3600:
3607:
3620:
649:; the foramen on the quadratojugal; a large quadratic foramen; and the absence of an external fenestra in the lower jaw. However, they also noticed more derived euornithopod traits, such as the form of the greater and anterior trochanter of the thighbone, although the premaxilla not reaching the lacrimal, the high jaw joint and the premaxilla being on the same level as the maxilla, were again basal traits. The fused mandibular symphysis might indicate a relation with the
3627:
3613:
136:
118:
477:
488:
2777:
500:
438:
and it is thought that an enormous volcanic eruption occurred burying everything within a fifty to sixty mile radius. The holotype contains a compressed skull and a partial postcranial skeleton. The second specimen, IVPP V 12530, was referred to the species. It consists of a better preserved, perhaps
455:
was a small bipedal herbivore. Because the specimens are juvenile it is hard to ascertain the adult size. The holotype is 71.1 centimetres (28.0 in) long with a 35.6 centimetres (14.0 in) long tail, nearly half the length of the complete animal. The length of the skull is 6.3 centimetres
507:
Both skulls are incomplete and had to be partially restored. The referred specimens shows an extremely large orbit compared to the skull size, with a length of 40% of the total skull length. Its snout is short, also comprising 40% of the skull length. Both traits are indicative of a
2734:"New small-bodied ornithopods (Dinosauria, Neornithischia) from the Early Cretaceous Wonthaggi Formation (Strzelecki Group) of the Australian-Antarctic rift system, with revision of Qantassaurus intrepidus Rich and Vickers-Rich, 1999"
560:. The longest is metatarsal III with a length of 5.5 centimetres (2.2 in); it is placed more anteriorly than the other metatarsals. Metatarsal I is more posterior and its upper part is transversely reduced to a splint.
523:
has at least thirteen teeth. Although the back, maxillary, teeth are fan-shaped like those of a plant eater, the front premaxillary teeth are narrower and longer, more like a carnivorous dinosaur. This may mean that
2689:
Han, Feng-Lu; Paul M. Barrett; Richard J. Butler; Xing Xu (2012). "Postcranial anatomy of
Jeholosaurus shangyuanensis (Dinosauria, Ornithischia) from the Lower Cretaceous Yixian Formation of China".
421:
3853:
2629:
Makovicky, Peter J.; Brandon M. Kilbourne; Rudyard W. Sadleir; Mark A. Norell (2011). "A new basal ornithopod (Dinosauria, Ornithischia) from the Late
Cretaceous of Mongolia".
653:. Very basal ornithischian traits included the presence of six teeth in the premaxilla with only a short toothless sector in front and a short hiatus with the maxillary teeth.
552:
slightly lower than the greater trochanter and a third as narrow as the latter. They are separated by a shallow cleft. Between the condyles of the lower femur the (anterior)
3951:
2578:
Chang, S.-C.; Gao, K.-Q.; Chang, Z.-F.; Jourdan, F. (2017). "New chronostratigraphic constraints on the Yixian
Formation with implications for the Jehol Biota".
4010:
460:
are 5.9 centimetres (2.3 in) each. The forelimbs are 25.4 centimetres (10.0 in) long and the hindlimbs are 33 centimetres (13 in). The
2671:
Zheng, X-.T., You, H.-L., Xu, X. and Dong, Z.-M. (2009). "An Early
Cretaceous heterodontosaurid dinosaur with filamentous integumentary structures."
532:, eating both plants and animals. The deeply inset ventral margin in the maxilla suggests fleshy cheeks may have been present. The nasals have large
3938:
4015:
2810:
2568:
Xu, Wang and You, 2000. A primitive ornithopod from the Early
Cretaceous Yixian Formation of Liaoning. Vertebrata PalAsiatica 38(4)318-325.
3599:
3619:
3606:
4020:
3626:
4030:
3612:
135:
3570:
2803:
4005:
2898:
369:
245:
556:
groove is absent. The fourth trochanter is pendant and located in a rather high position. The foot has four
439:
smaller, skull and some neck vertebrae. Both specimens represent juvenile or at least subadult individuals.
405:
3878:
2994:
572:
4025:
3977:
3247:
2796:
275:
3925:
3515:
3270:
2748:
2698:
2638:
2587:
349:
252:
3547:
3115:
1584:
1196:
714:
646:
509:
373:
3832:
3559:
2714:
2654:
1308:
541:
130:
3982:
3916:
2733:
377:
3964:
3956:
3800:
3145:
2158:
639:
549:
361:
353:
117:
3969:
434:, about 126-125 million years old. The layers consist of fluvial sandstone interspersed with
3379:
2782:
2756:
2706:
2646:
2595:
2362:
1878:
545:
537:
427:
325:
30:
548:
have been preserved, their exact number is unknown. The femur is bowed and has an anterior
3822:
3527:
3395:
2883:
2858:
2064:
1847:
1806:
1454:
961:
937:
880:
650:
2752:
2702:
2642:
2591:
3837:
3817:
3768:
3693:
3478:
3388:
3326:
3291:
3211:
2339:
2225:
2208:
2175:
1990:
1792:
1645:
1552:
792:
634:
568:
533:
393:
315:
225:
212:
476:
3999:
3785:
3735:
3493:
3464:
3450:
3434:
3319:
3312:
3204:
3124:
3048:
3041:
3009:
2386:
2313:
1949:
1916:
1715:
1660:
1414:
1339:
1295:
1020:
854:
764:
642:
576:
68:
2718:
2658:
3827:
3778:
3721:
3707:
3700:
3655:
3471:
3354:
3333:
3236:
3103:
3095:
3071:
3025:
2960:
2947:
2910:
2819:
2326:
2259:
2093:
1570:
1173:
1158:
1037:
691:
676:
630:
619:
611:
607:
592:
584:
483:
The curled position raises the question as to whether this animal died in a burrow.
381:
357:
196:
487:
2710:
2650:
2599:
595:; and the claw of the third toe being longer than the other third toe phalanges.
3910:
3756:
3728:
3686:
3679:
3672:
3504:
3403:
3368:
3305:
3259:
3184:
3063:
3033:
2986:
2978:
2299:
2078:
1732:
1684:
1631:
1439:
1373:
1271:
1247:
1221:
892:
830:
806:
780:
588:
333:
43:
1150:
The following cladogram was based on analysis by Zheng and colleagues in 2009.
