645:
could be either four or five. The case of the maxillary teeth is similar, with one side preserving 10 tooth positions with space for more and the other indicating a minimum of 11 teeth. The true number of maxillary teeth was likely higher, but erosion makes it impossible to determine this fact until better material is found. The teeth towards the front of the maxilla are closely spaced, which may suggest that there was too little space for the teeth of the upper and lower jaw to interlock. Again, this may change with the discovery of better material, as the type specimen does not preserve the bone that would be situated between the individual teeth. Regardless of the state of the early maxillary teeth, some degree of interlocking was present in
4352:
4347:
129:
3507:
142:
521:, and is present due to the fact that the front of the orbits forms a narrow point that extends up until the eleventh tooth of the maxilla. The orbits are also noted for being large, with their individual diameters being much greater than the distance between them. Overall the orbits have an uneven shape, with the anterior segment being pointed, while the main part of the orbits is more circular in shape. This shape is partially caused by how the
517:, as in the latter the fourth dentary tooth slides into a nearly enclosed hole in the bone rather than an open notch. Behind this notch, the maxillae swell laterally, forming a boss around the location where the fifth maxillary tooth would sit. The eyesockets, also known as orbits, are located comparably far towards the front to the skull, at least relative to other crocodilians. This condition is similar to that found in
461:
3512:
1928:
630:
1943:, with the closest geographically being the many isolated teeth and osteoderms that make up a part of the mass death assemblage of the Golden Fleece Locality. Although the fact that they were found in isolation means that they cannot be confidently assigned to the genus, Yates and Pledge nonetheless argue that they are likely to represent
2029:, the group appears to have experienced a major faunal turnover leading into the Pliocene, with much of the Miocene diversity going extinct after what was likely a brief period of especially intense aridity. While mekosuchines managed to bounce back after conditions briefly became wetter again, the case of the Tirari
1953:
from other localities within the
Mampuwordu Sand Member, which would indicate that this genus survived for an unusually long time. However, today it is generally accepted that these fossils were either reported with faulty provenance or were simply reworked from older layers. Even beyond the scenario
474:
possessed a relatively short and roughly triangular skull that broadly resembles that of today's crocodiles, indicating that it was a generalist predator. Additionally, the snout is described as platyrostral, effectively meaning that it was flattened. However, the precise degree to which this applies
508:
contribute. The region where the premaxilla transitions to the maxilla is marked by a prominent notch where the jaw is constricted to make space for the fourth tooth of the lower jaw, something commonly seen in crocodilians. While this condition can be observed in a variety of mekosuchines, it does
652:
Even less is known about the tooth crowns, of which only three are known. All were part of the posterior teeth of their respective sections and, accordingly, are short, blunt and broadly rounded. The teeth are compressed side to side and have smooth cutting surfaces. However, there are some better
435:
to be exposed to the elements and thus damaging it significantly. On top of this weathering of the surface, the fossil underwent significant compression during preservation, causing it to appear much flatter than it would have in life. The Golden Fleece
Locality, where the fossils were discovered,
1919:
was discovered is interpreted to have been a watering hole at the time the crocodilian was alive. The scattered and disassociated remains of turtles and crocodiles alike have been interpreted as a single mass death assemblage that may have been formed when the water dried up, leaving its aquatic
644:
suffered, the exact number of teeth is difficult to determine. Three teeth were preserved in the left premaxilla, with the missing portion before them indicating more were present. However, it remains uncertain whether this region housed one or two teeth, meaning the number of premaxillary teeth
599:
Since the mandibular remains found alongside the holotype skull are poorly preserved, little information could be gathered from them. The most notable aspect of the mandible is that the flat underside is offset from the side by a nearly 90ยฐ bend. This is most similar to mandibles that have been
2033:
represents the first sign of crocodylines replacing mekosuchines in the long run. While this shift is undeniable, the precise reason for the change remains unknown. While it is possible that competition between the two was a factor, given their similar generalist morphology, there is no direct
451:
word for "spear" (kalthi) and frons, meaning forehead, a name chosen in reference to the animal's elongated frontal process. The species epithet is a combination of the Latin "aurum" for gold and "vellus" meaning fleece, chosen to reflect the locality where the fossil has been discovered.
653:
preserved teeth that were collected from the same locality as the holotype. While referral of these teeth remains tentative, Yates and Pledge argue that the nature of the Golden Fleece locality, having been a dried up pool of low-diversity, makes it very likely that they too belonged to
2058:
that continued onward throughout the later part of the
Neogene and Quaternary. This would suggest that while the group managed to survive the early stages of this change in climate during the Miocene, they failed to adapt to the changing conditions. Following this line of thought,
529:, which are situated before the prefrontals, are elongated and irregularly triangular in shape. Thick ridges extend from the lacrimal's contribution to the orbits until the pointed tip of the bones. These ridges are the result of a distinct change in slope, unlike in the
596:, though exposed on the dorsal surface like in the majority of mekosuchines, is narrow, and only makes up a small part of the skull table. The supraoccipital is described as semilunate in shape, whereas it is triangular in most other mekosuchines.
2366:
Yates, A.M.; Ristevski, J.; Salisbury, S.W. (2023). "The last Baru (Crocodylia, Mekosuchinae): a new species of 'cleaver-headed crocodile' from central
Australia and the turnover of crocodylians during the Late Miocene in Australia".
621:. They range in shape from square to rectangular, with keels that range from being low to tall. As in other crocodilians, they are ornamented with a series of pits, save for the region where the osteoderms overlap with one another.
562:
this is taken to an extreme as the anterior process extends much further down the rostrum as is typical, accounting for about 64% of the frontal's entire length. Additionally, the space between the eyes is much wider than in
757:
and the dwarf forms. Its shifting position was explained as being the result of the poor preservation, subsequently limited number of usable characters and the incongruent results these characters yielded.
440:, the fact that the locality is thought to have been a mass death site has been used to argue that they likely represent a single species. As a result of this, the material was tentatively referred to
322:
was recovered from the
Mampuwordu Sand Member, which underlies the younger sediments of the Pompapillina Member. This is significant, as the latter preserves some of the earliest records of the genus
475:
is difficult to determine as the holotype skull was badly crushed during preservation, greatly exaggerating its form in profile view. Adam Yates and
Neville Pledge speculate that the skull of
773:, two other mekosuchines that exhibit a more typical crocodilian morphology. An even more recent paper by Ristevski and colleagues from 2023 proposes a different placement. In their results,
504:
are longer than they are wide, contrasting with those of its closest relatives. The nares themselves are almost entirely surrounded by the premaxillae, save for a small section where the
436:
also yielded a variety of isolated remains, mostly consisting of teeth, osteoderms and vertebrae. While the lack of overlap means that none can be definitely proven to have been those of
1962:, Yates and Pledge argue that their overall similar generalist nature means that they would have been unlikely to coexist. The final crocodilian known from the Mampuwordu Sand Member is
592:. Though small, the fenestrae are largely unobscured by the surrounding bones overhanging it, with only the parietal slightly overhanging the rear-most corner of the fenestrae. The
584:. The massive distance between the fenestrae is unusual for mekosuchines and only one other species is noted for having similar proportions, that being an unspecified species of
483:
given that the former preserves a distinct swelling above the fifth maxillary tooth that would be absent in an animal with a very deep (altirostral) skull similar to
356:
have broadly similar skull forms, with both being interpreted as generalist semi-aquatic predators much like many of today's crocodiles. Though far from large,
1988:
would have clearly occupied a different ecological niche within the
Pliocene Lake Eyre Basin and would thus not have been in competition with the semi-aquatic
761:
However, later studies did recover better resolved trees that somewhat clarified the position of this animal within the family. Lee and Yates (2018) recovered
2420:
2009:. Based on the dating of this member, the genus entered the Lake Eyre Basin during the middle Pliocene (3.9โ3.4 mya), possibly overlapping with the range of
2240:"The biochronology and palaeobiogeography of Baru (Crocodylia: Mekosuchinae) based on new specimens from the Northern Territory and Queensland, Australia"
4548:
4502:
4435:
1909:
was most likely a semi-aquatic generalist predator living in the waterways of what is now South
Australia, sharing its habitat with turtles such as
554:. As is typical for crocodilians, the frontal is divided into two segments, the posterior one of them is broad and makes up the space between the
336:
or simply moved into the region after the niche was left empty by the extinction of the local mekosuchines. Should the later be the case, then
657:. These teeth correspond well to more anterior dentition, being taller and more pointed, but less compressed, with a gentle inward curvature.
