1323:. Instead the specialized features of the frill- the large winglike hornlets on the margins of the frill, and the reduced ones on the back- show that it represents a side branch in chasmosaurine evolution. No fossils from this lineage are known from later rocks, consistent with a pattern of high turnover- species rapidly evolved, and became extinct- as seen in other horned dinosaurs in the latest Cretaceous. Its precise relationships to other members of the group are unclear, but given its relatively early appearance and the lack of features clearly linking it to later members of the Chasmosaurinae, it seems to represent a deep divergence within Chasmosaurinae.
2261:
2267:
118:
1454:
520:
2817:
136:
490:
2847:
1332:
2823:
447:
476:
A referred specimen, YPM VPPU 023262, overlaps the holotype in two areas, including the distal blade of the squamosal, and the postorbital horns. It resembles the holotype in several features, including the shape of the marginal hornlets and the concave lateral margin of the distal blade. Critically,
1281:
Despite the fragmentary nature of the fossils, they can be identified as a distinct species. Frills of horned dinosaurs are highly distinctive, with different species having distinct arrangements of the epoccipital hornlets. From species to species, hornlets differed in their number, arrangement,
473:(located above the eyes) horns are moderately elongate, inclined forward and outwards, and have an unusual teardrop-shaped cross section.Other fragmentary specimens are known from the same area, which preserve distinctive features of the frill.
1297:
is the oldest known member of the
Chasmosaurinae, but already shows many of the distinctive features of the group including the long, concave squamosals and pointed epioccipitals. Along with the presence of specialized centrosaurines such as
1308:, appears) and the middle Campanian, when Chasmosaurinae and Centrosaurinae show up in the fossil record. This period spans about 15 million years- a relatively short period in terms of dinosaur evolution, suggesting rapid evolution.
477:
it shares the same unusual horn shape seen in the holotype, supporting referral of the two fossils to the same species, and it helps to provide additional information on the anatomy of
2978:
1529:
Longrich, N. R. (2014). "The horned dinosaurs
Pentaceratops and Kosmoceratops from the upper Campanian of Alberta and implications for dinosaur biogeography".
1282:
size, and shape, and the bones of the frill, the squamosal and parietal, differ as well. This probably reflects rapid evolution of the frill in response to
527:, holotype. A, dorsal view, B, ventral view, C, posterior view. The epoccipital hornlets are very low and fused to the back of the frill, forming low keels
2927:
2998:
2973:
1304:
at the same time, it suggests rapid evolution of the horned dinosaurs between the
Turonian (when the oldest horned dinosaur in North America,
3003:
2260:
481:. It shows that the animal had large scallops on the frill's proximal margins, and a very broad, platelike bar in the center of the frill.
1582:
2266:
461:
YPM VPPU 022404 consists of an incomplete skull including the horns, parts of the frills, and fragments from the back of the frill.
382:
2840:
1467:
1286:, similar to how sexual selection creates a large variety of display feathers in living dinosaurs such as birds of paradise.
2993:
2816:
135:
2983:
2988:
806:
1315:
is currently the oldest known chasmosaurine, it does not seem to be ancestral to later horned dinosaurs such as
1673:
1575:
2905:
2968:
1791:
2867:
1348:
395:
274:
2940:
2850:
1700:
301:
117:
2822:
2914:
1538:
278:
1956:
1568:
1459:
1368:
1355:. The specimen was collected in brown/green mudstone and gray/yellow sandstone which dates to the
519:
2676:
2626:
2058:
1511:
1363:
period, approximately 78 million years ago. This specimen is housed at the Yale
Peabody Museum.
130:
2945:
2759:
2697:
2660:
2134:
2021:
1389:
409:
2789:
2587:
2412:
2114:
1809:
1546:
1503:
1283:
1001:
493:
416:
335:
2683:
2635:
2594:
2530:
2433:
2384:
2086:
2010:
1965:
1839:
1825:
1784:
1658:
1642:
1426:
1360:
1300:
1034:
569:
540:
511:
on the lateral margins of the frill are large near the front, but small towards the back.
30:
1542:
2690:
2649:
2580:
2567:
2538:
2523:
2493:
2454:
2440:
2357:
2318:
2197:
2107:
2072:
2065:
1945:
1895:
1818:
1798:
1432:
1396:
1061:
1047:
555:
254:
212:
1494:, a New Horned Dinosaur from the Middle Campanian Judith River Formation of Montana".