660:
analyses have recovered a basal position in the (Eu)ornithopoda. The following
408:
refers to the geographical area of
Shangyuan where the fossil site is located.
3749:
3742:
3536:
3485:
3457:
3442:
3418:
3361:
3340:
3218:
3197:
3173:
3166:
3152:
3138:
3087:
3079:
3055:
2772:
2402:
2242:
2127:
1973:
1899:
1772:
1397:
1208:
906:
750:
580:
557:
529:
516:
322:
88:
53:
3901:
3714:
3347:
3131:
3017:
2113:
1322:
661:
657:
329:
147:
93:
37:
2732:
Herne, Matthew C.; Nair, Jay P.; Evans, Alistair R.; Tait, Alan M. (2019).
499:
3895:
3665:
3159:
3002:
2867:
2843:
2049:
1607:
1233:
867:
737:
553:
457:
412:
397:
318:
180:
167:
83:
78:
63:
58:
48:
2761:
610:, as is shown by its ornithischian four-pronged pelvic structure with a
3943:
3793:
3225:
2849:
615:
520:
98:
73:
638:. Using the comparative method, they pointed out some similarities to
2837:
431:
157:
3872:
498:
486:
475:
465:
461:
401:
396:"; Jehol is the name of a historical province situated in western
365:
311:
3930:
3411:
435:
3876:
3642:
2945:
2830:
2792:
2788:
579:
length; a quadratojugal less than 30% of skull height; a
614:
bone pointing downward and backwards, parallel to the
3854:
List of stratigraphic units with ornithischian tracks
629:
to any family, limiting themselves to a placement as
3885:
3846:
3810:
3766:
3653:
3558:
3535:
3526:
3503:
3378:
3289:
3258:
3235:
3183:
3114:
2958:
2909:
2882:
1558:The cladogram below results from analysis by Herne
2580:Palaeogeography, Palaeoclimatology, Palaeoecology
1535:found a similar topology to that of Makovicky
2804:
8:
464:is 9 centimetres (3.5 in) long and the
368:, and named and described the same year by
3873:
3650:
3639:
3532:
3297:
3189:
2970:
2966:
2955:
2942:
2888:
2879:
2827:
2811:
2797:
2789:
2611:
2609:
1539:, 2011, in 2012, with a clade composed of
116:
20:
2760:
456:(2.5 in) and the lower jaws of the
2564:
2562:
2560:
2558:
2556:
2554:
2552:
2548:
536:dorsolaterally and a midline fossa. No
426:. It was found in a layer of the early
2617:The Princeton Field Guide to Dinosaurs
591:almost to the posterior border of the
7:
2619:, Princeton University Press p. 275
2378:
2354:
2332:
2305:
2200:
2150:
2143:
2119:
2109:
2085:
2056:
1965:
1941:
1891:
1884:
1822:
1798:
1788:
1764:
1757:
1707:
1700:
1676:
1652:
1623:
1599:
1577:
1565:
1389:
1314:
1287:
1263:
1239:
1214:
1189:
1165:
1153:
1012:
953:
929:
922:
898:
873:
846:
822:
798:
756:
729:
707:
683:
671:
664:was based on analysis by Makovicky
468:is 10.7 centimetres (4.2 in).
4011:Early Cretaceous dinosaurs of Asia
2691:Journal of Vertebrate Paleontology
2631:Journal of Vertebrate Paleontology
419:, on which the genus is based, is
14:
587:body length; a dentary extending
3625:
3618:
3611:
3605:
3598:
2775:
134:
567:include: enlarged laterodorsal
328:. It is thought to have been a
625:The describers did not assign
563:Some distinguishing traits of
1:
4016:Fossil taxa described in 2000
2711:10.1080/02724634.2012.694385
2651:10.1080/02724634.2011.557114
2600:10.1016/j.palaeo.2017.09.026
618:, while a forward-pointing
386:Jeholosaurus shangyuanensis
288:Jeholosaurus shangyuanensis
4047:
384:and only known species is
16:Extinct genus of dinosaurs
3649:
3638:
3595:
3300:
3192:
2973:
2969:
2954:
2941:
2891:
2878:
2826:
2400:
2383:
2376:
2359:
2352:
2337:
2330:
2310:
2303:
2256:
2239:
2222:
2205:
2198:
2172:
2155:
2148:
2141:
2124:
2117:
2107:
2090:
2083:
2061:
2054:
1987:
1970:
1963:
1946:
1939:
1913:
1896:
1889:
1882:
1844:
1827:
1820:
1803:
1796:
1786:
1769:
1762:
1755:
1729:
1712:
1705:
1698:
1681:
1674:
1657:
1650:
1628:
1621:
1604:
1597:
1582:
1575:
1452:
1437:
1411:
1394:
1387:
1370:
1353:
1336:
1319:
1312:
1292:
1285:
1268:
1261:
1244:
1237:
1219:
1212:
1194:
1187:
1170:
1163:
1034:
1017:
1010:
975:
958:
951:
934:
927:
920:
903:
896:
878:
871:
851:
844:
827:
820:
803:
796:
778:
761:
754:
734:
727:
712:
705:
688:
681:
481:Skeleton of Jeholosaurus.
281:
274:
131:Scientific classification
129:
124:
115:
23:
1551:. They named this clade
430:, dating from the early
348:specimens were found in
2741:Journal of Paleontology
622:supports the abdomen.