2319:
2046:, another generalist, inhabited Australia until the late Pleistocene, though it is unclear whether or not this genus coexisted with
580:. The fenestrae are widely spaced, much wider than each fenestra is long, with the interfenestral bar being formed entirely by the
4346:
2413:
4351:
3679:
3506:
1968:, which has been identified based on a maxillary fragment discovered in the Woodard Quarry. Though initially identified as a
3725:
3594:
1392:
2189:
Ristevski, J.; Willis, P.M.A.; Yates, A.M.; White, M.A.; Hart, L.J.; Stein, M.D.; Price, G.J.; Salisbury, S.W. (2023).
2406:
1614:
328:
in
Australia, which would eventually go on to replace mekosuchines. It is currently unclear whether or not the Tirari
3586:
2753:
141:
3807:
2315:"Tip-dating and homoplasy: reconciling the shallow molecular divergences of modern gharials with their long fossil"
3779:
785:, akin to some of the results found by Yates and Pledge. More specifically, one of their results suggested that
3425:
2746:
715:
struggled with its relationship to other mekosuchines, though Yates and Pledge ruled out the possibility that
3511:
3800:
1661:
1560:
571:
4286:
4060:
4390:
4310:
4084:
3793:
1499:
2191:"Migrations, diversifications and extinctions: the evolutionary history of crocodyliforms in Australasia"
2021:, would have come at a crucial point in the history of mekosuchines. Though highly successful during the
589:
525:
contribute to the orbital margin, as these bones form prominent flanges that extend into the orbits. The
4543:
4515:
4448:
4270:
4263:
4076:
4068:
3952:
2589:
2509:
1587:
4294:
1972:, later research has shown that the tall snout and ziphodont teeth are indistinguishable from those of
558:
and part of the skull table. The anterior process meanwhile is generally long and slender, however, in
1996:
428:
311:
4489:
4422:
3786:
3475:
3468:
3418:
3216:
2376:
2202:
2126:
1911:
1533:
1141:
128:
3454:
3181:
919:
883:
530:
723:
of crocodilian, making a position as a mekosuchine the most probable scenario. In their analysis,
4318:
4227:
4219:
3945:
3223:
3202:
3195:
2856:
2732:
2536:
2289:
2220:
2158:
2150:
1939:
A variety of crocodilian remains have been collected from the region that yielded the fossils of
1641:
533:, which has preorbital ridges raised above the rest of the skull. In general, the ridges seen on
136:
4520:
4453:
669:
measure around 44.7 cm (17.6 in) long from the broken snout tip to the remains of the
4234:
3733:
3700:
3461:
3188:
2993:
2348:
2271:
2142:
2018:
395:
749:
and all other mekosuchines to it being a derived member of
Mekosuchini, the clade containing
4212:
3447:
3310:
3273:
3209:
3104:
2970:
2384:
2338:
2328:
2261:
2251:
2210:
2134:
745:
in their results varied between it being a very basal animal outside of the clade formed by
649:
based on the presence of a large reception pit located between the sixth and seventh teeth.
421:
209:
196:
2115:"A Pliocene mekosuchine (Eusuchia: Crocodilia) from the Lake Eyre Basin of South Australia"
4302:
4278:
4241:
4161:
3909:
3636:
3395:
3380:
3345:
3288:
3158:
3014:
2959:
2776:
2760:
863:
522:
403:
399:
2003:
but do preserve what is generally thought to have been a generalist species of the genus
17:
2380:
2206:
2130:
2067:
entered the Lake Eyre Basin, with the latter simply moving into the newly opened niche.
617:
are also known from the Golden Fleece locality, though it is unsure if they belonged to
4175:
4153:
3895:
3887:
3845:
3643:
3578:
3564:
3437:
3150:
3134:
3063:
3021:
2979:
2951:
2929:
2886:
2821:
2808:
2783:
2768:
2739:
2670:
2626:
2611:
2343:
2314:
2266:
2239:
1949:
1231:
829:
733:
693:
593:
576:
362:
341:
4537:
4373:
4205:
4198:
4183:
4168:
4131:
4117:
4029:
4014:
4006:
3923:
3902:
3873:
3866:
3859:
3749:
3663:
3651:
3601:
3571:
3556:
3542:
3324:
3295:
3265:
3127:
3096:
3083:
3071:
3055:
3037:
3000:
2900:
2724:
2702:
2224:
2162:
2055:
1969:
1378:
980:
670:
581:
526:
345:
79:
1920:
inhabitants to die. This hypothesis is supported by the presence of a thin layer of
460:
4480:
4413:
4368:
4191:
4124:
4037:
4022:
3989:
3978:
3930:
3712:
3622:
3609:
3529:
3408:
3359:
3352:
3280:
3142:
3119:
2944:
2936:
2907:
2849:
2842:
2717:
2602:
2494:
2469:
1853:
1415:
1342:
1043:
1016:
815:
738:
551:
294:
261:
235:
2215:
2190:
2138:
727:
position varied greatly between trees, with its uncertain nature creating a large
4146:
4139:
3937:
3916:
3880:
3852:
3831:
3373:
3366:
3338:
3331:
3317:
3258:
3111:
3047:
3007:
2986:
2914:
2893:
2835:
2709:
2677:
2114:
497:
304:
301:
54:
2042:
to extinction by outcompeting it. This is further complicated by the fact that
1927:
629:
4100:
3964:
3769:
3693:
3686:
3672:
3251:
2868:
2648:
2548:
2460:
2430:
2005:
1204:
699:
614:
505:
368:
324:
219:
99:
64:
2146:
1947:
due to the nature of the site. More anomalous is the discovery of remains of
4050:
3388:
3171:
2655:
2521:
2022:
1921:
1733:
610:
555:
373:
315:
153:
104:
48:
2352:
2333:
2275:
1999:, specifically those of the Pompapillina Member, lack remains referable to
4474:
4407:
4253:
3741:
3302:
2922:
2662:
2478:
2454:
1964:
1780:
1251:
728:
602:
485:
425:
407:
308:
173:
94:
89:
74:
69:
59:
41:
2154:
4507:
4440:
2641:
2398:
2256:
2026:
1463:
777:
typically occupies a very basal position within Mekosuchinae alongside
417:
291:
183:
109:
84:
2388:
492:
The tip of the snout appears to have been differently proportioned in
3238:
2448:
2290:"Crikey! Massive prehistoric croc emerges from South East Queensland"
789:
may have been the basalmost member of the group, whereas another saw
432:
163:
4384:
567:
owing to the greater width of the posterior section of the frontal.
