513:. The CLAMP results after multiple linear regressions for Republic gave a mean annual temperature of approximately 8.0 °C (46.4 °F), while the LMA gave 9.2 ± 2.0 °C (48.6 ± 3.6 °F). This is lower than the mean annual temperature estimates given for the coastal Puget Group, which is estimated to have been between 15–18.6 °C (59.0–65.5 °F). The bioclimatic analysis for Republic suggests mean annual precipitation amounts of 115 ± 39 cm (45 ± 15 in).
121:
485:
480:
range between 0.7–1.2 km (0.43–0.75 mi) higher than the coastal forests. This is consistent with the paleoelevation estimates for the lake systems, which range between 1.1–2.9 km (1,100–2,900 m), which is similar to the modern elevation 0.8 km (0.50 mi), but higher.
581:
has "massive" genital stipes which sport rounded tops and a small dentical in the center. Both the male and female have rather thick short legs. The female's body is slightly longer than the male's at 5.5 mm (0.22 in) with a 1.31 mm (0.052 in) alitrunk. The small petiole is
582:
similar to the male's, with a subtrapizoidal outline in side profile. The female's head is rectangular, slightly longer than wide and has rounded rear margins above the eye sockets. As with the male's antennae, the female's antennae have scapes curved in the basal area.
348:
Dlussky and
Rasnitsyn noted in the type description that while the holotype male and paratype female were found in separate outcrops of the Klondike Mountain Formation, they felt the two shared enough similarities to be considered from the same species.
808:
Greenwood, D.R.; Archibald, S.B.; Mathewes, R.W; Moss, P.T. (2005). "Fossil biotas from the
Okanagan Highlands, southern British Columbia and northeastern Washington State: climates and ecosystems across an Eocene landscape".
451:
climate, in which winter temperatures rarely dropped low enough for snow, and which were seasonably equitable. The
Okanagan highlands paleoforest surrounding the lakes have been described as precursors to the modern
440:, having distinct seasonal temperature swings which dipped below freezing in the winters. However further study has shown the lake system was surrounded by a warm temperate ecosystem that likely had a
642:
Moss, P. T.; Greenwood, D. R.; Archibald, S. B. (2005). "Regional and local vegetation community dynamics of the Eocene
Okanagan Highlands (British Columbia – Washington State) from palynology".
782:
Wolfe, J.A.; Wehr, W.C. (1987). Middle Eocene dicotyledonous plants from
Republic, northeastern Washington (Report). Bulletin. Vol. 1597. United States Geological Survey. pp. 1–25.
1050:
1209:
412:. The formation preserves an upland lake system surrounded by a mixed conifer–broadleaf forest with nearby volcanism. The pollen flora has notable elements of
322:
886:
Dlussky, G. M.; Rasnitsyn, A. P. (1999). "Two new species of aculeate hymenopterans (Vespida=Hymenoptera) from the Middle Eocene of the United States".
1130:
1037:
1194:
756:
1199:
453:
329:, while the paratype was from site B4599 north of Republic at Mt Elizabeth. The fossils were described by myrmecologists Gennady Dlussky and
1214:
948:
Perfilieva, K. S. (2021). "Distribution and
Differentiation of Fossil Oecophylla (Hymenoptera: Formicidae) Species by Wing Imprints".
846:"Modernisation of the Hymenoptera: ants, bees, wasps, and sawflies of the early Eocene Okanagan Highlands of western North America"
519:
is one of four ants described from in and around
Republic. Two other fossils have been placed as members of the subfamily
312:
510:
405:
319:
1204:
905:
Perfilieva, K. S.; Dubovikoff, D. A.; Dlussky, G. M. (2017). "Miocene Ants (Hymenoptera, Formicidae) from Crimea".
120:
1189:
612:
398:
613:"Ants (Hymenoptera: Formicidae) of Formation Green River and some other Middle Eocene deposits of North America"
701:"The Ants (Hymenoptera: Formicidae) of Bol'shaya Svetlovodnaya (Late Eocene of Sikhote-Alin, Russian Far East)"
680:
Barden, P. (2017). "Fossil ants (Hymenoptera: Formicidae): ancient diversity and the rise of modern lineages".
