1270:(ray-finned) fishes were first described form the basin by Sven Nilsson in 1827, with further studies being conducted by paleontologist John W. Davis in 1890 and by paleontologist Mohamad Bazzi and colleagues in 2016. Though shark teeth have been recovered from the basin since early times, the shark and ray fauna was first studied in detail by paleontologist Mikael Siverson in the 20th century, who in a series of publications from 1989 to 2016, some together with other authors, identified a vast number of species.
251:
1724:
1349:
2506:
119:
1697:. The last comprehensive review of the plesiosaur fauna in the Kristianstad Basin was done by paleontologist Per-Ove Persson in the 1960s and his taxonomy is still used with caution, pending a much-needed new review. In particular, the elasmosaurid fossil material lacks important diagnostical features and might not be identifiable to the genus level. Elasmosaurids recognized as being present in the basin, per Persson's taxonomy, include two species of the large
1320:
1528:
1329:
1519:
1850:
916:
923:
285:
276:
267:
258:
3695:
1983:
348:
339:
330:
321:
312:
303:
294:
721:
1675:
930:
1340:
1209:
210:
2492:
2412:
1448:, recovered at Ă…sen and Ullstorp. The discovery of the leptoceratopsid remains were scientifically significant as they represented the first record of ceratopsians in Europe and also contradicted the previously prevailing hypothesis that more primitive members of that group preferred semi-arid and arid environments rather far away from coastal regions.
1840:, presumably occupied the top of the food web. The largest sharks presumably fed on most of the animals present in the basin, including sea turtles, smaller mosasaurs, plesiosaurs and other sharks. Bite marks from sharks are relatively common on reptile bones in the Kristianstad Basin. The overall most common genus of shark found is the lamniform
1064:
1052:
1040:
1028:
1016:
1004:
992:
980:
968:
956:
944:
1264:
geologist and mineralogist Assar
Hadding in 1919, Alf Lundegren in 1934, paleontologist Gustaf Troedsson in 1946 and 1954, paleontologist Richard Hägg in 1954 and geologist and paleontologist Fritz Brotzen in 1960. Further studies on the invertebrate fauna, as with the more publicized vertebrate fossils, have continued into the 21st century.
2455:
nektonic predators (such as cephalopods and fish), which then in turn attracted larger predators. It is probable that the richness of the environment also made Ivö Klack a nursery and feeding ground for migratory species, similar to rocky shores today. The discovery of the basin's only crocodylomorph at Ivö Klack might indicate that
1110:. Ivö Klack was once the shores of a small and rocky Cretaceous island and is one of the most extensively excavated sites in the basin, having been excavated since quarrying activities started in 1888. Though quarrying stopped in the late 1960s, the site continues to be excavated by paleontologists and amateurs.
1306:, from 1954 to 1996. More recent work on marine reptiles has been conducted primarily by Mikael Siverson and paleontologist Johan Lindgren (especially on mosasaurs) and paleontologists Elisabeth Einarsson, Benjamin P. Kear, Sven Sachs and Torsten M. Scheyer (especially on plesiosaurs and sea turtles).
2479:
environment there, similar to the environments preferred by their modern relatives. The sharks and rays at Ă…sen probably fed on fish and invertebrates, which occur with less diversity (though still large numbers) at Ă…sen than elsewhere. They might have primarily fed on soft-bodied invertebrates, less
2474:
mosasaurs have been found at Ă…sen, which suggests that the area offered some protection against predation by larger mosasaurs and other predators, possibly due to the murky waters produced by some nearby river system. This murky setting also seems to have been preferred by the many species of benthic
2454:
Fossil remains of large marine reptiles are also especially common at Ivö Klack. The high diversity of sharks and mosasaurs recovered at Ivö Klack shows that large predators frequented the rocky coastline. There must have been a productive ecosystem, with a diverse invertebrate fauna attracting small
2064:
Gastropod fossils are very rare, probably on account of their shells being easily broken and thus failing to be preserved as fossils. Though found throughout the basin, gastropods are mainly known from Ivö Klack, where the 17 species identified represent one of the most diverse gastropod faunas found
1887:). The rajiforms, adapted to colder temperatures, were the most common rays during the late Campanian, whereas sawfish and guitarfish had been common during the early Campanian. Local chimaerids, adapted to eat hard-shelled organisms with their flattened and crushing dental plates, include the genera
2402:
and deeper offshore environments. There were also shallow and protected coastal bays as well as coastal waters that were significantly deeper. Some structures within the rocks of the Basin, combined with the often fragmented and broken condition of the fossils recovered, indicate that the
Cretaceous
1481:
The recovered dinosaurs all represent animals with lengths less than three meters (9 ft), but this does not necessarily mean that larger dinosaurs were absent; it is equally likely that only small animals were transported out into the sea where they could be fossilized and preserved. When large
1259:
The various limestone and kaolin quarries throughout the basin were described in detail by geologist and paleontologist Karl A. Grönwall in 1915, followed by further descriptions by Alf
Lundegren in 1931, geologist Lars Bjerning in 1947, geologist Per H. Lundegårdh in 1971, geologists Jan Bergström
1602:
fauna of west-central
Alabama. The mosasaurs were large, carnivorous reptiles at the top of the food chain and since all the Campanian species are known from several sites, it is likely that all of them lived in the entire basin. The high diversity of mosasaurs in the basin can be explained by the
1284:
in 1916. Later studies on marine reptile fossils were also conducted by J. Christian Moberg in 1884 and 1885, Bernhard
Lundgren in 1888, Anders H. Hennig in 1910, Alf Lundegren in 1934, Gustaf Troedsson in 1946 and 1954 and prominently in a series of publications of paleontologist Per-Ove Persson,
1263:
Important historical studies conducted on the basin's many invertebrate taxa include those by
Bernhard Lundgren (in 1876, 1885 and 1895), paleontologist J. Christian Moberg in 1884 and 1885, geologist Anders H. Hennig in 1892, 1894 and 1904 (together with paleontologist Alfred Elis Törnebohm),
2397:
Most of the area preserved in the
Kristianstad Basin was a shallow marine inner shelf environment, as indicated by the present invertebrate fauna (which has been compared to modern faunas). Most of the water was probably less than 40 meters (131 ft) deep, but there were a wide range of
2365:
with several low islands and a number of small peninsulas. Remnants of these islands and peninsulas remain today in the form of rocky hills and mounts throughout northeastern Skåne, such as the Ivö Klack site, Fjälkinge backe, Kjugekull, Oppmannaberget, Vångaberget, Västanåberget and
1473:
The dinosaur fossils recovered represent the remains of specimens transported out into the sea during floods or storms. Despite being fragmentary and few in number, they are scientifically important as they represent some of the few remains of the poorly known dinosaur fauna of the
2439:, are significantly more common at Ivö Klack than they are in other sites, such as Ignaberga. The prominence of large sharks in the area probably derives from large sharks requiring large prey, and large marine reptiles being common at Ivö Klack as well. There are
2247:
1431:") was described in 2007 and further fragmentary fossils of other groups were recovered during excavations from 2010 to 2013. The local dinosaur fossils have mainly been researched by Johan Lindgren and paleontologist Jan Rees, alongside several colleagues.
1260:
and Naz Ahmed Shaikh in 1980 and geologists Mikael Erlström and Jan
Gabrielson in 1985 and 1992. Throughout the 20th century, quarrying was most intense at Åsen, Axeltorp and Ivö Klack, most of it conducted by the companies Höganäs keramik and Ifö-verken.
461:
80.5 million years ago). Fossils from these sites have been collected since the 18th century, but most of the excavations have taken place through commercial quarrying in the 20th century and paleontological expeditions in the 20th and 21st centuries.
2000:
The
Kristianstad Basin preserves a rich invertebrate fauna. Latest early Campanian-age deposits at Ivö Klack alone have in addition to about 40 vertebrate species yielded more than 200 distinct species of invertebrates. Groups represented include
1399:
Fossils of non-avian dinosaurs are exceptionally rare in
Scandinavia, and the Kristianstad Basin is one of only two locations in Sweden where dinosaur fossils have been recovered. The only other site of known dinosaur fossils in Sweden is the
1478:, which was an isolated landmass during the Late Cretaceous. It is also possible that the recovered dinosaurs were not from the landmass itself, but from the rocky islands of the adjacent archipelago in the Kristianstad Basin itself.