575:more than 25% maximum
573:quadratojugal fenestra
540:are preserved. Though
519:has six teeth and the
504:
496:
484:
267:J. shangyuanensis
4021:Taxa named by Xu Xing
3978:Paleobiology Database
502:
490:
479:
4031:Ornithischian genera
583:with almost 150% of
340:History of discovery
125:Skull (IVPP V12530)
3116:Heterodontosauridae
2762:10.1017/jpa.2018.95
2753:2019JPal...93..543H
2703:2012JVPal..32.1370H
2643:2011JVPal..31..626M
2592:2017PPP...487..399C
1585:Heterodontosauridae
1197:Heterodontosauridae
715:Heterodontosauridae
647:antorbital fenestra
392:means "lizard from
388:. The generic name
3833:Hypsilophodontidae
3571:Pachycephalosauria
3560:Pachycephalosauria
2615:Paul, G.S., 2010,
1309:Hypsilophodontidae
542:cervical vertebrae
505:
497:
485:
424: V 12529
3993:
3992:
3965:Open Tree of Life
3879:Taxon identifiers
3870:
3869:
3866:
3865:
3862:
3861:
3801:Serendipaceratops
3634:
3633:
3593:
3592:
3589:
3588:
3585:
3584:
3581:
3580:
3285:
3284:
3281:
3280:
3146:Heterodontosaurus
2937:
2936:
2933:
2932:
2929:
2928:
2539:
2538:
2530:
2529:
2521:
2520:
2512:
2511:
2503:
2502:
2494:
2493:
2485:
2484:
2476:
2475:
2467:
2466:
2458:
2457:
2449:
2448:
2440:
2439:
2431:
2430:
2422:
2421:
2413:
2412:
2289:
2288:
2280:
2279:
2271:
2270:
2187:
2186:
2159:Macrogryphosaurus
2038:
2037:
2029:
2028:
2020:
2019:
2011:
2010:
2002:
2001:
1928:
1927:
1868:
1867:
1859:
1858:
1744:
1743:
1528:
1527:
1519:
1518:
1510:
1509:
1501:
1500:
1492:
1491:
1483:
1482:
1474:
1473:
1465:
1464:
1426:
1425:
1147:
1146:
1138:
1137:
1129:
1128:
1120:
1119:
1111:
1110:
1102:
1101:
1093:
1092:
1084:
1083:
1075:
1074:
1066:
1065:
1057:
1056:
1048:
1047:
999:
998:
990:
989:
362:Liaoning Province
303:
302:
255:
4038:
3986:
3985:
3973:
3972:
3960:
3959:
3947:
3946:
3934:
3933:
3921:
3920:
3919:
3906:
3905:
3904:
3874:
3651:
3640:
3629:
3623:
3622:
3616:
3615:
3609:
3603:
3602:
3533:
3380:Thescelosauridae
3298:
3190:
2971:
2967:
2956:
2943:
2889:
2880:
2873:
2872:
2828:
2813:
2806:
2799:
2790:
2785:
2783:Dinosaurs portal
2780:
2779:
2778:
2767:
2766:
2764:
2738:
2729:
2723:
2722:
2697:(6): 1370β1395.
2686:
2680:
2669:
2663:
2662:
2626:
2620:
2613:
2604:
2603:
2575:
2569:
2566:
2379:
2363:Muttaburrasaurus
2355:
2333:
2306:
2201:
2151:
2144:
2120:
2110:
2086:
2057:
1966:
1942:
1892:
1885:
1879:Thescelosauridae
1823:
1799:
1789:
1765:
1758:
1708:
1701:
1677:
1653:
1624:
1600:
1578:
1566:
1390:
1315:
1288:
1264:
1240:
1215:
1190:
1166:
1154:
1013:
954:
930:
923:
899:
874:
847:
823:
799:
757:
730:
708:
684:
672:
620:prepubic process
546:caudal vertebrae
428:Yixian Formation
290:
286:
244:
237:
224:
211:
195:
139:
138:
120:
110:
40:
31:Early Cretaceous
29:Temporal range:
21:
4046:
4045:
4041:
4040:
4039:
4037:
4036:
4035:
4006:Thescelosaurids
3996:
3995:
3994:
3989:
3981:
3976:
3968:
3963:
3955:
3950:
3942:
3937:
3929:
3924:
3915:
3914:
3909:
3900:
3899:
3894:
3881:
3871:
3858:
3842:
3823:Phytodinosauria
3806:
3762:
3645:
3630:
3617:
3610:
3597:
3577:
3554:
3528:Marginocephalia
3522:
3499:
3396:Changchunsaurus
3374:
3294:
3277:
3254:
3231:
3179:
3110:
2963:
2950:
2925:
2905:
2899:Avemetatarsalia
2884:Avemetatarsalia
2874:
2859:Avemetatarsalia
2833:
2832:
2822:
2817:
2781:
2776:
2774:
2771:
2770:
2736:
2731:
2730:
2726:
2688:
2687:
2683:
2670:
2666:
2628:
2627:
2623:
2614:
2607:
2577:
2576:
2572:
2567:
2550:
2545:
2540:
2531:
2522:
2513:
2504:
2495:
2486:
2477:
2468:
2459:
2450:
2441:
2432:
2423:
2414:
2290:
2281:
2272:
2188:
2065:Marginocephalia
2039:
2030:
2021:
2012:
2003:
1929:
1869:
1860:
1848:Changchunsaurus
1745:
1549:Changchunsaurus
1529:
1520:
1511:
1502:
1493:
1484:
1475:
1466:
1455:Marginocephalia
1427:
1148:
1139:
1130:
1121:
1112:
1103:
1094:
1085:
1076:
1067:
1058:
1049:
1000:
991:
962:Changchunsaurus
881:Marginocephalia
651:Marginocephalia
601:
474:
450:
445:
342:
321:from the Early
316:neornithischian
299:
292:
284:
283:
270:
243:
235:
222:
209:
193:
133:
111:
109:
108:
107:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
35:
34:
27:
