31:
265:
Dr. Howard was born in 1952 and developed an early interest in biology and animal science. She pursued higher education in the United States, earning her Ph.D. from the
University of Maryland, College Park. Her academic and research pursuits set the stage for a career dedicated to animal conservation
299:
Dr. Howard's influence extends beyond her immediate achievements. She was a mentor and inspiration to many in the field of reproductive physiology, training the next generation of scientists who continue to advance conservation science. Her pioneering spirit and dedication to preserving biodiversity
274:
Dr. Howard began her career at the
Smithsonian's National Zoo, where she quickly became a leading figure in the zoo's reproductive physiology program. She focused on using artificial insemination and other reproductive technologies to breed endangered species, adapting techniques initially developed
295:
The techniques and protocols developed by Dr. Howard have had a profound and lasting impact on wildlife conservation. Her work has helped increase the populations of several endangered species and has provided a foundation for future conservation efforts. The methodologies she pioneered continue to
223:
In 1992, alongside the
Nashville Zoo President Rick Schwartz, she conducted the first successful clouded leopard artificial insemination. "If we lose this species," Howard says of the clouded leopards, "that means we probably lose other species in the forest. We may lose the forest all together. It
282:
In addition to her work with clouded leopards, Dr. Howard was instrumental in the black-footed ferret breeding program at the
National Zoo. During the 1980s, the black-footed ferret population had dwindled to just 18 individuals. Under her supervision, over 500 ferret kits were born, significantly
308:
Despite her significant contributions to animal conservation, Dr. JoGayle Howard's work is not widely recognized. She died in 2011 from melanoma, but her legacy lives on through the ongoing efforts of the scientists and conservationists she inspired. Dr. Howard's work remains highly influential,
256:
was a pioneering reproductive physiologist known for her groundbreaking work in the conservation of endangered species. As a researcher at the
Smithsonian's National Zoo, she adapted human infertility treatments to help breed endangered animals, making significant contributions to the field of
231:
in China. This study helped identify the causes of poor reproduction, which led to a rise in the population of giant pandas in breeding centers and zoos over the next ten years. “I’m not an emotional person,” she once told a reporter. “But it’s pretty cool when you realize you’re putting these
278:
One of Dr. Howard's most notable achievements was the first successful artificial insemination of a clouded leopard in 1992. This milestone was crucial for the conservation of this endangered species, showcasing the potential of assisted reproductive technologies in wildlife conservation.
286:
Dr. Howard's expertise was also pivotal in the birth of Tai Shan, the first surviving giant panda cub born at the
National Zoo. In 2005, she performed the artificial insemination of the panda Mei Xiang, leading to this significant achievement in panda conservation.
224:
may affect the human race eventually. I think most people want to just see these animals and want their future generations to see these animals. They are unique and nothing's going to replace a clouded leopard.""
528:
901:
487:
193:
Originally a veterinarian, Howard later became an expert in animal breeding, receiving the nickname "Sperm Queen".” She used common human infertility treatments, like
886:
896:
209:
123:
891:
236:
906:
733:
911:
632:
378:
30:
220:
breeding program at The
National Zoo. With her aid more than 500 kits were born and 150 artificial inseminations were performed.
35:
JoGayle Howard with a clouded leopard cub born via artificial insemination, 1992, Smithsonian
Institution Archives, SIA2017-023033
560:
309:
highlighting the critical role of scientific innovation in preserving endangered species and maintaining ecological balance.
507:
881:
403:
442:
92:
Successfully bred critically endangered black-footed ferret; survey of giant pandas in China from 1998 through 2000.
194:
183:
179:
76:
244:
198:
187:
81:
856:
876:
871:
330:
Member of the
American Society of Andrology; served as a member of the executive council (1993–1996)
408:
217:
175:
163:
821:
772:
716:
Lindburg, Donald; Baragona, Karen (2004-08-23). "17. Biomedical Survey of
Captive Giant Pandas".
202:
151:
725:
467:
296:
be used in breeding programs worldwide, demonstrating the enduring value of her contributions.