651:
beds in the upper parts of the strata, dating to the early Late Campanian through the earliest Maastrichtian. Dominant sediment types from the Early Campanian are oyster banks, calcarenites and calcareous, glauconitic quartz sands.
1404:, where they were first announced to be present by researchers in 2022. Though theropod teeth were reported from Ivö Klack by Per-Ove Persson in the 1950s, these have since been identified as fish teeth, probably from the genus
605:
that mark the basin's south-western boundary. To the south-east, the basin's margin is marked by the Baltic Sea while the basin's northern margin is more diffuse, with several small outliers of Cretaceous-age sediments.
1461:. The first fossil of a carnivorous dinosaur in Sweden was discovered by high school student Clarence Lagerström in 2015 at the Ugnsmunnarna site on Ivö island. After comparisons with the same bone in the Australian
2312:
In addition to the aforementioned dinosaurs and the freshwater turtle, further land-dwelling and amphibious animals have also been discovered in the basin. The basin preserves the fossils of the crocodylomorph
1709:, which measured around 4–5 metres in length (13.1–16.4 ft). The polycotylid fossils have not been identified down to the genus level, but they likely came from animals resembling the North American genus
1828:. All shark species found in the Kristianstad Basin were active predators, but many of them only fed on smaller food items such as bony fish and various invertebrates. Large nektonic lamniform sharks, such as
1116:, a partly overgrown cliff section located on the island Ivö, about three kilometers south of Ivö Klack. The rocks at Ugnsmunnarna were deposited in a deeper-water part surrounding the Cretaceous Ivö island.
1482:
dinosaurs are excluded, the dinosaur fauna preserved in the Kristianstad Basin resembles that of Campanian–Maastrichtian dinosaur-bearing formations in Canada, which also include small ornithopods (such as
2534:
1128:, a now disused limestone quarry adjacent to the northern end of Linderödsåsen. With the exception of abundant fragmentary invertebrate fossils, few other fossil remains have been found at Maltesholm.
655:
Accessible strata exposed in the basin range in age from Early or early Middle Santonian to earliest Maastrichtian. A majority of the fossil sites only expose strata of latest Early Campanian age (
2323:
are long and thin, suggesting a diet mostly composed of fish, but its teeth are unusually robust, meaning that it might also have fed on shelled invertebrates or larger animals. Fossils of small
3264:
2188:. Belemnites were a basic part of the food web and likely served as prey for many of the vertebrates in the basin, such as fish, plesiosaurs and the smaller mosasaurs. Crustaceans include both
1256:
studies were also conducted throughout the 19th and early 20th centuries, prominently by paleontologist and geologist Bernhard Lundgren in 1888 and geologist Alf Lundegren in 1931 and 1934.
1444:, it is likely that the Swedish leptoceratopsid represents a new taxon. Leptoceratopsid fossils include four maxillary teeth, a dentary tooth, two caudal vertebrae and one right manual
2793:"Palaeoenvironments, palaeoecology and palaeobiogeography of Late Cretaceous (Campanian) faunas from the Kristianstad Basin, southern Sweden, with applications for science education"
1603:
dissimilarity in dentition and body size between the species, meaning that they would not have competed with each other for food. The smallest and most common mosasaurs found are
1715:. Because plesiosaurs are found in all the different environments believed to have existed in the Kristianstad Basin, they are thought to have been present in the entire basin.
416:
in the south. The basin's northern boundary is more diffuse and there are several outlying portions of Cretaceous-age sediments. During the Cretaceous, the region was a shallow
3403:
2544:
1563:
present. In addition to these extinct groups, turtle remains are abundant in the Kristianstad Basin, though only two taxa have been identified so far; an indeterminate
1451:
Two left pedal phalanxes recovered at Ă…sen are probably attributable to two different small Ornithopod dinosaurs, with one of the phalanxes resembling the same bone in
1276:
fossils, found at Ivetofta, were first described from the basin by Sven Nilsson in 1835. Nilsson's work was followed by further investigations by physicist and chemist
1651:. After the latest early Campanian, the mosasaur fauna in the basin declined in diversity; going from the six genera present to just two by the middle late Campanian (
1770:(swimming freely in the body of water) or nectobenthic (active just above the sea floor). In addition to the lamniforms, other groups were present, including other
1970:, known for its long and thin teeth. With the exception of the pycnodontiforms, adapted to feed on shell-bearing organisms with their flat and crushing teeth, and
2529:
2539:
2475:
sharks and rays recovered at Ă…sen. Particularly, rays are significantly more common at Ă…sen than elsewhere. It is probable that they preferred the murky and
1232:
at Ivö Klack and Ignaberga in 1725. Most of the early fossil finds were belemnite fossils, descriptions of which were published by physician and naturalist
1410:. Fossil material conclusively identified as dinosaurian was recovered only relatively recently, with the earliest discovery being two teeth from a small
3739:
1794:, which went extinct in the early Campanian, before the time when a majority of the local fossil fauna lived. Squalomorphs were represented by smaller
3391:
1766:
sharks present are the most diverse Late Cretaceous lamniform fauna yet discovered. Most of the sharks in the basin were lamniforms and were either
1586:
The Swedish latest Early Campanian mosasaur fauna is one of the most taxon-rich assemblages of mosasaurs known, rivalled only by the Maastrichtian "
2037:. The most species-rich group by far is the bivalves, accounting for more than 70 of the 200 species at Ivö Klack. Common bivalves include various
1194:, an overgrown quarry which in addition to Campanian rocks also exposes approximately three meters (9 ft) of earliest Maastrichtian strata.
3744:
3729:
3013:"Food resources and habitat selection of a diverse vertebrate fauna from the upperlower Campanian of the Kristianstad Basin, southern Sweden"
2923:"Stratigraphical distribution of Campanian and Maastrichtian mosasaurs in Sweden – evidence of an intercontinental marine extinction event?"
250:
1503:
1428:
2279:. Fossil wood has been recovered from late Middle Santonian to earliest Campanian-age deposits at Ă…sen, representing the tree genera
1146:
shark teeth suggests that the localion was near an ancient river system. One of the most extensively excavated sites in the basin.
558:. Most of this weathering, and the uneven topography of the basement, is due to the warm and moist climate experienced during the
3769:
1122:, the Ignaberga site compromises two limestone quarries situated along the southern margin of the basin created by Nävlingeåsen.
3774:
1742:. Fragmentary pterosaur fossils (though currently unpublished) have been found in earliest Campanian-age deposits at Ullstorp.
3172:
Poropat, Stephen F.; Einarsson, Elizabeth; Lindgren, Johan; Bazzi, Mohamad; Lagerstam, Clarence; Kear, Benjamin P. (2015).
3105:
1470:, the bone, an incomplete right tibia, could confidently be identified as coming from a small non-avian theropod dinosaur.
3678:
1387:
dinosaur, whose fragmentary remains were identified as from a non-avian theropod through comparisons with the Australian
3764:
3173:
2403:
environment was a high-energy environment, where the water was fast-moving and agitated, created by waves and currents.
2216:. Decapod fossils are primarily just the claws, as they are more easily preserved than other portions of the crustacean
566:. The basement is overlaid by approximately 250 metres (820 feet) of mainly shallow water or marine sediments from the
2459:
preferred to live in coastal waters, where it could lay its egg on adjacent land, rest and heat up, similar to modern
3269:
Dollo, 1909 from the Upper Campanian of Ciply (Belgium) and a review of the Upper Cretaceous plesiosaurs from Europe"
2821:"Diversity and Ecology of a Middle Campanian (Late Cretaceous) Marine Reptile Assemblage from SkĂĄne, Southern Sweden"
3749:
3734:
3702:
3384:
465:
The Early Campanian deposits of the Kristianstad Basin preserve fossils of a diverse array of organisms, including
3238:
234:
2820:
2792:
2263:
Fossil plants that grew in the coastal areas surrounding the basin and on the islands dotted throughout include
3759:
3754:
3710:
3706:
3698:
3400:
3377:
2423:
have been found at Ă…sen, likely using the shallow and murky water there as protection against larger predators.
1918:
have also been identified based on numerous fossil vertebrae, teeth and scales. Local representatives included
1249:
2577:"Ett mystiskt ryggradsdjursfossil frĂĄn Ă…sen och dess koppling till den skĂĄnska, krittida ryggradsdjursfaunan"
2447:
are known to patrol around small islands inhabited by seals, possibly a behavior also present in the similar
2086:
Brachiopods, superficially similar to bivalves, are represented by numerous genera, the most prominent being
1555:
The warm shallow seas of the Kristianstad Basin were inhabited by a variety of marine reptiles, with several
3635:
3587:
3564:
2644:
1595:
1401:
614:
2643:
Lindgren, Johan; Currie, Philip J.; Siverson, Mikael; Rees, Jan; Cedeström, Peter; Lindgren, Filip (2007).