570:
578:
525:
338:
326:
992:
549:
1156:
1076:
354:
219:
1117:
1024:
957:
914:
818:
651:
409:
239:
469:
973:
930:
865:
736:
1. Early Eocene
Lagerstätten of the Okanagan Highlands (British Columbia and Washington State)".
330:
308:
275:
115:
1161:
1081:
359:
1143:
1063:
1055:
574:
367:
1148:
1068:
278:, United States. The genus is currently not placed into any ant subfamily, being treated as
965:
922:
857:
826:
783:
732:
Archibald, S.; Greenwood, D.; Smith, R.; Mathewes, R.; Basinger, J. (2011). "Great
Canadian
712:
659:
506:
502:
484:
473:
315:
717:
700:
961:
918:
822:
765:
Journal of the
Faculty of Science, Hokkaido University. Series 4, Geology and Mineralogy
655:
280:
1184:
1178:
977:
845:
457:
441:
296:
64:
934:
869:
444:
437:
417:
271:
254:
1108:
1015:
535:. The fourth species was first described as a member of the carpenter ant genus
531:
520:
489:
465:
448:
172:
39:
969:
926:
543:
537:
261:
84:
49:
477:
433:
152:
132:
89:
33:
1102:
1009:
566:
429:
402:
375:
364:
342:
304:
300:
107:
79:
74:
59:
54:
44:
861:
577:
attached to a curved scape approximately as long as the head width. The
565:
has a 4.8 mm (0.19 in) body with 2.05 mm (0.081 in)
1135:
1042:
498:
94:
69:
436:. Wolfe and Tanai (1987) interpreted the forest climate to have been
461:
425:
162:
142:
986:
844:
Archibald, S. B.; Rasnitsyn, A.; Brothers, D.; Mathewes, R. (2018).
830:
663:
788:
483:
413:
337:
as a reference to the Klondike Mountain Formation, and coined the
257:
1122:
1029:
990:
456:
of Eastern North America and Eastern Asia. Based on the fossil
421:
182:
606:
604:
602:
600:
598:
596:
594:
501:
for the Klondike Mountain Formation have been derived from
699:
Dlussky, G.M.; Rasnitsyn, A.P.; Perfilieva, K.S. (2015).
1092:
999:
761:(maples) in the Cenozoic of Western North America"
503:climate leaf analysis multivariate program (CLAMP)
803:
801:
799:
881:
879:
472:of Western Washington, which are described as
757:"Systematics, Phylogeny, and Distribution of
573:of subtrapizoidal outline. The antennae have
382:females have heads wider than long, while in
8:
675:
673:
345:honoring Pat White who found the holotype.
987:
637:
635:
633:
460:the lakes were higher and cooler than the
318:collection. The holotype was found at the
20:
787:
716:
270:. The species is solely known from the
590:
541:but was moved to the weaver ant genus
529:and one identified to the form genus
454:temperate broadleaf and mixed forests
7:
1210:Prehistoric insects of North America
718:10.23885/1814-3326-2015-11-1-131-152
325:site B4131, which is designated the
307:female, number SR 94-04-24 and both
611:Dlussky, G.; Rasnitsyn, A. (2002).
397:is known from two locations in the
811:Canadian Journal of Earth Sciences
644:Canadian Journal of Earth Sciences
14:
464:coastal forests preserved in the
378:and differing head proportions.
333:(2002) who picked the genus name
705:Caucasian Entomological Bulletin
420:, and distinct trace amounts of
264:with a single described species
119:
755:Wolfe, J.A.; Tanai, T. (1987).
353:males differ from the Ypresian
274:sediments exposed in northeast
386:the head is longer than wide.
