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Jenny Clack

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Clack is best known for her extensive body of work on early tetrapods, much of which redefined how paleontologists conceived of the evolution of limbs and other features associated with tetrapods' transition from other lobe-finned fishes. She began her career working primarily on the ear of early
353:", a major gap in the fossil record of early tetrapods and one that she subsequently began to fill in collaboration with other paleontologists. Clack also undertook extensive fieldwork expeditions in order to search for further remains of early tetrapods. In 1987, during an expedition to 275:
in 1970. In 1978, she accepted an invitation from Alec Panchen to obtain a Ph.D. from the same university, at the encouragement of her then new boyfriend Rob Clack. They married in 1980. Her doctorate was completed in 1984. She also held a Graduate Certificate in Museum Studies from the
1137:"Clack, Prof. Jennifer Alice, (born 3 Nov. 1947), Curator in Vertebrate Palaeontology, 2005–15, and Professor, 2006–15, now Professor Emeritus of Vertebrate Palaeontology, University Museum of Zoology, Cambridge; Fellow, Darwin College, Cambridge, 1997–2015, now Emeritus" 3140: 3095:
Clack, Jennifer Alice; Challands, Thomas James; Smithson, Timothy Richard; Smithson, Keturah Zoe (2 November 2018). "Newly recognized Famennian lungfishes from East Greenland reveal tooth plate diversity and blur the Devonian–Carboniferous boundary".
1003:. The university described her as "an internationally preeminent palaeontologist whose research has profoundly changed the understanding of the origin of terrestrial vertebrate life." Also in 2013, she was awarded the T Neville George Medal by the 408: 3296:
Clack, Jennifer A.; Bennett, Carys E.; Carpenter, David K.; Davies, Sarah J.; Fraser, Nicholas C.; Kearsey, Timothy I.; Marshall, John E. A.; Millward, David; Otoo, Benjamin K. A.; Reeves, Emma J.; Ross, Andrew J. (5 December 2016).
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Clack, Jennifer A.; Bennett, Carys E.; Carpenter, David K.; Davies, Sarah J.; Fraser, Nicholas C.; Kearsey, Timothy I.; Marshall, John E. A.; Millward, David; Otoo, Benjamin K. A.; Reeves, Emma J.; Ross, Andrew J. (5 December 2016).
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by the University of Cambridge, and took the title Professor of Vertebrate Palaeontology. She retired in 2015 and became Emeritus Professor of Vertebrate Palaeontology at the museum. In 1997, Clack was elected a
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book, Clack also co-authored a volume of the Handbuch der Paläoherpetologie series on early tetrapods with Andrew Milner in 2015 and co-edited a volume on the evolution of hearing in 2016.
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and is one of the most published vertebrate paleontologists in Nature, arguably the leading scientific journal in the world, with over 15 papers in that journal alone. In addition to her
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Clément, Gaël; Ahlberg, Per E.; Blieck, Alain; Blom, Henning; Clack, Jennifer A.; Poty, Edouard; Thorez, Jacques; Janvier, Philippe (2004). "Devonian tetrapod from western Europe".
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Clack, J. A. (1983). "The stapes of the Coal Measures embolomere Pholiderpeton scutigerum Huxley (Amphibia: Anthracosauria) and otic evolution in early tetrapods".
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Challands, Tom J.; Smithson, Timothy R.; Clack, Jennifer A.; Bennett, Carys E.; Marshall, John E. A.; Wallace-Johnson, Sarah M.; Hill, Henrietta (11 March 2019).
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Clack, J. A.; Ahlberg, P. E.; Finney, S. M.; Dominguez Alonso, P.; Robinson, J.; Ketcham, R. A. (2003). "A uniquely specialized ear in a very early tetrapod".
212:, specifically studying the "fish to tetrapod" transition: the origin, evolutionary development and radiation of early tetrapods and their relatives among the 4294: 3514:
Clack, J A (1 April 2003). "A new baphetid (stem tetrapod) from the Upper Carboniferous of Tyne and Wear, U.K., and the evolution of the tetrapod occiput".
