade 208:Clade 192:Clade 179:Clade 1823:GBIF 1465:see 1438:see 1284:ISBN 1263:ISBN 1210:PMID 1110:, . 1029:Two 483:and 400:The 362:and 274:1974 44:PreꞒ 1810:EoL 1259:464 1200:doi 1146:doi 1134:494 1112:doi 489:), 1886:: 1864:: 1851:: 1838:: 1825:: 1812:: 1797:: 1782:: 1407:: 1398:: 1389:: 1236:32 1234:. 1208:. 1198:. 1186:. 1182:. 1144:. 1132:. 1088:. 1073:^ 521:. 404:, 354:, 272:, 226:: 210:: 194:: 181:: 94:Pg 38:Ma 33:, 1546:? 1350:e 1343:t 1336:v 1290:) 1282:( 1271:. 1216:. 1202:: 1194:: 1188:4 1152:. 1148:: 1140:: 1114:: 256:† 243:† 230:† 214:† 198:† 99:N 89:K 84:J 79:T 74:P 69:C 64:D 59:S 54:O 49:Ꞓ

Index

Late Cretaceous
Ma
PreꞒ

O
S
D
C
P
T
J
K
Pg
N

Mongolian Natural History Museum
Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Dinosauria
Ornithischia
Neornithischia
Pachycephalosauria
Pachycephalosauridae
Homalocephale
Maryanska
Osmolska
1974

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