1762:
are relatively common fossil finds. In particular, the basin was home to a highly diverse shark fauna; the
1615:, both of which reached lengths of 2–4 metres (6.6–13.1 ft). Among the larger Campanian mosasaurs are
626:
176:
2398:
environments present. These environments included rocky and sandy beach areas, drowned river valleys and
3549:
2094:
2067:
922:
3645:
3602:
3559:
3369:
1228:. The earliest known fossil discoveries within the basin were made by the physician and paleontologist
1142:. The topography of the basement rock, the presence of floodplain sediments and the recovery of fossil
915:
3526:
3490:
3328:
2212:
1723:
497:
species). The Kristianstad Basin has also yielded fossils of several varieties of reptiles, including
3640:
3592:
3470:
2519:
2358:
1591:
641:
610:
424:
386:
136:
3650:
2875:"Late Cretaceous (Campanian) actinopterygian fishes from the Kristianstad Basin of southern Sweden"
2511:
2299:
2136:
2088:
2079:
1974:, able to feed on relatively large prey, the ray-finned fish found likely all fed on smaller fish.
1245:
3450:
2698:
2604:
2576:
2293:
2222:
1348:
598:
594:
413:
409:
3582:
3531:
3495:
3465:
3460:
3193:
3086:
2942:
2894:
2666:
2605:"Early Campanian mosasaurs (Reptilia; Mosasauridae) from the Kristianstad Basin, southern Sweden"
2329:
2305:
1825:
1784:
1747:
1440:, but due to the great geographical distance and the lack of some ridges present in the teeth of
617:. The sediments within the basin are dominated by fine- to coarse-grained sandy biocalcarenites (
547:
2234:
2228:
1790:
3329:"Taphonomy and palaeoecology of the gastropod fauna from a Late Cretaceous rocky shore, Sweden"
2922:
1901:
1883:
1319:
1075:
118:
3348:
2524:
2444:
2375:
2334:
2142:
1817:
1421:
1358:
1233:
633:
2699:"Palaeospinacid selachians from the Late Cretaceous of the Kristianstad Basin, SkĂĄne, Sweden"
2160:
3482:
3442:
3340:
3309:
3185:
3078:
3065:
Poropat, Stephen F.; Einarsson, Elisabeth; Lindgren, Johan; Bazzi, Mohamad (December 2015).
3024:
2934:
2886:
2656:
2497:
2051:
1951:
1927:
1711:
1599:
1277:
1253:
1229:
1213:
563:
382:
370:
132:
1527:
1518:
1328:
450:
to the earliest Maastrichtian. A majority of the fossil sites only expose strata of latest
3779:
3541:
2391:
2379:
2319:, which lived alongside the coastlines of the mainland and the small islands. The jaws of
2276:
2268:
2251:
1807:
1775:
1657:
1466:
1389:
1298:
1280:
in 1837, archaeologist and bibliographer Johan Henrik Schröder in 1885 and paleontologist
792:
602:
506:
451:
443:
406:
31:
1083:
Most of the sites within the Kristianstad Basin are the result of commercial kaolin clay/
238:
1849:
3627:
3574:
3518:
1956:
1915:
1799:
1690:
1490:
1453:
1406:
1373:
1273:
1267:
2361:
within the Kristianstad Basin remained very shallow, and its northern parts formed an
1982:
1434:
The morphology of the teeth recovered from the leptoceratopsid roughly corresponds to
1188:, a series of five small quarries (one of which remains active) close to Nävlingeåsen.
3723:
3597:
3554:
3197:
3090:
3066:
3043:
2946:
2898:
2874:
2848:
2670:
2661:
2315:
2255:
1942:
1779:
1694:
1679:
1648:
1475:
1436:
1416:
1379:
1363:
1302:
873:
637:
575:
543:
535:
439:
402:
394:
77:
39:
27:
3174:"Late Cretaceous dinosaurian remains from the Kristianstad Basin of southern Sweden"
3067:"Late Cretaceous dinosaurian remains from the Kristianstad Basin of southern Sweden"
1179:
2440:
2399:
2324:
2246:
2100:
2042:
1992:
1836:
1821:
1812:
1795:
1771:
1729:
1699:
1682:
1629:
1564:
1560:
1484:
1445:
1290:
1143:
799:
710:
644:
498:
230:
3344:
3313:
3028:
1689:
Plesiosaurs were represented by two groups in the Kristianstad Basin; long-necked
720:
3411:
3135:
2467:
2428:
2371:
2362:
2345:
2217:
2206:
2130:
2106:
2065:
in an ancient rocky shore environment. Genera identified include, among others,
1937:
1873:
1854:
1830:
1763:
1759:
1705:
1674:
1623:
1617:
1462:
1411:
1368:
1225:
1169:
785:
618:
579:
555:
486:
428:
417:
162:
52:
2238:. Polychate fossils are typically trace fossils; fossilized burrows and nests.
609:
During the Late Cretaceous, the basement of the basin was subjected to several
3433:
2938:
2487:
2460:
2367:
2165:
2119:
2034:
2026:
2018:
2014:
2010:
2002:
1932:
1867:
1862:
1842:
1803:
1735:
1663:
1635:
1573:
1568:
1541:
1424:
1420:, being found at Ivö Klack in 2001. More comprehensive material referred to a
1357:
Dinosaur fossils recovered in the Kristianstad Basin include the remains of a
1294:
1281:
1165:
891:
567:
539:
490:
470:
398:
378:
97:
62:
3352:
2470:
during the Campanian. Teeth and vertebrae of small-sized (probably juvenile)
2164:. Cephalopods are represented by the abundant belemnites and the less common
3668:
2476:
2339:
2264:
2154:
2148:
2057:
1987:
1961:
1907:
1895:
1739:
1611:
1605:
1579:
1384:
1208:
1103:
1099:
1084:
763:
725:
698:
630:
622:
587:
583:
571:
447:
435:
421:
166:
102:
46:
35:
2825:
Independent Project at the Department of Earth Sciences, Uppsala University
1339:
1220:
The Kristianstad Basin is one of the most prolific and historically famous
2411:
1158:, a natural cave located on the southwestern slope of the Balsberget hill.
3616:
3509:
3422:
2387:
2287:
2193:
2189:
2111:
2006:
1966:
1889:
1755:
1671:), possibly the result of an intercontinental mosasaur extinction event.
1556:
1537:
1286:
1221:
1139:
1135:
666:"-strata, is exceptionally rich in vertebrate fossils and diversity. The
559:
531:
514:
510:
482:
478:
390:
197:
123:
Kaolinbrottet, one of the quarries at Ivö Klack in the Kristianstad Basin
92:
87:
72:
67:
57:
2197:
2124:
2115:
2046:
2030:
1923:
1878:
729:
667:
551:
474:
107:
82:
3212:
3130:
2357:
During the transgressions experienced during the Late Cretaceous, the
670:
of the Kristianstad Basin, based on the local belemnite fossils, are:
2122:, which occur in many different genera, the most common of which are
2073:
2038:
1767:
1577:, but it is now considered more likely that they represent the genus
502:
215:
3189:
3082:
2890:
2427:
Ivö Klack was a small island during the Campanian. Teeth from large
1212:
Contemporary pencil drawing of Swedish physician and paleontologist
1107:
3297:
3012:
2410:
2281:
2245:
2022:
1981:
1848:
1751:
1722:
1673:
1207:
719:
648:
593:
The surface of the basement slopes to the south and is cut off by
494:
466:
180:
1216:, who first discovered fossils in the Kristianstad Basin in 1725.
3276:
Bulletin de l'Institut Royal des Sciences Naturelles de Belgique
2873:
Bazzi, Mohamad; Einarsson, Elisabeth; Kear, Benjamin P. (2015).
2383:
2272:
2201:
1947:
3373:
2849:"Late Cretaceous and Danian neoselachians from southern Sweden"
2535:
List of stratigraphic units with indeterminate dinosaur fossils
3298:"An early Campanian rocky shore at Ivö Klack, southern Sweden"
1919:
1571:. The sea turtle remains were initially referred to the genus
590:
age, overlaid by marine strata of latest Early Campanian age.