17:
12:
11:
5:
4044:
4042:
4034:
4033:
4028:
4023:
4018:
4013:
4008:
3998:
3997:
3991:
3990:
3988:
3987:
3974:
3961:
3948:
3935:
3922:
3907:
3891:
3889:
3883:
3882:
3877:
3868:
3867:
3864:
3863:
3860:
3859:
3857:
3856:
3850:
3848:
3847:Other articles
3844:
3843:
3841:
3840:
3838:Jeholosauridae
3835:
3830:
3825:
3820:
3818:Ornithoscelida
3814:
3812:
3808:
3807:
3805:
3804:
3797:
3790:
3782:
3774:
3772:
3769:Incertae sedis
3764:
3763:
3761:
3760:
3753:
3746:
3739:
3732:
3725:
3718:
3711:
3704:
3697:
3694:Ferganocephale
3690:
3683:
3676:
3669:
3661:
3659:
3647:
3646:
3643:
3636:
3635:
3632:
3631:
3596:
3594:
3591:
3590:
3587:
3586:
3583:
3582:
3579:
3578:
3576:
3575:
3574:
3573:
3564:
3562:
3556:
3555:
3553:
3552:
3551:
3550:
3541:
3539:
3530:
3524:
3523:
3521:
3520:
3519:
3518:
3509:
3507:
3501:
3500:
3498:
3497:
3490:
3482:
3479:Thescelosaurus
3475:
3468:
3461:
3454:
3447:
3439:
3431:
3423:
3415:
3408:
3400:
3392:
3389:Albertadromeus
3384:
3382:
3376:
3375:
3373:
3372:
3365:
3358:
3351:
3344:
3337:
3330:
3327:Kulindadromeus
3323:
3316:
3309:
3301:
3295:
3292:Neornithischia
3290:
3287:
3286:
3283:
3282:
3279:
3278:
3276:
3275:
3274:
3273:
3264:
3262:
3256:
3255:
3253:
3252:
3251:
3250:
3241:
3239:
3233:
3232:
3230:
3229:
3222:
3215:
3212:Scutellosaurus
3208:
3201:
3193:
3187:
3181:
3180:
3178:
3177:
3170:
3163:
3156:
3149:
3142:
3135:
3128:
3120:
3118:
3112:
3111:
3109:
3108:
3100:
3092:
3084:
3076:
3068:
3060:
3052:
3045:
3038:
3030:
3022:
3014:
3006:
2999:
2991:
2983:
2974:
2964:
2959:
2952:
2951:
2946:
2939:
2938:
2935:
2934:
2931:
2930:
2927:
2926:
2924:
2923:
2922:
2921:
2915:
2913:
2907:
2906:
2904:
2903:
2902:
2901:
2892:
2886:
2876:
2875:
2871:
2870:
2861:
2852:
2846:
2840:
2831:
2824:
2823:
2818:
2816:
2815:
2808:
2801:
2793:
2787:
2786:
2769:
2768:
2747:(3): 543β584.
2724:
2681:
2679:(19): 333-336.
2664:
2637:(3): 626β640.
2621:
2605:
2570:
2547:
2546:
2544:
2541:
2537:
2536:
2533:
2532:
2528:
2527:
2524:
2523:
2519:
2518:
2515:
2514:
2510:
2509:
2506:
2505:
2501:
2500:
2497:
2496:
2492:
2491:
2488:
2487:
2483:
2482:
2479:
2478:
2474:
2473:
2470:
2469:
2465:
2464:
2461:
2460:
2456:
2455:
2452:
2451:
2447:
2446:
2443:
2442:
2438:
2437:
2434:
2433:
2429:
2428:
2425:
2424:
2420:
2419:
2416:
2415:
2411:
2410:
2407:
2406:
2399:
2396:
2395:
2392:
2391:
2382:
2377:
2375:
2372:
2371:
2368:
2367:
2358:
2353:
2351:
2348:
2347:
2344:
2343:
2340:Rhabdodontidae
2336:
2331:
2329:
2323:
2322:
2319:
2318:
2309:
2304:
2302:
2296:
2295:
2292:
2291:
2287:
2286:
2283:
2282:
2278:
2277:
2274:
2273:
2269:
2268:
2265:
2264:
2255:
2252:
2251:
2248:
2247:
2238:
2235:
2234:
2231:
2230:
2226:Leaellynasaura
2221:
2218:
2217:
2214:
2213:
2209:Galleonosaurus
2204:
2199:
2197:
2194:
2193:
2190:
2189:
2185:
2184:
2181:
2180:
2176:Gasparinisaura
2171:
2168:
2167:
2164:
2163:
2154:
2149:
2147:
2142:
2140:
2137:
2136:
2133:
2132:
2123:
2118:
2116:
2108:
2106:
2103:
2102:
2099:
2098:
2089:
2084:
2082:
2074:
2073:
2070:
2069:
2060:
2055:
2053:
2045:
2044:
2041:
2040:
2036:
2035:
2032:
2031:
2027:
2026:
2023:
2022:
2018:
2017:
2014:
2013:
2009:
2008:
2005:
2004:
2000:
1999:
1996:
1995:
1991:Thescelosaurus
1986:
1983:
1982:
1979:
1978:
1969:
1964:
1962:
1959:
1958:
1955:
1954:
1945:
1940:
1938:
1935:
1934:
1931:
1930:
1926:
1925:
1922:
1921:
1912:
1909:
1908:
1905:
1904:
1895:
1890:
1888:
1883:
1881:
1875:
1874:
1871:
1870:
1866:
1865:
1862:
1861:
1857:
1856:
1853:
1852:
1843:
1840:
1839:
1836:
1835:
1826:
1821:
1819:
1816:
1815:
1812:
1811:
1802:
1797:
1795:
1793:Jeholosauridae
1787:
1785:
1782:
1781:
1778:
1777:
1768:
1763:
1761:
1756:
1754:
1751:
1750:
1747:
1746:
1742:
1741:
1738:
1737:
1728:
1725:
1724:
1721:
1720:
1711:
1706:
1704:
1699:
1697:
1694:
1693:
1690:
1689:
1680:
1675:
1673:
1670:
1669:
1666:
1665:
1656:
1651:
1649:
1646:Neornithischia
1641:
1640:
1637:
1636:
1627:
1622:
1620:
1617:
1616:
1613:
1612:
1603:
1598:
1596:
1593:
1592:
1589:
1588:
1581:
1576:
1574:
1564:
1553:Jeholosauridae
1526:
1525:
1522:
1521:
1517:
1516:
1513:
1512:
1508:
1507:
1504:
1503:
1499:
1498:
1495:
1494:
1490:
1489:
1486:
1485:
1481:
1480:
1477:
1476:
1472:
1471:
1468:
1467:
1463:
1462:
1459:
1458:
1451:
1448:
1447:
1444:
1443:
1436:
1433:
1432:
1429:
1428:
1424:
1423:
1420:
1419:
1410:
1407:
1406:
1403:
1402:
1393:
1388:
1386:
1383:
1382:
1379:
1378:
1369:
1366:
1365:
1362:
1361:
1352:
1349:
1348:
1345:
1344:
1335:
1332:
1331:
1328:
1327:
1318:
1313:
1311:
1305:
1304:
1301:
1300:
1291:
1286:
1284:
1281:
1280:
1277:
1276:
1267:
1262:
1260:
1257:
1256:
1253:
1252:
1243:
1238:
1236:
1230:
1229:
1226:
1225:
1218:
1213:
1211:
1205:
1204:
1201:
1200:
1193:
1188:
1186:
1183:
1182:
1179:
1178:
1169:
1164:
1162:
1152:
1145:
1144:
1141:
1140:
1136:
1135:
1132:
1131:
1127:
1126:
1123:
1122:
1118:
1117:
1114:
1113:
1109:
1108:
1105:
1104:
1100:
1099:
1096:
1095:
1091:
1090:
1087:
1086:
1082:
1081:
1078:
1077:
1073:
1072:
1069:
1068:
1064:
1063:
1060:
1059:
1055:
1054:
1051:
1050:
1046:
1045:
1042:
1041:
1033:
1030:
1029:
1026:
1025:
1016:
1011:
1009:
1006:
1005:
1002:
1001:
997:
996:
993:
992:
988:
987:
984:
983:
974:
971:
970:
967:
966:
957:
952:
950:
947:
946:
943:
942:
933:
928:
926:
921:
919:
916:
915:
912:
911:
902:
897:
895:
889:
888:
885:
884:
877:
872:
870:
864:
863:
860:
859:
850:
845:
843:
840:
839:
836:
835:
826:
821:
819:
816:
815:
812:
811:
802:
797:
795:
793:Neornithischia
789:
788:
785:
784:
777:
774:
773:
770:
769:
760:
755:
753:
747:
746:
743:
742:
733:
728:
726:
723:
722:
719:
718:
711:
706:
704:
701:
700:
697:
696:
687:
682:
680:
670:
635:incertae sedis
600:
597:
569:nasal foramina
473:
470:
449:
446:
444:
441:
344:The first two
341:
338:
301:
300:
293:
279:
278:
272:
271:
263:
261:
257:
256:
233:
229:
228:
226:Jeholosauridae
220:
216:
215:
213:Neornithischia
207:
200:
199:
191:
184:
183:
178:
171:
170:
165:
161:
160:
155:
151:
150:
145:
141:
140:
127:
126:
122:
121:
113:
112:
104:
103:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
41:
28:
15:
13:
10:
9:
6:
4:
3:
2:
4043:
4032:
4029:
4027:
4024:
4022:
4019:
4017:
4014:
4012:
4009:
4007:
4004:
4003:
4001:
3984:
3979:
3975:
3971:
3966:
3962:
3958:
3953:
3949:
3945:
3940:
3936:
3932:
3927:
3923:
3918:
3912:
3908:
3903:
3897:
3893:
3892:
3890:
3888:
3884:
3880:
3875:
3855:
3852:
3851:
3849:
3845:
3839:
3836:
3834:
3831:
3829:
3826:
3824:
3821:
3819:
3816:
3815:
3813:
3809:
3803:
3802:
3798:
3796:
3795:
3791:
3788:
3787:
3786:Dravidosaurus
3783:
3781:
3780:
3776:
3775:
3773:
3771:
3770:
3765:
3759:
3758:
3754:
3752:
3751:
3747:
3745:
3744:
3740:
3738:
3737:
3736:Syngonosaurus
3733:
3731:
3730:
3726:
3724:
3723:
3719:
3717:
3716:
3712:
3710:
3709:
3705:
3703:
3702:
3698:
3696:
3695:
3691:
3689:
3688:
3684:
3682:
3681:
3677:
3675:
3674:
3670:
3668:
3667:
3663:
3662:
3660:
3658:
3657:
3652:
3648:
3641:
3637:
3628:
3624:
3621:
3614:
3608:
3601:
3572:
3568:
3567:
3566:
3565:
3563:
3561:
3557:
3549:
3545:
3544:
3543:
3542:
3540:
3538:
3534:
3531:
3529:
3525:
3517:
3513:
3512:
3511:
3510:
3508:
3506:
3502:
3496:
3495:
3494:Zephyrosaurus
3491:
3488:
3487:
3483:
3481:
3480:
3476:
3474:
3473:
3469:
3467:
3466:
3465:Oryctodromeus
3462:
3460:
3459:
3455:
3453:
3452:
3451:Nevadadromeus
3448:
3445:
3444:
3440:
3437:
3436:
3435:Koreanosaurus
3432:
3429:
3428:
3424:
3421:
3420:
3416:
3414:
3413:
3409:
3406:
3405:
3401:
3398:
3397:
3393:
3391:
3390:
3386:
3385:
3383:
3381:
3377:
3371:
3370:
3366:
3364:
3363:
3359:
3357:
3356:
3352:
3350:
3349:
3345:
3343:
3342:
3338:
3336:
3335:
3331:
3329:
3328:
3324:
3322:
3321:
3320:Hypsilophodon
3317:
3315:
3314:
3313:Hexinlusaurus
3310:
3308:
3307:
3303:
3302:
3299:
3296:
3293:
3288:
3272:
3268:
3267:
3266:
3265:
3263:
3261:
3257:
3249:
3245:
3244:
3243:
3242:
3240:
3238:
3234:
3228:
3227:
3223:
3221:
3220:
3216:
3214:
3213:
3209:
3207:
3206:
3205:Scelidosaurus
3202:
3200:
3199:
3195:
3194:
3191:
3188:
3186:
3182:
3176:
3175:
3171:
3169:
3168:
3164:
3162:
3161:
3157:
3155:
3154:
3150:
3148:
3147:
3143:
3141:
3140:
3136:
3134:
3133:
3129:
3127:
3126:
3125:Abrictosaurus