2684:
2017:
in Australia, likely arriving after having migrated there via the
1926:
1177:
720:
687:
628:
501:
459:
448:
297:
4494:
4427:
2054:
and mekosuchines in general proceeded to suffer from the ongoing
640:
Given the extensive damage that the underside of the holotype of
609:
Only a handful of postcranial remains have been found, including
431:. The fossil was discovered upside down, causing the area of the
3757:
2619:
1760:
1063:
539:
4388:
3527:
2806:
2587:
2441:
2402:
570:
The skull table is flat and trapezoid in shape with elliptical
348:. A point in favour of the competition hypothesis is that both
685:
still exceeded the total body length of "dwarf forms" such as
673:. While this is far from the size of larger mekosuchines like
479:
may have been moderately deep, making it similar to that of
360:
was nonetheless bigger than many other mekosuchines such as
731:
at the base of the group that contains Mekosuchinae minus
1958:
fossils were wrongfully attributed to the same units as
741:
to all other taxa). The different positions occupied by
1905:
Based on its platyrostral, short and triangular skull,
424:, was recovered from the Mampuwordu Sand Member of the
1931:
The gulf snapping turtle may have appeared alongside
2195:
Alcheringa: An Australasian Journal of Palaeontology
1784:
1764:
1737:
1665:
1645:
1618:
1591:
1564:
1537:
1503:
1467:
1433:
1396:
1255:
1235:
1208:
1181:
1145:
1125:
1068:
1047:
1020:
984:
964:
923:
887:
867:
833:
268:
4464:
4397:
4251:
4099:
4048:
3988:
3962:
3829:
3767:
3711:
3662:
3620:
3540:
3435:
3406:
3237:
3169:
3082:
3035:
2969:
2866:
2819:
2694:
2601:
2547:
2520:
2493:
613:that provide no further valuable information. Some
2313:Michael S. Y. Lee; Adam M. Yates (27 June 2018).
2113:Yates, Adam M.; Pledge, Neville S. (2017-01-02).
2414:
376:, meaning it contains only a single species,
8:
4385:
4109:
3998:
3970:
3837:
3717:
3628:
3548:
3537:
3524:
3243:
3088:
3079:
2878:
2874:
2827:
2816:
2803:
2598:
2584:
2526:
2499:
2490:
2438:
2421:
2407:
2399:
127:
31:
2342:
2332:
2265:
2255:
2214:
496:when compared to other mekosuchines. The
394:were discovered on the western shores of
2108:
2106:
2104:
2102:
2100:
2076:
1894:
665:The preserved portions of the skull of
550:, and the one it was named for, is its
2184:
2182:
2180:
2178:
2176:
2174:
2172:
2098:
2096:
2094:
2092:
2090:
2088:
2086:
2084:
2082:
2080:
1365:
802:
7:
1976:and clearly distinct from those of
1915:. The Golden Fleece Locality where
1752:
1725:
1633:
1606:
1579:
1552:
1525:
1518:
1491:
1455:
1448:
1421:
1411:
1384:
1374:
1367:
1223:
1196:
1169:
1117:
1110:
1035:
1008:
952:
945:
938:
911:
855:
848:
821:
811:
804:
546:The most characteristic feature of
340:may have simply been the victim of
2320:Proceedings of the Royal Society B
2119:Journal of Vertebrate Paleontology
447:The generic name derives from the
25:
4549:Prehistoric pseudosuchian genera
4350:
4345:
3510:
3505:
1912:Elseya sp. (cf. E. lavarockorum)
140:
2063:could have disappeared before
2050:. A second hypothesis is that
1995:Much younger sediments of the
1:
2216:10.1080/03115518.2023.2201319
2139:10.1080/02724634.2017.1244540
500:are narrow and the external
27:Extinct genus of crocodylian
1615:Mekosuchus whitehunterensis
4565:
793:share this placement with
737:(which was recovered as a
719:could belong to any other
574:similar to those found in
4466:Kalthifrons aurivellensis
4363:
4343:
4112:
4001:
3973:
3840:
3720:
3631:
3551:
3536:
3523:
3503:
3246:
3091:
2881:
2877:
2830:
2815:
2802:
2597:
2583:
2529:
2502:
2489:
2437:
1851:
1777:
1757:
1750:
1730:
1723:
1658:
1638:
1631:
1611:
1604:
1584:
1577:
1557:
1550:
1530:
1523:
1516:
1496:
1489:
1460:
1453:
1446:
1430:Kalthifrons aurivellensis
1426:
1419:
1409:
1389:
1382:
1372:
1340:
1248:
1228:
1221:
1201:
1194:
1174:
1167:
1138:
1122:
1115:
1108:
1060:
1040:
1033:
1013:
1006:
977:
957:
950:
943:
936:
916:
909:
880:
860:
853:
846:
826:
819:
809:
379:Kalthifrons aurivellensis
273:Kalthifrons aurivellensis
267:
260:
137:Scientific classification
135:
126:
34:
18:Kalthifrons aurivellensis
3680:"Crocodylus" gariepensis
3953:"Tomistoma" lusitanicum
3726:"Crocodylus" megarhinus
3595:"Crocodylus" megarhinus
1662:Mekosuchus inexpectatus
1561:Trilophosuchus rackhami
1393:"Crocodylus" megarhinus
765:as a close relative of
572:supratemporal fenestrae
2369:Papers in Paleontology
2334:10.1098/rspb.2018.1071
1936:
1500:Australosuchus clarkae
637:
468:
278:Yates and Pledge, 2017
255:Yates and Pledge, 2017
4516:Paleobiology Database
4449:Paleobiology Database
1930:
1924:underlying the sand.
1588:Volia athollandersoni
632:
463:
332:directly outcompeted
318:. More specifically,
4228:"Tomistoma" coppensi
4220:"Tomistoma" cairense
3946:"Tomistoma" cairense
3587:"Crocodylus" affinis
2754:"Crocodylus" affinis
2238:Yates, A.M. (2017).
2013:. The appearance of
386:Discovery and naming
4235:"Tomistoma" dowsoni
2381:2023PPal....9E1523Y
2207:2023Alch...47..370R
2131:2017JVPal..37E4540Y
1124:Bullock Creek taxon
920:Kambara murgonensis
884:Kambara implexidens
633:Size comparison of
590:Alcoota fossil site
531:Saltwater Crocodile
416:), consisting of a
3808:C. thorbjarnarsoni
2857:Listrognathosuchus
2733:Brachyuranochampsa
2257:10.7717/peerj.3458
1937:
1642:Mekosuchus sanderi
711:Early research on
638:
537:resemble those of
469:
464:Holotype skull of
4531:
4530:
4391:Taxon identifiers
4382:
4381:
4359:
4358:
4341:
4340:
4337:
4336:
4333:
4332:
4329:
4328:
4095:
4094:
3825:
3824:
3821:
3820:
3817:
3816:
3780:C. anthropophagus
3519:
3518:
3501:
3500:
3497:
3496:
3493:
3492:
3489:
3488:
3485:
3484:
3233:
3232:
3031:
3030:
2994:Eurycephalosuchus
2798:
2797:
2794:
2793:
2747:"Crocodylus" acer
2579:
2578:
2575:
2574:
2571:
2570:
2567:
2566:
2389:10.1002/spp2.1523
2019:Malay Archipelago
1898:
1897:
1891:
1890:
1882:
1881:
1873:
1872:
1864:
1863:
1840:
1839:
1831:
1830:
1822:
1821:
1813:
1812:
1804:
1803:
1795:
1794:
1712:
1711:
1703:
1702:
1694:
1693:
1685:
1684:
1676:
1675:
1478:
1477:
1362:
1361:
1353:
1352:
1329:
1328:
1320:
1319:
1311:
1310:
1302:
1301:
1293:
1292:
1284:
1283:
1275:
1274:
1266:
1265:
1156:
1155:
1097:
1096:
1088:
1087:
1079:
1078:
995:
994:
898:
897:
509:help distinguish
396:Lake Palankarinna
283:
282:
256:
16:(Redirected from
4556:
4524:
4523:
4511:
4510:
4498:
4497:
4485:
4484:
4483:
4457:
4456:
4444:
4443:
4431:
4430:
4418:
4417:
4416:
4386:
4354:
4349:
4311:G. pachyrhynchus
4213:Siquisiquesuchus
4110:
3999:
3971:
3838:
3718:
3629:
3549:
3538:
3525:
3514:
3509:
3476:C. wannlangstoni
3469:C. venezuelensis
3311:Globidentosuchus
3274:Centenariosuchus
3244:
3105:Allognathosuchus
3089:
3080:
2971:Orientalosuchina
2879:
2875:
2828:
2817:
2804:
2599:
2585:
2527:
2500:
2491:
2484:
2483:
2439:
2423:
2416:
2409:
2400:
2393:
2392:
2363:
2357:
2356:
2346:
2336:
2310:
2304:
2303:
2301:
2300:
2286:
2280:
2279:
2269:
2259:
2235:
2229:
2228:
2218:
2186:
2167:
2166:
2110:
1997:Tirari Formation
1786:
1766:
1753:
1739:
1726:
1667:
1647:
1634:
1620:
1607:
1593:
1580:
1566:
1553:
1539:
1526:
1519:
1505:
1492:
1469:
1456:
1449:
1435:
1422:
1412:
1398:
1385:
1375:
1368:
1257:
1237:
1224:
1210:
1197:
1183:
1170:
1147:
1127:
1118:
1111:
1070:
1049:
1036:
1022:
1009:
986:
966:
953:
946:
939:
925:
912:
889:
869:
856:
849:
835:
822:
812:
805:
800:
799:
429:Tirari Formation
312:Tirari Formation
270:
254:
247:
234:
210:Archosauriformes
197:Archosauromorpha
145:
144:
131:
121:
51:
40:Temporal range:
32:
21:
4564:
4563:
4559:
4558:
4557:
4555:
4554:
4553:
4534:
4533:
4532:
4527:
4519:
4514:
4506:
4501:
4493:
4488:
4479:
4478:
4473:
4460:
4452:
4447:
4439:
4434:
4426:
4421:
4412:
4411:
4406:
4393:
4383:
4378:
4355:
4325:
4287:G. curvirostris
4264:G. bengawanicus
4247:
4242:Toyotamaphimeia
4162:Hesperogavialis
4103:
4091:
4044:
3992:
3984:
3967:
3958:
3910:Megadontosuchus
3834:
3813:
3801:C. palaeindicus
3763:
3707:
3658:
3637:Aldabrachampsus
3625:
3616:
3545:
3532:
3515:
3481:
3455:C. brevirostris
3431:
3402:
3396:Wannaganosuchus
3381:Stangerochampsa
3346:Orthogenysuchus
3289:Chinatichampsus
3229:
3165:
3159:Wannaganosuchus
3078:
3040:
3027:
3015:Orientalosuchus
2965:
2960:Stangerochampsa
2871:
2862:
2824:
2811:
2790:
2777:Prodiplocynodon
2761:Portugalosuchus
2690:
2593:
2563:
2543:
2516:
2485:
2444:
2443:
2433:
2427:
2397:
2396:
2365:
2364:
2360:
2312:
2311:
2307:
2298:
2296:
2288:
2287:
2283:
2237:
2236:
2232:
2188:
2187:
2170:
2125:(1): e1244540.
2112:
2111:
2078:
2073:
1903:
1892:
1883:
1874:
1865:
1841:
1832:
1823:
1814:
1805:
1796:
1713:
1704:
1695:
1686:
1677:
1479:
1363:
1354:
1330:
1321:
1312:
1303:
1294:
1285:
1276:
1267:
1157:
1098:
1089:
1080:
996:
899:
864:Kambara taraina
709:
663:
627:
565:Australosuchus,
458:
422:mandibular rami
404:Lake Eyre Basin
400:South Australia
390:The remains of
388:
372:. The genus is
352:and the Tirari
307:known from the
279:
276:
253:
245:
232:
139:
122:
120:
119:
118:
117:
112:
107:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
46:
45:
38:
28:
23:
22:
15:
12:
11:
5:
4562:
4560:
4552:
4551:
4546:
4536:
4535:
4529:
4528:
4526:
4525:
4512:
4499:
4486:
4470:
4468:
4462:
4461:
4459:
4458:
4445:
4432:
4419:
4403:
4401:
4395:
4394:
4389:
4380:
4379:
4377:
4376:
4371:
4364:
4361:
4360:
4357:
4356:
4344:
4342:
4339:
4338:
4335:
4334:
4331:
4330:
4327:
4326:
4324:
4323:
4315:
4307:
4299:
4291:
4283:
4275:
4267:
4259:
4257:
4249:
4248:
4246:
4245:
4238:
4231:
4224:
4216:
4209:
4202:
4195:
4188:
4180:
4176:Maomingosuchus
4172:
4165:
4158:
4154:Harpacochampsa
4150:
4143:
4136:
4128:
4121:
4113:
4107:
4097:
4096:
4093:
4092:
4090:
4089:
4081:
4077:T. lusitanicum
4073:
4065:
4061:T. calaritanum
4056:
4054:
4046:
4045:
4043:
4042:
4034:
4026:
4019:
4011:
4002:
3996:
3986:
3985:
3983:
3982:
3974:
3968:
3963:
3960:
3959:
3957:
3956:
3949:
3942:
3934:
3927:
3920:
3913:
3906:
3899:
3896:Maomingosuchus
3892:
3888:Leptorrhamphus
3884:
3877:
3870:
3863:
3856:
3849:
3846:Dollosuchoides
3841:
3835:
3830:
3827:
3826:
3823:
3822:
3819:
3818:
3815:
3814:
3812:
3811:
3804:
3797:
3794:C. falconensis
3790:
3783:
3775:
3773:
3765:
3764:
3762:
3761:
3754:
3746:
3738:
3730:
3721:
3715:
3709:
3708:
3706:
3705:
3697:
3690:
3683:
3676:
3668:
3666:
3660:
3659:
3657:
3656:
3648:
3644:Dzungarisuchus
3640:
3632:
3626:
3621:
3618:
3617:
3615:
3614:
3606:
3598:
3591:
3583:
3579:Australosuchus
3575:
3568:
3565:Antecrocodylus
3561:
3552:
3546:
3541:
3534:
3533:
3528:
3521:
3520:
3517:
3516:
3504:
3502:
3499:
3498:
3495:
3494:
3491:
3490:
3487:
3486:
3483:
3482:
3480:
3479:
3472:
3465:
3458:
3451:
3443:
3441:
3433:
3432:
3430:
3429:
3426:M. latrubessei
3422:
3414:
3412:
3404:
3403:
3401:
3400:
3392:
3385:
3377:
3370:
3363:
3356:
3349:
3342:
3335:
3328:
3321:
3314:
3307:
3299:
3292:
3285:
3277:
3270:
3262:
3255:
3247:
3241:
3235:
3234:
3231:
3230:
3228:
3227:
3220:
3213:
3206:
3199:
3192:
3185:
3177:
3175:
3167:
3166:
3164:
3163:
3155:
3151:Procaimanoidea
3147:
3139:
3135:Hassiacosuchus
3131:
3124:
3116:
3108:
3101:
3092:
3086:
3077:
3076:
3068:
3064:Menatalligator
3060:
3052:
3043:
3041:
3036:
3033:
3032:
3029:
3028:
3026:
3025:
3022:Protoalligator
3018:
3011:
3004:
2997:
2990:
2983:
2980:Dongnanosuchus
2975:
2973:
2967:
2966:
2964:
2963:
2956:
2952:Procaimanoidea
2948:
2941:
2933:
2930:Hassiacosuchus
2926:
2919:
2911:
2904:
2897:
2890:
2887:Albertochampsa
2882:
2872:
2867:
2864:
2863:
2861:
2860:
2853:
2846:
2839:
2831:
2825:
2822:Alligatoroidea
2820:
2813:
2812:
2809:Alligatoroidea
2807:
2800:
2799:
2796:
2795:
2792:
2791:
2789:
2788:
2784:Planocraniidae
2780:
2773:
2769:Pristichampsus
2765:
2757:
2750:
2743:
2740:Charactosuchus
2736:
2729:
2721:
2714:
2706:
2698:
2696:
2692:
2691:
2689:
2688:
2681:
2674:
2671:Trilophosuchus
2667:
2659:
2652:
2645:
2638:
2631:
2627:Harpacochampsa
2623:
2616:
2612:Australosuchus
2607:
2605:
2595:
2594:
2588:
2581:
2580:
2577:
2576:
2573:
2572:
2569:
2568:
2565:
2564:
2562:
2561:
2560:
2559:
2553:
2551:
2545:
2544:
2542:
2541:
2540:
2539:
2530:
2524:
2518:
2517:
2515:
2514:
2513:
2512:
2503:
2497:
2487:
2486:
2482:
2481:
2472:
2463:
2457:
2451:
2442:
2435:
2434:
2428:
2426:
2425:
2418:
2411:
2403:
2395:
2394:
2358:
2305:
2281:
2230:
2201:(4): 370โ415.
