187:
66:
41:
3424:
239:, which includes bats that have prominent nostrils splaying out in opposite directions. The pelage of the species is gender dimorphic, males are notably darker and fawn-coloured, females are predominantly buff-yellow lightened by a drab colour at the base of the hair.
3484:
253:
in 1993. A review of the systematics two years later elevated the status of the group to species and distinguished island populations previously described as
3499:
3465:
3318:
2976:
3344:
3087:
2952:
2943:
417:
2869:
369:"Systematic review of Nyctimene cephalotes and N. albiventer (Chiroptera : Pteropodidae) in the Maluku and Sulawesi regions, Indonesia"
1281:
1272:
2801:
1326:
3494:
2887:
2837:
1371:
879:
515:
2819:
1290:
3078:
3370:
2878:
2783:
2774:
2307:
1299:
897:
551:
3504:
2484:
1681:
1380:
1344:
1317:
614:
259:. The holotype was collected in 1992 in a mist net, close to sea level, at the edge of forest near a lake, on an island in the
3388:
2810:
2475:
1362:
1353:
665:
3349:
2030:
1881:
1845:
1732:
1335:
1037:
888:
542:
3458:
2792:
2765:
1854:
971:
843:
596:
587:
3133:
3383:
3179:
1441:
1308:
656:
578:
560:
524:
938:
3489:
2828:
2379:
2111:
1976:
1504:
929:
632:
533:
410:
65:
3096:
2644:
1563:
264:
343:(Chiroptera: Pteropodidae) from the Banda, Kai and Aru Islands, Maluku, Indonesia–implications for biogeography"
3509:
2433:
2183:
1872:
1863:
1773:
1404:
1046:
1028:
780:
756:
2388:
2093:
2057:
995:
852:
3451:
3170:
2280:
2012:
1940:
1821:
1797:
1723:
1613:
1522:
1513:
1471:
1159:
1126:
3245:
2617:
2612:
2289:
2192:
2075:
1967:
1480:
1135:
811:
790:
623:
2361:
2343:
2003:
1604:
1554:
1193:
605:
403:
186:
160:
3283:
3212:
1240:
861:
3305:
3188:
2689:
2671:
2370:
2334:
2219:
2147:
2138:
1913:
1764:
255:
249:
308:
2698:
2560:
2406:
2316:
2201:
1985:
1663:
30:
2635:
2542:
2397:
2325:
2256:
2129:
2039:
1949:
1922:
1595:
1019:
60:
3396:
3292:
3046:
3037:
2911:
2626:
2415:
2271:
2246:
2228:
2174:
2165:
2156:
2120:
2021:
1994:
1690:
1672:
1218:
498:
260:
235:
176:
137:
3435:
3401:
3362:
3013:
2653:
2599:
2451:
2442:
2424:
2352:
2298:
2262:
2210:
2084:
2048:
1931:
1699:
870:
723:
303:
2662:
2551:
2237:
2066:
1747:
1413:
1395:
1183:
1094:
1070:
947:
732:
45:
395:
3151:
2102:
1958:
463:
285:
3478:
3357:
3203:
3028:
3004:
2926:
2726:
2684:
962:
771:
647:
294:
214:
50:
3297:
3161:
2994:
2967:
2902:
2738:
2466:
1641:
1578:
1495:
1454:
747:
693:
485:
368:
338:
3375:
3331:
3277:
3061:
2748:
2512:
1109:
1061:
912:
222:
3268:
2525:
1836:
1714:
1537:
1174:
826:
469:
3431:
3114:
2575:
1812:
1231:
1150:
1085:
1010:
706:
77:
3262:
3124:
1896:
1788:
1654:
1255:
986:
802:
457:
451:
445:
439:
267:, Colin Keast, who assisted in the preparation of the specimen's skulls.
218:
97:
3323:
2852:
427:
210:
127:
3423:
3336:
2594:
107:
87:
3239:
263:. The epithet of the species refers to an honorary assistant at the
390:
Mammal
Species of the World: A Taxonomic and Geographic Reference
3310:
3243:
2723:
2509:
1638:
1438:
1215:
690:
482:
399:
392:. Third Edition. The Johns Hopkins University Press, Baltimore
117:
3439:
233:
Keast's tube-nosed fruit bat is a species in the genus
337:
Kitrchener, D.J.; Packer, W.C.; Maryandto, I. (1993).
3252:
3201:
3159:
3150:
3122:
3113:
3059:
3026:
3002:
2993:
2965:
2924:
2900:
2850:
2746:
2737:
2573:
2523:
2464:
1894:
1834:
1810:
1786:
1745:
1712:
1652:
1576:
1535:
1493:
1452:
1393:
1253:
1229:
1172:
1148:
1107:
1083:
1059:
1008:
984:
960:
910:
824:
800:
769:
745:
704:
645:
496:
367:Kitchener, D.J.; Packer, W.C; Suyanto, A. (1995).
247:The taxon was first described as a subspecies of
332:
330:
328:
326:
362:
360:
309:10.2305/IUCN.UK.2020-3.RLTS.T136441A21983677.en
284:Helgen, K.; Hutson, A.M.; Tsang, S.M. (2020).
