423:
the rear edge are much higher than the fourteen on the front edge, which cover only the nearly horizontal upper part of the front edge. The tooth base, though very wide, is constricted. The base is asymmetrical, with the right side in front view protruding much further than the left side; because it is not known whether it is a left or right tooth, it cannot be established what is the inner and what the outer side. The denticles are separated by "blood grooves", and "blood pits" are also present. The back denticles point obliquely upwards but have no hooked upper corners.
151:
1246:
130:
1223:
478:
412:
422:
The tooth is two millimeters tall and very recurved, with a strongly convex front edge and a nearly vertical back edge. The tooth is stout, with a maximum fore-aft length of 1.9 millimeters. Both edges are serrated, showing low rectangular denticles (individual serrations). The twelve denticles on
403:, about 151 million years old. The taxon is thus present in stratigraphic zone 6 of the Morrison Formation. The specimen consists of a single maxillary (upper cheek) tooth crown (the root is missing). It cannot be determined whether it represents a left or a right tooth.
439:
based on the anatomy of the tooth. At that time, it was the oldest known troodontid, and is the first of that group discovered from the
Jurassic. A troodontid presence in the Jurassic was predicted by the standard theory regarding the
444:, claiming that birds and troodontids are closely related (Chure suggested that they may even have been sister taxa, though this is not currently supported). Within this context,
1311:
1357:
1298:
1362:
658:
552:, a new dinosaur from the Morrison Formation (Upper Jurassic) of Dinosaur National Monument; the oldest troodontid (Theropoda: Maniraptora)."
1239:
504:
449:
1367:
651:
345:
150:
755:
523:
Chure, D.J. and Britt, B.B., 1993, "New data on theropod dinosaurs from the Late
Jurassic Morrison Fm. (MF)",
373:
1266:
1324:
644:
340:
reported the discovery of small theropod remains, found by screenwashing large amounts of earth in the
1352:
1249:
809:
392:
341:
337:
333:
1020:
782:
396:
145:
1329:
1222:
1316:
1008:
920:
898:
497:
491:
1207:
1186:
964:
483:
459:
416:
304:
452:
argument, that such a close relationship were unlikely because the then oldest known bird,
1179:
1071:
1044:
1000:
942:
740:
463:
441:
380:, who, in the early twentieth century, excavated the Dinosaur National Monument quarry.
1150:
1037:
979:
972:
949:
927:
891:
1346:
582:
Brochu, C.A., and Norell, M.A., 2000, "Temporal congruence and the origin of birds",
454:
377:
369:
357:
349:
291:
80:
41:
466:; later also non-fragmentary Jurassic troodontid material was reported, from China.
1171:
1157:
1143:
1128:
1112:
1080:
1052:
956:
906:
877:
855:
821:
667:
436:
400:
372:κοπάριον, "small surgical knife", in reference to the small size of the tooth. The
311:
308:
260:
234:
623:
coelurosaurian dinosaurs and their implications for understanding avian origins",
17:
1136:
1101:
1094:
1029:
993:
986:
913:
794:
129:
55:
1289:
1164:
706:
697:
473:
208:
100:
65:
1200:
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767:
715:
221:
162:
105:
49:
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691:
384:
301:
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195:
182:
137:
95:
90:
75:
70:
60:
411:
1303:
1193:
1060:
724:
110:
85:
603:
troodontid theropod from China with long feathers on the metatarsus",
571:
Jurassic West: The
Dinosaurs of the Morrison Formation and Their World
685:
172:
1260:
410:
287:
353:
315:
1264:
678:
640:
636:
356:. In 1994, Chure named and described a unique tooth as the
1273:
1111:
1070:
1019:
875:
820:
793:
766:
739:
324:which is known only from a single isolated tooth.
599:Hu D., Hou L., Zhang L. and Xu X., 2009, "A pre-
501:- Another troodontid reported from the Jurassic.
652:
8:
544:
542:
540:
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536:
1261:
883:
872:
799:
772:
745:
736:
675:
659:
645:
637:
128:
31:
573:. Indiana University Press. pp. 327–329.