393:, the southernmost province of Sweden. The basin extends from
3011:
Sørensen, Anne Mehlin; Surlyk, Finn; Lindgren, Johan (2013).
1172:, the sediments penetrated are of latest Early Campanian age.
1590:
shales" fauna of southwestern Niger, the late Maastrichtian
1545:
from the Kristianstad Basin (left) and a reconstruction of
910:
Map of prominent fossil sites within the Kristianstad Basin
3106:"Unika fyndet: Tre nya dinosaurier upptäckta i Billesholm"
2333:, have also been discovered. Fossils of amphibious birds,
434:
Though the sediments in the basin range in age from the
2645:"The First Neoceratopsian Dinosaur Remains From Europe"
1639:, which surpassed 10 metres (32.8 ft) in length.
534:, the southernmost province of Sweden, extending from
3327:
Sørensen, Anne Mehlin; Surlyk, Finn (1 August 2011).
2443:
fossils from Ivö Klack with shark bite marks. Modern
1594:
of the southern Netherlands, the early Maastrichtian
3239:"Gymnasieelev hittade Sveriges första rovdinosaurie"
3044:"Maastrichtian squaloid sharks from southern Sweden"
2337:, have also been found, representing the two genera
621:
that contain fossils) and more or less consolidated
3667:
3626:
3615:
3573:
3540:
3517:
3508:
3481:
3441:
3432:
3421:
3410:
2545:
List of fossiliferous stratigraphic units in Sweden
226:
221:
203:
192:
187:
172:
158:
153:
142:
128:
21:
3178:Geological Society of London, Special Publications
2110:. The local echinoderm fauna, which also included
578:. In addition to the aquatic sediments, there are
530:The Kristianstad Basin is located in northeastern
513:faunas in the world and some of the few non-avian
2220:. Corals are primarily represented by the genera
1703:as well as one or two species of the local genus
550:of the basin is crystalline and deeply weathered
2879:Geological Society, London, Special Publications
1178:, a natural cliff section located near the lake
1087:exploitation. The most prominent sites include:
446:, ranging in age from the Early or early Middle
16:Cretaceous geological formation in SkĂĄne, Sweden
2466:The Ă…sen site is believed to have been a murky
1598:fauna of southern Belgium and the contemporary
1240:included the first illustrations of the common
3071:Geological Society London Special Publications
2581:Examensarbeten I Geologi Vid Lunds Universitet
1152:, an abandoned and overgrown limestone quarry.
629:beds are also common, typically consisting of
3385:
3131:"Dinosauriefynd i SkĂĄne skriver om historien"
2530:Lists of dinosaur-bearing stratigraphic units
2480:likely to be preserved in the fossil record.
1204:Local paleontological and geological research
8:
3296:Surlyk, Finn; Sørensen, Anne Mehlin (2010).
3263:Bardet, Nathalie; Godefroit, Pascal (1995).
2540:List of mosasaur-bearing stratigraphic units
1857:dental parts found in the Kristianstad Basin
3404:fossiliferous stratigraphic units in Sweden
2703:Dissertations in Geology at Lund University
2609:Dissertations in Geology at Lund University
1806:. Among the non-lamniform galeomorphs were
1106:quarry on the northern slope of the island
662:80.5 million years ago). This strata, the "
3623:
3514:
3438:
3429:
3418:
3392:
3378:
3370:
2786:
2784:
2782:
2780:
2778:
2776:
2774:
2772:
2770:
2768:
2766:
2764:
2762:
2760:
2758:
2756:
2754:
2752:
2750:
2748:
2746:
2744:
2742:
2740:
2738:
2736:
2734:
2732:
2378:and local plant life included low-growing
2168:. Five genera of belemnites are recorded;
1457:and the other resembling the same bone in
442:, the only accessible strata are from the
347:
338:
329:
320:
311:
302:
293:
284:
275:
266:
257:
117:
18:
2868:
2866:
2730:
2728:
2726:
2724:
2722:
2720:
2718:
2716:
2714:
2712:
2660:
3291:
3289:
3167:
3165:
3163:
3161:
3159:
3157:
3155:
3153:
3006:
3004:
3002:
3000:
2998:
2996:
2994:
2992:
2990:
2988:
2986:
2984:
2982:
2980:
2978:
2976:
1248:in 1821 and zoologist and archaeologist
672:
2974:
2972:
2970:
2968:
2966:
2964:
2962:
2960:
2958:
2956:
2842:
2840:
2838:
2836:
2834:
2638:
2570:
2568:
2566:
2564:
2562:
2560:
2556:
2916:
2914:
2912:
2910:
2908:
2814:
2812:
2810:
2808:
2806:
2692:
2690:
2688:
2686:
2684:
2682:
2680:
2636:
2634:
2632:
2630:
2628:
2626:
2624:
2622:
2620:
2618:
2598:
2596:
2594:
2592:
2590:
1782:. Among the hybodonts were the genera
1567:(a freshwater softshell turtle) and a
493:and fish (including a large amount of
509:, as well as one of the most diverse
7:
1504:Paleobiota of the Kristianstad Basin
1252:in 1826, 1827, 1835 and 1857. Early
1098:, an abandoned and partly overgrown
2419:mosasaurs in shallow water. Young
2407:Paleoecology of Ivö Klack and Åsen
1727:Reconstruction of the large shark
405:in the west and ends with the two
14:
3740:Upper Cretaceous Series of Europe
1964:). The most common fish found is
1383:(bottom-left) as well as a small
1285:who described several species of
3694:
3693:
2662:10.1111/j.1475-4983.2007.00690.x
2504:
2490:
2196:(the group that contains modern
1661:) and two by the Maastrichtian (
1526:
1517:
1488:) and leptoceratopsids (such as
1347:
1338:
1327:
1318:
1062:
1050:
1038:
1026:
1014:
1002:
990:
978:
966:
954:
942:
928:
921:
914:
831:Gonioteuthis quadrata scaniensis
743:Gonioteuthis westfalicagranulata
346:
337:
328:
319:
310:
301:
292:
283:
274:
265:
256:
249:
208:
3211:Olsson, Caroline (2001-09-02).
2259:found in the Kristianstad Basin
2242:Terrestrial and amphibious life
1996:found in the Kristianstad Basin
1:
3745:Geologic formations of Sweden
3679:Sub-Mesozoic hilly peneplains
3345:10.1016/j.cretres.2011.03.003
3314:10.1016/j.cretres.2010.07.006
3029:10.1016/j.cretres.2013.02.002
2791:Einarsson, Elisabeth (2018).
2204:), represented by the genera
1627:, the 8-metre (26.2 ft)
881:
813:
771:
656:
542:, in the east to the town of
455:
401:, in the east to the town of
146:
3730:Sedimentary basins of Europe
3237:Andersson, Ki (2015-01-01).
2853:Lund Publications in Geology
846:Belemnellocamax balsvikensis
781:Gonioteuthis granulaquadrata
3104:Melin, Simon (2022-10-13).
1685:from the Kristianstad Basin
1242:Belemnellocamax mammillatus
823:Belemnellocamax mammillatus
664:Belemnellocamax mammillatus
355:Kristianstad Basin (Sweden)
3796:
1536:Fossil tooth of the giant
1501:
570:, ranging in age from the
481:(notably large numbers of
3691:
3093:– via ResearchGate.
3042:Siverson, Mikael (1993).
2939:10.1080/11035890401262221
2847:Siverson, Mikael (1993).
2697:Siverson, Mikael (1989).
2575:Hajny, Cassandra (2014).
856:
841:
762:
732:in the Kristianstad Basin
718:
705:
697:
647:. There are also several
244:
235:Kristianstad Municipality
116:
26:
3213:"Dinosauriefynd i SkĂĄne"
2921:Lindgren, Johan (2004).
2603:Lindgren, Johan (1998).
2415:Reconstruction of three
2353:Depositional environment
1244:belemnites), naturalist
1094:, historically known as
3770:Shallow marine deposits
2819:Källsten, Lena (2015).
