35:
272:
In 2020, Guadanucci summarized the status of the subfamily at that time. Most of the members that had been studied were scattered across the family tree of the
Theraphosidae. Guadanucci considered that no further taxonomic changes should be considered until there had been a comprehensive sampling of
764:
Foley, Saoirse; Lüddecke, Tim; Cheng, Dong-Qiang; Krehenwinkel, Henrik; Künzel, Sven; Longhorn, Stuart J.; Wendt, Ingo; von Wirth, Volker; Tänzler, Rene; Vences, Miguel & Piel, William H. (2019). "Tarantula phylogenomics: A robust phylogeny of deep theraphosid clades inferred from transcriptome
235:
said that it could be "nothing but a sign of immaturity". Young spiders have only fine hairs (setae) rather than scopulae. In successive moults, scopulae replace the setae in stages, so divisions at first present may disappear when the scopulae are complete at full maturity. In spite of these
269:, and another group were placed in a new subfamily, Schismatothelinae. The remaining genera were distributed across the theraphosid phylogeny. This result has been confirmed by other studies, one of which also suggested that the newly defined subfamily Schismatothelinae was not monophyletic.
194:
was proposed in 2014. One of the authors of that proposal subsequently said that no further taxonomic changes should be considered until there had been a more comprehensive sampling of the subfamily. As of
January 2021, the status of the Ischnocolinae remains unresolved.
231:" (apart from this character .. the Ischnocoleae have hardly any common characteristics and they form a weakly homogeneous set). Doubts were expressed early on as to the validity of the divided tarsal scopulae as a defining characteristic. In 1895,
236:
concerns, genera were continually added to the subfamily, so that in 1986, for example, Andrew M. Smith said the subfamily included 49 genera, although he regarded it as "ripe for sweeping revision". In his 1985 monograph on the
957:
Mirza, Z.A.; Sanap, R.V. & Bhosale, H. (2014). "Preliminary review of Indian
Eumenophorinae (Araneae: Theraphosidae) with description of a new genus and five new species from the Western Ghats".
263:
study was published that confirmed the non-monophyly of
Ischnocolinae when broadly defined. A small number of genera could be retained in much reduced subfamily, Ischnocolinae
225:
in 1892. Following
Ausserer's characterization of the genus, Simon used the divided tarsal scopulae as the defining characteristic of the group. However, Simon noted that "
833:
667:
366:
Species in several genera were placed in the newly created
Schismatothelinae. Subsequently some species were transferred to other genera.
510:
Also in 2014, Mirza, Sanap & Bhosale further revised some Indian taxa associated with the
Ischnocolinae, described the new genus
221:
of the third and fourth legs as a key characteristic. The subfamily
Ischnocolinae was first described (as the group Ischnocoleae) by
256:. His key to the subfamilies of the Theraphosidae split genera that had been placed in the subfamily among three informal groups.
245:
173:
729:
Guadanucci, J.P.L. (2014). "Theraphosidae phylogeny: relationships of the 'Ischnocolinae' genera (Araneae, Mygalomorphae)".
34:
451:
Genera suggested to be excluded from the subfamily
Ischnocolinae as they were unresolved across the phylogeny include:
1023:
228:à part ce caractère .. les Ischnocoleae n'ont guère de particularités communes et ils forment un ensemble peu homogène
181:
567:
Ausserer, A. (1871). "Beiträge zur
Kenntniss der Arachniden-Familie der Territelariae Thorell (Mygalidae Autor)".
218:
214:
169:
166:
274:
260:
177:
1018:
232:
176:(i.e. potentially inherited from an ancestor by multiple descendant groups), and both morphological and
820:. Zoological Monographs, vol 6. Vol. 6. Cham: Springer International Publishing. pp. 77–91.
966:
913:
774:
622:"Notes on the identity of some of the types of Mygalomorphae in the collection of the British Museum"
939:
839:
798:
746:
29:
994:
882:
829:
790:
663:
532:
495:
984:
974:
929:
921:
874:
821:
816:
Guadanucci, J.P.L. (2020). "Ischnocolinae and Schismatothelinae". In PĂ©rez-Miles, F. (ed.).