729:
698:
690:
614:
606:
536:
413:
324:
Distinguished Research and Scientist Award from the American Association of Zoo Veterinarians
811:
760:
721:
680:
596:
213:
147:
62:
784:
159:
143:
865:
816:
799:
321:
Ulysses S. Seal Conservation Award from the American Association of Zoo Veterinarians
174:
Howard was born on May 20, 1951, in Dallas, Texas. She graduated with a doctorate in
825:
529:"JoGayle Howard, a National Zoo scientist who helped breed giant pandas, dies at 59"
404:"JoGayle Howard, a National Zoo scientist who helped breed giant pandas, dies at 59"
345:
Featured Conservation Scientist (British Airways Exhibit at Millennium Dome, London)
564:
601:
584:
749:
228:
685:
668:
851:
694:
610:
540:
417:
846:
488:"The Smithsonian Mourns: Dr. JoGayle Howard, Wildlife Biologist (1951-2011)"
240:
139:
800:"Do Ranger Stations Deter Poaching Activity in National Parks in Thailand?"
702:
618:
283:
contributing to the recovery of the species from the brink of extinction.
361:
Do Ranger Stations Deter Poaching Activity in National Parks in Thailand?
182:
from University of Maryland in 1989. Dr. Howard received her D.V.M. from
764:
757:
Recovery of the Black-footed Ferret: Progress and Continuing Challenges
47:
857:
Smithsonian’s ‘Women of Science’ Comics Star Real-Life Superheroines
318:
Research Career Award (SERCA) from the National Institutes of Health
155:
750:"Use of Reproductive Technology for Black-footed Ferret Recovery"
327:
Recovery Champion Award from the U.S. Fish & Wildlife Service
798:
Jenks, Kate E.; Howard, Jogayle; Leimgruber, Peter (2012).
250:
She died from cancer on March 5, 2011, in Washington D.C.
235:
Howard oversaw the project to artificially inseminate the
583:
Smithsonian Conservation Biology Institute (2011-05-01).
100:
2010 U.S. Fish & Wildlife Service "Recovery Champion"
633:"JoGayle Howard Obituary (2011) The Dallas Morning News"
379:"JoGayle Howard Obituary (2011) The Dallas Morning News"
333:
Vice-president, Women in Andrology Committee (1999–2000)
852:
Leopard Lifesaver: Smithsonian Scientist JoGayle Howard
508:"JoGayle Howard, Brilliant Scientist and Pioneer, Dies"
443:"JoGayle Howard, Brilliant Scientist and Pioneer, Dies"
468:"In Memoriam: JoGayle Howard, D.V.M., Ph.D. 1951–2011"
561:"In Memoriam JoGayle Howard, D.V.M., Ph.D. 1951–2011"
342:
Research Achievement (US Fish & Wildlife Service)
585:"In MemoriamJoGayle Howard, D.V.M., Ph.D. 1951–2011"
336:
President, Women in Andrology Committee (2000–2001)
119:
111:
96:
88:
70:
55:
40:
21:
257:theriogenology—the study of animal reproduction.
115:Veterinary science, zoology, reproductive biology
339:Dr. Howard delivered the Buckeye Lecture in 1997
178:from Texas A&M in 1980. She earned a PhD in
138:(May 20, 1951 – March 5, 2011) was an American
74:Sunset High School, Dallas, TX. Class of 1969
8:
210:Smithsonian Conservation Biology Institute
124:Smithsonian Conservation Biology Institute
29:
18:
902:20th-century American non-fiction writers
815:
726:10.1525/california/9780520238671.003.0031
684:
600:
243:, and produced the first surviving cub,
370:
355:Giant Pandas: Biology and Conservation
887:20th-century American women scientists
780:
770:
718:Giant Pandas: Biology and Conservation
662:
660:
658:
656:
654:
652:
7:
578:
576:
574:
437:
435:
433:
232:animals back where they should be.”
897:20th-century American women writers
358:Recovery of the Black-footed Ferret
227:From 1998 to 2000, Howard surveyed
720:. University of California Press.