2168:
2075:
2074:
2072:
2069:
2034:evidence that
1982:Australosuchus
1980:. Unlike with
1956:Australosuchus
1950:Australosuchus
1902:
1899:
1896:
1895:
1893:
1889:
1888:
1885:
1884:
1880:
1879:
1876:
1875:
1871:
1870:
1867:
1866:
1862:
1861:
1858:
1857:
1850:
1847:
1846:
1843:
1842:
1838:
1837:
1834:
1833:
1829:
1828:
1825:
1824:
1820:
1819:
1816:
1815:
1811:
1810:
1807:
1806:
1802:
1801:
1798:
1797:
1793:
1792:
1789:
1788:
1776:
1773:
1772:
1769:
1768:
1756:
1751:
1749:
1746:
1745:
1742:
1741:
1734:Paludirex spp.
1729:
1724:
1722:
1719:
1718:
1715:
1714:
1710:
1709:
1706:
1705:
1701:
1700:
1697:
1696:
1692:
1691:
1688:
1687:
1683:
1682:
1679:
1678:
1674:
1673:
1670:
1669:
1657:
1654:
1653:
1650:
1649:
1637:
1632:
1630:
1627:
1626:
1623:
1622:
1610:
1605:
1603:
1600:
1599:
1596:
1595:
1583:
1578:
1576:
1573:
1572:
1569:
1568:
1556:
1551:
1549:
1546:
1545:
1542:
1541:
1529:
1524:
1522:
1517:
1515:
1512:
1511:
1508:
1507:
1495:
1490:
1488:
1485:
1484:
1481:
1480:
1476:
1475:
1472:
1471:
1459:
1454:
1452:
1447:
1445:
1442:
1441:
1438:
1437:
1425:
1420:
1418:
1410:
1408:
1405:
1404:
1401:
1400:
1388:
1383:
1381:
1373:
1371:
1366:
1364:
1360:
1359:
1356:
1355:
1351:
1350:
1347:
1346:
1339:
1336:
1335:
1332:
1331:
1327:
1326:
1323:
1322:
1318:
1317:
1314:
1313:
1309:
1308:
1305:
1304:
1300:
1299:
1296:
1295:
1291:
1290:
1287:
1286:
1282:
1281:
1278:
1277:
1273:
1272:
1269:
1268:
1264:
1263:
1260:
1259:
1247:
1244:
1243:
1240:
1239:
1232:Trilophosuchus
1227:
1222:
1220:
1217:
1216:
1213:
1212:
1200:
1195:
1193:
1190:
1189:
1186:
1185:
1173:
1168:
1166:
1163:
1162:
1159:
1158:
1154:
1153:
1150:
1149:
1137:
1134:
1133:
1130:
1129:
1121:
1116:
1114:
1109:
1107:
1104:
1103:
1100:
1099:
1095:
1094:
1091:
1090:
1086:
1085:
1082:
1081:
1077:
1076:
1073:
1072:
1059:
1056:
1055:
1052:
1051:
1039:
1034:
1032:
1029:
1028:
1025:
1024:
1012:
1007:
1005:
1002:
1001:
998:
997:
993:
992:
989:
988:
976:
973:
972:
969:
968:
956:
951:
949:
944:
942:
937:
935:
932:
931:
928:
927:
915:
910:
908:
905:
904:
901:
900:
896:
895:
892:
891:
879:
876:
875:
872:
871:
859:
854:
852:
847:
845:
842:
841:
838:
837:
830:Australosuchus
825:
820:
818:
810:
808:
803:
779:Australosuchus
734:Australosuchus
708:
705:
694:Trilophosuchus
662:
659:
626:
623:
594:supraoccipital
577:Trilophosuchus
527:lacrimal bones
515:Australosuchus
457:
454:
387:
384:
363:Trilophosuchus
342:global cooling
281:
280:
277:
265:
264:
258:
257:
243:
239:
238:
230:
223:
222:
217:
213:
212:
207:
200:
199:
194:
187:
186:
181:
177:
176:
171:
167:
166:
161:
157:
156:
151:
147:
146:
133:
132:
124:
123:
115:
114:
113:
108:
103:
98:
93:
88:
83:
78:
73:
68:
63:
58:
53:
52:
39:
26:
24:
14:
13:
10:
9:
6:
4:
3:
2:
4561:
4550:
4547:
4545:
4542:
4541:
4539:
4522:
4517:
4513:
4509:
4504:
4500:
4496:
4491:
4487:
4482:
4476:
4472:
4471:
4469:
4467:
4463:
4455:
4450:
4446:
4442:
4437:
4433:
4429:
4424:
4420:
4415:
4409:
4405:
4404:
4402:
4400:
4396:
4392:
4387:
4375:
4374:Gryposuchinae
4372:
4370:
4366:
4365:
4362:
4353:
4348:
4321:
4320:
4316:
4313:
4312:
4308:
4305:
4304:
4300:
4297:
4296:
4292:
4289:
4288:
4284:
4281:
4280:
4276:
4273:
4272:
4268:
4266:
4265:
4261:
4260:
4258:
4256:
4255:
4250:
4244:
4243:
4239:
4237:
4236:
4232:
4230:
4229:
4225:
4222:
4221:
4217:
4215:
4214:
4210:
4208:
4207:
4206:Rhamphosuchus
4203:
4201:
4200:
4199:Piscogavialis
4196:
4194:
4193:
4189:
4186:
4185:
4184:Paratomistoma
4181:
4178:
4177:
4173:
4171:
4170:
4169:Ikanogavialis
4166:
4164:
4163:
4159:
4156:
4155:
4151:
4149:
4148:
4144:
4142:
4141:
4137:
4134:
4133:
4132:Gavialosuchus
4129:
4127:
4126:
4122:
4120:
4119:
4118:Aktiogavialis
4115:
4114:
4111:
4108:
4106:
4102:
4098:
4087:
4086:
4085:T. taiwanicum
4082:
4079:
4078:
4074:
4071:
4070:
4066:
4063:
4062:
4058:
4057:
4055:
4053:
4052:
4047:
4040:
4039:
4035:
4032:
4031:
4030:Paratomistoma
4027:
4025:
4024:
4020:
4017:
4016:
4015:Gavialosuchus
4012:
4009:
4008:
4007:Brasilosuchus
4004:
4003:
4000:
3997:
3995:
3994:sensu stricto
3991:
3987:
3981:
3980:
3976:
3975:
3972:
3969:
3966:
3961:
3955:
3954:
3950:
3948:
3947:
3943:
3940:
3939:
3935:
3933:
3932:
3928:
3926:
3925:
3924:Paratomistoma
3921:
3919:
3918:
3914:
3912:
3911:
3907:
3905:
3904:
3903:Maroccosuchus
3900:
3898:
3897:
3893:
3890:
3889:
3885:
3883:
3882:
3878:
3876:
3875:
3874:Gunggamarandu
3871:
3869:
3868:
3867:Gavialosuchus
3864:
3862:
3861:
3860:Ferganosuchus
3857:
3855:
3854:
3850:
3848:
3847:
3843:
3842:
3839:
3836:
3833:
3828:
3810:
3809:
3805:
3803:
3802:
3798:
3796:
3795:
3791:
3789:
3788:
3784:
3782:
3781:
3777:
3776:
3774:
3772:
3771:
3766:
3760:
3759:
3755:
3752:
3751:
3750:Tzaganosuchus
3747:
3744:
3743:
3739:
3736:
3735:
3731:
3728:
3727:
3723:
3722:
3719:
3716:
3714:
3710:
3703:
3702:
3698:
3696:
3695:
3691:
3689:
3688:
3684:
3682:
3681:
3677:
3675:
3674:
3670:
3669:
3667:
3665:
3664:Osteolaeminae
3661:
3654:
3653:
3652:Oxysdonsaurus
3649:
3646:
3645:
3641:
3639:
3638:
3634:
3633:
3630:
3627:
3624:
3619:
3612:
3611:
3607:
3604:
3603:
3602:Jiangxisuchus
3599:
3597:
3596:
3592:
3589:
3588:
3584:
3581:
3580:
3576:
3574:
3573:
3572:Astorgosuchus
3569:
3567:
3566:
3562:
3559:
3558:
3557:Albertosuchus
3554:
3553:
3550:
3547:
3544:
3543:Crocodyloidea
3539:
3535:
3531:
3526:
3522:
3513:
3508:
3478:
3477:
3473:
3471:
3470:
3466:
3464:
3463:
3462:C. gasparinae
3459:
3457:
3456:
3452:
3450:
3449:
3445:
3444:
3442:
3440:
3439:
3434:
3428:
3427:
3423:
3421:
3420:
3416:
3415:
3413:
3411:
3410:
3405:
3398:
3397:
3393:
3391:
3390:
3386:
3383:
3382:
3378:
3376:
3375:
3371:
3369:
3368:
3364:
3362:
3361:
3357:
3355:
3354:
3350:
3348:
3347:
3343:
3341:
3340:
3336:
3334:
3333:
3329:
3327:
3326:
3325:Kuttanacaiman
3322:
3320:
3319:
3315:
3313:
3312:
3308:
3305:
3304:
3300:
3298:
3297:
3296:Culebrasuchus
3293:
3291:
3290:
3286:
3283:
3282:
3278:
3276:
3275:
3271:
3268:
3267:
3266:Brachychampsa
3263:
3261:
3260:
3256:
3254:
3253:
3249:
3248:
3245:
3242:
3240:
3236:
3226:
3225:
3221:
3219:
3218:
3217:A. prenasalis
3214:
3212:
3211:
3207:
3205:
3204:
3200:
3198:
3197:
3193:
3191:
3190:
3186:
3184:
3183:
3179:
3178:
3176:
3174:
3173:
3168:
3161:
3160:
3156:
3153:
3152:
3148:
3145:
3144:
3140:
3137:
3136:
3132:
3130:
3129:
3128:Chrysochampsa
3125:
3122:
3121:
3117:
3114:
3113:
3109:
3107:
3106:
3102:
3099:
3098:
3097:Akanthosuchus
3094:
3093:
3090:
3087:
3085:
3084:Alligatorinae
3081:
3074:
3073:
3072:Sajkanosuchus
3069:
3066:
3065:
3061:
3058:
3057:
3056:Lianghusuchus
3053:
3050:
3049:
3045:
3044:
3042:
3039:
3038:Alligatoridae
3034:
3024:
3023:
3019:
3017:
3016:
3012:
3010:
3009:
3005:
3003:
3002:
3001:Jiangxisuchus
2998:
2996:
2995:
2991:
2989:
2988:
2984:
2982:
2981:
2977:
2976:
2974:
2972:
2968:
2962:
2961:
2957:
2954:
2953:
2949:
2947:
2946:
2942:
2939:
2938:
2934:
2932:
2931:
2927:
2925:
2924:
2920:
2917:
2916:
2912:
2910:
2909:
2905:
2903:
2902:
2901:Brachychampsa
2898:
2896:
2895:
2891:
2889:
2888:
2884:
2883:
2880:
2876:
2873:
2870:
2865:
2859:
2858:
2854:
2852:
2851:
2847:
2845:
2844:
2840:
2838:
2837:
2833:
2832:
2829:
2826:
2823:
2818:
2814:
2810:
2805:
2801:
2786:
2785:
2781:
2779:
2778:
2774:
2771:
2770:
2766:
2763:
2762:
2758:
2756:
2755:
2751:
2749:
2748:
2744:
2742:
2741:
2737:
2735:
2734:
2730:
2727:
2726:
2725:Borealosuchus
2722:
2720:
2719:
2715:
2712:
2711:
2707:
2705:
2704:
2703:Albertosuchus
2700:
2699:
2697:
2693:
2687:
2686:
2682:
2680:
2679:
2675:
2673:
2672:
2668:
2665:
2664:
2660:
2658:
2657:
2653:
2651:
2650:
2646:
2644:
2643:
2639:
2637:
2636:
2632:
2629:
2628:
2624:
2622:
2621:
2617:
2614:
2613:
2609:
2608:
2606:
2604:
2600:
2596:
2591:
2586:
2582:
2557:
2556:
2555:
2554:
2552:
2550:
2546:
2538:
2534:
2533:
2532:
2531:
2528:
2525:
2523:
2519:
2511:
2507:
2506:
2505:
2504:
2501:
2498:
2496:
2492:
2488:
2480:
2476:
2473:
2471:
2467:
2464:
2462:
2458:
2456:
2452:
2450:
2446:
2445:
2440:
2436:
2432:
2424:
2419:
2417:
2412:
2410:
2405:
2404:
2401:
2390:
2386:
2382:
2378:
2374:
2370:
2362:
2359:
2354:
2350:
2345:
2340:
2335:
2330:
2326:
2322:
2321:
2316:
2309:
2306:
2295:
2291:
2285:
2282:
2277:
2273:
2268:
2263:
2258:
2253:
2249:
2245:
2241:
2234:
2231:
2226:
2222:
2217:
2212:
2208:
2204:
2200:
2196:
2192:
2185:
2183:
2181:
2179:
2177:
2175:
2173:
2169:
2164:
2160:
2156:
2152:
2148:
2144:
2140:
2136:
2132:
2128:
2124:
2120:
2116:
2109:
2107:
2105:
2103:
2101:
2099:
2097:
2095:
2093:
2091:
2089:
2087:
2085:
2083:
2081:
2077:
2070:
2068:
2066:
2062:
2057:
2056:aridification
2053:
2049:
2045:
2041:
2037:
2032:
2028:
2024:
2020:
2016:
2012:
2008:
2007:
2002:
1998:
1993:
1991:
1987:
1983:
1979:
1975:
1971:
1970:sebecosuchian
1967:
1966:
1961:
1957:
1952:
1951:
1946:
1942:
1934:
1929:
1925:
1923:
1918:
1914:
1913:
1908:
1900:
1887:
1886:
1878:
1877:
1869:
1868:
1860:
1859:
1856:
1855:
1849:
1848:
1845:
1844:
1836:
1835:
1827:
1826:
1818:
1817:
1809:
1808:
1800:
1799:
1791:
1790:
1787:
1783:
1782:
1781:Quinkana spp.