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Pierce, Stephanie E.; Ahlberg, Per E.; Hutchinson, John R.; Molnar, Julia L.; Sanchez, Sophie; Tafforeau, Paul; Clack, Jennifer A. (13 January 2013).
1459: 3421:"A new genus of Devonian tetrapod from North-East Greenland, with new information on the lower jaw of Ichthyostega: A NEW GENUS OF DEVONIAN TETRAPOD" 3189:
Richards, Kelly R.; Sherwin, Janet E.; Smithson, Timothy R.; Bennion, Rebecca F.; Davies, Sarah J.; Marshall, John E. A.; Clack, Jennifer A. (2017).
473:, a set of documentaries about scientists who have made important discoveries. This may be viewed on YouTube. Clack was honored by her peers with a 4269: 189: 33: 3236:
Smithson, Timothy R.; Browne, Michael A. E.; Davies, Sarah J.; Marshall, John E. A.; Millward, David; Walsh, Stig A.; Clack, Jennifer A. (2017).
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Ahlberg, Per E.; Clack, Jennifer A.; Luks̆evic̆s, Ervīns (1996). "Rapid braincase evolution between Panderichthys and the earliest tetrapods".
1606:"A Tournaisian (earliest Carboniferous) conglomerate-preserved non-marine faunal assemblage and its environmental and sedimentological context" 305: 224: 152: 2604:
Callier, V.; Clack, J. A.; Ahlberg, P. E. (17 April 2009). "Contrasting Developmental Trajectories in the Earliest Known Tetrapod Forelimbs".
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Otoo, Benjamin K. A.; Clack, Jennifer A.; Smithson, Timothy R.; Bennett, Carys E.; Kearsey, Timothy I.; Coates, Michael I. (3 October 2018).
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Clack, Jennifer A. (2011). "A new microsaur from the early carboniferous (Viséan) of East Kirkton, Scotland, showing soft tissue evidence".
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Clack, Jennifer A.; Bennett, Carys E.; Davies, Sarah J.; Scott, Andrew C.; Sherwin, Janet E.; Smithson, Timothy R. (3 January 2019).
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Clack supervised many graduate students who went on to pursue successful careers in paleontology and evolutionary biology, including
4279: 1015: 338: 272: 228: 235:, where she devoted her career to studying the early development of tetrapods, the "four-legged" animals said to have evolved from 345:
tetrapods and later expanded to more broadly addressing the osteology and evolution of tetrapods. Together with Michael Coates (
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Clack, J. A. (1993). "Silvanerpeton miripedes, a new anthracosauroid from the Viséan of East Kirkton, West Lothian, Scotland".
1171:"Fossils, function and phylogeny: Papers on early vertebrate evolution in honour of Professor Jennifer A. Clack – Introduction" 384:
Most recently, she led a major consortium project (TW:eed) investigating some exciting new fossils from Northumberland and the
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in 1970. Additional surveys in 1998 led to the collection of substantial new material, including what is now recognized as
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Ahlberg, Per Erik; Clack, Jennifer A.; Blom, Henning (2005). "The axial skeleton of the Devonian tetrapod Ichthyostega".
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Over the course of her lengthy career, Clack published in some of the most notable scientific journals, including
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period; this project has produced numerous publications furthering the understanding of early tetrapod evolution.
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Johanson, Zerina; Jeffery, Jonathan; Challands, Tom; Pierce, Stephanie; Clack, Jennifer (27 October 2020).
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Clack, Jennifer A. (1998). "A new Early Carboniferous tetrapod with a mélange of crown-group characters".
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Beznosov, Pavel A.; Clack, Jennifer A.; Lukševičs, Ervīns; Ruta, Marcello; Ahlberg, Per Erik (2019).
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Coates, M. I.; Clack, J. A. (1991). "Fish-like gills and breathing in the earliest known tetrapod".
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Coates, Michael I.; Clack, Jennifer A. (1995). "Romer's gap: Tetrapod origins and terrestriality".