3775:Paleontology in Sweden
3565:Vilhelmsfält Formation
2424:
2260:
1997:
1858:
1743:
1686:
1647:were likely the local
1596:Ciply Phosphatic Chalk
1414:, similar to those of
1217:
1138:, presently used as a
904:Prominent fossil sites
858:Belemnitella mucronata
827:Belemnitella mucronata
755:Gonioteuthis granulata
733:
728:species and belemnite
375:Kristianstadsbassängen
3267:Plesiosaurus houzeaui
2414:
2249:
1985:
1852:
1726:
1677:
1211:
723:
713:westfalica westfalica
526:Geological background
2520:List of fossil sites
2327:lizards, similar to
1960:, similar to modern
1946:, similar to modern
1861:Fossil rays include
1592:Maastricht Formation
1236:in 1752 (whose work
842:No precise estimate
706:No precise estimate
387:geological formation
183:, oyster banks, sand
137:geological formation
3765:Maastrichtian Stage
3333:Cretaceous Research
3302:Cretaceous Research
3017:Cretaceous Research
2512:Paleontology portal
2118:, was dominated by
1199:History of research
674:
517:known from Sweden.
28:Stratigraphic range
3603:Röddinge Formation
3583:Djupadal Formation
3560:Röddinge Formation
3550:Mariedal Formation
3456:Kristianstad Basin
2445:great white sharks
2425:
2370:. The climate was
2335:hesperornithiforms
2261:
1998:
1859:
1748:cartilaginous fish
1744:
1687:
1367:(top-left), small
1218:
734:
673:
367:Kristianstad Basin
22:Kristianstad Basin
3750:Cretaceous Sweden
3735:Geology of Sweden
3717:
3716:
3687:
3686:
3663:
3662:
3659:
3658:
3646:Kågeröd Formation
3636:Höganäs Formation
3611:
3610:
3588:Höganäs Formation
3504:
3503:
3471:Köpinge Sandstone
2525:Geology of Sweden
1916:ray-finned fishes
1826:synechodontiforms
1693:and short-necked
1402:Höganäs Formation
901:
900:
582:deposits of late
546:in the west. The
363:
362:
3787:
3697:
3696:
3671:
3624:
3619:
3527:Annero Formation
3515:
3491:Annero Formation
3439:
3430:
3425:
3419:
3414:
3394:
3387:
3380:
3371:
3364:
3363:
3361:
3359:
3324:
3318:
3317:
3293:
3284:
3283:
3273:
3260:
3254:
3253:
3251:
3250:
3234:
3228:
3227:
3225:
3224:
3208:
3202:
3201:
3169:
3148:
3147:
3145:
3144:
3127:
3121:
3120:
3118:
3117:
3101:
3095:
3094:
3062:
3056:
3055:
3039:
3033:
3032:
3008:
2951:
2950:
2918:
2903:
2902:
2870:
2861:
2860:
2844:
2829:
2828:
2816:
2801:
2800:
2797:Litholund Theses
2788:
2707:
2706:
2694:
2675:
2674:
2664:
2640:
2613:
2612:
2600:
2585:
2584:
2572:
2514:
2509:
2508:
2507:
2500:
2498:Dinosaurs portal
2495:
2494:
2493:
2431:sharks, such as
2380:flowering plants
2277:flowering plants
2275:and low-growing
2213:Protocallianassa
2055:) and the genus
1712:Dolichorhynchops
1678:Fossil teeth of
1600:Mooreville Chalk
1530:
1521:
1377:(top-right) and
1351:
1342:
1331:
1322:
1310:Dinosaur fossils
1278:Wilhelm Hisinger
1254:sedimentological
1246:Göran Wahlenberg
1230:Magnus Bromelius
1224:fossil sites in
1214:Magnus Bromelius
1077:
1071:
1066:
1065:
1059:
1054:
1053:
1047:
1042:
1041:
1035:
1030:
1029:
1023:
1018:
1017:
1011:
1006:
1005:
999:
994:
993:
987:
982:
981:
975:
970:
969:
963:
958:
957:
951:
946:
945:
939:
932:
931:
925:
918:
886:
883:
818:
815:
776:
773:
675:
661:
658:
574:to the earliest
564:Early Cretaceous
460:
457:
438:to the earliest
389:in northeastern
383:structural basin
350:
349:
341:
340:
332:
331:
323:
322:
314:
313:
305:
304:
296:
295:
287:
286:
278:
277:
269:
268:
260:
259:
253:
214:
212:
211:
148:
133:Structural basin
121:
112:
49:
19:
3795:
3794:
3790:
3789:
3788:
3786:
3785:
3784:
3760:Campanian Stage
3755:Santonian Stage
3720:
3719:
3718:
3713:
3683:
3669:
3655:
3617:
3607:
3569:
3536:
3500:
3477:
3423:
3412:
3406:
3398:
3368:
3367:
3357:
3355:
3326:
3325:
3321:
3295:
3294:
3287:
3271:
3262:
3261:
3257:
3248:
3246:
3236:
3235:
3231:
3222:
3220:
3210:
3209:
3205:
3190:10.1144/SP434.8
3171:
3170:
3151:
3142:
3140:
3129:
3128:
3124:
3115:
3113:
3103:
3102:
3098:
3083:10.1144/SP434.8
3064:
3063:
3059:
3041:
3040:
3036:
3010:
3009:
2954:
2920:
2919:
2906:
2891:10.1144/SP434.5
2872:
2871:
2864:
2846:
2845:
2832:
2818:
2817:
2804:
2790:
2789:
2710:
2696:
2695:
2678:
2642:
2641:
2616:
2602:
2601:
2588:
2574:
2573:
2558:
2553:
2510:
2505:
2503:
2496:
2491:
2489:
2486:
2409:
2392:deciduous trees
2355:
2269:deciduous trees
2244:
2178:Belemnellocamax
1980:
1952:pachycormiforms
1928:pycnodontiforms
1881:and rajiforms (
1808:bullhead sharks
1721:
1669:Plioplatecarpus
1658:Plioplatecarpus
1553:
1552:
1551:
1550:
1533:
1532:
1531:
1523:
1522:
1511:
1509:Marine reptiles
1506:
1500:
1467:Australovenator
1422:leptoceratopsid
1397:
1396:
1395:
1394:
1393:(bottom-right).
1390:Australovenator
1359:leptoceratopsid
1354:
1353:
1352:
1344:
1343:
1334:
1333:
1332:
1324:
1323:
1312:
1268:Actinopterygian
1206:
1201:
1168:in the town of
1134:, an abandoned
1081:
1080:
1079:
1078:
1074:
1072:
1069:
1067:
1063:
1060:
1057:
1055:
1051:
1048:
1045:
1043:
1039:
1036:
1033:
1031:
1027:
1024:
1021:
1019:
1015:
1012:
1009:
1007:
1003:
1000:
997:
995:
991:
988:
985:
983:
979:
976:
973:
971:
967:
964:
961:
959:
955:
952:
949:
947:
943:
940:
937:
935:
933:
929:
926:
919:
911:
906:
884:
816:
793:Belemnellocamax
774:
659:
538:, a bay in the
528:
523:
507:crocodylomorphs
458:
452:Early Campanian
444:Late Cretaceous
397:, a bay in the
359:
358:
357:
356:
353:
352:
351:
343:
342:
334:
333:
325:
324:
316:
315:
307:
306:
298:
297:
289:
288:
280:
279:
271:
270:
262:
261:
209:
207:
124:
111:
110:
105:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
45:86.3–71.3
44:
43:
32:Late Cretaceous
17:
12:
11:
5:
3793:
3791:
3783:
3782:
3777:
3772:
3767:
3762:
3757:
3752:
3747:
3742:
3737:
3732:
3722:
3721:
3715:
3714:
3692:
3689:
3688:
3685:
3684:
3682:
3681:
3675:
3673:
3665:
3664:
3661:
3660:
3657:
3656:
3654:
3653:
3651:Mine Formation
3648:
3643:
3641:Höör Sandstone
3638:
3632:
3630:
3621:
3613:
3612:
3609:
3608:
3606:
3605:
3600:
3595:
3593:Höör Sandstone
3590:
3585:
3579:
3577:
3571:
3570:
3568:
3567:
3562:
3557:
3552:
3546:
3544:
3538:
3537:
3535:
3534:
3529:
3523:
3521:
3512:
3506:
3505:
3502:
3501:
3499:
3498:
3493:
3487:
3485:
3479:
3478:
3476:
3475:
3474:
3473:
3463:
3458:
3453:
3447:
3445:
3436:
3427:
3416:
3408:
3407:
3399:
3397:
3396:
3389:
3382:
3374:
3366:
3365:
3339:(4): 472–479.
3319:
3308:(6): 567–576.
3285:
3255:
3229:
3203:
3149:
3122:
3096:
3077:(1): 231–239.