3122:
3121:
3119:
3117:
3113:
3106:
3105:
3101:
3098:
3097:
3093:
3090:
3089:
3085:
3082:
3081:
3077:
3074:
3073:
3069:
3066:
3065:
3061:
3058:
3057:
3053:
3051:
3050:
3049:Lesothosaurus
3046:
3044:
3043:
3042:Laquintasaura
3039:
3036:
3035:
3031:
3028:
3027:
3023:
3020:
3019:
3015:
3012:
3011:
3010:Eucoelophysis
3007:
3005:
3004:
3000:
2997:
2996:
2992:
2989:
2988:
2984:
2981:
2980:
2976:
2975:
2972:
2968:
2965:
2962:
2957:
2953:
2949:
2944:
2940:
2919:
2918:
2917:
2916:
2914:
2912:
2908:
2900:
2896:
2895:
2894:
2893:
2890:
2887:
2885:
2881:
2877:
2869:
2865:
2862:
2860:
2856:
2853:
2851:
2847:
2845:
2841:
2839:
2835:
2834:
2829:
2825:
2821:
2814:
2809:
2807:
2802:
2800:
2795:
2794:
2791:
2784:
2773:
2763:
2758:
2754:
2750:
2746:
2742:
2735:
2728:
2725:
2720:
2716:
2712:
2708:
2704:
2700:
2696:
2692:
2685:
2682:
2678:
2674:
2668:
2665:
2660:
2656:
2652:
2648:
2644:
2640:
2636:
2632:
2625:
2622:
2618:
2612:
2610:
2606:
2601:
2597:
2593:
2589:
2585:
2581:
2574:
2571:
2565:
2563:
2561:
2559:
2557:
2555:
2553:
2549:
2542:
2535:
2534:
2526:
2525:
2517:
2516:
2508:
2507:
2499:
2498:
2490:
2489:
2481:
2480:
2472:
2471:
2463:
2462:
2454:
2453:
2445:
2444:
2436:
2435:
2427:
2426:
2418:
2417:
2409:
2408:
2405:
2404:
2398:
2397:
2394:
2393:
2390:
2389:
2388:
2387:Tenontosaurus
2381:
2380:
2374:
2373:
2370:
2369:
2366:
2365:
2364:
2357:
2356:
2350:
2349:
2346:
2345:
2342:
2341:
2335:
2334:
2328:
2325:
2324:
2321:
2320:
2317:
2316:
2315:
2314:Hypsilophodon
2308:
2307:
2301:
2298:
2297:
2294:
2293:
2285:
2284:
2276:
2275:
2267:
2266:
2263:
2262:
2261:
2254:
2253:
2250:
2249:
2246:
2245:
2244:
2237:
2236:
2233:
2232:
2229:
2228:
2227:
2220:
2219:
2216:
2215:
2212:
2211:
2210:
2203:
2202:
2196:
2195:
2192:
2191:
2183:
2182:
2179:
2178:
2177:
2170:
2169:
2166:
2165:
2162:
2161:
2160:
2153:
2152:
2146:
2145:
2139:
2138:
2135:
2134:
2131:
2130:
2129:
2122:
2121:
2115:
2112:
2111:
2105:
2104:
2101:
2100:
2097:
2096:
2095:
2088:
2087:
2081:
2080:
2076:
2075:
2072:
2071:
2068:
2067:
2066:
2059:
2058:
2052:
2051:
2047:
2046:
2043:
2042:
2034:
2033:
2025:
2024:
2016:
2015:
2007:
2006:
1998:
1997:
1994:
1993:
1992:
1985:
1984:
1981:
1980:
1977:
1976:
1975:
1968:
1967:
1961:
1960:
1957:
1956:
1953:
1952:
1951:
1950:Zephyrosaurus
1944:
1943:
1937:
1936:
1933:
1932:
1924:
1923:
1920:
1919:
1918:
1917:Koreanosaurus
1911:
1910:
1907:
1906:
1903:
1902:
1901:
1894:
1893:
1887:
1886:
1880:
1877:
1876:
1873:
1872:
1864:
1863:
1855:
1854:
1851:
1850:
1849:
1842:
1841:
1838:
1837:
1834:
1833:
1832:
1825:
1824:
1818:
1817:
1814:
1813:
1810:
1809:
1808:
1801:
1800:
1794:
1791:
1790:
1784:
1783:
1780:
1779:
1776:
1775:
1774:
1767:
1766:
1760:
1759:
1753:
1752:
1749:
1748:
1740:
1739:
1736:
1735:
1734:
1727:
1726:
1723:
1722:
1719:
1718:
1717:
1716:Hexinlusaurus
1710:
1709:
1703:
1702:
1696:
1695:
1692:
1691:
1688:
1687:
1686:
1679:
1678:
1672:
1671:
1668:
1667:
1664:
1663:
1662:
1661:Lesothosaurus
1655:
1654:
1648:
1647:
1643:
1642:
1639:
1638:
1635:
1634:
1633:
1626:
1625:
1619:
1618:
1615:
1614:
1611:
1610:
1609:
1602:
1601:
1595:
1594:
1591:
1590:
1587:
1586:
1580:
1579:
1573:
1572:
1568:
1567:
1563:
1561:
1556:
1554:
1550:
1546:
1542:
1538:
1534:
1524:
1523:
1515:
1514:
1506:
1505:
1497:
1496:
1488:
1487:
1479:
1478:
1470:
1469:
1461:
1460:
1457:
1456:
1450:
1449:
1446:
1445:
1442:
1441:
1435:
1434:
1431:
1430:
1422:
1421:
1418:
1417:
1416:
1415:Zephyrosaurus
1409:
1408:
1405:
1404:
1401:
1400:
1399:
1392:
1391:
1385:
1384:
1381:
1380:
1377:
1376:
1375:
1368:
1367:
1364:
1363:
1360:
1359:
1358:
1351:
1350:
1347:
1346:
1343:
1342:
1341:
1340:Hypsilophodon
1334:
1333:
1330:
1329:
1326:
1325:
1324:
1317:
1316:
1310:
1307:
1306:
1303:
1302:
1299:
1298:
1297:
1296:Hexinlusaurus
1290:
1289:
1283:
1282:
1279:
1278:
1275:
1274:
1273:
1266:
1265:
1259:
1258:
1255:
1254:
1251:
1250:
1249:
1242:
1241:
1235:
1232:
1231:
1228:
1227:
1224:
1223:
1217:
1216:
1210:
1207:
1206:
1203:
1202:
1199:
1198:
1192:
1191:
1185:
1184:
1181:
1180:
1177:
1176:
1175:
1168:
1167:
1161:
1160:
1156:
1155:
1151:
1143:
1142:
1134:
1133:
1125:
1124:
1116:
1115:
1107:
1106:
1098:
1097:
1089:
1088:
1080:
1079:
1071:
1070:
1062:
1061:
1053:
1052:
1044:
1043:
1040:
1039:
1032:
1031:
1028:
1027:
1024:
1023:
1022:
1021:Hypsilophodon
1015:
1014:
1008:
1007:
1004:
1003:
995:
994:
986:
985:
982:
981:
980:
973:
972:
969:
968:
965:
964:
963:
956:
955:
949:
948:
945:
944:
941:
940:
939:
932:
931:
925:
924:
918:
917:
914:
913:
910:
909:
908:
901:
900:
894:
891:
890:
887:
886:
883:
882:
876:
875:
869:
866:
865:
862:
861:
858:
857:
856:
855:Hexinlusaurus
849:
848:
842:
841:
838:
837:
834:
833:
832:
825:
824:
818:
817:
814:
813:
810:
809:
808:
801:
800:
794:
791:
790:
787:
786:
783:
782:
776:
775:
772:
771:
768:
767:
766:
765:Lesothosaurus
759:
758:
752:
749:
748:
745:
744:
741:
740:
739:
732:
731:
725:
724:
721:
720:
717:
716:
710:
709:
703:
702:
699:
698:
695:
694:
693:
686:
685:
679:
678:
674:
673:
669:
667:
663:
659:
654:
652:
648:
644:
643:Euornithopoda
641:
637:
636:
632:
628:
623:
621:
617:
613:
609:
608:ornithischian
605:
598:
596:
594:
590:
586:
582:
578:
577:quadratojugal
574:
570:
566:
561:
559:
555:
551:
547:
543:
539:
535:
531:
527:
522:
518:
513:
511:
501:
494:
489:
482:
478:
471:
469:
467:
463:
459:
454:
447:
442:
440:
437:
433:
429:
425:
423:
418:
414:
409:
407:
406:specific name
403:
400:and northern
399:
395:
391:
387:
383:
379:
375:
371:
367:
363:
359:
355:
351:
347:
339:
337:
335:
331:
327:
324:
320:
317:
313:
309:
308:
297:
291:
289:
280:
277:
276:Binomial name
273:
269:
268:
262:
259:
258:
254:
250:
247:
242:
241:
234:
231:
230:
227:
221:
218:
217:
214:
208:
205:
202:
201:
198:
192:
189:
186:
185:
182:
179:
176:
173:
172:
169:
166:
163:
162:
159:
156:
153:
152:
149:
146:
143:
142:
137:
132:
128:
123:
119:
114:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
39:
36:126β125
32:
26:
22:
19:
4026:Yixian fauna
3917:Jeholosaurus
3887:Jeholosaurus
3886:
3828:Silesauridae
3799:
3792:
3784:
3779:Ajkaceratops
3777:
3767:
3755:
3748:
3741:
3734:
3727:
3722:Notoceratops
3720:
3713:
3708:Gongbusaurus
3706:
3701:Geranosaurus
3699:
3692:
3685:
3678:
3671:
3664:
3656:Nomina dubia
3654:
3604:
3492:
3484:
3477:
3472:Parksosaurus
3470:
3463:
3456:
3449:
3441:
3433:
3427:Jeholosaurus
3426:
3425:
3417:
3410:
3402:
3394:
3387:
3367:
3360:
3355:Sanxiasaurus
3353:
3346:
3339:
3334:Minimocursor
3332:
3325:
3318:
3311:
3304:
3248:Ankylosauria
3237:Ankylosauria
3224:
3217:
3210:
3203:
3196:
3172:
3165:
3158:
3151:
3144:
3137:
3130:
3123:
3104:Technosaurus
3102:
3096:Soumyasaurus
3094:
3086:
3078:
3072:Pisanosaurus
3070:
3062:
3054:
3047:
3040:
3032:
3026:Ignotosaurus
3024:
3016:
3008:
3001:
2993:
2985:
2977:
2961:Ornithischia
2948:Ornithischia
2911:Ornithischia
2863:
2854:
2820:Ornithischia
2744:
2740:
2727:
2694:
2690:
2684:
2676:
2672:
2667:
2634:
2630:
2624:
2616:
2583:
2579:
2573:
2401:
2385:
2384:
2361:
2360:
2338:
2327:Iguanodontia
2312:
2311:
2260:Diluvicursor
2258:
2257:
2241:
2240:
2224:
2223:
2207:
2206:
2174:
2173:
2157:
2156:
2126:
2125:
2094:Parksosaurus
2092:
2091:
2077:
2063:
2062:
2048:
1989:
1988:
1972:
1971:
1948:
1947:
1915:
1914:
1898:
1897:
1846:
1845:
1831:Jeholosaurus
1830:
1829:
1828:
1805:
1804:
1771:
1770:
1731:
1730:
1714:
1713:
1683:
1682:
1659:
1658:
1644:
1630:
1629:
1606:
1605:
1583:
1571:Ornithischia
1569:
1559:
1557:
1548:
1544:
1541:Jeholosaurus
1540:
1536:
1532:
1530:
1453:
1438:
1413:
1412:
1396:
1395:
1372:
1371:
1357:Jeholosaurus
1356:
1355:
1354:
1338:
1337:
1321:
1320:
1294:
1293:
1270:
1269:
1246:
1245:
1220:
1195:
1174:Pisanosaurus
1172:
1171:
1159:Ornithischia
1157:
1149:
1038:Iguanodontia
1035:
1019:
1018:
979:Jeholosaurus
978:
977:
976:
960:
959:
936:
935:
905:
904:
879:
853:
852:
829:
828:
805:
804:
779:
763:
762:
736:
735:
713:
692:Pisanosaurus
690:
689:
677:Ornithischia
675:
665:
655:
633:
631:Ornithischia
627:Jeholosaurus
626:
624:
604:Jeholosaurus
603:
602:
585:premaxillary
565:Jeholosaurus
564:
562:
526:Jeholosaurus
525:
514:
512:individual.