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Clack was born on 3 November 1947, the only child of Ernest and Alice Agnew. She was brought up in
3631: 1518:"A fish and tetrapod fauna from Romer's Gap preserved in Scottish Tournaisian floodplain deposits" 1210: 1110: 4222: 4171: 4098: 4081: 4076: 3720: 3676: 3582: 3488: 3337: 3218: 3168: 3121: 3077: 3050:"A lungfish survivor of the end-Devonian extinction and an Early Carboniferous dipnoan radiation" 2965: 2886: 2835: 2816:. Springer Handbook of Auditory Research. Vol. 59. Cham: Springer International Publishing. 2645: 2586: 2535: 2401: 2344: 2278: 2215: 2152: 2095: 2036: 1985: 1926: 1860: 1806: 1755: 1704: 1586: 1498: 959: 875: 843: 532: 452: 3968: 3885: 4202: 4192: 3893: 3574: 3539: 3401: 3329: 3321: 3269: 3210: 3160: 3113: 3069: 3030: 3012: 2878: 2825: 2769: 2751: 2712: 2694: 2637: 2629: 2578: 2570: 2527: 2519: 2466: 2448: 2393: 2385: 2336: 2328: 2270: 2262: 2207: 2199: 2144: 2136: 2087: 2079: 2028: 2020: 1977: 1969: 1918: 1910: 1852: 1798: 1747: 1696: 1688: 1645: 1627: 1578: 1539: 1490: 1482: 1414: 1320: 1192: 1136: 1055: 996: 908: 404: 289: 3874: 2297: 2170:
Ahlberg, Per E.; Clack, Jennifer A.; Lukševičs, Ervīns; Blom, Henning; Zupiņš, Ivars (2008).
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Clack, Jennifer A.; Ahlberg, Erik (2004). Arratia, G.; Wilson, M.V.H.; Cloutier, R. (eds.).
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Neenan, James M.; Ruta, Marcello; Clack, Jennifer A.; Rayfield, Emily J. (22 April 2014).
993: 326: 2665:"Earliest Carboniferous tetrapod and arthropod faunas from Scotland populate Romer's Gap" 2171: 3694: 3527: 3436: 3387: 3191:"Diverse and durophagous: Early Carboniferous chondrichthyans from the Scottish Borders" 3000: 2953: 2866: 2853:
Coates, M. I.; Clack, J. A. (1990). "Polydactyly in the earliest known tetrapod limbs".
2680: 2617: 2507: 2436: 2421:"Life history of the stem tetrapod Acanthostega revealed by synchrotron microtomography" 2316: 2250: 2187: 2067: 1957: 1898: 1840: 1786: 1735: 1676: 1400: 4217: 4197: 4028: 4023: 3983: 3025: 2984: 2764: 2731: 2707: 2664: 2461: 2420: 1640: 1605: 1316: 1026: 621: 448: 354: 350: 313: 3190: 3049: 1558: 4253: 4003: 3963: 3953: 3920: 3680: 3586: 3445: 3420: 3222: 3172: 3125: 2969: 2539: 1864: 1590: 1409: 1384: 515: 393: 256: 240: 3341: 3081: 2839: 2649: 2590: 1502: 422:
had more than five toes per foot: the Upper Devonian tetrapods from East Greenland,
4187: 3998: 2890: 2405: 2348: 2282: 2219: 2156: 2099: 2040: 1989: 1810: 1759: 1708: 576: 441: 430: 424: 368: 362: 220:, published in 2002 (second edition, 2012) and written with the layperson in mind. 201: 135: 3065: 2961: 2234: 1930: 3835:"Professor Jenny Clack awarded the Palaeontological Association's Lapworth Medal" 3238:"A new Mississippian tetrapod from Fife, Scotland, and its environmental context" 2942:"A New Look At Carboniferous Rhizodontid Humeri (Sarcopterygii; Tetrapodomorpha)" 1085: 4207: 4159: 4132: 4115: 3958: 1141: 474: 389: 2663:
Smithson, T. R.; Wood, S. P.; Marshall, J. E. A.; Clack, J. A. (5 March 2012).
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Earth and Environmental Science Transactions of the Royal Society of Edinburgh
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Earth and Environmental Science Transactions of the Royal Society of Edinburgh
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Earth and Environmental Science Transactions of the Royal Society of Edinburgh
2821: 2732:"Feeding biomechanics in Acanthostega and across the fish–tetrapod transition" 2515: 2419:
Sanchez, Sophie; Tafforeau, Paul; Clack, Jennifer A.; Ahlberg, Per E. (2016).