554:Brigham Young University Geology Studies
516:
318:. It contains the single named species
619:Xu X., Ma Q. & Hu D., 2010, "Pre-
7:
1358:Dinosaurs of the Morrison Formation
584:Journal of Vertebrate Paleontology
525:Journal of Vertebrate Paleontology
25:
1245:
1244:
1221:
476:
462:, i.e. the Troodontidae and the
149:
569:Foster, J. (2007). "Appendix."
505:Timeline of troodontid research
458:, lived much earlier than the
1:
1363:Fossil taxa described in 1994
448:could be used to refute the
1384:
346:Dinosaur National Monument
27:Extinct genus of dinosaurs
1234:
1219:
886:
871:
852:
802:
775:
748:
735:
674:
266:
259:
146:Scientific classification
144:
136:
127:
34:
625:Chinese Science Bulletin
548:Chure, D. J. (1994). "
419:
1325:Paleobiology Database
414:
399:dating from the late
1368:Paleontology in Utah
393:Brushy Basin Member
391:, was found in the
364:. The generic name
360:of the new species
1021:Jinfengopteryginae
550:Koparion douglassi
420:
397:Morrison Formation
362:Koparion douglassi
321:Koparion douglassi
269:Koparion douglassi
18:Koparion douglassi
1340:
1339:
1267:Taxon identifiers
1258:
1257:
1230:
1229:
1217:
1216:
1009:Halszkaraptorinae
921:Hesperornithoides
899:Archaeornithoides
867:
866:
863:
862:
848:
847:
844:
843:
840:
839:
498:Hesperornithoides
492:List of dinosaurs
279:
278:
255:
16:(Redirected from
1375:
1333:
1332:
1320:
1319:
1307:
1306:
1294:
1293:
1292:
1262:
1248:
1247:
1225:
1187:Stenonychosaurus
965:Polyodontosaurus
884:
873:
800:
773:
746:
737:
730:
729:
676:
661:
654:
647:
638:
631:
617:
611:
597:
591:
580:
574:
567:
561:
546:
531:
521:
486:
484:Dinosaurs portal
481:
480:
479:
460:Deinonychosauria
450:temporal paradox
417:life restoration
300:, from the late
253:
246:
233:
154:
153:
132:
122:
52:
40:Temporal range:
32:
21:
1383:
1382:
1378:
1377:
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1373:
1372:
1343:
1342:
1341:
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1328:
1323:
1315:
1310:
1302:
1297:
1288:
1287:
1282:
1269:
1259:
1254:
1226:
1213:
1180:Saurornithoides
1107:
1072:Sinovenatorinae
1066:
1045:Liaoningvenator
1015:
1001:Anchiornithidae
943:Liaoningvenator
880:
859:
836:
816:
789:
762:
756:Avemetatarsalia
741:Avemetatarsalia
731:
681:
680:
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665:
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581:
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513:
482:
477:
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464:Dromaeosauridae
442:origin of birds
431:Chure assigned
429:
409:
368:comes from the
330:
275:
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38:
28:
23:
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15:
12:
11:
5:
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1215:
1214:
1212:
1211:
1204:
1197:
1190:
1183:
1176:
1168:
1161:
1154:
1151:Latenivenatrix
1147:
1140:
1133:
1125:
1117:
1115:
1109:
1108:
1106:
1105:
1098:
1091:
1084:
1076:
1074:
1068:
1067:
1065:
1064:
1057:
1049:
1041:
1038:Jinfengopteryx
1034:
1025:
1023:
1017:
1016:
1014:
1013:
1005:
997:
990:
983:
976:
973:Sinornithoides
969:
961:
953:
950:Papiliovenator
946:
939:
931:
928:Jianianhualong
924:
917:
910:
903:
895:
892:Albertavenator
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502:
494:
488:
487:
471:
468:
428:
427:Classification
425:
408:
405:
329:
326:
292:coelurosaurian
277:
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26:
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14:
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669:
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648:
643:
642:
639:
629:
626:
622:
621:Archaeopteryx
616:
613:
609:
606:
602:
601:Archaeopteryx
596:
593:
589:
585:
579:
576:
572:
566:
563:
559:
555:
551:
545:
543:
541:
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510:
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500:
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489:
485:
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469:
467:
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461:
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455:Archaeopteryx
451:
447:
443:
438:
434:
426:
424:
418:
415:Hypothetical
413:
406:
404:
402:
398:
394:
390:
386:
381:
379:
378:Earl Douglass
375:
374:specific name
371:
370:Ancient Greek
367:
363:
359:
358:type specimen
355:
351:
350:Uintah County
347:
343:
339:
335:
327:
325:
323:
322:
317:
313:
310:
306:
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126:
112:
107:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
51:
43:
42:Late Jurassic
37:
33:
30:
19:
1274:
1206:
1199:
1192:
1185:
1178:
1172:Philovenator
1170:
1163:
1158:Linhevenator
1156:
1149:
1144:Hypnovenator
1142:
1135:
1129:Byronosaurus
1127:
1120:
1113:Troodontinae
1100:
1093:
1086:
1081:Daliansaurus
1079:
1059:
1053:Philovenator
1051:
1043:
1036:
1028:
1007:
999:
992:
985:
978:
971:
963:
957:Paronychodon
955:
948:
941:
934:
933:
926:
919:
912:
907:Geminiraptor
905:
897:
890:
878:Troodontidae
856:Troodontidae
854:
822:Troodontidae
720:
711:
702:
668:Troodontidae
627:
624:
620:
615:
607:
604:
600:
595:
587:
583:
578:
570:
565:
557:
553:
549:
527:
524:
519:
496:
453:
445:
437:Troodontidae
432:
430:
421:
401:Kimmeridgian
388:
382:
365:
361:
342:Rainbow Park
338:Brooks Britt
334:Daniel Chure
331:
320:
319:
309:Kimmeridgian
282:
281:
280:
268:
267:
261:Type species
248:
247:
235:Troodontidae
215:
202:
189:
35:
29:
1353:Troodontids
1137:Gobivenator
1102:Sinusonasus
1095:Sinovenator
994:Xixiasaurus
987:Tochisaurus
914:Harenadraco
810:Maniraptora
795:Maniraptora
630:: 3971–3977
407:Description
274:Chure, 1994
254:Chure, 1994
1347:Categories
1238:See also:
1165:Pectinodon
831:see below↓
707:Dinosauria
698:Sauropsida
511:References
387:specimen,
209:Saurischia
196:Dinosauria
1208:Zanabazar
1201:Urbacodon
1122:Borogovia
783:Theropoda
768:Theropoda
716:Theropoda
684:Kingdom:
610:: 640-643
590:: 197-200
446:Koparion
389:DINO 3353
332:In 1993,
328:Discovery
290:of small
222:Theropoda
169:Kingdom:
163:Eukaryota
48:151
1290:Q1821811
1284:Wikidata
1275:Koparion
1250:Category
1240:Timeline
935:Koparion
692:Chordata
690:Phylum:
686:Animalia
560:: 11-15.
530:(3): 30A
470:See also
433:Koparion
385:holotype
366:Koparion
302:Jurassic
298:dinosaur
295:theropod
283:Koparion
249:Koparion
228:Family:
183:Chordata
179:Phylum:
173:Animalia
159:Domain:
138:Holotype
36:Koparion
1317:1141797
1304:4821488
1194:Troodon
1061:Tamarro
725:Paraves
696:Class:
435:to the
395:of the
376:honors
241:Genus:
117:↓
605:Nature
314:), of
305:Period
140:tooth
1330:68411
1312:IRMNG
1030:Almas
980:Talos
721:Clade
712:Clade
703:Clade
344:near
312:stage
288:genus
286:is a
216:Clade
203:Clade
190:Clade
1299:GBIF
808:see
781:see
754:see
383:The
354:Utah
336:and
316:Utah
56:PreꞒ
1088:Mei
608:461
348:in
1349::
1327::
1314::
1301::
1286::
723::
714::
705::
628:55
588:20
586:,
558:40
556:,
535:^
528:13
352:,
218::
205::
192::
106:Pg
50:Ma
44:,
1175:?
1132:?
1056:?
1048:?
1033:?
1012:?
1004:?
968:?
960:?
938:?
902:?
660:e
653:t
646:v
307:(
245:†
232:†
111:N
101:K
96:J
91:T
86:P
81:C
76:D
71:S
66:O
61:Ꞓ
20:)
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