506:
493:Jeholosaurus
492:
480:
453:Jeholosaurus
452:
451:
420:
417:Jeholosaurus
416:
415:specimen of
410:
390:Jeholosaurus
389:
385:
374:Wang Xioalin
358:Beipiao City
346:Jeholosaurus
345:
343:
307:Jeholosaurus
306:
305:
304:
295:
287:
282:
266:
265:
248:
240:Jeholosaurus
239:
238:
203:
197:Ornithischia
187:
174:
25:Jeholosaurus
24:
18:
3911:Wikispecies
3757:Trimucrodon
3729:Siluosaurus
3687:Fabrosaurus
3680:Craspedodon
3673:Bienosaurus
3516:Ornithopoda
3505:Ornithopoda
3404:Changmiania
3369:Yandusaurus
3306:Agilisaurus
3271:Stegosauria
3260:Stegosauria
3185:Thyreophora
3064:Lutungutali
3034:Kwanasaurus
2987:Asilisaurus
2979:Amanasaurus
2586:: 399β406.
2300:Clypeodonta
2079:Ornithopoda
1733:Yandusaurus
1685:Agilisaurus
1632:Thyreophora
1440:Ornithopoda
1374:Yandusaurus
1272:Agilisaurus
1248:Stormbergia
1222:Thyreophora
893:Ornithopoda
831:Agilisaurus
807:Stormbergia
781:Thyreophora
589:posteriorly
558:metatarsals
503:Restoration
443:Description
330:herbivorous
4000:Categories
3811:Other taxa
3750:Tichosteus
3743:Tatisaurus
3548:Ceratopsia
3537:Ceratopsia
3486:Yueosaurus
3458:Orodromeus
3443:Nanosaurus
3362:Xiaosaurus
3341:Nanosaurus
3219:Yuxisaurus
3198:Emausaurus
3174:Tianyulong
3167:Pegomastax
3153:Lycorhinus
3139:Fruitadens
3088:Silesaurus
3080:Sacisaurus
3056:Lewisuchus
2920:see belowβ
2868:Dinosauria
2850:Sauropsida
2543:References
2403:Dryomorpha
2243:Anabisetia
2128:Talenkauen
1974:Yueosaurus
1900:Orodromeus
1773:Nanosaurus
1398:Orodromeus
1209:Genasauria
907:Orodromeus
751:Genasauria
645:: a small
581:predentary
550:trochanter
530:omnivorous
517:premaxilla
334:ornithopod
323:Cretaceous
181:Dinosauria
3715:Laosaurus
3348:Phyllodon
3132:Echinodon
3018:Gamatavus
2836:Kingdom:
2114:Elasmaria
1323:Othnielia
662:cladogram
658:cladistic
599:Phylogeny
538:palpebrae
491:Skull of
378:You Hailu
260:Species:
154:Kingdom:
148:Eukaryota
3896:Wikidata
3666:Alocodon
3644:See also
3160:Manidens
3003:Eocursor
2995:Diodorus
2844:Chordata
2842:Phylum:
2838:Animalia
2719:86754247
2659:86257644
2050:Cerapoda
1608:Eocursor
1562:, 2019.
1234:Cerapoda
868:Cerapoda
738:Eocursor
668:, 2011.
554:extensor
534:foramina
510:juvenile
458:mandible
398:Liaoning
354:Lujiatun
319:dinosaur
219:Family:
168:Chordata
164:Phylum:
158:Animalia
144:Domain:
3970:4129093
3957:1229533
3944:4823034
3931:4532026
3902:Q137694
3794:Jakapil
3226:Jakapil
2848:Class:
2749:Bibcode
2699:Bibcode
2639:Bibcode
2588:Bibcode
616:ischium
593:angular
521:maxilla
472:Anatomy
370:Xu Xing
285:†
264:†
232:Genus:
105:↓
2717:
2673:Nature
2657:
1560:et al.
1547:, and
1537:et al.
1533:et al.
666:et al.
656:Later
606:is an
432:Aptian
404:. The
380:. The
332:small
326:Period
298:, 2000
296:et al.
249:et al.
3983:68236
3952:IRMNG
2864:Clade
2855:Clade
2737:(PDF)
2715:S2CID
2655:S2CID
640:basal
612:pubis
466:tibia
462:femur
402:Hebei
394:Jehol
366:China
356:near
312:genus
310:is a
204:Clade
188:Clade
175:Clade
3939:GBIF
3569:see
3546:see
3514:see
3419:Haya
3412:Fona
3269:see
3246:see
2897:see
1807:Haya
1545:Haya
1531:Han
938:Haya
571:; a
544:and
528:was
515:The
448:Size
436:tuff
422:IVPP
413:type
411:The
382:type
376:and
350:2000
253:2000
44:Preκ
3926:EoL
2757:doi
2707:doi
2677:458
2647:doi
2596:doi
2584:487
1036:to
352:at
314:of
294:Xu
4002::
3980::
3967::
3954::
3941::
3928::
3913::
3898::
2866::
2857::
2755:.
2745:93
2743:.
2739:.
2713:.
2705:.
2695:32
2693:.
2675:,
2653:.
2645:.
2635:31
2633:.
2608:^
2594:.
2582:.
2551:^
1555:.
1543:,
372:,
364:,
360:,
336:.
251:,
246:Xu
206::
190::
177::
94:Pg
38:Ma
33:,
3789:?
3489:?
3446:?
3438:?
3430:?
3422:?
3407:?
3399:?
3107:?
3099:?
3091:?
3083:?
3075:?
3067:?
3059:?
3037:?
3029:?
3021:?
3013:?
2998:?
2990:?
2982:?
2812:e
2805:t
2798:v
2765:.
2759::
2751::
2721:.
2709::
2701::
2661:.
2649::
2641::
2602:.
2598::
2590::
495:.
236:β
223:β
210:β
194:β
99:N
89:K
84:J
79:T
74:P
69:C
64:D
59:S
54:O
49:κ
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