3459:
411:
8:
3466:
3452:
3240:
3156:
3119:
2999:
2743:
2734:
2720:
2520:
2506:
1649:
1635:
1449:
1435:
1226:
1212:
701:
687:
493:
479:
418:
404:
396:
185:
39:
20:
307:
3088:East African little collared fruit bat (
373:Records of the Western Australian Museum
347:Records of the Western Australian Museum
209:) is a species of megabat in the family
2977:Veldkamp's dwarf epauletted fruit bat (
276:
3485:IUCN Red List near threatened species
2944:Hayman's dwarf epauletted fruit bat (
2802:Lesser Angolan epauletted fruit bat (
388:D.E. Wilson & D.M. Reeder, 2005:
7:
3420:
3418:
2953:Peters' dwarf epauletted fruit bat (
2870:Buettikofer's epauletted fruit bat (
1372:New Britain naked-backed fruit bat (
880:Nusatenggara short-nosed fruit bat (
516:Broad-striped tube-nosed fruit bat (
2820:East African epauletted fruit bat (
1291:Philippine naked-backed fruit bat (
1282:Beaufort's naked-backed fruit bat (
1273:Andersen's naked-backed fruit bat (
295:IUCN Red List of Threatened Species
225:. It was named after Colin Keast.
194:Keast's tube-nosed fruit bat range
3438:. You can help Knowledge (XXG) by
2308:Little golden-mantled flying fox (
1327:Solomon's naked-backed fruit bat (
1300:Halmahera naked-backed fruit bat (
898:Indonesian short-nosed fruit bat (
552:Round-eared tube-nosed fruit bat (
14:
2888:Franquet's epauletted fruit bat (
2838:Wahlberg's epauletted fruit bat (
2485:Sulawesi stripe-faced fruit bat (
1682:Giant golden-crowned flying fox (
1381:Greenish naked-backed fruit bat (
1345:Moluccan naked-backed fruit bat (
1318:Sulawesi naked-backed fruit bat (
615:Philippine tube-nosed fruit bat (
24:Keast's tube-nosed fruit bat
3422:
2811:Ethiopian epauletted fruit bat (
2476:Mindoro stripe-faced fruit bat (
1363:Western naked-backed fruit bat (
1354:Panniet naked-backed fruit bat (
666:Steadfast tube-nosed fruit bat (
64:
3500:Taxa named by Darrell Kitchener
2784:Peters's epauletted fruit bat (
2775:Ansell's epauletted fruit bat (
2031:New Ireland masked flying fox (
1882:New Georgian monkey-faced Bat (
1846:Bougainville monkey-faced Bat (
1733:Small white-winged flying fox (
1336:Lesser naked-backed fruit bat (
889:Greater short-nosed fruit bat (
543:Mountain tube-nosed fruit bat (
2793:Gambian epauletted fruit bat (
2766:Angolan epauletted fruit bat (
1855:Guadalcanal monkey-faced Bat (
844:Lesser short-nosed fruit bat (
597:Demonic tube-nosed fruit bat (
588:Malaita tube-nosed fruit bat (
570:Keast's tube-nosed fruit bat (
1:
3079:São Tomé collared fruit bat (
1309:Biak naked-backed fruit bat (
657:Lesser tube-nosed fruit bat (
579:Island tube-nosed fruit bat (
561:Dragon tube-nosed fruit bat (
525:Common tube-nosed fruit bat (
179:, Packer, and Maryanto, 1993.
3069:(Little collared fruit bats)
2829:Minor epauletted fruit bat (
2380:Philippine gray flying fox (
1977:Bismarck masked flying fox (
1038:Ratanaworabhan's fruit bat (
633:Umboi tube-nosed fruit bat (
201:Keast's tube-nosed fruit bat
3097:Little collared fruit bat (
1755:(Straw-coloured fruit bats)
1564:New Caledonia blossom bat (
972:Fischer's pygmy fruit bat (
3526:
3417:
2434:New Caledonia flying fox (
2184:Black-bearded flying fox (
1873:Montane monkey-faced Bat (
1864:Greater monkey-faced Bat (
1774:Straw-coloured fruit bat (
1405:Sulawesi harpy fruit bat (
1047:White-collared fruit bat (
1029:Javan tailless fruit bat (
939:Rickart's dyak fruit bat (
781:Spotted-winged fruit bat (
757:Mindanao pygmy fruit bat (
3495:Mammals described in 1993
3171:Short-palated fruit bat (
2733:
2719:
2519:
2505:
2281:Geelvink Bay flying fox (
2112:Livingstone's fruit bat (
2013:Large Samoan flying fox (
1941:Small Samoan flying fox (
1822:Small-toothed fruit bat (
1798:Fijian monkey-faced Bat (
1724:White-winged flying fox (
1648:
1634:
1614:Moss-forest blossom bat (
1514:Black-bellied fruit bat (
1505:Fardoulis's blossom bat (
1472:Long-tongued nectar bat (
1462:(Long-tongued fruit bats)
1448:
1434:
1225:
1211:
1127:Greater musky fruit bat (
930:Brooks's dyak fruit bat (
700:
686:
534:Pallas's tube-nosed bat (
492:
478:
434:
426:Extant species of family
265:Western Australian Museum
193:
184:
166:
159:
61:Scientific classification
59:
37:
28:
23:
3430:This article related to
3134:D'Anchieta's fruit bat (
2645:Leschenault's rousette (
2290:Grey-headed flying fox (
2193:Black-eared flying fox (
2076:Ontong Java flying fox (
1968:Ashy-headed flying fox (
1545:(Long-tailed fruit bats)
1481:Long-tongued fruit bat (
1263:(Bare-backed fruit bats)
1136:Lesser musky fruit bat (
812:Black-capped fruit bat (
624:Eastern tube-nosed Bat (
3180:Campo-Ma’an fruit bat (
2389:Temminck's flying fox (
2362:Little red flying fox (
2344:Madagascan flying fox (
2094:Andersen's flying fox (
2058:Gilliard's flying fox (
2004:Spectacled flying fox (
1605:Halmahera blossom bat (
1555:Long-tailed fruit bat (
996:Salim Ali's fruit bat (
853:Horsfield's fruit bat (
606:Lesser tube-nosed bat (
506:(tube-nosed fruit bats)
3505:Bats of Southeast Asia
2934:(Dwarf epauleted bats)
2756:(Epauleted fruit bats)
2690:Long-haired rousette (
2672:Bare-backed rousette (
2371:Seychelles fruit bat (
2335:Rodrigues flying fox (
2220:Mauritian flying fox (
2148:Big-eared flying fox (
1914:Admiralty flying fox (
1765:Madagascan fruit bat (
1523:Woodford's fruit bat (
1160:Blanford's fruit bat (
834:(Dog-faced fruit bats)
791:Balionycteris seimundi
302:: e.T136441A21983677.