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Earth and Environmental Science Transactions of the Royal Society of Edinburgh
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Earth and Environmental Science Transactions of the Royal Society of Edinburgh
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Clack, Jennifer A.; Ahlberg, Per E.; Blom, Henning; Finney, Sarah M. (2012).
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Clack, J. A. (2 April 2004). "PALEONTOLOGY: Enhanced: From Fins to Fingers".
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Pierce, Stephanie E.; Clack, Jennifer A.; Hutchinson, John R. (23 May 2012).
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CLACK, JENNIFER A.; AHLBERG, PER E.; BLOM, HENNING; FINNEY, SARAH M. (2012).
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On 17 July 2014, she was awarded an honorary Doctor of Science degree by the
467:). In April 2012 she was featured in an episode of the BBC television series 3772: 3372:"Lungfish diversity in Romer's Gap: reaction to the end-Devonian extinction" 3317: 2904: 2689: 2625: 2566: 1478: 958:
Clack died on 26 March 2020 at the age of 72, after a five-year battle with
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following up on field notes of researchers who had collected material of
236: 209: 3299:"Phylogenetic and environmental context of a Tournaisian tetrapod fauna" 3008: 2489:"Morphology of the earliest reconstructable tetrapod Parmastega aelidae" 2444: 2324: 2258: 2195: 2075: 1965: 1906: 1663:
Clack, J. A. (1989). "Discovery of the earliest-known tetrapod stapes".
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by Cambridge, taking the title Professor of Vertebrate Palaeontology.
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Smithson, Timothy R.; Richards, Kelly R.; Clack, Jennifer A. (2016).
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had eight (compared to the six toes of the Russian Devonian tetrapod
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A new stem tetrapod from the Early Carboniferous of Northern Ireland
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Clack, Jennifer A.; Fay, Richard R; Popper, Arthur N., eds. (2016).
2380: 2363: 2131: 2114: 2016: 3864: 3837:. University of Cambridge - Department of Zoology. 18 December 2015 1848: 3747:"University to bestow five honorary degrees at 515th Convocation" 3889: 3880:
Jenny Clack, Paleontologist: The First Vertebrate Walks on Land
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Clack is probably best known for discovering that the earliest
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Ruta, Marcello; Ahlberg, Per E.; Smithson, Timothy R. (2018).
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lobe-finned fishes and colonised the freshwater swamps of the
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English paleontologist and evolutionary biologist (1947–2020)
2172:"Ventastega curonica and the origin of tetrapod morphology" 1340:
Bulletin du Muséum National d'Histoire Naturelle, Section C
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Gaining Ground: the Origin and Early Evolution of Tetrapods
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Gaining Ground: the Origin and Early Evolution of Tetrapods
3483:. MĂĽnchen: Verlag Dr. Friedrich Pfeil. pp. 309–320. 1079: 1077: 2736:
Proceedings of the Royal Society B: Biological Sciences
2298:"Vertebral architecture in the earliest stem tetrapods" 1111:"Congregation of the Regent House on 9 December 2000" 4320:
Fellows of the American Academy of Arts and Sciences
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Biographical Memoirs of Fellows of the Royal Society
1050: 1048: 1046: 1044: 1042: 4231: 4180: 3931: 357:, Clack and her team discovered the remains of the 145: 127: 109: 89: 81: 69: 50: 23: 3655:"Jennifer A. Clack. 3 November 1947—26 March 2020" 200:; 3 November 1947 – 26 March 2020) was an English 3139:Smithson, Timothy R.; Clack, Jennifer A. (2017). 1557:Smithson, Timothy R.; Clack, Jennifer A. (2017). 4315:Members of the Royal Swedish Academy of Sciences 3608:. Department of Zoology, University of Cambridge 1433:"The TWeed project: Evolution's Missing Chapter" 3860:Jenny Clack on the Cambridge University website 3777:University of Cambridge - Department of Zoology 2669:Proceedings of the National Academy of Sciences 539:Beznosov, Clack, LukševiÄŤs, Ruta & Ahlberg 409:Proceedings of the National Academy of Sciences 3630:. National Academy of Sciences. Archived from 2113:Ahlberg, Per Erik; Clack, Jennifer A. (2006). 1358:"Professor Jennifer Clack (Doctor of Science)" 1243:. No. 6396. 23 September 2015. p. 12 3901: 3721:"Featured Scientists | Your Inner Fish | PBS" 1117:. No. 5831. 13 December 2000. p. 27 8: 2983:Ahlberg, Per E.; Clack, Jennifer A. (2020). 1217:. No. 5718. 5 November 1997. p. 24 1145:. Oxford University Press. 1 December 2020. 481:Taxa named by Jennifer Clack and colleagues 208:. She specialised in the early evolution of 3745:Koppes, Steve; Allen, Susie (15 May 2013). 271:. She received a B.Sc. in Zoology from the 3908: 3894: 3886: 444:was not an ancestral trait for tetrapods. 42: 20: 3670: 3444: 3395: 3263: 3253: 3024: 2763: 2706: 2688: 2460: 2379: 2130: 1639: 1621: 1533: 1408: 1305:Zoological Journal of the Linnean Society 1292:. No. 6036. 17 May 2006. p. 17. 1186: 567:Clack, Challands, Smithon & Smithson 337:. On 1 October 2006, Clack was awarded a 3821:"Six new members elected to the Academy" 3653:Ahlberg, P. E.; Smithson, T. R. (2021). 1062:. Oxford University Press. November 2014 978:, the first woman to achieve the honor. 479: 325:; since 1 October 2015, she has been an 288:. On 9 December 2000, she was awarded a 3801:. University of Leicester. 10 July 2014 3602:"Professor Jenny Clack, FRS, 1947-2020" 1364:. University of Leicester. 17 July 2014 1038: 3496: 3486: 1883:"An early tetrapod from 'Romer's Gap'" 992:On 15 June 2013, Clack was awarded an 153:Cambridge University Museum of Zoology 4310:21st-century British women scientists 4275:Alumni of the University of Leicester 3365: 3363: 3361: 3359: 3357: 3355: 3353: 3351: 3291: 3289: 3287: 3285: 3283: 3184: 3182: 2482: 2480: 1876: 1874: 1822: 1820: 987:American Academy of Arts and Sciences 7: 4290:Fellows of Darwin College, Cambridge 1164: 1162: 1160: 1151:10.1093/ww/9780199540884.013.U249891 4295:Female fellows of the Royal Society 3054:Journal of Systematic Palaeontology 2364:"The origin of terrestrial hearing" 349:), Clack defined what is known as " 333:in Vertebrate Palaeontology at the 231:of Vertebrate Palaeontology at the 3865:Home page - Jennifer "Jenny" Clack 3516:Canadian Journal of Earth Sciences 2946:Journal of Vertebrate Paleontology 1317:10.1111/j.1096-3642.1983.tb01163.x 1237:"College Notices – Darwin College" 850:Clack, Ahlberg, Blöm & Finney 14: 1016:Royal Swedish Academy of Sciences 273:University of Newcastle upon Tyne 216:. She is best known for her book 4305:People educated at Bolton School 3600:Aucott, Rachel (26 March 2020). 