3057:
3034:
2952:
2933:(2): 221–229.
2904:
2862:
2830:
2802:
2708:
2676:
2655:(4): 929–937.
2614:
2586:
2555:
2554:
2552:
2549:
2548:
2547:
2542:
2537:
2532:
2527:
2522:
2516:
2515:
2501:
2485:
2482:
2408:
2405:
2354:
2351:
2252:crocodylomorph
2243:
2240:
1979:
1976:
1972:Protosphyraena
1957:Protosphyraena
1720:
1717:
1649:apex predators
1633:and the giant
1535:
1534:
1525:
1524:
1516:
1515:
1514:
1513:
1512:
1510:
1507:
1502:Main article:
1499:
1498:Fossil content
1496:
1491:Unescoceratops
1454:Thescelosaurus
1429:Kristianasaura
1407:Protosphyraena
1374:Thescelosaurus
1356:
1355:
1346:
1345:
1337:
1336:
1335:
1326:
1325:
1317:
1316:
1315:
1314:
1313:
1311:
1308:
1299:crocodylomorph
1274:Marine reptile
1234:Kilian Stobæus
1205:
1202:
1200:
1197:
1196:
1195:
1189:
1183:
1173:
1159:
1153:
1147:
1129:
1123:
1117:
1111:
1073:
1068:
1061:
1056:
1049:
1044:
1037:
1032:
1025:
1020:
1013:
1008:
1001:
996:
989:
984:
977:
972:
965:
960:
953:
948:
941:
934:
927:
920:
913:
912:
909:
908:
907:
905:
902:
899:
898:
896:
888:
879:
876:
870:
869:
867:
863:
862:
860:
855:
851:
850:
848:
843:
840:
839:Earliest late
836:
835:
833:
820:
811:
807:
806:
804:
778:
769:
766:
760:
759:
757:
752:
748:
747:
745:
740:
736:
735:
717:
715:
707:
704:
701:
695:
694:
691:
688:
685:
682:
679:
638:bivalve shells
615:transgressions
527:
524:
522:
519:
361:
360:
354:
345:
344:
336:
335:
327:
326:
318:
317:
309:
308:
300:
299:
291:
290:
282:
281:
273:
272:
264:
263:
255:
254:
248:
247:
246:
245:
242:
241:
228:
224:
223:
219:
218:
205:
201:
200:
196:North-eastern
194:
190:
189:
185:
184:
174:
170:
169:
160:
156:
155:
151:
150:
149:250 m (820 ft)
144:
140:
139:
130:
126:
125:
122:
114:
113:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
50:
24:
23:
15:
13:
10:
9:
6:
4:
3:
2:
3792:
3781:
3778:
3776:
3773:
3771:
3768:
3766:
3763:
3761:
3758:
3756:
3753:
3751:
3748:
3746:
3743:
3741:
3738:
3736:
3733:
3731:
3728:
3727:
3725:
3712:
3708:
3704:
3700:
3690:
3680:
3677:
3676:
3674:
3672:
3666:
3652:
3649:
3647:
3644:
3642:
3639:
3637:
3634:
3633:
3631:
3629:
3625:
3622:
3620:
3614:
3604:
3601:
3599:
3598:Rya Formation
3596:
3594:
3591:
3589:
3586:
3584:
3581:
3580:
3578:
3576:
3572:
3566:
3563:
3561:
3558:
3556:
3555:Rya Formation
3553:
3551:
3548:
3547:
3545:
3543:
3539:
3533:
3530:
3528:
3525:
3524:
3522:
3520:
3516:
3513:
3511:
3507:
3497:
3494:
3492:
3489:
3488:
3486:
3484:
3480:
3472:
3469:
3468:
3467:
3464:
3462:
3459:
3457:
3454:
3452:
3449:
3448:
3446:
3444:
3440:
3437:
3435:
3431:
3428:
3426:
3420:
3417:
3415:
3409:
3405:
3402:
3395:
3390:
3388:
3383:
3381:
3376:
3375:
3372:
3354:
3350:
3346:
3342:
3338:
3334:
3330:
3323:
3320:
3315:
3311:
3307:
3303:
3299:
3292:
3290:
3286:
3281:
3277:
3270:
3268:
3259:
3256:
3244:
3240:
3233:
3230:
3218:
3214:
3207:
3204:
3199:
3195:
3191:
3187:
3183:
3179:
3175:
3168:
3166:
3164:
3162:
3160:
3158:
3156:
3154:
3150:
3138:
3137:
3132:
3126:
3123:
3111:
3107:
3100:
3097:
3092:
3088:
3084:
3080:
3076:
3072:
3068:
3061:
3058:
3053:
3049:
3048:Palaeontology
3045:
3038:
3035:
3030:
3026:
3022:
3018:
3014:
3007:
3005:
3003:
3001:
2999:
2997:
2995:
2993:
2991:
2989:
2987:
2985:
2983:
2981:
2979:
2977:
2975:
2973:
2971:
2969:
2967:
2965:
2963:
2961:
2959:
2957:
2953:
2948:
2944:
2940:
2936:
2932:
2928:
2924:
2917:
2915:
2913:
2911:
2909:
2905:
2900:
2896:
2892:
2888:
2884:
2880:
2876:
2869:
2867:
2863:
2858:
2854:
2850:
2843:
2841:
2839:
2837:
2835:
2831:
2826:
2822:
2815:
2813:
2811:
2809:
2807:
2803:
2798:
2794:
2787:
2785:
2783:
2781:
2779:
2777:
2775:
2773:
2771:
2769:
2767:
2765:
2763:
2761:
2759:
2757:
2755:
2753:
2751:
2749:
2747:
2745:
2743:
2741:
2739:
2737:
2735:
2733:
2731:
2729:
2727:
2725:
2723:
2721:
2719:
2717:
2715:
2713:
2709:
2704:
2700:
2693:
2691:
2689:
2687:
2685:
2683:
2681:
2677:
2672:
2668:
2663:
2658:
2654:
2650:
2649:Palaeontology
2646:
2639:
2637:
2635:
2633:
2631:
2629:
2627:
2625:
2623:
2621:
2619:
2615:
2610:
2606:
2599:
2597:
2595:
2593:
2591:
2587:
2583:(in Swedish).