1775:
1774:
1771:
1770:
1767:
1763:
1762:
1755:
1754:
1748:
1747:
1744:
1743:
1740:
1736:
1735:
1728:
1727:
1721:
1720:
1717:
1716:
1708:
1707:
1699:
1698:
1690:
1689:
1681:
1680:
1672:
1671:
1668:
1664:
1663:
1656:
1655:
1652:
1651:
1648:
1644:
1643:
1636:
1635:
1629:
1628:
1625:
1624:
1621:
1617:
1616:
1609:
1608:
1602:
1601:
1598:
1597:
1594:
1590:
1589:
1582:
1581:
1575:
1574:
1571:
1570:
1567:
1563:
1562:
1555:
1554:
1548:
1547:
1544:
1543:
1540:
1536:
1535:
1534:"Baru" huberi
1528:
1527:
1521:
1520:
1514:
1513:
1510:
1509:
1506:
1502:
1501:
1494:
1493:
1487:
1486:
1483:
1482:
1474:
1473:
1470:
1466:
1465:
1458:
1457:
1451:
1450:
1444:
1443:
1440:
1439:
1436:
1432:
1431:
1424:
1423:
1417:
1414:
1413:
1407:
1406:
1403:
1402:
1399:
1395:
1394:
1387:
1386:
1380:
1379:Crocodyloidea
1377:
1376:
1370:
1369:
1358:
1357:
1349:
1348:
1345:
1344:
1338:
1337:
1334:
1333:
1325:
1324:
1316:
1315:
1307:
1306:
1298:
1297:
1289:
1288:
1280:
1279:
1271:
1270:
1262:
1261:
1258:
1254:
1253:
1246:
1245:
1242:
1241:
1238:
1234:
1233:
1226:
1225:
1219:
1218:
1215:
1214:
1211:
1207:
1206:
1199:
1198:
1192:
1191:
1188:
1187:
1184:
1180:
1179:
1172:
1171:
1165:
1164:
1161:
1160:
1152:
1151:
1148:
1144:
1143:
1142:"Baru" huberi
1136:
1135:
1132:
1131:
1128:
1120:
1119:
1113:
1112:
1106:
1105:
1102:
1101:
1093:
1092:
1084:
1083:
1075:
1074:
1071:
1066:
1065:
1058:
1057:
1054:
1053:
1050:
1046:
1045:
1038:
1037:
1031:
1030:
1027:
1026:
1023:
1019:
1018:
1011:
1010:
1004:
1003:
1000:
999:
991:
990:
987:
983:
982:
981:Pallimnarchus
975:
974:
971:
970:
967:
963:
962:
955:
954:
948:
947:
941:
940:
934:
933:
930:
929:
926:
922:
921:
914:
913:
907:
906:
903:
902:
894:
893:
890:
886:
885:
878:
877:
874:
873:
870:
866:
865:
858:
857:
851:
850:
844:
843:
840:
839:
836:
832:
831:
824:
823:
817:
814:
813:
807:
806:
801:
798:
796:
792:
788:
784:
780:
776:
772:
768:
767:Pallimnarchus
764:
759:
756:
752:
748:
744:
740:
736:
735:
730:
726:
722:
718:
714:
706:
704:
702:
701:
696:
695:
690:
689:
684:
680:
676:
672:
671:quadrate bone
668:
660:
658:
656:
650:
648:
643:
636:
631:
624:
622:
620:
616:
612:
607:
605:
604:
597:
595:
591:
587:
583:
579:
578:
573:
568:
566:
561:
557:
553:
549:
544:
542:
541:
536:
532:
528:
524:
520:
516:
512:
507:
503:
499:
495:
490:
488:
487:
482:
478:
473:
467:
462:
455:
453:
450:
445:
443:
439:
434:
430:
427:
423:
419:
415:
414:
409:
405:
401:
397:
393:
385:
383:
381:
380:
375:
371:
370:
365:
364:
359:
355:
351:
347:
346:aridification
343:
339:
335:
331:
327:
326:
321:
317:
313:
310:
306:
303:
299:
296:
293:
289:
288:
275:
274:
266:
263:
259:
252:
251:
244:
241:
240:
237:
231:
228:
225:
224:
221:
218:
215:
214:
211:
208:
205:
202:
201:
198:
195:
192:
189:
188:
185:
182:
179:
178:
175:
172:
169:
168:
165:
162:
159:
158:
155:
152:
149:
148:
143:
138:
134:
130:
125:
111:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
50:
43:
37:
33:
30:
19:
4544:Mekosuchinae
4465:
4398:
4369:Brevirostres
4319:G. papuensis
4317:
4309:
4301:
4293:
4285:
4277:
4271:G. breviceps
4269:
4262:
4252:
4240:
4233:
4226:
4218:
4211:
4204:
4197:
4192:Penghusuchus
4190:
4182:
4174:
4167:
4160:
4152:
4145:
4138:
4130:
4125:Dadagavialis
4123:
4116:
4104:
4083:
4075:
4067:
4059:
4049:
4038:Thecachampsa
4036:
4028:
4023:Melitosaurus
4021:
4013:
4005:
3993:
3990:Tomistominae
3979:Sacacosuchus
3977:
3951:
3944:
3936:
3931:Thecachampsa
3929:
3922:
3915:
3908:
3901:
3894:
3886:
3879:
3872:
3865:
3858:
3851:
3844:
3806:
3799:
3792:
3787:C. checchiai
3785:
3778:
3768:
3756:
3748:
3740:
3732:
3724:
3713:Crocodylinae
3699:
3692:
3685:
3678:
3671:
3650:
3642:
3635:
3623:Crocodylidae
3610:Mekosuchinae
3608:
3600:
3593:
3585:
3577:
3570:
3563:
3555:
3530:Longirostres
3474:
3467:
3460:
3453:
3448:C. australis
3446:
3436:
3424:
3417:
3409:Melanosuchus
3407:
3394:
3387:
3379:
3372:
3365:
3360:Paranasuchus
3358:
3353:Paranacaiman
3351:
3344:
3337:
3330:
3323:
3316:
3309:
3301:
3294:
3287:
3281:Ceratosuchus
3279:
3272:
3264:
3257:
3250:
3222:
3215:
3208:
3201:
3194:
3187:
3182:A. hailensis
3180:
3170:
3157:
3149:
3143:Navajosuchus
3141:
3133:
3126:
3120:Ceratosuchus
3118:
3110:
3103:
3095:
3070:
3062:
3054:
3046:
3020:
3013:
3006:
2999:
2992:
2985:
2978:
2958:
2950:
2945:Navajosuchus
2943:
2937:Leidyosuchus
2935:
2928:
2921:
2913:
2908:Ceratosuchus
2906:
2899:
2892:
2885:
2855:
2850:Leidyosuchus
2848:
2843:Diplocynodon
2841:
2834:
2782:
2775:
2767:
2759:
2752:
2745:
2738:
2731:
2723:
2718:Asiatosuchus
2716:
2708:
2701:
2683:
2676:
2669:
2661:
2654:
2647:
2640:
2634:
2633:
2625:
2618:
2610:
2603:Mekosuchinae
2592:crocodilians
2510:Pseudosuchia
2495:Pseudosuchia
2474:
2470:Pseudosuchia
2465:
2431:crocodilians
2372:
2368:
2361:
2324:
2318:
2308:
2297:. Retrieved
2294:ScienceDaily
2293:
2284:
2247:
2243:
2233:
2198:
2194:
2122:
2118:
2064:
2060:
2051:
2047:
2043:
2039:
2035:
2030:
2014:
2010:
2004:
2001:Kalthifrons,
2000:
1994:
1989:
1985:
1981:
1977:
1973:
1963:
1959:
1955:
1948:
1944:
1940:
1938:
1932:
1916:
1910:
1906:
1904:
1901:Paleobiology
1854:Crocodylidae
1852:
1779:
1778:
1759:
1758:
1732:
1731:
1660:
1659:
1640:
1639:
1613:
1612:
1586:
1585:
1559:
1558:
1532:
1531:
1498:
1497:
1464:Kambara spp.