782:
738:
697:
633:
600:
222:
162:
131:
970:
917:
778:
165:
based the group, which he noted was only weakly homogeneous, on the presence of divided
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489:
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Verhandlungen der Kaiserlich-Königlichen Zoologisch-Botanischen Gesellschaft in Wien
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249:
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116:
66:
46:
998:
886:
794:
686:"The spider infraorder Mygalomorphae (Araneae): cladistics and systematics"
456:
387:
285:
According to Guadanucci in 2014, a much reduced subfamily, Ischnocolinae
86:
934:
439:
765:
data sheds light on the prickly issue of urticating setae evolution".
742:
96:
56:
701:
265:
190:
857:
Guadanucci, J.P.L. & Weinmann, D. (2014). "The spider genera
900:
Guadanucci, J.P.L. & Weinmann, D. (2015). "Description of
322:
of those included; the genus was recovered as non-monophyletic
660:
The Tarantula : Classification and Identification Guide
599:. Vol. 1 (2nd ed.). Paris: Roret. p. 133ff.
289:, was monophyletic. It included species from five genera:
904:
gen. nov. (Araneae, Theraphosidae, Schismatothelinae)".
180:
phylogenetic studies have shown that, as traditionally
580:
Scopula an Tarsus III und IV sehr deutlich getheilt
690:Bulletin of the American Natural History Museum
244:considered the divided tarsal scopulae to be a
188:. A much more narrowly defined Ischnocolinae
8:
906:Studies on Neotropical Fauna and Environment
578:
226:
248:character and the Ischnocolinae to be non-
172:. This feature was later considered to be
20:
988:
978:
933:
604:
865:Karsch (Mygalomorphae, Theraphosidae)".
724:
722:
720:
718:
559:
415:, plus an unidentified related species)
383:, plus an unidentified related species)
626:Annals and Magazine of Natural History
767:Molecular Phylogenetics and Evolution
593:Simon, E. (1892). "1. Ischnocoleae".
7:
516:, and treated them in the subfamily
213:. Ausserer used the division of the
14:
662:. London: Fitzgerald Publishing.
161:(family Theraphosidae). In 1892,
596:Histoire naturelle des araignées
33:
474:(included under junior synonym
252:, treating it as Theraphosidae
157:are a problematic subfamily of
1:
926:10.1080/01650521.2015.1110309
980:10.1371/journal.pone.0087928
826:10.1007/978-3-030-48644-0_3
787:10.1016/j.ympev.2019.106573
1040:
879:10.11646/zootaxa.3795.3.3
684:Raven, Robert J. (1985).
658:Smith, Andrew M. (1986).
638:10.1080/00222939508680262
493:(maybe closely allied to
144:
139:
30:Scientific classification
28:
23:
431:, originally treated as
411:, originally treated as
379:, originally treated as
606:10.5962/bhl.title.51973
184:, the subfamily is not
16:Subfamily of tarantulas
579:
275:molecular phylogenetic
261:molecular phylogenetic
227:
620:Pocock, R.I. (1895).
818:New World Tarantulas
971:2014PLoSO...987928M
918:2015SNFE...50..221G
779:2019MolPE.14006573F
273:the subfamily in a
1024:Spider subfamilies
499:but not indicated)
445:S. longibulbi
433:Hemiercus modestus
381:Holothele colonica
336:I. valentinus
207:erected the genus
835:978-3-030-48643-3
743:10.1111/zsc.12065
731:Zoologica Scripta
669:978-0-9510939-0-0
533:Neoheterophrictus
496:Neoheterophrictus
397:G. atlantica
332:I. ignoratus
300:A. rufescens
151:
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320:H. culebrae
312:H. longipes
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632:(93): 223–230.
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561:
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429:S. modesta
425:S. lineata
413:Holothele incei
393:G. cerrado
360:T. nitidum
316:H. rondoni
283:
201:
129:
32:
17:
12:
11:
5:
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1035:
1027:
1026:
1021:
1011:
1010:
1005:
1004:
949:
912:(3): 221–228.