3446:10.1111/j.1475-4983.2011.01117.x 2115:"A firm step from water to land" 1410:10.1111/j.1475-4983.2011.01117.x 3462:Special Papers in Palaeontology 2814:Evolution of the Vertebrate Ear 1025:'s most prestigious award, the 970:In 2008, Clack was awarded the 834:Smithson, Richards & Clack 820:Smithson, Richards & Clack 806:Smithson, Richards & Clack 792:Smithson, Richards & Clack 778:Smithson, Richards & Clack 764:Smithson, Richards & Clack 747:Smithson, Richards & Clack 730:Smithson, Richards & Clack 255:, England. She was educated at 4270:Alumni of Newcastle University 3306:Nature Ecology & Evolution 2792:Dr. Friedrich Pfeil Publishing 1467:Nature Ecology & Evolution 1: 4285:British women paleontologists 3066:10.1080/14772019.2019.1572234 2962:10.1080/02724634.2020.1813150 1290:Cambridge University Reporter 1241:Cambridge University Reporter 1215:Cambridge University Reporter 1115:Cambridge University Reporter 1084:Clack, Rob (6 October 2009). 1056:"CLACK, Prof. Jennifer Alice" 1005:Geological Society of Glasgow 981:In 2009, Clack was elected a 329:. From 2000 to 2005, she was 4325:Fellows of the Royal Society 3628:"Daniel Giraud Elliot Medal" 1023:Palaeontological Association 976:National Academy of Sciences 306:University Museum of Zoology 3753:. The University of Chicago 2362:Clack, Jennifer A. (2015). 983:Fellow of the Royal Society 4341: 4300:Scientists from Manchester 2989:Royal Society Open Science 2948:. e1813150 (3): e1813150. 2923:"Beautiful Minds: YouTube" 972:Daniel Giraud Elliot Medal 386:Borders Region of Scotland 304:In 1981, Clack joined the 3571:10.1017/s0263593300006179 3207:10.1017/s1755691018000166 3157:10.1017/S1755691018000075 2822:10.1007/978-3-319-46661-3 2516:10.1038/s41586-019-1636-y 1575:10.1017/s1755691018000075 1437:National Museums Scotland 1269:. University of Cambridge 1267:Darwin College, Cambridge 1188:10.1017/S1755691019000057 1086:"Jenny Clack – Biography" 463:), and Matthew Friedman ( 457:University College London 323:Darwin College, Cambridge 223:Clack was curator at the 171: 163:Darwin College, Cambridge 120: 41: 4280:English palaeontologists 4239:William, Prince of Wales 3875:Interview on PBS website 247:Early life and education 3318:10.1038/s41559-016-0002 3242:Papers in Palaeontology 3098:Papers in Palaeontology 2690:10.1073/pnas.1117332109 2626:10.1126/science.1167542 2567:10.1126/science.1096415 1479:10.1038/s41559-016-0002 1012:University of Leicester 941:Silvanerpeton miripedes 925:Eucritta melanolimnetes 773:Xylognathus macrustenus 335:University of Cambridge 310:University of Cambridge 294:University of Cambridge 286:University of Cambridge 278:University of Leicester 233:University of Cambridge 158:University of Cambridge 103:University of Cambridge 99:University of Leicester 3870:Profile on PBS website 3672:10.1098/rsbm.2021.0008 2748:10.1098/rspb.2013.2689 1263:"Master & fellows" 465:University of Michigan 440:). This suggests that 392:stage of the earliest 206:evolutionary biologist 1881:Clack, J. A. (2002). 