275:for human infertility treatments.
208:In 1980, she began working at the
14:
892:20th-century American scientists
817:10.1111/j.1744-7429.2012.00869.x
907:Smithsonian Institution people
1:
602:10.1095/biolreprod.111.092502
667:Pukazhenthi, Budhan (2011).
300:have left a lasting legacy.
186:in 1980, and Ph.D. from the
912:21st-century American women
79:, Veterinary Science, 1980
928:
686:10.2164/jandrol.111.013680
527:Brown, Emma (2011-03-08).
447:Smithsonian's National Zoo
402:Brown, Emma (2011-03-09).
266:and reproductive science.
254:JoGayle Howard (1952-2011)
129:
104:
28:
261:Early Life and Education
184:Texas A&M University
77:Texas A&M University
589:Biology of Reproduction
313:Awards and recognitions
201:, and was a pioneer in
195:artificial insemination
180:reproductive physiology
59:March 5, 2011 (aged 59)
841:Multimedia appearances
304:Recognition and Legacy
270:Career and Innovations
199:In vitro fertilisation
188:University of Maryland
82:University of Maryland
136:JoGayle Dillon Howard
23:JoGayle Dillon Howard
673:Journal of Andrology
164:black-footed ferrets
146:specializing in the
882:American zoologists
847:Nature’s Matchmaker
765:10.3133/sir20055293
669:"Dr JoGayle Howard"
409:The Washington Post
218:black-footed ferret
176:veterinary medicine
759:. pp. 28–36.
203:electroejaculation
152:endangered species
16:American zoologist
735:978-0-520-93016-2
291:Impact on Society
239:’s female panda,
133:
132:
106:Scientific career
84:, doctorate, 1989
919:
830:
829:
819:
795:
789:
788:
782:
778:
776:
768:
754:
746:
740:
739:
713:
707:
706:
688:
664:
647:
646:
644:
643:
629:
623:
622:
604:
595:(5): 1072–1073.
580:
569:
568:
563:. Archived from
557:
551:
550:
548:
547:
524:
518:
517:
515:
514:
504:
498:
497:
495:
494:
484:
478:
477:
475:
474:
464:
458:
457:
455:
454:
439:
428:
427:
425:
424:
399:
393:
392:
390:
389:
375:
216:She oversaw the
214:Washington, D.C.
160:clouded leopards
148:captive breeding
144:theriogenologist
63:Washington, D.C.
33:
19:
927:
926:
922:
921:
920:
918:
917:
916:
862:
861:
838:
833:
797:
796:
792:
779:
769:
752:
748:
747:
743:
736:
715:
714:
710:
666:
665:
650:
641:
639:
631:
630:
626:
582:
581:
572:
559:
558:
554:
545:
543:
533:Washington Post
526:
525:
521:
512:
510:
506:
505:
501:
492:
490:
486:
485:
481:
472:
470:
466:
465:
461:
452:
450:
441:
440:
431:
422:
420:
401:
400:
396:
387:
385:
377:
376:
372:
368:
352:
315:
306:
293:
272:
263:
172:
80:
75:
71:Alma mater
66:
60:
51:
45:
36:
24:
17:
12:
11:
5:
925:
923:
915:
914:
909:
904:
899:
894:
889:
884:
879:
874:
864:
863:
860:
859:
854:
849:
843:
842:
837:
836:External links
834:
832:
831:
810:(6): 826–833.
790:
781:|journal=
741:
734:
708:
679:(4): 345–347.
648:
624:
570:
567:on 2015-09-23.