2582:
2578:
2571:
2569:
2567:
2565:
2563:
2561:
2557:
2550:
2546:
2543:
2541:
2538:
2536:
2533:
2531:
2528:
2526:
2523:
2521:
2518:
2517:
2513:
2502:
2499:
2488:
2483:
2481:
2478:
2473:
2469:
2464:
2462:
2458:
2457:Aigialosuchus
2452:
2450:
2446:
2442:
2438:
2434:
2430:
2422:
2418:
2413:
2406:
2404:
2401:
2395:
2393:
2389:
2385:
2381:
2377:
2373:
2369:
2364:
2360:
2352:
2350:
2348:
2347:
2342:
2341:
2336:
2332:
2331:
2326:
2322:
2321:Aigialosuchus
2318:
2317:
2316:Aigialosuchus
2310:
2308:
2307:
2302:
2301:
2300:Silvianthemum
2296:
2295:
2290:
2289:
2284:
2283:
2278:
2274:
2270:
2266:
2258:
2257:
2256:Aigialosuchus
2253:
2250:Teeth of the
2248:
2241:
2239:
2237:
2236:
2231:
2230:
2225:
2224:
2219:
2215:
2214:
2209:
2208:
2203:
2199:
2195:
2191:
2187:
2183:
2179:
2175:
2171:
2167:
2163:
2162:
2157:
2156:
2151:
2150:
2145:
2144:
2139:
2138:
2137:Echinogalerus
2133:
2132:
2127:
2126:
2121:
2117:
2113:
2109:
2108:
2103:
2102:
2097:
2096:
2091:
2090:
2084:
2082:
2081:
2076:
2075:
2070:
2069:
2062:
2060:
2059:
2054:
2053:
2048:
2044:
2040:
2036:
2032:
2028:
2024:
2020:
2016:
2012:
2008:
2004:
1995:
1994:
1990:of the genus
1989:
1984:
1978:Invertebrates
1977:
1975:
1973:
1969:
1968:
1963:
1959:
1958:
1953:
1949:
1945:
1944:
1943:Pachyrhizodus
1939:
1935:
1934:
1929:
1925:
1921:
1917:
1912:
1910:
1909:
1904:
1903:
1898:
1897:
1892:
1891:
1886:
1885:
1880:
1876:
1875:
1870:
1869:
1864:
1856:
1851:
1847:
1845:
1844:
1839:
1838:
1833:
1832:
1827:
1823:
1822:carpet sharks
1819:
1818:ground sharks
1815:
1814:
1809:
1805:
1801:
1800:hexanchiforms
1797:
1793:
1792:
1787:
1786:
1781:
1777:
1773:
1769:
1765:
1761:
1757:
1753:
1749:
1741:
1738:
1737:
1732:
1731:
1725:
1718:
1716:
1714:
1713:
1708:
1707:
1702:
1701:
1696:
1692:
1691:elasmosaurids
1684:
1681:
1676:
1672:
1670:
1666:
1665:
1660:
1659:
1654:
1650:
1646:
1642:
1638:
1637:
1632:
1631:
1626:
1625:
1620:
1619:
1614:
1613:
1608:
1607:
1601:
1597:
1593:
1589:
1584:
1582:
1581:
1576:
1575:
1570:
1566:
1562:
1558:
1548:
1544:
1543:
1539:
1529:
1520:
1508:
1505:
1497:
1495:
1493:
1492:
1487:
1486:
1479:
1477:
1476:Baltic Shield
1471:
1469:
1468:
1464:
1460:
1459:Hypsilophodon
1456:
1455:
1449:
1447:
1443:
1442:Leptoceratops
1439:
1438:
1437:Leptoceratops
1432:
1430:
1426:
1423:
1419:
1418:
1417:Hypsilophodon
1413:
1409:
1408:
1403:
1392:
1391:
1386:
1382:
1381:
1380:Hypsilophodon
1376:
1375:
1370:
1366:
1365:
1364:Leptoceratops
1360:
1350:
1341:
1330:
1321:
1309:
1307:
1305:
1304:
1303:Aigialosuchus
1300:
1296:
1292:
1288:
1283:
1279:
1275:
1271:
1269:
1265:
1261:
1257:
1255:
1251:
1247:
1243:
1239:
1235:
1231:
1227:
1223:
1215:
1210:
1203:
1198:
1193:
1190:
1187:
1184:
1181:
1177:
1174:
1171:
1167:
1163:
1160:
1157:
1154:
1151:
1148:
1145:
1141:
1137:
1133:
1130:
1127:
1124:
1121:
1118:
1115:
1112:
1109:
1105:
1101:
1097:
1093:
1090:
1089:
1088:
1086:
1076:
924:
917:
903:
897:
895:
893:
889:
880:
877:
875:
874:Maastrichtian
872:
871:
868:
865:
864:
861:
859:
853:
852:
849:
847:
844:
838:
837:
834:
832:
828:
824:
821:
812:
810:Latest early
809:
808:
805:
803:
801:
796:
794:
789:
787:
782:
779:
770:
767:
765:
761:
758:
756:
753:
750:
749:
746:
744:
741:
738:
737:
731:
727:
722:
716:
714:
712:
708:
703:Early–middle
702:
700:
696:
692:
689:
687:Belemnite(s)
686:
683:
680:
677:
676:
671:
669:
665:
653:
650:
646:
645:clastic rocks
643:
639:
635:
632:
628:
627:Conglomeratic
624:
620:
616:
612:
607:
604:
600:
596:
595:Linderödsåsen
591:
589:
585:
581:
577:
576:Maastrichtian
573:
569:
565:
561:
557:
553:
549:
545:
541:
537:
533:
525:
520:
518:
516:
512:
508:
504:
500:
496:
492:
488:
484:
480:
476:
472:
468:
463:
453:
449:
445:
441:
440:Maastrichtian
437:
432:
430:
426:
423:
419:
415:
411:
410:Linderödsåsen
408:
404:
400:
396:
392:
388:
384:
380:
376:
372:
368:
252:
243:
240:
236:
232:
229:
225:
220:
217:
206:
202:
199:
195:
191:
186:
182:
178:
177:Conglomerates
175:
171:
168:
164:
161:
157:
152:
145:
141:
138:
134:
131:
127:
120:
115:
109:
104:
99:
94:
89:
84:
79:
74:
69:
64:
59:
54:
48:
41:
40:Maastrichtian
37:
33:
29:
25:
20:
3455:
3451:BĂĄstad Basin
3356:. Retrieved
3336:
3332:
3322:
3305:
3301:
3279:
3275:
3266:
3258:
3247:. Retrieved
3245:(in Swedish)
3242:
3232:
3221:. Retrieved
3219:(in Swedish)
3216:
3206:
3181:
3177:
3141:. Retrieved
3139:(in Swedish)
3134:
3125:
3114:. Retrieved
3112:(in Swedish)
3109:
3099:
3074:
3070:
3060:
3051:
3047:
3037:
3020:
3016:
2930:
2926:
2882:
2878:
2856:
2852:
2824:
2796:
2702:
2652:
2648:
2608:
2580:
2471:
2465:
2461:crocodilians
2456:
2453:
2449:Cretoxyrhina
2448:
2436:
2433:Cretoxyrhina
2432:
2426:
2420:
2416:
2396:
2356:
2344:
2338:
2328:
2320:
2314:
2311:
2304:
2298:
2294:Scandianthus
2292:
2286:
2280:
2262:
2254:
2233:
2227:
2223:Leptophyllia
2221:
2211:
2205:
2186:Gonioteuthis
2185:
2182:Belemnitella
2181:
2177:
2173:
2169:
2159:
2153:
2147:
2141:
2135:
2129:
2123:
2105:
2101:Rhynchonella
2099:
2093:
2087:
2085:
2078:
2072:
2066:
2063:
2056:
2050:
1999:
1993:Gonioteuthis
1991:
1971:
1965:
1955:
1941:
1931:
1913:
1906:
1900:
1894:
1888:
1882:
1872:
1866:
1860:
1841:
1837:Cretoxyrhina
1835:
1829:
1813:Heterodontus
1811:
1789:
1783:
1780:squalomorphs
1745:
1734:
1733:attacking a
1730:Cretoxyrhina
1728:
1710:
1704:
1700:Elasmosaurus
1698:
1695:polycotylids
1688:
1668:
1662:
1656:
1653:Prognathodon
1652:
1644:
1641:Prognathodon
1640:
1634:
1630:Prognathodon
1628:
1622:
1616:
1610:
1604:
1587:
1585:
1578:
1572:
1554:
1546:
1540:
1489:
1485:Parksosaurus
1483:
1480:
1472:
1465:
1463:megaraptoran
1458:
1452:
1450:
1441:
1435:
1433:
1415:
1405:
1398:
1388:
1378:
1372:
1362:
1301:
1272:
1266:
1262:
1258:
1250:Sven Nilsson
1241:
1237:
1219:
1191:
1185:
1175:
1161:
1155:
1149:
1131:
1125:
1119:
1114:Ugnsmunnarna
1113:
1095:
1091:
1082:
1034:Ugnsmunnarna
890:
866:Middle late
857:
845:
830:
826:
822:
800:Belemnitella
798:
791:
784:
780:
754:
742:
739:Late middle
724:Some of the
711:Gonioteuthis
709:
681:Subdivision
663:
654:
619:calcarenites
608:
599:Nävlingeåsen
592:
586:to earliest
529:
464:
433:
414:Nävlingeåsen
374:
366:
364:
239:SkĂĄne County
231:Kristianstad
222:Type section
3532:Vomb Trough
3496:Vomb Trough
3466:Vomb Trough
3461:Särdal Beds
3243:SVT Nyheter
3217:Aftonbladet
3184:: 231–239.
3136:Aftonbladet
3110:SVT Nyheter
2885:: 277–292.
2468:river mouth
2372:subtropical
2363:archipelago
2346:Hesperornis
2330:Araeosaurus
2325:scincomorph
2306:Actinocalyx
2218:exoskeleton
2207:Callianassa
2170:Actinocamax
2131:Echinocorys
2120:sea urchins
2107:Terebratula
2043:inoceramids
2035:polychaetes
2027:crustaceans
2019:echinoderms
2015:brachiopods
2003:cephalopods
1874:Squatirhina
1831:Squalicorax
1804:angelsharks
1796:squaliforms
1785:Polyacrodus
1772:galeomorphs
1706:Scanisaurus
1683:plesiosaurs
1680:polycotylid
1624:Hainosaurus
1618:Platecarpus
1561:plesiosaurs
1425:ceratopsian
1371:similar to
1369:ornithopods
1361:similar to
1295:sea turtles
1291:plesiosaurs
1226:Scandinavia
854:Early late
795:grossouvrei
786:Actinocamax
684:Time (mya)
642:terrigenous
640:and coarse
611:regressions
580:delta plain
556:Precambrian
499:plesiosaurs
491:sea urchins
471:brachiopods
429:archipelago
418:subtropical
163:Calcarenite
3724:Categories
3703:Cretaceous
3282:: 179–186.