1021:In 2018, she won the 1001:University of Chicago 713:Clack & Smithson 697:Clack & Smithson 680:Clack & Smithson 663:Clack & Smithson 646:Clack & Smithson 593:Whitropus longicalcus 584:Smithson & Clack 347:University of Chicago 259:(Girls' Division), a 999:(DSc) degree by the 882:Clack & Ahlberg 860:Kirktonecta milnerae 801:Ballagadus caustrimi 707:Aytonerpeton microps 548:Limanichthys fraseri 523:Ahlberg & Clack 477:published in 2019. 388:which date from the 292:(ScD) degree by the 178:Jennifer Alice Clack 140:Evolutionary biology 94:Newcastle University 55:Jennifer Alice Agnew 4121:Henning Sirringhaus 3701:. The Royal Society 3634:on 29 December 2010 3528:2003CaJES..40..483C 3437:2012Palgy..55...73C 3388:2016Palgy..59...29S 3009:10.1098/rsos.192117 3001:2020RSOS....792117A 2954:2020JVPal..40E3150J 2867:1990Natur.347...66C 2681:2012PNAS..109.4532S 2618:2009Sci...324..364C 2508:2019Natur.574..527B 2445:10.1038/nature19354 2437:2016Natur.537..408S 2325:10.1038/nature11825 2317:2013Natur.494..226P 2259:10.1038/nature11124 2251:2012Natur.486..523P 2196:10.1038/nature06991 2188:2008Natur.453.1199A 2182:(7199): 1199–1204. 2076:10.1038/nature03893 2068:2005Natur.437..137A 1966:10.1038/nature01904 1958:2003Natur.425...65C 1907:10.1038/nature00824 1899:2002Natur.418...72C 1841:1998Natur.394...66C 1787:1996Natur.381...61A 1736:1991Natur.352..234C 1677:1989Natur.342..425C 1401:2012Palgy..55...73C 501:Rossichthys clackae 482: 461:Flinders University 4223:Steven D. Tanksley 4099:Richard Passingham 4082:Arthur B. McDonald 4035:Christopher Hunter 3823:. 3 February 2014. 3397:10.1111/pala.12203 2742:(1781): 20132689. 1623:10.7717/peerj.5972 1535:10.1111/pala.12395 960:endometrial cancer 876:Occidens portlocki 844:Ymeria denticulata 815:Coccovedus celatus 562:Celsiodon ahlbergi 533:Parmastega aelidae 480: 455:), Paul Upchurch ( 453:Uppsala University 359:Devonian tetrapods 214:lobe-finned fishes 4247: 4246: 4203:Roger D. Kornberg 4193:H. Robert Horvitz 4150:Jonathan Tennyson 4059:Dimitris Kioussis 3699:Fellows Directory 3606:www.zoo.cam.ac.uk 3255:10.1002/spp2.1086 3110:10.1002/spp2.1242 3060:(21): 1825–1846. 2831:978-3-319-46659-0 2788:"Basal Tetrapoda" 2675:(12): 4532–4537. 2612:(5925): 364–367. 2502:(7779): 527–531. 2431:(7620): 408–411. 2374:(7542): 168–169. 2311:(7436): 226–229. 2245:(7404): 523–526. 2125:(7085): 748–749. 2062:(7055): 137–140. 2011:(6973): 412–413. 1730:(6332): 234–236. 1671:(6248): 425–427. 997:Doctor of Science 951: 950: 944:gen. et sp. nov. 928:gen. et sp. nov. 912:gen. et sp. nov. 909:Pederpes finneyae 896:gen. et sp. nov. 879:gen. et sp. nov. 863:gen. et sp. nov. 847:gen. et sp. nov. 817:gen. et sp. nov. 789:gen. et sp. nov. 775:gen. et sp. nov. 710:gen. et sp. nov. 694:gen. et sp. nov. 677:gen. et sp. nov. 660:gen. et sp. nov. 643:gen. et sp. nov. 607:Deltodus tubineus 595:gen. et sp. nov. 581:gen. et sp. nov. 564:gen. et sp. nov. 553:Challands et al. 550:gen. et sp. nov. 536:gen. et sp. nov. 520:gen. et sp. nov. 290:Doctor of Science 225:Museum of Zoology 175: 174: 122:Scientific career 4332: 4144:Jesper Svejstrup 4127:Maurice Skolnick 4053:Jonathan Keating 4041:Graham Hutchings 4014:Nicholas Harberd 3944:Ross J. Anderson 3939:Robert Ainsworth 3910: 3903: 3896: 3887: 3847: 3846: 3844: 3842: 3831: 3825: 3824: 3817: 3811: 3810: 3808: 3806: 3795: 3789: 3788: 3786: 3784: 3769: 3763: 3762: 3760: 3758: 3742: 3736: 3735: 3733: 3731: 3717: 3711: 3710: 3708: 3706: 3695:"Jennifer Clack" 3691: 3685: 3684: 3674: 3650: 3644: 3643: 3641: 3639: 3624: 3618: 3617: 3615: 3613: 3597: 3591: 3590: 3565:(3–4): 369–376. 