489:
2962:
2768:
2747:
2717:
2708:
2608:
2502:
2447:
2426:
2398:
2391:
2377:
2370:
2176:
2165:
2158:
2100:
2093:
2079:
2037:
1972:
1938:
1909:
1862:
1515:
1405:
1380:
984:
931:
862:
740:
535:
500:
435:
68:
2782:
2733:
2726:
2642:
2615:
2601:
2509:
2482:
2475:
2468:
2405:
2297:
2284:
2215:
2204:
2189:
2151:
2143:
2125:
2030:
1996:
1979:
1931:
1924:
1880:
1685:
1633:
1422:
1376:
1305:
823:
707:
621:
538:. It is more primitive than most genera within the subfamily with the exception of
241:
196:
1550:
2890:
2899:
2775:
2669:
2552:
2516:
2339:
2051:
1916:
1438:
1411:
1099:
470:
428:
43:
2545:
2461:
2311:
2044:
1832:
1766:
1753:
1712:
1591:
1449:
1372:
1085:
466:
413:
402:
228:
88:
53:
1385:
1356:
508:
504:
431:
147:
93:
37:
1331:
544:. The below cladogram follows Longrich (2015), who named a new species of
2884:
2346:
2332:
2325:
1888:
1624:
1615:
1507:
1401:
1340:
458:
405:
180:
167:
83:
78:
63:
58:
48:
507:(bony projections) on the rear edge of the frill, the epiparietals. The
2932:
2740:
1902:
1846:
1417:
1352:
420:
399:
98:
73:
446:
1609:
157:
2861:
1330:
518:
488:
445:
2919:
465:
shows a distinctive combination of characters, not seen in other
503:) has a broad midline bar, a rounded caudal margin, and reduced
2865:
2282:
1751:
1602:
1564:
359:
350:
1560:
344:
362:
368:
341:
383:
374:
356:
338:
548:, and included nearly all species of chasmosaurine.
371:
365:
347:
2874:
2758:
2716:
2707:
2659:
2625:
2566:
2492:
2356:
2295:
2214:
2175:
2133:
2124:
2020:
1994:
1955:
1860:
1808:
1764:
1711:
1684:
1657:
353:
1496:Bulletin of the Peabody Museum of Natural History
450:Squamosals of the type and referred specimens of
423:, United States. Like other horned dinosaurs,
1485:
1483:
1576:
8:
2979:Late Cretaceous dinosaurs of North America
2862:
2713:
2572:
2362:
2303:
2292:
2279:
2181:
2130:
2002:
1872:
1868:
1776:
1772:
1761:
1748:
1690:
1663:
1654:
1599:
1583:
1569:
1561:
523:Posterior of the frill (parietal bone) of
116:
20:
1347:YPM VPPU 022404 were recovered from the
1479:
7:
1128:
1106:
1077:
1053:
1026:
976:
969:
923:
916:
869:
857:
798:
763:
756:
732:
651:
628:
616:
609:
585:
561:
551:
14:
2846:
2845:
2821:
2815:
2265:
2259:
1468:Timeline of ceratopsian research
1452:
334:
134:
1:
2999:Ceratopsians of North America
2974:Fossil taxa described in 2013
1551:10.1016/j.cretres.2014.06.011
3004:Late Cretaceous ceratopsians
947:Almond Formation chasmosaur
3020:
16:Extinct genus of dinosaurs
2835:
2813:
2575:
2365:
2306:
2291:
2278:
2256:
2184:
2005:
1875:
1871:
1779:
1775:
1760:
1747:
1693:
1666:
1653:
1598:
1490:Longrich, N. R. (2013). "
1148:
1133:
1126:
1111:
1104:
1082:
1075:
1058:
1051:
1031:
1024:
998:
981:
974:
967:
945:
928:
921:
914:
889:
877:Kosmoceratops richardsoni
874:
867:
855:
820:
807:Pentaceratops sternbergii
803:
796:
785:Williams Fork chasmosaur
783:
768:
761:
754:
737:
730:
704:
671:
656:
649:
633:
626:
614:
607:
590:
583:
566:
559:
434:. It is the oldest known
307:
300:
131:Scientific classification
129:
124:
115:
23:
771:Pentaceratops aquilonius
1792:Micropachycephalosaurus
1394:, the pachycephalosaur
674:Chasmosaurus irvinensis
1349:Judith River Formation
1336:
528:
496:
454:
2941:Paleobiology Database
1397:Colepiocephale lambei
1334:
522:
492:
449:
408:. It lived around 78
2994:Ornithischian genera
1508:10.3374/014.054.0103
398:horned face") is an
2906:Judiceratops tigris
2876:Judiceratops tigris
1957:Archaeoceratopsidae
1543:2014CrRes..51..292L
1531:Cretaceous Research
1492:Judiceratops tigris
1460:Paleontology portal
1339:The remains of the
314:Judiceratops tigris
2984:Fossils of Montana
2677:Crittendenceratops
2627:Albertaceratopsini
1421:, and with fellow
1371:with bony fishes,
1337:
659:Chasmosaurus belli
529:
497:
455:
412:, during the Late
2956:
2955:
2868:Taxon identifiers
2859:
2858:
2831:
2830:
2811:
2810:
2807:
2806:
2803:
2802:
2799:
2798:
2760:Pachyrhinosaurini
2698:Yehuecauhceratops
2661:Nasutoceratopsini
2562:
2561:
2274:
2273:
2254:
2253:
2250:
2249:
2246:
2245:
2242:
2241:
2238:
2237:
2234:
2233:
2135:Protoceratopsidae
2022:Leptoceratopsidae
1990:
1989:
1856:
1855:
1743:
1742:
1739:
1738:
1735:
1734:
1731:
1730:
1390:Brachylophosaurus
1278:
1277:
1269:
1268:
1260:
1259:
1251:
1250:
1242:
1241:
1233:
1232:
1224:
1223:
1215:
1214:
1206:
1205:
1197:
1196:
1188:
1187:
1179:
1178:
1170:
1169:
1161:
1160:
1013:
1012:
956:
955:
903:
902:
844:
843:
835:
834:
719:
718:
693:
692:
684:
683:
410:million years ago
325:
324:
281:
3011:
2989:Campanian genera
2949:
2948:
2936:
2935:
2923:
2922:
2910:
2909:
2908:
2895:
2894:
2893:
2863:
2849:
2848:
2826:
2825:
2819:
2790:Pachyrhinosaurus
2714:
2588:Machairoceratops
2573:
2413:Coahuilaceratops
2363:
2304:
2293:
2280:
2269:
2264:
2263:
2182:
2131:
2115:Zhuchengceratops
2003:
1873:
1869:
1810:Chaoyangsauridae
1777:
1773:
1762:
1749:
1691:
1664:
1655:
1648:
1647:
1600:
1585:
1578:
1571:
1562:
1555:
1554:
1526:
1520:
1519:
1487:
1462:
1457:
1456:
1455:
1369:paleoenvironment
1351:in Hill County,
1284:sexual selection
1129:
1107:
1078:
1054:
1027:
1002:Coahuilaceratops
977:
970:
924:
917:
870:
858:
799:
764:
757:
733:
652:
629:
617:
610:
586:
562:
552:
494:Life restoration
390:
386:
381:
380:
377:
376:
373:
370:
367:
364:
361:
358:
355:
352:
349:
346:
343:
340:
316:
312:
273:
266:
253:
240:
227:
211:
195:
139:
138:
120:
110:
40:
29:Temporal range:
21:
3019:
3018:
3014:
3013:
3012:
3010:
3009:
3008:
2959:
2958:
2957:
2952:
2944:
2939:
2931:
2926:
2918:
2913:
2904:
2903:
2898:
2889:
2888:
2883:
2870:
2860:
2855:
2827:
2820:
2795:
2754:
2703:
2684:Furcatoceratops
2655:
2636:Albertaceratops
2621:
2595:Menefeeceratops
2558:
2531:Tatankaceratops
2488:
2434:Mercuriceratops
2385:Arrhinoceratops
2352:
2300:
2287:
2270:
2258:
2230:
2210:
2171:
2120:
2087:Montanoceratops
2016:
2011:Graciliceratops
1999:
1986:
1966:Archaeoceratops
1951:
1865:
1852:
1840:Xuanhuaceratops
1826:Hualianceratops
1804:
1785:Albalophosaurus
1769:
1756:
1727:
1707:
1680:
1674:Avemetatarsalia
1659:Avemetatarsalia
1649:
1643:Marginocephalia
1605:
1604:
1594:
1589:
1559:
1558:
1528:
1527:
1523:
1489:
1488:
1481:
1476:
1458:
1453:
1451:
1448:
1427:Albertaceratops
1361:Late Cretaceous
1329:
1301:Albertaceratops
1292:
1279:
1270:
1261:
1252:
1243:
1234:
1225:
1216:
1207:
1198:
1189:
1180:
1171:
1162:
1035:Arrhinoceratops
1014:
957:
904:
845:
836:
720:
694:
685:
570:Mercuriceratops
541:Mercuriceratops
517:
487:
444:
419:in what is now
388:
384:
337:
333:
321:
318:
310:
309:
296:
272:
264:
251:
238:
225:
209:
193:
133:
111:
109:
108:
107:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
35:
34:
31:Late Cretaceous
27:
17:
12:
11:
5:
3017:
3015:
3007:
3006:
3001:
2996:
2991:
2986:
2981:
2976:
2971:
2969:Chasmosaurines
2961:
2960:
2954:
2953:
2951:
2950:
2937:
2924:
2911:
2896:
2880:
2878:
2872:
2871:
2866:
2857:
2856:
2854:
2853:
2843:
2836:
2833:
2832:
2829:
2828:
2814:
2812:
2809:
2808:
2805:
2804:
2801:
2800:
2797:
2796:
2794:
2793:
2786:
2779:
2772:
2764:
2762:
2756:
2755:
2753:
2752:
2744:
2737:
2730:
2722:
2720:
2711:
2705:
2704:
2702:
2701:
2694:
2691:Nasutoceratops
2687:
2680:
2673:
2665:
2663:
2657:
2656:
2654:
2653:
2650:Medusaceratops
2646:
2639:
2631:
2629:
2623:
2622:
2620:
2619:
2612:
2605:
2598:
2591:
2584:
2581:Diabloceratops
2576:
2570:
2568:Centrosaurinae
2564:
2563:
2560:
2559:
2557:
2556:
2549:
2542:
2539:Titanoceratops
2535:
2527:
2524:Regaliceratops
2520:
2513:
2506:
2498:
2496:
2494:Triceratopsini
2490:
2489:
2487:
2486:
2479:
2472:
2465:
2458:
2455:Sierraceratops
2451:
2444:
2441:Navajoceratops
2437:
2430:
2423:
2416:
2409:
2402:
2395:
2388:
2381:
2374:
2366:
2360:
2358:Chasmosaurinae
2354:
2353:
2351:
2350:
2343:
2336:
2329:
2322:
2319:Brachyceratops
2315:
2307:
2301:
2296:
2289:
2288:
2283:
2276:
2275:
2272:
2271:
2257:
2255:
2252:
2251:
2248:
2247:
2244:
2243:
2240:
2239:
2236:
2235:
2232:
2231:
2229:
2228:
2227:
2226:
2220:
2218:
2212:
2211:
2209:
2208:
2201:
2198:Turanoceratops
2194:
2185:
2179:
2173:
2172:
2170:
2169:
2162:
2155:
2148:
2139:
2137:
2128:
2122:
2121:
2119:
2118:
2111:
2108:Unescoceratops
2104:
2097:
2090:
2083:
2076:
2073:Ischioceratops
2069:
2066:Gryphoceratops
2062:
2055:
2048:
2041:
2034:
2026:
2024:
2018:
2017:
2015:
2014:
2006:
2000:
1995:
1992:
1991:
1988:
1987:
1985:
1984:
1976:
1969:
1961:
1959:
1953:
1952:
1950:
1949:
1946:Sasayamagnomus
1942:
1935:
1928:
1921:
1913:
1906:
1899:
1896:Auroraceratops
1892:
1885:
1876:
1866:
1861:
1858:
1857:
1854:
1853:
1851:
1850:
1843:
1836:
1829:
1822:
1819:Chaoyangsaurus
1814:
1812:
1806:
1805:
1803:
1802:
1799:Psittacosaurus
1795:
1788:
1780:
1770:
1765:
1758:
1757:
1752:
1745:
1744:
1741:
1740:
1737:
1736:
1733:
1732:
1729:
1728:
1726:
1725:
1724:
1723:
1717:
1715:
1709:
1708:
1706:
1705:
1704:
1703:
1694:
1688:
1682:
1681:
1679:
1678:
1677:
1676:
1667:
1661:
1651:
1650:
1646:
1645:
1636:
1627:
1618:
1612:
1603:
1596:
1595:
1590:
1588:
1587:
1580:
1573:
1565:
1557:
1556:
1521:
1478:
1477:
1475:
1472:
1471:
1470:
1464:
1463:
1447:
1444:
1433:Medusaceratops