1:
1195:Fossil taxa described in 2002
620:Russian Entomological Journal
476:ecosystems. Estimates of the
390:Distribution and paleoecology
313:Stonerose Interpretive Center
303:male, number SR 94-05-07 and
1200:Hymenoptera of North America
320:Klondike Mountain Formations
1215:Klondike Mountain Formation
406:Klondike Mountain Formation
1231:
507:leaf margin analysis (LMA)
288:History and classification
970:10.1134/S003103012101010X
927:10.1134/S0031030117040098
850:The Canadian Entomologist
399:Eocene Okanagan Highlands
238:Dlussky & Rasnitsyn,
225:
218:
116:Scientific classification
114:
23:
295:was identified from two
950:Paleontological Journal
907:Paleontological Journal
888:Paleontological Journal
526:Propalosoma gutierrezae
499:mean annual temperature
474:lowland tropical forest
401:, both outcrops of the
494:
355:Green River Formations
1157:Paleobiology Database
1077:Paleobiology Database
487:
374:female has a shorter
311:are preserved in the
16:Extinct genus of ants
962:2021PalJ...55...76P
919:2017PalJ...51..391P
862:10.4039/tce.2017.59
823:2005CaJES..42..167G
682:Myrmecological News
656:2005CaJES..42..187M
550:Oecophylla kraussei
470:Chuckanut Formation
368:reproductive stipes
309:compression fossils
561:The known male of
495:
445:upper microthermal
331:Alexandr Rasnitsyn
316:paleoentomological
260:in the ant family
1205:Fossil ant genera
1172:
1171:
1144:Open Tree of Life
1094:Klondikia whiteae
1064:Open Tree of Life
993:Taxon identifiers
738:Geoscience Canada
575:filiform funiculi
563:Klondikia whiteae
517:Klondikia whiteae
497:Estimates of the
449:lower mesothermal
395:Klondikia whiteae
360:Eoformica pinguis
293:Klondikia whiteae
267:Klondikia whiteae
246:
245:
232:Klondikia whiteae
1222:
1190:Ypresian insects
1165:
1164:
1152:
1151:
1139:
1138:
1126:
1125:
1113:
1112:
1111:
1085:
1084:
1072:
1071:
1059:
1058:
1046:
1045:
1033:
1032:
1020:
1019:
1018:
988:
982:
981:
945:
939:
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902:
896:
895:
883:
874:
873:
841:
835:
834:
805:
794:
793:
791:
779:
773:
772:
752:
746:
745:
729:
723:
722:
720:
696:
690:
689:
677:
668:
667:
639:
628:
627:
617:
608:
509:of the Republic
408:in and north of
363:in notably more
339:specific epithet
276:Washington state
234:
230:
194:
124:
123:
106:
36:
29:Temporal range:
21:
1230:
1229:
1225:
1224:
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1220:
1219:
1175:
1174:
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1168:
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1134:
1129:
1121:
1116:
1107:
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1075:
1067:
1062:
1054:
1049:
1041:
1036:
1028:
1023:
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1008:
995:
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947:
946:
942:
904:
903:
899:
885:
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877:
843:
842:
838:
831:10.1139/e04-100
807:
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797:
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776:
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753:
749:
731:
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698:
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693:
679:
678:
671:
664:10.1139/E04-095
641:
640:
631:
615:
610:
609:
592:
588:
559:
392:
290:
242:
236:
228:
227:
214:
211:K. whiteae
192:
118:
110:
105:
104:
103:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
31:
30:
27:
17:
12:
11:
5:
1228:
1226:
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1212:
1207:
1202:
1197:
1192:
1187:
1177:
1176:
1170:
1169:
1167:
1166:
1153:
1140:
1127:
1114:
1098:
1096:
1090:
1089:
1087:
1086:
1073:
1060:
1047:
1034:
1021:
1005:
1003:
997:
996:
991:
984:
983:
940:
913:(4): 391–401.
897:
875:
856:(2): 205–257.
836:
817:(2): 167–185.
795:
774:
747:
724:
711:(1): 131–152.
691:
669:
650:(2): 187–204.
629:
589:
587:
584:
558:
555:
523:, the species
478:paleoelevation
391:
388:
297:type specimens
289:
286:
281:incertae sedis
253:is an extinct
244:
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15:
13:
10:
9:
6:
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3:
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1201:
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1128:
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1039:
1035:
1031:
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1007:
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959:
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951:
944:
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932:
928:
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876:
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837:
832:
828:
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816:
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804:
802:
800:
796:
790:
789:10.3133/b1597
785:
778:
775:
770:
766:
762:
760:
751:
748:
744:(4): 155–164.