552:
519:
499:
479:
459:
429:
394:
369:
367:
364:
363:
362:
359:
356:
351:
348:
347:
346:
343:
340:
337:
334:
331:
328:
325:
322:
319:
314:
311:
305:
302:
292:
289:
271:
268:
262:
259:
171:
168:
131:
130:
127:
126:
121:
117:
116:
113:
109:
108:
102:
101:
98:
94:
93:
90:
89:Known for
86:
85:
72:
68:
67:
61:
57:
53:
52:
46:
42:
38:
37:
34:
26:
25:
22:
15:
13:
10:
9:
6:
4:
3:
2:
924:
913:
910:
908:
905:
903:
900:
898:
895:
893:
890:
888:
885:
883:
880:
878:
875:
873:
870:
869:
867:
858:
855:
853:
850:
848:
845:
844:
840:
839:
835:
827:
823:
818:
813:
809:
805:
801:
794:
791:
786:
774:
766:
762:
758:
751:
745:
742:
737:
731:
727:
723:
719:
712:
709:
704:
700:
696:
692:
687:
682:
678:
674:
670:
663:
661:
659:
657:
655:
653:
649:
638:
634:
628:
625:
620:
616:
612:
608:
603:
598:
594:
590:
586:
579:
577:
575:
571:
566:
562:
556:
553:
542:
538:
534:
530:
523:
520:
509:
503:
500:
489:
483:
480:
469:
463:
460:
448:
444:
438:
436:
434:
430:
419:
415:
411:
410:
405:
398:
395:
384:
380:
374:
371:
365:
360:
357:
354:
353:
349:
344:
341:
338:
335:
332:
329:
326:
323:
320:
317:
316:
312:
310:
303:
301:
297:
290:
288:
284:
280:
276:
269:
267:
260:
258:
255:
251:
248:
246:
242:
238:
233:
230:
225:
221:
219:
215:
211:
206:
204:
200:
196:
191:
189:
185:
181:
177:
169:
167:
165:
161:
157:
153:
149:
145:
141:
137:
128:
125:
122:
118:
114:
110:
107:
103:
99:
95:
91:
87:
83:
78:
73:
69:
64:
58:
54:
50:, Texas, U.S.
49:
43:
39:
32:
27:
20:
807:
803:
793:
756:
744:
717:
711:
676:
672:
640:. Retrieved
636:
627:
592:
588:
565:the original
555:
544:. Retrieved
532:
522:
511:. Retrieved
502:
491:. Retrieved
482:
471:. Retrieved
462:
451:. Retrieved
449:. 2011-03-27
446:
421:. Retrieved
407:
397:
386:. Retrieved
382:
373:
350:Publications
307:
298:
294:
285:
281:
277:
273:
264:
253:
252:
249:
237:National Zoo
234:
229:giant pandas
226:
222:
207:
205:techniques.
192:
173:
135:
134:
120:Institutions
105:
44:May 20, 1951
877:2011 deaths
872:1951 births
247:, in 2005.
866:Categories
804:Biotropica
642:2022-01-20
637:Legacy.com
546:2020-02-29
513:2020-02-29
493:2020-02-29
473:2020-02-29
453:2022-01-20
423:2020-02-29
388:2022-01-20
383:Legacy.com
366:References
783:ignored (
773:cite book
695:1939-4640
611:0006-3363
541:0190-8286
418:0190-8286
241:Mei Xiang
190:in 1989.
170:Biography
140:zoologist
826:83482638
703:21474784
619:21649962
245:Tai Shan
154:such as
824:
732:
701:
693:
617:
609:
539:
416:
162:, and
156:pandas
112:Fields
97:Awards
65:, U.S.
48:Dallas
822:S2CID
753:(PDF)
785:help
730:ISBN
699:PMID
691:ISSN
615:PMID
607:ISSN
537:ISSN
414:ISSN
197:and
142:and
56:Died
41:Born
812:doi
761:doi
722:doi
681:doi
597:doi
212:in
150:of
868::
820:.
808:44
806:.
802:.
777::
775:}}
771:{{
755:.
728:.
697:.
689:.
677:32
675:.
671:.
651:^
635:.
613:.
605:.
593:84
591:.
587:.
573:^
535:.
531:.
445:.
432:^
412:.
406:.
381:.
166:.
158:,
828:.
814::
787:)
767:.
763::
738:.
724::
705:.
683::
645:.
621:.
599::
549:.
516:.
496:.
476:.
456:.
426:.
391:.
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