3554: 3548: 3547: 3511: 3505: 3504: 3498: 3494: 3492: 3484: 3476: 3470: 3469: 3457: 3451: 3450: 3448: 3416: 3410: 3409: 3399: 3367: 3346: 3345: 3303: 3293: 3278: 3277: 3267: 3257: 3233: 3227: 3226: 3186: 3177: 3176: 3136: 3130: 3129: 3092: 3086: 3085: 3045: 3039: 3038: 3028: 2980: 2974: 2973: 2937: 2931: 2930: 2919: 2913: 2912: 2901: 2895: 2894: 2875:10.1038/347066a0 2850: 2844: 2843: 2809: 2803: 2802: 2800: 2798: 2784: 2778: 2777: 2767: 2727: 2721: 2720: 2710: 2692: 2660: 2654: 2653: 2601: 2595: 2594: 2550: 2544: 2543: 2493: 2484: 2475: 2474: 2464: 2416: 2410: 2409: 2383: 2359: 2353: 2352: 2302: 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4329: 4250: 4249: 4248: 4243: 4227: 4213:Joseph Stiglitz 4176: 4138:Malcolm Stevens 4111:Wolfram Schultz 3994:James Gimzewski 3969:Michel ChrĂ©tien 3949:Michael Ashfold 3927: 3914: 3856: 3851: 3850: 3840: 3838: 3833: 3832: 3828: 3819: 3818: 3814: 3804: 3802: 3797: 3796: 3792: 3782: 3780: 3771: 3770: 3766: 3756: 3754: 3744: 3743: 3739: 3729: 3727: 3719: 3718: 3714: 3704: 3702: 3693: 3692: 3688: 3652: 3651: 3647: 3637: 3635: 3626: 3625: 3621: 3611: 3609: 3599: 3598: 3594: 3556: 3555: 3551: 3536:10.1139/e02-065 3513: 3512: 3508: 3495: 3485: 3478: 3477: 3473: 3459: 3458: 3454: 3418: 3417: 3413: 3369: 3368: 3349: 3301: 3295: 3294: 3281: 3235: 3234: 3230: 3188: 3187: 3180: 3138: 3137: 3133: 3094: 3093: 3089: 3047: 3046: 3042: 2982: 2981: 2977: 2939: 2938: 2934: 2921: 2920: 2916: 2903: 2902: 2898: 2861:(6288): 66–69. 2852: 2851: 2847: 2832: 2811: 2810: 2806: 2796: 2794: 2786: 2785: 2781: 2729: 2728: 2724: 2662: 2661: 2657: 2603: 2602: 2598: 2561:(5667): 57–58. 2552: 2551: 2547: 2491: 2486: 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90:Alma mater 74: 65:3 November 1947 60: 58: 57: 56: 37: 28: 26: 17: 12: 11: 5: 4338: 4336: 4328: 4327: 4322: 4317: 4312: 4307: 4302: 4297: 4292: 4287: 4282: 4277: 4272: 4267: 4262: 4252: 4251: 4245: 4244: 4242: 4241: 4235: 4233: 4229: 4228: 4226: 4225: 4220: 4218:Rashid Sunyaev 4215: 4210: 4205: 4200: 4198:Thomas Kailath 4195: 4190: 4184: 4182: 4178: 4177: 4175: 4174: 4169: 4162: 4157: 4152: 4147: 4140: 4135: 4130: 4123: 4118: 4113: 4108: 4105:Guy Richardson 4101: 4096: 4089: 4084: 4079: 4074: 4069: 4066:Stephen Larter 4062: 4055: 4050: 4047:Peter Isaacson 4043: 4038: 4031: 4029:Christine Holt 4026: 4024:Brian Hemmings 4021: 4016: 4011: 4006: 4001: 3996: 3991: 3986: 3981: 3976: 3971: 3966: 3961: 3956: 3951: 3946: 3941: 3935: 3933: 3929: 3928: 3915: 3913: 3912: 3905: 3898: 3890: 3884: 3883: 3882:on Vimeo, 2012 3877: 3872: 3867: 3862: 3855: 3854:External links 3852: 3849: 3848: 3826: 3812: 3790: 3764: 3751:U Chicago News 3737: 3712: 3686: 3645: 3619: 3592: 3549: 3522:(4): 483–498. 3506: 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Index

FRS
FLS

Newcastle University
University of Leicester
University of Cambridge
Zoology
Palaeontology
Evolutionary biology
Cambridge University Museum of Zoology
University of Cambridge
Darwin College, Cambridge
FRS
FLS
palaeontologist
evolutionary biologist
tetrapods
lobe-finned fishes
Museum of Zoology
Professor
University of Cambridge
Devonian
Carboniferous
Manchester
Bolton School
private school
Bolton
Lancashire
University of Newcastle upon Tyne
University of Leicester

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