743:
739:
735:
728:
725:
719:
714:
710:
706:
702:
695:
692:
687:
683:
676:
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552:
551:
546:
545:
540:
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534:
533:
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527:
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508:
505:analysis and
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459:
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381:
377:
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336:
332:
328:
327:type locality
324:
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283:
282:
277:
273:
269:
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233:
224:
221:
220:Binomial name
217:
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109:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
41:
35:
26:
22:
19:
1093:
1000:
956:(1): 76–89.
953:
949:
943:
910:
906:
900:
891:
887:
853:
849:
839:
814:
810:
777:
768:
764:
758:
750:
741:
737:
734:Lagerstätten
733:
727:
708:
704:
694:
685:
681:
647:
643:
623:
619:
562:
560:
548:
542:
536:
530:
524:
516:
515:
496:
488:
438:microthermal
418:golden larch
394:
393:
383:
379:
371:
370:, while the
358:
350:
347:
334:
292:
291:
279:
272:Early Eocene
266:
265:
255:hymenopteran
249:
248:
247:
231:
226:
210:
209:
196:
195:
24:
18:
771:(1): 1–246.
557:Description
532:Myrmeciites
521:Myrmeciinae
490:Myrmeciites
466:Puget Group
365:sclerotized
173:Hymenoptera
32:49.42
1179:Categories
894:: 546–549.
626:: 411–436.
586:References
569:and small
544:Oecophylla
538:Camponotus
511:paleoflora
384:K. whiteae
380:E. pinguis
372:K. whiteae
351:K. whiteae
262:Formicidae
183:Formicidae
153:Arthropoda
1109:Q19612741
1016:Q19862601
1001:Klondikia
978:232070811
553:in 2017.
335:Klondikia
250:Klondikia
204:Species:
197:Klondikia
139:Kingdom:
133:Eukaryota
25:Klondikia
1123:31359206
1103:Wikidata
1030:25139159
1010:Wikidata
935:90536477
870:90017208
567:alitrunk
410:Republic
403:Ypresian
376:alitrunk
357:species
343:matronym
305:paratype
301:holotype
179:Family:
149:Phylum:
143:Animalia
129:Domain:
108:Ypresian
1149:6281942
1136:8555462
1069:4583926
1056:1140933
1043:4679036
958:Bibcode
915:Bibcode
819:Bibcode
688:: 1–30.
652:Bibcode
571:petiole
430:cypress
229:†
208:†
189:Genus:
169:Order:
163:Insecta
159:Class:
101:↓
1162:256056
1082:255948
976:
933:
868:
579:gaster
462:coeval
458:biotas
432:, and
426:spruce
299:, the
1051:IRMNG
974:S2CID
931:S2CID
866:S2CID
616:(PDF)
442:mesic
414:birch
341:as a
258:genus
1185:Ants
1131:GBIF
1038:GBIF
759:Acer
493:sp.
468:and
434:palm
416:and
323:UWBM
240:2002
40:PreꞒ
1118:EoL
1025:EoL
966:doi
923:doi
858:doi
854:150
827:doi
784:doi
713:doi
660:doi
547:as
447:to
422:fir
1181::
1159::
1146::
1133::
1120::
1105::
1079::
1066::
1053::
1040::
1027::
1012::
972:.
964:.
954:55
952:.
929:.
921:.
911:51
909:.
892:33
890:.
878:^
864:.
852:.
848:.
825:.
815:42
813:.
798:^
769:22
767:.
763:.
742:38
740:.
709:11
707:.
703:.
686:24
684:.
672:^
658:.
648:42
646:.
632:^
624:11
622:.
618:.
593:^
428:,
424:,
284:.
90:Pg
34:Ma
980:.
968::
960::
937:.
925::
917::
872:.
860::
833:.
829::
821::
792:.
786::
721:.
715::
666:.
662::
654::
193:†
95:N
85:K
80:J
75:T
70:P
65:C
60:D
55:S
50:O
45:Ꞓ
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