1462:
1461:
1429:
1428:
1427:
1416:Mekosuchinae
1391:
1390:
1343:Longirostres
1341:
1250:
1249:
1230:
1229:
1203:
1202:
1176:
1175:
1140:
1139:
1123:
1062:
1061:
1044:Baru darrowi
1042:
1041:
1017:Baru wickeni
1015:
1014:
979:
978:
960:
959:
958:
918:
917:
882:
881:
862:
861:
828:
827:
816:Mekosuchinae
794:
790:
786:
782:
778:
774:
770:
766:
762:
760:
754:
750:
746:
742:
739:sister taxon
732:
725:Kalthifrons'
724:
716:
712:
710:
698:
692:
686:
682:
678:
674:
666:
664:
654:
651:
646:
641:
639:
634:
618:
608:
601:
600:referred to
598:
585:
575:
569:
564:
559:
552:frontal bone
547:
545:
540:Baru wickeni
538:
534:
519:Baru darrowi
518:
514:
510:
493:
491:
484:
480:
476:
471:
470:
465:
446:
441:
437:
412:
411:
391:
389:
378:
377:
367:
361:
357:
353:
349:
337:
333:
329:
323:
319:
295:monospecific
286:
285:
284:
272:
271:
262:Type species
249:
248:
236:Mekosuchinae
226:
203:
190:
35:
29:
4399:Kalthifrons
4295:G. leptodus
4147:Hanyusuchus
4140:Gryposuchus
4069:T. gaudense
3938:Tienosuchus
3917:Ocepesuchus
3881:Kentisuchus
3853:Dollosuchus
3832:Gavialoidea
3374:Purussaurus
3367:Protocaiman
3339:Necrosuchus
3332:Mourasuchus
3318:Gnatusuchus
3259:Bottosaurus
3224:A. thomsoni
3203:A. munensis
3196:A. mefferdi
3112:Arambourgia
3048:Balanerodus
3008:Krabisuchus
2987:Eoalligator
2915:Deinosuchus
2894:Arambourgia
2836:Deinosuchus
2710:Arenysuchus
2678:Ultrastenos
2635:Kalthifrons
2061:Kalthifrons
2052:Kalthifrons
2040:Kalthifrons
2011:Kalthifrons
1990:Kalthifrons
1978:Kalthifrons
1960:Kalthifrons
1945:Kalthifrons
1941:Kalthifrons
1933:Kalthifrons
1917:Kalthifrons
1907:Kalthifrons
961:Kalthifrons
791:Kalthifrons
787:Kalthifrons
775:Kalthifrons
763:Kalthifrons
743:Kalthifrons
717:Kalthifrons
713:Kalthifrons
683:Kalthifrons
667:Kalthifrons
655:Kalthifrons
647:Kalthifrons
642:Kalthifrons
635:Kalthifrons
619:Kalthifrons
560:Kalthifrons
548:Kalthifrons
535:Kalthifrons
523:prefrontals
511:Kalthifrons
506:nasal bones
498:premaxillae
494:Kalthifrons
477:Kalthifrons
472:Kalthifrons
466:Kalthifrons
456:Description
442:Kalthifrons
438:Kalthifrons
392:Kalthifrons
358:Kalthifrons
350:Kalthifrons
338:Kalthifrons
334:Kalthifrons
320:Kalthifrons
305:crocodylian
302:mekosuchine
287:Kalthifrons
250:Kalthifrons
36:Kalthifrons
4538:Categories
4481:Q117466787
4414:Q109546662
4367:See also:
4105:sensu lato
4101:Gavialinae
3965:Gavialidae
3770:Crocodylus
3694:Rimasuchus
3687:Euthecodon
3673:Brochuchus
3419:M. fisheri
3252:Acresuchus
3239:Caimaninae
3189:A. mcgrewi
2869:Globidonta
2649:Mekosuchus
2558:see belowโ
2549:Crocodilia
2461:Sauropsida
2299:2021-01-09
2071:References
2065:Crocodylus
2048:Crocodylus
2036:Crocodylus
2031:Crocodylus
2015:Crocodylus
2006:Crocodylus
1205:Mekosuchus
700:Mekosuchus
615:osteoderms
556:eyesockets
413:SAM P35062
410:(specimen
369:Mekosuchus
354:Crocodylus
330:Crocodylus
325:Crocodylus
220:Crocodilia
4303:G. lewisi
4279:G. browni
4051:Tomistoma
3389:Tsoabichi
3210:A. olseni
3172:Alligator
2656:Paludirex
2537:Neosuchia
2522:Neosuchia
2447:Kingdom:
2250:: e3458.
2225:258878554
2163:133338958
2147:0272-4634
2044:Paludirex
2023:Oligocene
1761:Baru spp.
707:Phylogeny
679:Paludirex
625:Dentition
611:vertebrae
588:from the
374:monotypic
316:Australia
160:Kingdom:
154:Eukaryota
47:3.9
4495:47446643
4475:Wikidata
4428:61146877
4408:Wikidata
4254:Gavialis
3742:Quinkana
3303:Eocaiman
2923:Eocaiman
2663:Quinkana
2479:Eusuchia
2455:Chordata
2453:Phylum:
2449:Animalia
2429:Extinct
2353:30051855
2327:(1881).
2276:28649471
2155:44866024
1986:Quinkana
1974:Quinkana
1965:Quinkana
1922:selenite
1252:Quinkana
751:Quinkana
729:polytomy
603:Quinkana
582:parietal
486:Quinkana
426:Pliocene
420:and two
408:holotype
309:Pliocene
184:Reptilia
174:Chordata
170:Phylum:
164:Animalia
150:Domain:
42:Pliocene
4508:9447114
4441:9511709
3734:Kinyang
3701:Kinyang
2642:Kambara
2459:Class:
2377:Bibcode
2344:6030529
2267:5482264
2203:Bibcode
2127:Bibcode
2027:Miocene
1067:Alcoota
795:Kambara
783:Kambara
747:Kambara
418:cranium
292:extinct
242:Genus:
216:Order:
180:Class:
116:↓
4521:347871
4454:347870
3438:Caiman
2695:Others
2351:
2341:
2274:
2264:
2223:
2161:
2153:
2145:
2038:drove
433:palate
406:. The
290:is an
2685:Volia
2590:Basal
2475:Clade
2466:Clade
2375:(5).
2244:PeerJ
2221:S2CID
2159:S2CID
2151:JSTOR
1954:that
1178:Volia
721:clade
688:Volia
513:from
502:nares
481:Baru,
449:Dieri
298:genus
227:Clade
204:Clade
191:Clade
4503:GBIF
4436:GBIF
3758:Voay
2620:Baru
2535:see
2508:see
2349:PMID
2272:PMID
2143:ISSN
2025:and
1064:Baru
781:and
771:Baru
769:and
755:Baru
697:and
677:and
675:Baru
661:Size
586:Baru
366:and
344:and
55:Pre๊
4490:EoL
4423:EoL
2385:doi
2339:PMC
2329:doi
2325:285
2262:PMC
2252:doi
2211:doi
2135:doi
402:'s
398:in
314:of
300:of
4540::
4518::
4505::
4492::
4477::
4451::
4438::
4425::
4410::
2477::
2468::
2383:.
2371:.
2347:.
2337:.
2323:.
2317:.
2292:.
2270:.
2260:.
2246:.
2242:.
2219:.
2209:.
2199:47
2197:.
2193:.
2171:^
2157:.
2149:.
2141:.
2133:.
2123:37
2121:.
2117:.
2079:^
1992:.
1984:,
797:.
753:,
703:.
691:,
681:,
606:.
543:.
489:.
444:.
382:.
229::
206::
193::
105:Pg
49:Ma
44:,
4322:?
4314:?
4306:?
4298:?
4290:?
4282:?
4274:?
4223:?
4187:?
4179:?
4157:?
4135:?
4088:?
4080:?
4072:?
4064:?
4041:?
4033:?
4018:?
4010:?
3941:?
3891:?
3753:?
3745:?
3737:?
3729:?
3704:?
3655:?
3647:?
3613:?
3605:?
3590:?
3582:?
3560:?
3399:?
3384:?
3306:?
3284:?
3269:?
3162:?
3154:?
3146:?
3138:?
3123:?
3115:?
3100:?
3075:?
3067:?
3059:?
3051:?
2955:?
2940:?
2918:?
2787:?
2772:?
2764:?
2728:?
2713:?
2666:?
2630:?
2615:?
2422:e
2415:t
2408:v
2391:.
2387::
2379::
2373:9
2355:.
2331::
2302:.
2278:.
2254::
2248:5
2227:.
2213::
2205::
2165:.
2137::
2129::
1935:.
1785:โ
1765:โ
1738:โ
1666:โ
1646:โ
1619:โ
1592:โ
1565:โ
1538:โ
1504:โ
1468:โ
1434:โ
1397:โ
1256:โ
1236:โ
1209:โ
1182:โ
1146:โ
1126:โ
1069:โ
1048:โ
1021:โ
985:โ
965:โ
924:โ
888:โ
868:โ
834:โ
269:โ
246:โ
233:โ
110:N
100:K
95:J
90:T
85:P
80:C
75:D
70:S
65:O
60:๊
20:)
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