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Langstonia

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4056: 1069:, using its pressure force to retain and drown his victims is not viable on land; these crocodylomorphs also had more elongated limbs, stiff back, ideal to hold the weight and give strength to run. Langston had already suggested that these teeth were indicative that this animal had more terrestrial habits than those of existing species and the Miocene contemporary crocodylians, and in the modern world only could be vaguely compared with the less aquatic habits of the caimans of the genus 2643: 1018: 142: 124: 3316: 482: 3888: 3335: 5016: 3881: 831:: The largest fragment dental crown reported of La Venta, locality V-4528. Reaches up to 20.5 mm. high, but its base is much wider than other teeth, reaching 23.8 mm. Apart from its size, three times that of the teeth of the holotype of the species, this tooth is characterized by a blunt, relatively straight edges and a more oval profile tranverasal fastened with small saws. It may not correspond to 3895: 4108: 4090: 3903: 5028: 3328: 4072: 4063: 4081: 4117: 4099: 4135: 3322: 4128: 1079:, pp. 134–135) Busbey suggested the fact that his skull was high implied that these animals seek not to make pressure during the bite indefinitely, but to produce cuts then back quickly so that the bleeding weakens their prey slowly up to devour, similar to what has been thought to be the case for the extant 920:
rib attached, with a wide, rectangular neural spine; it measured 10.7 cm high by 5.5 cm wide, while it is estimated that the sacral ribs should be 10 cm wide at its rear end. The other is an anterior caudal vertebrae, perhaps the first, with incomplete neural spine, with a broad neural
782:
specimen. Their teeth are quite compressed and has a first side facing forward tooth, the second is directed upwards and the third also slopes, while others are vertical; the first teeth have a wide separation between them (diastema). Symphysis extending beyond the sixth tooth, while the bone surface
1036:
with jaws and teeth even more flattened, although its general proportions are reminiscent of this species, which suggests that their skull would be higher and laterally flattened, in contrast to modern crocodilians, which generally have a horizontally flattened skull with conical teeth. Teeth are of
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in that fitted the fourth lower tooth that exceeded the 3.5 centimeters; has a narrow palate, lateral nasal aperture and without mild depression around. The dentary is high, with four sockets, two projecting forward and a third and fourth straight, the latter being the largest. The associated
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the zyphodont kind, with very flattened sides, slightly curved back and with serrated edges with small denticles (between 5-6 denticles per millimeter in the teeth of this genus) and also without any grooves on its surface. This type of teeth appears in the close relatives of sebecids, the
2331:
Carvalho, Ismar de Souza; Antunes Texeira, Vicente de Paula; Da Fonseca Ferraz, Mara Lúcia; Borges Ribeiro, Luiz Carlos; Martinelli, Agustín Guillermo; Macedo Neto, Francísco; Sertich, Joseph J. W.; Cardoso Cunha, Gabriel; Cardoso Cunha, Isabella; Fonseca Ferraz, Patrícia (2011).
1136:- indicate that the climate was warm and humid with heavy rainfall and with drought periods not extending beyond 3–4 months, consisting of watersheds, forests and certain adjacent areas of open grasslands where it could hunt down a large variety of species belonging to the 596:, it was also proportionally thinner, and had with more recurved teeth being laterally compressed. Additionally Langston referred a series of zyphodont teeth in the area found this species, with some teeth referred to as similar but classified generally to 2656: 854:: Includes a portion of the left anterior premaxillary, middle left maxillary fragment, a fragment of the left splenial bone, back of the left suspensory bone formed by parts of the quadratojugal, the 1157:
were likely apex predators in their environment, and as an effect they would be particularly susceptible to ecological changes that caused other lineages, particularly hoofed mammals (such as groups
809:: An elongated, slender and compressed laterally tooth, with a slight curvature, intermediate in size between 44561 and other teeth reported, with larger saws (3 per millimeter tooth; in 939:: A tooth of the Duke 106 locality. It is also incomplete, but its measurements (52 mm high by 30 mm wide) makes it the largest tooth reported for a sebecosuchian in Colombia. 778:, with six tooth sockets of which five have broken teeth, collected in 1945 in the locality V-4517. This element measures 189.0 millimeters in length, being 68% higher than the rate of 2542:
Prevosti, Francisco J.; Forasiepi, Analía; Zimicz, Natalia (2013). "The evolution of the Cenozoic terrestrial mammalian predator guild in South America: competition or replacement?".