892:
873:(3): 275–288.
863:Schismatothele
849:
834:
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737:(5): 508–518.
714:
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552:
551:
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547:Sahydroaraneus
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540:Plesiophrictus
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529:
526:Heterophrictus
518:Eumenophorinae
513:Sahydroaraneus
508:
507:
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490:Plesiophrictus
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254:incertae sedis
205:Anton Ausserer
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965:(2): e87928.
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314:(included as
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287:sensu stricto
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266:sensu stricto
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246:plesiomorphic
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238:Mygalomorphae
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191:sensu stricto
187:
183:
182:circumscribed
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174:plesiomorphic
171:
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155:Ischnocolinae
147:
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133:
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127:Ischnocolinae
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117:Theraphosidae
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107:Mygalomorphae
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27:
22:
19:
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952:
935:11449/161205
909:
905:
902:Neoholothele
901:
895:
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811:
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766:
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734:
730:
705:. Retrieved
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641:. Retrieved
629:
628:. Series 6.
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471:Dolichothele
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311:
305:
299:
295:Acanthopelma
293:
286:
284:
271:
264:
258:
253:
250:monophyletic
242:Robert Raven
223:Eugène Simon
208:
202:
189:
186:monophyletic
163:Eugène Simon
154:
152:
145:
126:
103:Infraorder:
18:
859:Euthycaelus
483:Heterothele
476:Oligoxystre
464:Chaetopelma
372:Euthycaelus
355:Trichopelma
343:Reichlingia
327:Ischnocolus
259:In 2014, a
233:R.I. Pocock
210:Ischnocolus
123:Subfamily:
77:Chelicerata
73:Subphylum:
1013:Categories
861:Simon and
773:: 106573.
707:2016-01-18
643:2021-01-28
583:", p. 127.
575:: 117–224.
554:References
159:tarantulas
146:See text.
67:Arthropoda
844:235056744
803:199389268
712:p. 116ff.
504:Nesiergus
307:Holothele
203:In 1872,
178:molecular
87:Arachnida
53:Kingdom:
47:Eukaryota
999:24551072
959:PLOS ONE
944:86207520
887:24870477
795:31374259
751:86484123
702:2246/955
457:Catumiri
388:Guyruita
215:scopulae
199:Taxonomy
170:scopulae
113:Family:
63:Phylum:
57:Animalia
43:Domain:
990:3925112
967:Bibcode
914:Bibcode
867:Zootaxa
775:Bibcode
648:p. 225.
440:Sickius
310:– only
277:study.
217:on the
140:Genera
97:Araneae
93:Order:
83:Class:
997:
987:
942:
885:
842:
832:
801:
793:
749:
674:p. 70.
666:
318:) and
281:Genera
167:tarsal
134:, 1892
940:S2CID
840:S2CID
799:S2CID
747:S2CID
219:tarsi
132:Simon
995:PMID
883:PMID
871:3795
830:ISBN
791:PMID
664:ISBN
153:The
985:PMC
975:doi
930:hdl
922:doi
875:doi
822:doi
783:doi
771:140
739:doi
698:hdl
694:182
634:doi
601:doi
1015::
993:.
983:.
973:.
961:.
938:.
928:.
920:.
910:50
908:.
881:.
869:.
838:.
828:.
797:.
789:.
781:.
769:.
745:.
735:43
733:.
717:^
696:.
692:.
688:.
630:16
624:.
573:21
571:.
520:.
427:,
395:,
334:,
240:,
1001:.
977::
969::
963:9
946:.
932::
924::
916::
889:.
877::
846:.
824::
805:.
785::
777::
753:.
741::
710:.
700::
672:.
646:.
636::
609:.
603::
577:"
478:)
447:)
443:(
435:)
423:(
407:(
399:)
391:(
375:(
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358:(
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346:(
338:)
330:(
302:)
298:(
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