2699:Madagascan rousette (
2618:Geoffroy's rousette (
2583:(rousette fruit bats)
2561:Philippine dawn bat (
2407:Vanikoro flying fox (
2317:Solomons flying fox (
2202:Caroline flying fox (
1986:Moluccan flying fox (
1664:Sulawesi flying fox (
339:"Taxonomic status of
3213:Zenker's fruit bat (
2879:Dobson's fruit bat (
2543:Greater nectar bat (
2398:Insular flying fox (
2326:Rennell flying fox (
2130:Okinawa flying fox (
2040:Nicobar flying fox (
1950:Vanuatu flying fox (
1923:Aldabra flying fox (
1596:Common blossom bat (
1241:Bulmer's fruit bat (
1020:Tailless fruit bat (
862:Peters's fruit bat (
256:Nyctimene cephalotes
250:Nyctimene albiventer
3189:Pohle's fruit bat (
3038:Azagnyi fruit bat (
2912:Hammer-headed Bat (
2701:R. madagascariensis
2636:Egyptian rousette (
2627:Sulawesi rousette (
2416:Kosrae flying fox (
2272:Masked flying fox (
2247:Ornate flying fox (
2229:Temotu flying fox (
2175:Indian flying fox (
2166:Mariana fruit bat (
2157:Lesser flying fox (
2139:Lyle's flying fox (
2121:Lombok flying fox (
2022:Ryukyu flying fox (
1995:Makira flying fox (
1735:D. microleucopterus
1691:Palawan fruit bat (
1673:Talaud flying fox (
31:Conservation status
3014:Angolan rousette (
2654:Linduan Rousette (
2620:R. amplexicaudatus
2600:Manado fruit bat (
2452:Dwarf flying fox (
2443:Pemba flying fox (
2425:Large flying fox (
2353:Samoa flying fox (
2299:Bonin flying fox (
2263:Pelew flying fox (
2211:Great flying fox (
2085:Small flying fox (
2049:Banks flying fox (
1932:Black flying fox (
1700:Sunda flying fox (
1117:(Musky fruit bats)
920:(Dayak fruit bats)
871:Minute fruit bat (
724:Borneo fruit bat (
714:(Pygmy fruit bats)
3490:Nyctimene (genus)
3447:
3446:
3412:
3411:
3397:Open Tree of Life
3246:Taxon identifiers
3237:
3236:
3233:
3232:
3229:
3228:
3225:
3224:
3182:C. campomaanensis
3146:
3145:
3109:
3108:
3072:
2989:
2988:
2937:
2863:
2759:
2715:
2714:
2711:
2710:
2663:Comoro rousette (
2586:
2552:Cave nectar bat (
2536:
2533:(Dawn fruit bats)
2501:
2500:
2497:
2496:
2238:Ceram fruit bat (
2067:Gray flying fox (
2006:P. conspicillatus
1907:
1758:
1630:
1629:
1626:
1625:
1589:
1548:
1465:
1430:
1429:
1426:
1425:
1414:Harpy fruit bat (
1266:
1219:Harpyionycterinae
1207:
1206:
1203:
1202:
1184:Swift fruit bat (
1120:
1095:Dusky fruit bat (
1073:O. cartilagonodus
1071:Luzon fruit bat (
948:Dayak fruit bat (
923:
900:C. titthaecheilus
837:
814:C. melanocephalus
733:Pygmy fruit bat (
717:
682:
681:
678:
677:
509:
261:Maluku (province)
198:
197:
54:
3517:
3468:
3461:
3454:
3426:
3419:
3405:
3404:
3392:
3391:
3379:
3378:
3366:
3365:
3353:
3352:
3340:
3339:
3327:
3326:
3314:
3313:
3301:
3300:
3288:
3287:
3286:
3284:Nyctimene keasti
3273:
3272:
3271:
3254:Nyctimene keasti
3241:
3157:
3120:
3081:M. brachycephala
3071:
3070:
3066:
3047:Woermann's bat (
3000:
2936:
2935:
2931:
2862:
2861:
2860:(Epauleted bats)
2857:
2758:
2757:
2753:
2744:
2735:
2721:
2585:
2584:
2580:
2535:
2534:
2530:
2521:
2507:
2373:P. seychellensis
2292:P. poliocephalus
2213:P. neohibernicus
2103:Kei flying fox (
1988:P. chrysoproctus
1959:Aru flying fox (
1916:P. admiralitatum
1906:
1905:
1901:
1757:
1756:
1752:
1650:
1636:
1588:
1587:
1583:
1566:N. neocaledonica
1547:
1546:
1542:
1464:
1463:
1459:
1450:
1436:
1265:
1264:
1260:
1227:
1213:
1119:
1118:
1114:
922:
921:
917:
836:
835:
831:
716:
715:
711:
702:
688:
508:
507:
503:
494:
480:
420:
413:
406:
397:
381:
380:
364:
355:
354:
334:
321:
320:
318:
316:
311:
288:Nyctimene keasti
281:
206:Nyctimene keasti
189:
172:
170:Nyctimene keasti
69:
68:
48:
43:
42:
21:
3525:
3524:
3520:
3519:
3518:
3516:
3515:
3514:
3510:Fruit bat stubs
3475:
3474:
3473:
3472:
3415:
3413:
3408:
3400:
3395:
3387:
3382:
3374:
3369:
3361:
3356:
3348:
3343:
3335:
3330:
3322:
3317:
3309:
3304:
3296:
3291:
3282:
3281:
3276:
3267:
3266:
3261:
3248:
3238:
3221:
3197:
3142:
3105:
3068:
3067:
3065:
3055:
3022:
2985:
2961:
2933:
2932:
2930:
2920:
2896:
2872:E. buettikoferi
2859:
2858:
2856:
2846:
2755:
2754:
2752:
2729:
2707:
2647:R. leschenaulti
2582:
2581:
2579:
2569:
2532:
2531:
2529:
2515:
2493:
2460:
2409:P. tuberculatus
2231:P. nitendiensis
2114:P. livingstonii
2060:P. gilliardorum
1903:
1902:
1900:
1890:
1830:
1806:
1782:
1754:
1753:
1751:
1741:
1708:
1644:
1622:
1585:
1584:
1582:
1572:
1544:
1543:
1541:
1531:
1489:
1461:
1460:
1458:
1444:
1422:
1396:Harpyionycteris
1389:
1356:D. pannietensis
1262:
1261:
1259:
1249:
1221:
1199:
1168:
1144:
1116:
1115:
1113:
1103:
1079:
1055:
1004:
980:
956:
919:
918:
916:
906:
882:C. nusatenggara
833:
832:
830:
820:
796:
765:
759:A. paucidentata
741:
713:
712:
710:
696:
674:
641:
505:
504:
502:
488:
474:
430:
424:
385:
384:
366:
365:
358:
336:
335:
324:
314:
312:
283:
282:
278:
273:
245:
231:
180:
174:
168:
155:
63:
55:
46:Near Threatened
44:
40:
33:
17:
12:
11:
5:
3523:
3521:
3513:
3512:
3507:
3502:
3497:
3492:
3487:
3477:
3476:
3471:
3470:
3463:
3456:
3448:
3445:
3444:
3427:
3410:
3409:
3407:
3406:
3393:
3380:
3367:
3354:
3341:
3328:
3315:
3302:
3289:
3274:
3258:
3256:
3250:
3249:
3244:
3235:
3234:
3231:
3230:
3227:
3226:
3223:
3222:
3220:
3219:
3209:
3207:
3199:
3198:
3196:
3195:
3186:
3177:
3167:
3165:
3154:
3152:Scotonycterini
3148:
3147:
3144:
3143:
3141:
3140:
3130:
3128:
3117:
3111:
3110:
3107:
3106:
3104:
3103:
3094:
3085:
3075:
3073:
3057:
3056:
3054:
3053:
3044:
3034:
3032:
3024:
3023:
3021:
3020:
3010:
3008:
2997:
2991:
2990:
2987:
2986:
2984:
2983:
2973:
2971:
2963:
2962:
2960:
2959:
2950:
2946:M. intermedius
2940:
2938:
2922:
2921:
2919:
2918:
2908:
2906:
2898:
2897:
2895:
2894:
2885:
2876:
2866:
2864:
2848:
2847:
2845:
2844:
2835:
2826:
2817:
2808:
2799:
2790:
2781:
2772:
2762:
2760:
2741:
2731:
2730:
2724:
2717:
2716:
2713:
2712:
2709:
2708:
2706:
2705:
2696:
2679:
2678:
2669:
2660:
2651:
2642:
2638:R. aegyptiacus
2633:
2624:
2607:
2606:
2589:
2587:
2571:
2570:
2568:
2567:
2558:
2549:
2539:
2537:
2517:
2516:
2510:
2503:
2502:
2499:
2498:
2495:
2494:
2492:
2491:
2482:
2478:S. mindorensis
2472:
2470:
2462:
2461:
2459:
2458:
2449:
2440:
2431:
2422:
2413:
2404:
2395:
2386:
2377:
2368:
2359:
2350:
2341:
2337:P. rodricensis
2332:
2323:
2314:
2305:
2296:
2287:
2278:
2269:
2260:
2253:
2244:
2235:
2226:
2217:
2208:
2199:
2190:
2186:P. melanopogon
2181:
2172:
2163:
2154:
2145:
2136:
2132:P. loochoensis
2127:
2123:P. lombocensis
2118:
2109:
2100:
2096:P. intermedius
2091:
2087:P. hypomelanus
2082:
2073:
2064:
2055:
2046:
2037:
2028:
2019:
2010:
2001:
1992:
1983:
1979:P. capistratus
1974:
1965:
1956:
1947:
1938:
1929:
1925:P. aldabrensis
1920:
1910:
1908:
1904:(flying foxes)
1892:
1891:
1889:
1888:
1879:
1870:
1861:
1852:
1842:
1840:
1832:
1831:
1829:
1828:
1818:
1816:
1808:
1807:
1805:
1804:
1794:
1792:
1784:
1783:
1781:
1780:
1771:
1761:
1759:
1743:
1742:
1740:
1739:
1730:
1726:D. leucopterus
1720:
1718:
1710:
1709:
1707:
1706:
1697:
1688:
1679:
1670:
1660:
1658:
1646:
1645:
1639:
1632:
1631:
1628:
1627:
1624:
1623:
1621:
1620:
1611:
1602:
1592:
1590:
1586:(blossom bats)
1574:
1573:
1571:
1570:
1561:
1551:
1549:
1533:
1532:
1530:
1529:
1520:
1511:
1501:
1499:
1491:
1490:
1488:
1487:
1478:
1468:
1466:
1446:
1445:
1442:Macroglossinae
1439:
1432:
1431:
1428:
1427:
1424:
1423:
1421:
1420:
1411:
1401:
1399:
1391:
1390:
1388:
1387:
1378:
1369:
1360:
1351:
1347:D. moluccensis
1342:
1333:
1324:
1315:
1306:
1297:
1288:
1279:
1269:
1267:
1251:
1250:
1248:
1247:
1237:
1235:
1223:
1222:
1216:
1209:
1208:
1205:
1204:
1201:
1200:
1198:
1197:
1190:
1180:
1178:
1170:
1169:
1167:
1166:
1156:
1154:
1146:
1145:
1143:
1142:
1133:
1123:
1121:
1105:
1104:
1102:
1101:
1091:
1089:
1081:
1080:
1078:
1077:
1067:
1065:
1057:
1056:
1054:
1053:
1044:
1035:
1026:
1016:
1014:
1006:
1005:
1003:
1002:
992:
990:
982:
981:
979:
978:
968:
966:
958:
957:
955:
954:
945:
936:
926:
924:
908:
907:
905:
904:
895:
886:
877:
868:
864:C. luzoniensis
859:
855:C. horsfieldii
850:
840:
838:
822:
821:
819:
818:
808:
806:
798:
797:
795:
794:
787:
777:
775:
767:
766:
764:
763:
753:
751:
743:
742:
740:
739:
730:
720:
718:
698:
697:
691:
684:
683:
680:
679:
676:
675:
673:
672:
663:
653:
651:
643:
642:
640:
639:
630:
621:
612:
603:
594:
590:N. malaitensis
585:
576:
567:
558:
549:
540:
531:
522:
512:
510:
490:
489:
483:
476:
475:
473:
472:
466:
464:Laurasiatheria
460:
454:
448:
442:
435:
432:
431:
425:
423:
422:
415:
408:
400:
394:
393:
383:
382:
356:
322:
275:
274:
272:
269:
244:
241:
230:
227:
196:
195:
191:
190:
182:
181:
175:
164:
163:
157:
156:
152:N. keasti
149:
147:
143:
142:
135:
131:
130:
125:
121:
120:
115:
111:
110:
105:
101:
100:
95:
91:
90:
85:
81:
80:
75:
71:
70:
57:
56:
38:
35:
34:
29:
26:
25:
16:Species of bat
15:
13:
10:
9:
6:
4:
3:
2:
3522:
3511:
3508:
3506:
3503:
3501:
3498:
3496:
3493:
3491:
3488:
3486:
3483:
3482:
3480:
3469:
3464:
3462:
3457:
3455:
3450:
3449:
3443:
3441:
3437:
3433:
3428:
3425:
3421:
3416:
3403:
3398:
3394:
3390:
3385:
3381:
3377:
3372:
3368:
3364:
3359:
3355:
3351:
3346:
3342:
3338:
3333:
3329:
3325:
3320:
3316:
3312:
3307:
3303:
3299:
3294:
3290:
3285:
3279:
3275:
3270:
3264:
3260:
3259:
3257:
3255:
3251:
3247:
3242:
3218:
3216:
3211:
3210:
3208:
3206:
3205:
3204:Scotonycteris
3200:
3194:
3192:
3187:
3185:
3183:
3178:
3176:
3174:
3169:
3168:
3166:
3164:
3163:
3158:
3155:
3153:
3149:
3139:
3137:
3132:
3131:
3129:
3127:
3126:
3121:
3118:
3116:
3112:
3102:
3100:
3095:
3093:
3091:
3086:
3084:
3082:
3077:
3076:
3074:
3064:
3063:
3058:
3052:
3050:
3045:
3043:
3041:
3036:
3035:
3033:
3031:
3030:
3029:Megaloglossus
3025:
3019:
3017:
3016:L. angolensis
3012:
3011:
3009:
3007:
3006:
3005:Lissonycteris
3001:
2998:
2996:
2992:
2982:
2980:
2979:N. veldkampii
2975:
2974:
2972:
2970:
2969:
2964:
2958:
2956:
2951:
2949:
2947:
2942:
2941:
2939:
2929:
2928:
2927:Micropteropus
2923:
2917:
2915:
2914:H. monstrosus
2910:
2909:
2907:
2905:
2904:
2899:
2893:
2891:
2886:
2884:
2882:
2877:
2875:
2873:
2868:
2867:
2865:
2855:
2854:
2849:
2843:
2841:
2836:
2834:
2832:
2827:
2825:
2823:
2818:
2816:
2814:
2809:
2807:
2805:
2800:
2798:
2796:
2791:
2789:
2787:
2782:
2780:
2778:
2773:
2771:
2769:
2768:E. angolensis
2764:
2763:
2761:
2751:
2750:
2745:
2742:
2740:
2736:
2732:
2728:
2727:Epomophorinae
2722:
2718:
2704:
2702:
2697:
2695:
2693:
2687:
2686:
2685:Stenonycteris
2681:
2680:
2677:
2675:
2674:R. spinalatus
2670:
2668:
2666:
2665:R. obliviosus
2661:
2659:
2657:
2656:R. linduensis
2652:
2650:
2648:
2643:
2641:
2639:
2634:
2632:
2630:
2629:R. celebensis
2625:
2623:
2621:
2615:
2614:
2609:
2608:
2605:
2603:
2597:
2596:
2591:
2590:
2588:
2578:
2577:
2572:
2566:
2564:
2559:
2557:
2555:
2550:
2548:
2546:
2541:
2540:
2538:
2528:
2527:
2522:
2518:
2514:
2508:
2504:
2490:
2488:
2483:
2481:
2479:
2474:
2473:
2471:
2469:
2468:
2463:
2457:
2455:
2450:
2448:
2446:
2445:P. voeltzkowi
2441:
2439:
2437:
2432:
2430:
2428:
2423:
2421:
2419:
2414:
2412:
2410:
2405:
2403:
2401:
2396:
2394:
2392:
2391:P. temminckii
2387:
2385:
2383:
2378:
2376:
2374:
2369:
2367:
2365:
2364:P. scapulatus
2360:
2358:
2356:
2351:
2349:
2347:
2342:
2340:
2338:
2333:
2331:
2329:
2324:
2322:
2320:
2315:
2313:
2311:
2306:
2304:
2302:
2297:
2295:
2293:
2288:
2286:
2284:
2279:
2277:
2275:
2274:P. personatus
2270:
2268:
2266:
2265:P. pelewensis
2261:
2259:
2258:
2254:
2252:
2250:
2245:
2243:
2241:
2236:
2234:
2232:
2227:
2225:
2223:
2218:
2216:
2214:
2209:
2207:
2205:
2204:P. molossinus
2200:
2198:
2196:
2191:
2189:
2187:
2182:
2180:
2178:
2173:
2171:
2169:
2164:
2162:
2160:
2155:
2153:
2151:
2146:
2144:
2142:
2137:
2135:
2133:
2128:
2126:
2124:
2119:
2117:
2115:
2110:
2108:
2106:
2101:
2099:
2097:
2092:
2090:
2088:
2083:
2081:
2079:
2074:
2072:
2070:
2065:
2063:
2061:
2056:
2054:
2052:
2047:
2045:
2043:
2038:
2036:
2034:
2029:
2027:
2025:
2024:P. dasymallus
2020:
2018:
2016:
2011:
2009:
2007:
2002:
2000:
1998:
1993:
1991:
1989:
1984:
1982:
1980:
1975:
1973:
1971:
1966:
1964:
1962:
1957:
1955:
1953:
1948:
1946:
1944:
1939:
1937:
1935:
1930:
1928:
1926:
1921:
1919:
1917:
1912:
1911:
1909:
1899:
1898:
1893:
1887:
1885:
1880:
1878:
1876:
1871:
1869:
1867:
1862:
1860:
1858:
1853:
1851:
1849:
1844:
1843:
1841:
1839:
1838:
1833:
1827:
1825:
1820:
1819:
1817:
1815:
1814:
1809:
1803:
1801:
1796:
1795:
1793:
1791:
1790:
1785:
1779:
1777:
1772:
1770:
1768:
1763:
1762:
1760:
1750:
1749:
1744:
1738:
1736:
1731:
1729:
1727:
1722:
1721:
1719:
1717:
1716:
1711:
1705:
1703:
1698:
1696:
1694:
1689:
1687:
1685:
1680:
1678:
1676:
1671:
1669:
1667:
1666:A. celebensis
1662:
1661:
1659:
1657:
1656:
1651:
1647:
1643:
1637:
1633:
1619:
1617:
1612:
1610:
1608:
1603:
1601:
1599:
1594:
1593:
1591:
1581:
1580:
1575:
1569:
1567:
1562:
1560:
1558:
1557:N. macdonaldi
1553:
1552:
1550:
1540:
1539:
1534:
1528:
1526:
1521:
1519:
1517:
1512:
1510:
1508:
1507:M. fardoulisi
1503:
1502:
1500:
1498:
1497:
1492:
1486:
1484:
1479:
1477:
1475:
1470:
1469:
1467:
1457:
1456:
1451:
1447:
1443:
1437:
1433:
1419:
1417:
1416:H. whiteheadi
1412:
1410:
1408:
1407:H. celebensis
1403:
1402:
1400:
1398:
1397:
1392:
1386:
1384:
1379:
1377:
1375:
1374:D. praedatrix
1370:
1368:
1366:
1361:
1359:
1357:
1352:
1350:
1348:
1343:
1341:
1339:
1334:
1332:
1330:
1325:
1323:
1321:
1316:
1314:
1312:
1307:
1305:
1303:
1298:
1296:
1294:
1289:
1287:
1285:
1280:
1278:
1276:
1271:
1270:
1268:
1258:
1257:
1252:
1246:
1244:
1239:
1238:
1236:
1234:
1233:
1228:
1224:
1220:
1214:
1210:
1196:
1195:
1194:T. suhaniahae
1191:
1189:
1187:
1186:T. nigrescens
1182:
1181:
1179:
1177:
1176:
1171:
1165:
1163:
1158:
1157:
1155:
1153:
1152:
1147:
1141:
1139:
1134:
1132:
1130:
1125:
1124:
1122:
1112:
1111:
1106:
1100:
1098:
1093:
1092:
1090:
1088:
1087:
1082:
1076:
1074:
1069:
1068:
1066:
1064:
1063:
1058:
1052:
1050:
1045:
1043:
1041:
1036:
1034:
1032:
1027:
1025:
1023:
1018:
1017:
1015:
1013:
1012:
1007:
1001:
999:
994:
993:
991:
989:
988:
983:
977:
975:
970:
969:
967:
965:
964:
963:Haplonycteris
959:
953:
951:
946:
944:
942:
937:
935:
933:
928:
927:
925:
915:
914:
909:
903:
901:
896:
894:
892:
887:
885:
883:
878:
876:
874:
869:
867:
865:
860:
858:
856:
851:
849:
847:
846:C. brachyotis
842:
841:
839:
829:
828:
823:
817:
815:
810:
809:
807:
805:
804:
799:
793:
792:
788:
786:
784:
779:
778:
776:
774:
773:
772:Balionycteris
768:
762:
760:
755:
754:
752:
750:
749:
744:
738:
736:
731:
729:
727:
722:
721:
719:
709:
708:
703:
699:
695:
689:
685:
671:
669:
664:
662:
660:
655:
654:
652:
650:
649:
648:Paranyctimene
644:
638:
636:
631:
629:
627:
622:
620:
618:
613:
611:
609:
604:
602:
600:
595:
593:
591:
586:
584:
582:
577:
575:
573:
568:
566:
564:
563:N. draconilla
559:
557:
555:
550:
548:
546:
541:
539:
537:
536:N. cephalotes
532:
530:
528:
527:N. albiventer
523:
521:
519:
514:
513:
511:
501:
500:
495:
491:
487:
481:
477:
471:
467:
465:
461:
459:
455:
453:
449:
447:
443:
441:
437:
436:
433:
429:
421:
416:
414:
409:
407:
402:
401:
398:
391:
387:
386:
378:
374:
370:
363:
361:
357:
353:(3): 399–417.
352:
348:
344:
342:
333:
331:
329:
327:
323:
310:
305:
301:
297:
296:
291:
289:
280:
277:
270:
268:
266:
262:
258:
257:
252:
251:
242:
240:
238:
237:
228:
226:
224:
220:
216:
212:
208:
207:
202:
192:
188:
183:
178:
173:
171:
165:
162:
161:Binomial name
158:
154:
153:
148:
145:
144:
141:
140:
136:
133:
132:
129:
126:
123:
122:
119:
116:
113:
112:
109:
106:
103:
102:
99:
96:
93:
92:
89:
86:
83:
82:
79:
76:
73:
72:
67:
62:
58:
52:
47:
36:
32:
27:
22:
19:
3440:expanding it
3429:
3414:
3253:
3214:
3202:
3190:
3181:
3172:
3162:Casinycteris
3160:
3136:P. anchietae
3135:
3123:
3098:
3089:
3080:
3060:
3049:M. woermanni
3048:
3039:
3027:
3015:
3003:
2995:Myonycterini
2978:
2968:Nanonycteris
2966:
2954:
2945:
2925:
2913:
2903:Hypsignathus
2901:
2890:E. franqueti
2889:
2880:
2871:
2851:
2840:E. wahlbergi
2839:
2830:
2821:
2812:
2803:
2795:E. gambianus
2794:
2786:E. crypturus
2785:
2776:
2767:
2747:
2739:Epomophorini
2700:
2691:
2682:
2673:
2664:
2655:
2646:
2637:
2628:
2619:
2610:
2601:
2592:
2574:
2562:
2553:
2544:
2524:
2486:
2477:
2467:Styloctenium
2465:
2454:P. woodfordi
2453:
2444:
2435:
2426:
2417:
2408:
2399:
2390:
2382:P. speciosus
2381:
2372:
2363:
2355:P. samoensis
2354:
2345:
2336:
2327:
2318:
2309:
2301:P. pselaphon
2300:
2291:
2282:
2273:
2264:
2257:P. pelagicus
2255:
2248:
2239:
2230:
2221:
2212:
2203:
2195:P. melanotus
2194:
2185:
2176:
2168:P. mariannus
2167:
2159:P. mahaganus
2158:
2149:
2140:
2131:
2122:
2113:
2104:
2095:
2086:
2077:
2068:
2059:
2050:
2041:
2032:
2023:
2014:
2005:
1996:
1987:
1978:
1969:
1960:
1952:P. anetianus
1951:
1943:P. allenorum
1942:
1933:
1924:
1915:
1895:
1883:
1874:
1866:P. flanneryi
1865:
1856:
1847:
1835:
1823:
1811:
1800:M. acrodonta
1799:
1787:
1775:
1767:E. dupreanum
1766:
1746:
1734:
1725:
1713:
1701:
1692:
1683:
1674:
1665:
1653:
1642:Pteropodinae
1615:
1607:S. carolinae
1606:
1598:S. australis
1597:
1579:Syconycteris
1577:
1565:
1556:
1536:
1525:M. woodfordi
1524:
1515:
1506:
1496:Melonycteris
1494:
1482:
1473:
1455:Macroglossus
1453:
1415:
1406:
1394:
1382:
1373:
1364:
1355:
1346:
1337:
1328:
1319:
1310:
1302:D. crenulata
1301:
1292:
1284:D. beauforti
1283:
1275:D. anderseni
1274:
1254:
1242:
1230:
1192:
1185:
1173:
1162:S. blanfordi
1161:
1149:
1137:
1128:
1108:
1096:
1084:
1072:
1060:
1048:
1039:
1030:
1022:M. ecaudatus
1021:
1009:
998:L. salimalii
997:
985:
973:
961:
950:D. spadiceus
949:
940:
931:
911:
899:
890:
881:
872:
863:
854:
845:
825:
813:
801:
789:
782:
770:
758:
748:Alionycteris
746:
734:
725:
705:
694:Cynopterinae
667:
658:
646:
635:N. vizcaccia
634:
626:N. robinsoni
625:
616:
607:
598:
589:
580:
571:
569:
562:
553:
544:
535:
526:
517:
497:
486:Nyctimeninae
462:Superorder:
456:Infraclass:
428:Pteropodidae
389:
376:
372:
350:
346:
340:
313:. Retrieved
299:
293:
287:
279:
254:
248:
246:
234:
232:
211:Pteropodidae
205:
204:
200:
199:
169:
167:
151:
150:
138:
128:Pteropodidae
18:
3332:iNaturalist
3278:Wikispecies
3191:C. ophiodon
3173:C. argynnis
3099:M. torquata
3062:Myonycteris
2955:M. pusillus
2813:E. labiatus
2749:Epomophorus
2513:Rousettinae
2487:S. wallacei
2427:P. vampyrus
2400:P. tonganus
2328:P. rennelli
2240:P. ocularis
2150:P. macrotis
2105:P. keyensis
2078:P. howensis
2051:P. fundatus
2042:P. faunulus
1997:P. cognatus
1970:P. caniceps
1961:P. aruensis
1715:Desmalopex
1702:A. mackloti
1693:A. leucotis
1516:M. melanops
1483:M. sobrinus
1293:D. chapmani
1243:A. bulmerae
1110:Ptenochirus
1062:Otopteropus
1049:M. wetmorei
1040:M. niphanae
1031:M. kusnotoi
974:H. fischeri
941:D. rickarti
913:Dyacopterus
783:B. maculata
726:A. aequalis
554:N. cyclotis
315:19 November
229:Description
223:Kai Islands
3479:Categories
3432:fruit bats
3215:S. zenkeri
3090:M. relicta
3040:M. azagnyi
2881:E. dobsoni
2822:E. minimus
2804:E. grandis
2777:E. anselli
2725:Subfamily
2692:R. lanosus
2563:E. robusta
2554:E. spelaea
2526:Eonycteris
2511:Subfamily
2436:P. vetulus
2418:P. ualanus
2319:P. rayneri
2310:P. pumilus
2249:P. ornatus
2069:P. griseus
2033:P. ennisae
1875:P. pulchra
1837:Pteralopex
1684:A. jubatus
1675:A. humilis
1640:Subfamily
1538:Notopteris
1474:M. minimus
1440:Subfamily
1383:D. viridis
1365:D. peronii
1329:D. inermis
1320:D. exoleta
1217:Subfamily
1175:Thoopterus
932:D. brooksi
873:C. minutus
827:Cynopterus
692:Subfamily
608:N. minutus
599:N. masalai
545:N. certans
484:Subfamily
470:Chiroptera
379:: 125–142.
271:References
221:, and the
118:Chiroptera
3115:Plerotini
2683:Subgenus
2613:Rousettus
2611:Subgenus
2602:R. bidens
2593:Subgenus
2576:Rousettus
2283:P. pohlei
2177:P. medius
1934:P. alecto
1857:P. atrata
1848:P. anceps
1824:N. frosti
1813:Neopteryx
1776:E. helvum
1616:S. hobbit
1311:D. emersa
1232:Aproteles
1151:Sphaerias
1129:P. jagori
1097:P. lucasi
1086:Penthetor
1011:Megaerops
891:C. sphinx
735:A. alecto
707:Aethalops
659:P. raptor
617:N. rabori
572:N. keasti
499:Nyctimene
438:Kingdom:
341:Nyctimene
236:Nyctimene
213:found in
177:Kitchener
146:Species:
139:Nyctimene
84:Kingdom:
78:Eukaryota
3389:13800199
3269:Q1765241
3263:Wikidata
3125:Plerotes
2831:E. minor
2545:E. major
2346:P. rufus
2222:P. niger
2141:P. lylei
1897:Pteropus
1789:Mirimiri
1655:Acerodon
1338:D. minor
1256:Dobsonia
1138:P. minor
987:Latidens
803:Chironax
668:P. tenax
581:N. major
518:N. aello
458:Eutheria
452:Mammalia
446:Chordata
444:Phylum:
440:Animalia
243:Taxonomy
219:Tanimbar
124:Family:
108:Mammalia
98:Chordata
94:Phylum:
88:Animalia
74:Domain:
51:IUCN 3.1
3402:3613459
3376:1004420
3324:7261741
3311:7242818
2853:Epomops
2015:P. coxi
1884:P. taki
1748:Eidolon
468:Order:
450:Class:
134:Genus:
114:Order:
104:Class:
49: (
3363:136441
3350:945977
2595:Boneia
3434:is a
3337:74797
3298:485LH
215:Babar
3436:stub
3358:IUCN
3345:ITIS
3319:GBIF
317:2021
300:2020
3384:MSW
3371:MDD
3306:EoL
3293:CoL
304:doi
3481::
3399::
3386::
3373::
3360::
3347::
3334::
3321::
3308::
3295::
3280::
3265::
2688::
2616::
2598::
377:17
375:.
371:.
359:^
351:16
349:.
345:.
325:^
298:.
292:.
217:,
3467:e
3460:t
3453:v
3442:.
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3193:)
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2017:)
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592:)
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520:)
419:e
412:t
405:v
319:.
306::
290:"
286:"
203:(
53:)
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