1328:
1325:
1291:
1288:
1276:
1275:
1272:
1271:
1267:
1266:
1263:
1262:
1258:
1257:
1254:
1253:
1249:
1248:
1245:
1244:
1240:
1239:
1236:
1235:
1231:
1230:
1227:
1226:
1222:
1221:
1218:
1217:
1213:
1212:
1209:
1208:
1204:
1203:
1200:
1199:
1195:
1194:
1191:
1190:
1186:
1185:
1182:
1181:
1177:
1176:
1173:
1172:
1168:
1167:
1164:
1163:
1159:
1158:
1155:
1154:
1147:
1144:
1143:
1140:
1139:
1132:
1127:
1125:
1122:
1121:
1118:
1117:
1110:
1105:
1103:
1095:
1094:
1091:
1090:
1081:
1076:
1074:
1071:
1070:
1067:
1066:
1062:Titanoceratops
1057:
1052:
1050:
1048:Triceratopsini
1044:
1043:
1040:
1039:
1030:
1025:
1023:
1020:
1019:
1016:
1015:
1011:
1010:
1007:
1006:
997:
994:
993:
990:
989:
980:
975:
973:
968:
966:
963:
962:
959:
958:
954:
953:
950:
949:
944:
941:
940:
937:
936:
927:
922:
920:
915:
913:
910:
909:
906:
905:
901:
900:
897:
896:
888:
885:
884:
881:
880:
873:
868:
866:
856:
854:
851:
850:
847:
846:
842:
841:
838:
837:
833:
832:
829:
828:
819:
816:
815:
812:
811:
802:
797:
795:
792:
791:
788:
787:
782:
779:
778:
775:
774:
767:
762:
760:
755:
753:
750:
749:
746:
745:
736:
731:
729:
726:
725:
722:
721:
717:
716:
713:
712:
703:
700:
699:
696:
695:
691:
690:
687:
686:
682:
681:
678:
677:
670:
667:
666:
663:
662:
655:
650:
648:
645:
644:
641:
640:
632:
627:
625:
615:
613:
608:
606:
603:
602:
599:
598:
589:
584:
582:
579:
578:
575:
574:
565:
560:
558:
556:Chasmosaurinae
550:
516:
515:Classification
513:
486:
483:
443:
440:
323:
322:
320:Longrich, 2013
319:
305:
304:
298:
297:
293:J. tigris
289:
287:
283:
282:
262:
258:
257:
255:Chasmosaurinae
249:
245:
244:
236:
232:
231:
223:
216:
215:
213:Neornithischia
207:
200:
199:
191:
184:
183:
178:
171:
170:
165:
161:
160:
155:
151:
150:
145:
141:
140:
127:
126:
122:
121:
113:
112:
104:
103:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
41:
28:
15:
13:
10:
9:
6:
4:
3:
2:
3016:
3005:
3002:
3000:
2997:
2995:
2992:
2990:
2987:
2985:
2982:
2980:
2977:
2975:
2972:
2970:
2967:
2966:
2964:
2947:
2942:
2938:
2934:
2929:
2925:
2921:
2916:
2912:
2907:
2901:
2897:
2892:
2886:
2882:
2881:
2879:
2877:
2873:
2869:
2864:
2852:
2844:
2842:
2838:
2837:
2834:
2824:
2818:
2792:
2791:
2787:
2785:
2784:
2780:
2778:
2777:
2773:
2771:
2770:
2769:Achelousaurus
2766:
2765:
2763:
2761:
2757:
2750:
2749:
2748:Styracosaurus
2745:
2743:
2742:
2738:
2736:
2735:
2731:
2729:
2728:
2724:
2723:
2721:
2719:
2718:Centrosaurini
2715:
2712:
2710:
2709:Eucentrosaura
2706:
2700:
2699:
2695:
2693:
2692:
2688:
2686:
2685:
2681:
2679:
2678:
2674:
2672:
2671:
2667:
2666:
2664:
2662:
2658:
2652:
2651:
2647:
2645:
2644:
2640:
2638:
2637:
2633:
2632:
2630:
2628:
2624:
2618:
2617:
2613:
2611:
2610:
2609:Wendiceratops
2606:
2604:
2603:
2599:
2597:
2596:
2592:
2590:
2589:
2585:
2583:
2582:
2578:
2577:
2574:
2571:
2569:
2565:
2555:
2554:
2550:
2548:
2547:
2543:
2541:
2540:
2536:
2533:
2532:
2528:
2526:
2525:
2521:
2519:
2518:
2514:
2512:
2511:
2507:
2505:
2504:
2503:Eotriceratops
2500:
2499:
2497:
2495:
2491:
2485:
2484:
2480:
2478:
2477:
2473:
2471:
2470:
2466:
2464:
2463:
2459:
2457:
2456:
2452:
2450:
2449:
2448:Pentaceratops
2445:
2443:
2442:
2438:
2436:
2435:
2431:
2429:
2428:
2427:Kosmoceratops
2424:
2422:
2421:
2417:
2415:
2414:
2410:
2408:
2407:
2403:
2401:
2400:
2399:Bravoceratops
2396:
2394:
2393:
2392:Bisticeratops
2389:
2387:
2386:
2382:
2380:
2379:
2378:Anchiceratops
2375:
2373:
2372:
2371:Agujaceratops
2368:
2367:
2364:
2361:
2359:
2355:
2349:
2348:
2344:
2342:
2341:
2337:
2335:
2334:
2330:
2328:
2327:
2323:
2321:
2320:
2316:
2314:
2313:
2309:
2308:
2305:
2302:
2299:
2294:
2290:
2286:
2281:
2277:
2268:
2262:
2224:
2223:
2222:
2221:
2219:
2217:
2213:
2207:
2206:
2202:
2200:
2199:
2195:
2192:
2191:
2187:
2186:
2183:
2180:
2178:
2177:Ceratopsoidea
2174:
2168:
2167:
2166:Protoceratops
2163:
2161:
2160:
2159:Breviceratops
2156:
2154:
2153:
2149:
2146:
2145:
2141:
2140:
2138:
2136:
2132:
2129:
2127:
2123:
2117:
2116:
2112:
2110:
2109:
2105:
2103:
2102:
2101:Udanoceratops
2098:
2096:
2095:
2094:Prenoceratops
2091:
2089:
2088:
2084:
2082:
2081:
2080:Leptoceratops
2077:
2075:
2074:
2070:
2068:
2067:
2063:
2061:
2060:
2056:
2054:
2053:
2049:
2047:
2046:
2042:
2040:
2039:
2038:Bainoceratops
2035:
2033:
2032:
2028:
2027:
2025:
2023:
2019:
2013:
2012:
2008:
2007:
2004:
2001:
1998:
1993:
1982:
1981:
1977:
1975:
1974:
1973:Helioceratops
1970:
1968:
1967:
1963:
1962:
1960:
1958:
1954:
1948:
1947:
1943:
1941:
1940:
1939:Mosaiceratops
1936:
1934:
1933:
1929:
1927:
1926:
1922:
1919:
1918:
1914:
1912:
1911:
1910:Koreaceratops
1907:
1905:
1904:
1900:
1898:
1897:
1893:
1891:
1890:
1886:
1883:
1882:
1878:
1877:
1874:
1870:
1867:
1864:
1863:Neoceratopsia
1859:
1849:
1848:
1844:
1842:
1841:
1837:
1835:
1834:
1830:
1828:
1827:
1823:
1821:
1820:
1816:
1815:
1813:
1811:
1807:
1801:
1800:
1796:
1794:
1793:
1789:
1787:
1786:
1782:
1781:
1778:
1774:
1771:
1768:
1763:
1759:
1755:
1750:
1746:
1721:
1720:
1719:
1718:
1716:
1714:
1710:
1702:
1698:
1697:
1696:
1695:
1692:
1689:
1687:
1683:
1675:
1671:
1670:
1669:
1668:
1665:
1662:
1660:
1656:
1652:
1644:
1640:
1637:
1635:
1631:
1628:
1626:
1622:
1619:
1617:
1613:
1611:
1607:
1606:
1601:
1597:
1593:
1586:
1581:
1579:
1574:
1572:
1567:
1566:
1563:
1552:
1548:
1544:
1540:
1536:
1532:
1525:
1522:
1517:
1513:
1509:
1505:
1501:
1497:
1493:
1486:
1484:
1480:
1473:
1469:
1466:
1465:
1461:
1450:
1445:
1443:
1441:
1440:
1435:
1434:
1429:
1428:
1424:
1420:
1419:
1414:
1413:
1408:
1407:
1406:Dromaeosaurus
1403:
1399:
1398:
1393:
1391:
1387:
1383:
1382:
1381:Champsosaurus
1378:
1377:Choristoderan
1374:
1370:
1366:
1362:
1358:
1354:
1350:
1346:
1342:
1333:
1326:
1324:
1322:
1321:Pentaceratops
1318:
1314:
1309:
1307:
1303:
1302:
1296:
1289:
1287:
1285:
1274:
1273:
1265:
1264:
1256:
1255:
1247:
1246:
1238:
1237:
1229:
1228:
1220:
1219:
1211:
1210:
1202:
1201:
1193:
1192:
1184:
1183:
1175:
1174:
1166:
1165:
1157:
1156:
1153:
1152:
1146:
1145:
1142:
1141:
1138:
1137:
1131:
1130:
1124:
1123:
1120:
1119:
1116:
1115:
1109:
1108:
1102:
1101:
1097:
1096:
1093:
1092:
1089:
1088:
1087:
1080:
1079:
1073:
1072:
1069:
1068:
1065:
1064:
1063:
1056:
1055:
1049:
1046:
1045:
1042:
1041:
1038:
1037:
1036:
1029:
1028:
1022:
1021:
1018:
1017:
1009:
1008:
1005:
1004:
1003:
996:
995:
992:
991:
988:
987:
986:
985:Bravoceratops
979:
978:
972:
971:
965:
964:
961:
960:
952:
951:
948:
943:
942:
939:
938:
935:
934:
933:
932:Anchiceratops
926:
925:
919:
918:
912:
911:
908:
907:
899:
898:
895:
894:sp. CMN 8301
893:
892:Kosmoceratops
887:
886:
883:
882:
879:
878:
872:
871:
865:
864:
863:Kosmoceratops
860:
859:
853:
852:
849:
848:
840:
839:
831:
830:
827:
826:
825:
818:
817:
814:
813:
810:
809:
808:
801:
800:
794:
793:
790:
789:
786:
781:
780:
777:
776:
773:
772:
766:
765:
759:
758:
752:
751:
748:
747:
744:
743:
742:
741:Agujaceratops
735:
734:
728:
727:
724:
723:
715:
714:
711:
710:
709:
702:
701:
698:
697:
689:
688:
680:
679:
676:
675:
669:
668:
665:
664:
661:
660:
654:
653:
647:
646:
643:
642:
639:
638:sp. CMN 2280
637:
631:
630:
624:
623:
619:
618:
612:
611:
605:
604:
601:
600:
597:
596:
595:
588:
587:
581:
580:
577:
576:
573:
572:
571:
564:
563:
557:
554:
553:
549:
547:
546:Pentaceratops
543:
542:
537:
536:chasmosaurine
533:
526:
521:
514:
512:
510:
506:
502:
501:parietal bone
495:
491:
484:
482:
480:
474:
472:
468:
464:
460:
453:
448:
441:
439:
437:
436:chasmosaurine
433:
430:
427:was a large,
426:
422:
418:
415:
411:
407:
404:
401:
397:
393:
392:
379:
331:
330:
317:
315:
306:
303:
302:Binomial name
299:
295:
294:
288:
285:
284:
280:
276:
271:
270:
263:
260:
259:
256:
250:
247:
246:
243:
237:
234:
233:
230:
224:
221:
218:
217:
214:
208:
205:
202:
201:
198:
192:
189:
186:
185:
182:
179:
176:
173:
172:
169:
166:
163:
162:
159:
156:
153:
152:
149:
146:
143:
142:
137:
132:
128:
123:
119:
114:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
39:
36:78–77.5
32:
26:
22:
19:
2875:
2788:
2783:Stellasaurus
2781:
2774:
2767:
2746:
2739:
2734:Coronosaurus
2732:
2727:Centrosaurus
2725:
2696:
2689:
2682:
2675:
2668:
2648:
2643:Lokiceratops
2641:
2634:
2616:Xenoceratops
2614:
2607:
2602:Sinoceratops
2600:
2593:
2586:
2579:
2551:
2544:
2537:
2529:
2522:
2515:
2510:Nedoceratops
2508:
2501:
2483:Vagaceratops
2481:
2476:Utahceratops
2474:
2469:Terminocavus
2467:
2460:
2453:
2446:
2439:
2432:
2425:
2420:Judiceratops
2419:
2418:
2411:
2406:Chasmosaurus
2404:
2397:
2390:
2383:
2376:
2369:
2345:
2338:
2331:
2324:
2317:
2310:
2298:Ceratopsidae
2285:Ceratopsidae
2216:Ceratopsidae
2205:Zuniceratops
2203:
2196:
2190:Ajkaceratops
2188:
2164:
2157:
2152:Bagaceratops
2150:
2144:Ajkaceratops
2142:
2126:Coronosauria
2113:
2106:
2099:
2092:
2085:
2078:
2071:
2064:
2057:
2050:
2043:
2036:
2031:Asiaceratops
2029:
2009:
1997:Euceratopsia
1980:Yamaceratops
1978:
1971:
1964:
1944:
1937:
1932:Microceratus
1930:
1925:Liaoceratops
1923:
1915:
1908:
1901:
1894:
1887:
1881:Ajkaceratops
1879:
1845:
1838:
1831:
1824:
1817:
1797:
1790:
1783:
1701:Ornithischia
1686:Ornithischia
1638:
1634:Ornithischia
1629:
1620:
1534:
1530:
1524:
1499:
1495:
1491:
1437:
1431:
1425:
1423:ceratopsians
1416:
1410:
1404:
1395:
1388:
1379:
1365:Judiceratops
1364:
1345:Judiceratops
1344:
1343:specimen of
1338:
1327:Paleoecology
1320:
1317:Chasmosaurus
1316:
1313:Judiceratops
1312:
1310:
1306:Zuniceratops
1299:
1295:Judiceratops
1294:
1293:
1280:
1150:
1149:
1135:
1134:
1114:T. utahensis
1113:
1112:
1098:
1084:
1083:
1060:
1059:
1033:
1032:
1000:
999:
983:
982:
946:
930:
929:
891:
890:
876:
875:
861:
824:Utahceratops
822:
821:
805:
804:
784:
770:
769:
739:
738:
708:Mojoceratops
706:
705:
673:
672:
658:
657:
636:Chasmosaurus
635:
634:
622:Chasmosaurus
620:
594:Judiceratops
593:
592:
591:
568:
567:
545:
539:
532:Judiceratops
531:
530:
525:Judiceratops
524:
498:
479:Judiceratops
478:
475:
463:Judiceratops
462:
456:
452:Judiceratops
451:
425:Judiceratops
424:
396:Judith River
329:Judiceratops
328:
327:
326:
313:
308:
292:
291:
269:Judiceratops
268:
267:
242:Ceratopsidae
219:
203:
197:Ornithischia
187:
174:
25:Judiceratops
24:
18:
2900:Wikispecies
2776:Einiosaurus
2670:Avaceratops
2553:Triceratops
2517:Ojoceratops
2340:Monoclonius
2052:Ferrisaurus
1917:Kulceratops
1537:: 292–308.
1439:Avaceratops
1412:Gorgosaurus
1367:shared its
1335:Restoration
1136:T. horridus
1100:Triceratops
534:is a basal
499:Its frill (
485:Description
471:postorbital
467:ceratopsids
429:quadrupedal
394:; meaning "
248:Subfamily:
2963:Categories
2839:See also:
2546:Torosaurus
2462:Spiclypeus
2312:Agathaumas
2225:see below↓
2045:Cerasinops
1833:Stenopelix
1767:Ceratopsia
1754:Ceratopsia
1722:see below↓
1713:Ceratopsia
1625:Dinosauria
1592:Ceratopsia
1474:References
1392:canadensis
1373:amphibians
1151:T. prorsus
1086:Torosaurus
509:osteoderms
505:osteoderms
414:Cretaceous
229:Ceratopsia
181:Dinosauria
2891:Q11878380
1608:Kingdom:
1516:129801786
1502:: 51–65.
1402:theropods
1386:hadrosaur
1359:stage of
1357:Campanian
1311:Although
1290:Evolution
442:Discovery
432:herbivore
286:Species:
154:Kingdom:
148:Eukaryota
2920:42418329
2885:Wikidata
2851:Category
2841:Timeline
2347:Polyonax
2333:Dysganus
2326:Ceratops
1889:Aquilops
1616:Chordata
1614:Phylum:
1610:Animalia
1446:See also
1341:holotype
459:holotype
406:dinosaur
275:Longrich
235:Family:
168:Chordata
164:Phylum:
158:Animalia
144:Domain:
2933:8469786
2741:Spinops
2059:Gremlin
1847:Yinlong
1539:Bibcode
1418:Troodon
1353:Montana
469:. The
421:Montana
400:extinct
391:-ə-tops
311:†
290:†
261:Genus:
105:↓
2946:266208
1514:
1400:, the
1384:, the
1375:, the
417:Period
403:horned
125:Frill
1639:Clade
1630:Clade
1621:Clade
1512:S2CID
387:-dee-
220:Clade
204:Clade
188:Clade
175:Clade
2928:GBIF
1699:see
1672:see
1436:and
1415:and
1319:and
457:The
389:SERR
279:2013
44:PreꞒ
2915:EoL
1903:Beg
1547:doi
1504:doi
385:JOO
2965::
2943::
2930::
2917::
2902::
2887::
1641::
1632::
1623::
1545:.
1535:51
1533:.
1510:.
1500:54
1498:.
1482:^
1442:.
1430:,
1409:,
438:.
360:ɛr
351:iː
345:uː
342:dʒ
277:,
222::
206::
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