3249:2020-12-30
3223:2020-12-30
3143:2022-10-13
3116:2022-10-13
3054:(1): 1–19.
2551:References
2441:plesiosaur
2437:Cretalamna
2368:Ryssberget
2359:inland sea
2235:Parasmilia
2229:Micrabacia
2174:Belemnella
2011:gastropods
1938:elopiforms
1933:Anomoeodus
1868:Rhinobatos
1863:guitarfish
1843:Carcharias
1791:Meristodon
1760:chimaerids
1750:, such as
1736:Pteranodon
1664:Mosasaurus
1645:Tylosaurus
1636:Tylosaurus
1588:Mosasaurus
1574:Osteopygis
1569:sea turtle
1565:trionychid
1547:Tylosaurus
1542:Tylosaurus
1412:Ornithopod
1293:alongside
1282:Carl Wiman
1166:drill core
1126:Maltesholm
1096:Blaksudden
1070:Maltesholm
894:lanceolata
892:Belemnella
623:sandstones
568:Cretaceous
544:Hässleholm
540:Baltic Sea
536:Hanöbukten
487:belemnites
425:inland sea
403:Hässleholm
399:Baltic Sea
395:Hanöbukten
379:Cretaceous
3358:1 January
3353:0195-6671
3198:130034135
3091:130034135
3023:: 85–92.
2947:128401131
2899:131738899
2671:129733977
2477:estuarine
2472:Clidastes
2429:lamniform
2421:Clidastes
2417:Clidastes
2376:temperate
2340:Baptornis
2190:barnacles
2166:ammonites
2155:Phymosoma
2149:Micraster
2068:Campanile
2058:Spondylus
2049:(such as
2031:bryozoans
1988:belemnite
1962:swordfish
1954:(such as
1940:(such as
1930:(such as
1908:Ischyodus
1902:Elasmodus
1896:Edaphodon
1884:Walteraja
1855:chimaerid
1810:(such as
1776:hybodonts
1764:lamniform
1746:Teeth of
1740:pterosaur
1612:Eonatator
1606:Clidastes
1580:Euclastes
1557:mosasaurs
1287:mosasaurs
1120:Ignaberga
1100:limestone
1092:Ivö Klack
1085:limestone
1058:Ignaberga
1046:Ivö Klack
878:Earliest
768:Earliest
764:Campanian
726:belemnite
699:Santonian
631:belemnite
588:Campanian
584:Santonian
572:Barremian
554:from the
515:dinosaurs
475:bryozoans
448:Santonian
436:Barremian
422:temperate
227:Named for
167:sandstone
154:Lithology
143:Thickness
36:Santonian
3711:Triassic
3707:Jurassic
3699:Mesozoic
3401:Mesozoic
2484:See also
2388:conifers
2288:Platanus
2265:conifers
2198:lobsters
2194:decapods
2143:Holaster
2116:crinoids
2112:starfish
2047:scallops
2007:bivalves
1967:Enchodus
1924:teleosts
1914:Several
1890:Amylodon
1768:nectonic
1538:mosasaur
1385:theropod
1297:and the
1238:Opuscula
1222:Mesozoic
1186:Ullstorp
1170:Bromölla
1162:Ivetofta
1156:Balsberg
1150:Axeltorp
1144:hybodont
1140:landfill
1136:clay pit
1010:Ullstorp
998:Ivetofta
986:Axeltorp
950:Balsberg
730:biozones
668:biozones
560:Jurassic
548:basement
511:mosasaur
483:bivalves
479:molluscs
188:Location
38:– Early
2400:neritic
2161:Salenia
2125:Cidaris
2080:Patella
2039:oysters
1986:Fossil
1879:sawfish
1853:Fossil
1549:(right)
1446:phalanx
1192:Balsvik
1180:Ivösjön
962:Balsvik
885:
817:
775:
660:
552:bedrock
521:Geology
503:turtles
459:
377:) is a
371:Swedish
204:Country
159:Primary
3780:Scania
3542:Middle
3351:
3196:
3089:
2945:
2897:
2669:
2089:Crania
2074:Nerita
2052:Pecten
2023:corals
1950:) and
1752:sharks
1104:kaolin
938:6miles
693:Image
634:rostra
603:horsts
601:, two
407:horsts
216:Sweden
213:
193:Region
3628:Upper
3575:Lower
3519:Upper
3483:Lower
3443:Upper
3272:(PDF)
3194:S2CID
3087:S2CID
2943:S2CID
2895:S2CID
2667:S2CID
2384:ferns
2282:Pinus
2273:ferns
2202:crabs
2095:Magas
1176:Kjuge
974:Kjuge
887:71.3
819:80.5
802:alpha
788:verus
777:83.5
751:Late
649:flint
532:SkĂĄne
495:shark
467:algae
454:age (
391:SkĂĄne
381:-age
198:SkĂĄne
181:flint
173:Other
3360:2021
3349:ISSN
2435:and
2390:and
2343:and
2303:and
2232:and
2210:and
2200:and
2192:and
2184:and
2158:and
2114:and
2104:and
2077:and
2033:and
1948:tuna
1920:gars
1905:and
1871:and
1834:and
1824:and
1802:and
1788:and
1778:and
1758:and
1756:rays
1719:Fish
1667:and
1655:and
1643:and
1609:and
1559:and
1289:and
1164:, a
1132:Ă…sen
1102:and
1022:Ă…sen
936:10km
690:Ref
678:Age
613:and
597:and
505:and
485:and
427:and
412:and
385:and
365:The
129:Type
53:Preęž’
3341:doi
3310:doi
3186:doi
3182:434
3079:doi
3075:434
3025:doi
2935:doi
2931:126
2927:GFF
2887:doi
2883:434
2857:110
2657:doi
2374:to
1936:),
1877:),
1816:),
1494:).
1108:Ivö
636:or
562:or
489:),
420:to
3726::
3709:•
3705:•
3701:•
3670:Pz
3618:Tr
3424:Mz
3413:Ph
3347:.
3337:32
3335:.
3331:.
3306:31
3304:.
3300:.
3288:^
3280:65
3278:.
3274:.
3241:.
3215:.
3192:.
3180:.
3176:.
3152:^
3133:.
3108:.
3085:.
3073:.
3069:.
3052:36
3050:.
3046:.
3021:42
3019:.
3015:.
2955:^
2941:.
2929:.
2925:.
2907:^
2893:.
2881:.
2877:.
2865:^
2855:.
2851:.
2833:^
2823:.
2805:^
2795:.
2711:^
2701:.
2679:^
2665:.
2653:50
2651:.
2647:.
2617:^
2607:.
2589:^
2579:.
2559:^
2463:.
2451:.
2394:.
2386:,
2382:,
2349:.
2309:.
2297:,
2291:,
2285:,
2271:,
2267:,
2226:,
2180:,
2176:,
2172:,
2152:,
2146:,
2140:,
2134:,
2128:,
2098:,
2092:,
2083:.
2071:,
2061:.
2045:,
2041:,
2029:,
2025:,
2021:,
2017:,
2013:,
2009:,
2005:,
1926:,
1922:,
1911:.
1899:,
1893:,
1846:.
1820:,
1798:,
1774:,
1754:,
1621:,
1583:.
1427:("
882:c.
829:,
825:,
814:c.
797:,
790:,
783:,
772:c.
657:c.
625:.
501:,
477:,
473:,
469:,
456:c.
431:.
373::
237:,
233:,
179:,
165:,
147:c.
135:,
103:Pg
47:Ma
42:,
34:,
30::
3510:J
3434:K
3393:e
3386:t
3379:v
3362:.
3343::
3316:.
3312::
3265:"
3252:.
3226:.
3200:.
3188::
3146:.
3119:.
3081::
3031:.
3027::
2949:.
2937::
2901:.
2889::
2859:.
2827:.
2799:.
2705:.
2673:.
2659::
2611:.
1865:(
1182:.
369:(
108:N
98:K
93:J
88:T
83:P
78:C
73:D
68:S
63:O
58:ęž’
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