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in Brazil. Includes a full left premaxilla with a tooth preserved, and a piece of left maxilla with two alveoli. By having very flattened and spaced teeth looks remarkably similar to
4145: 955:
right tooth, measures 20 mm in height at the crown and 12 on the base, while its root has a length equivalent to 2.5 times the height of the crown. Unlike other teeth of
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Langston, along with the Argentine paleontologist Zulma Gasparini, reported new remains of the species in a contribution on fossil crocodylomorphs of Colombia for the book
3967: 975:: A single tooth found in the area of Fitzcarrald Arch in the localities of Inuya and Mapuya in the Amazon region of Peru; its size and shape are indistinguishable from 895:, the front left tooth, part of the dorsal border of a rostrum and two vertebrae. The premaxillary measured 12.8 cm in length, is tall and thin compared to the 755:
Here is a list of specimens assigned to the species, with a brief description of these. They are grouped according to the institution in which they are preserved:
702:, in honor of its initial descriptor and his "(..)advances in knowledge of crocodylians of Tertiary Colombia in particular and South America in general", while 4372: 4127: 5113: 870:. These fragments were recovered in 1976 in the "Monkey Pod" locality of La Venta (Baraya Member of the Villavieja Formation) by Dr Lundelius Ernest Jr. 529:
named the "lechos de monos" (Monkey Beds) in 1945. Many of the remains discovered were then sent to the collections of the Museum of Paleontology at the
5143: 1028:
Due to the fragmentary remains generally known for this family, it's only possible to describe some general aspects of the appearance and biology of
2601:
Salas-Gismondi, R.; Antoine, P.-O.; Baby, P.; Brusset, S.; Benammi, M.; Espurt, N.; de Franceschi, D.; Pujos, F.; Tejada, J.; Urbina, M. (2007).
592:. Langston gave the taxon a new species considering that besides being larger, as the dentary fragment is 68% larger than the Argentine species, 5138: 3960: 2691: 933:: Tooth from the Duke 196 locality, is incomplete and worn, but is similar to the tooth UCMP 40186. It measures 39 mm high by 27 wide. 3930: 903:
in the premaxilla, from the edge of the second tooth to the fourth, which is particularly enlarged, being twice the third (reverse that in
4055: 622:
Arthur Busbey (1986) later brought new remains of the species also from the "Monkey Beds" of Huila in Colombia, this time a fragment of
2617: 530: 5019: 3953: 541:, in which he made a detailed analysis of several fossils of crocodylomorphs in Colombia, including the remains of other species as 2400: 1065:. These forms were land animals that used their teeth to cut and tear their prey, because the traditional method of capture of the 1075:, which have a higher muzzle and a higher degree of ossification around the eye socket, so that they partially resemble sebecids.( 927:: A tooth found in the Duke 91 locality. The crown of this is twice as high as it is wide, reaching 36 mm. high and 17 wide. 5108: 649:(1997). In this case it would be new specimens of teeth, a somewhat fuller premaxilla, an anterior fragment of dentary, and two 533:(UCMP), where they are still housed. The description of the specimens did not come until 1965, when the American paleontologist 841:: The lower teeth reported by Langston (1965). Recovered in the locality V-4421, measuring just 6.1 mm. wide at the base. 657:, which also were unknown to this group. The specimens discovered on this occasion were deposited in the Geological Museum of 5133: 615:
the presence of a huge cranial portion (area of the snout), referring it to this genus, and although its age coincides with
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Langston (Sebecia, Metasuchia) in the Middle Paleocene of the Itaboraí Basin, Rio de Janeiro State, Southeastern Brazil"
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Kay, R.; Madden, R. (1997). "Paleogeography and paleoecology". In Kay, R.; Madden, R.; Cifelli, R.; et al. (eds.).
735:(and where have been found fossils of other sebecids) has been classified as related to this species, under the name of 716:. Paolillo and Linares also clarified the status of the Peruvian fossil before assigned to the species, assigning it to 720:, although this does not rule out its presence in more southern areas; in the area of the Peruvian Amazon known as the 611:
Subsequently, Éric Buffetaut and Robert Hoffstetter (1977) reported from the Ipururo Formation in the Amazon region of
4238: 2373:
Langston, W. (1965). "Fossil crocodilians from Colombia and the Cenozoic History of the Crocodylia in South America".
2602: 1188:, a lineage of ziphodont terrestrial crocodylomorphs forming part of a separate lineage of the southern continent of 803:: Tooth of the Late Oligocene fauna found in Coyaima, locality V-4411. It measures 22.5 mm. height at the crown. 3887: 141: 4329: 4306: 3315: 825:: A fragment of the dental crown, found in the locality V-4528; it measured 16.3 mm at the base of the crown. 747:, suggesting that the latter is part of a lineage of sebecids with an extensive temporal and geographical record. 5123: 4188: 2684: 2256:"Evolution of South American mammalian predators (Borhyaenoidea): anatomical & palaeobiological implications" 965: 3334: 2444: 1429: 458:
was named in 2007 by Alfredo Paolillo and Omar Linares for fossils originally described by Langston in 1965 as
3880: 3894: 993: 794: 605: 5043: 4831: 4716: 4038: 638:), which were the earliest known for this family. Busbey also ruled out the Peruvian fossil belonging to 3902: 664:
Moreover, Alfredo Paolillo and Omar Linares described a new genus of a large Venezuelan Miocene sebecid,
5075: 4462: 4390: 4276: 4206: 4201: 4002: 2776: 1143:
The disappearance of the system of large rivers of the Amazon lake system and the gradual uplift of the
584:; thus extending the time range of this genus and the family, hitherto known only from remains of the 543: 451: 340: 5023: 4672: 4251: 4071: 502: 721: 4924: 4916: 4772: 4319: 4098: 3327: 2677: 2507:"A new sebecid mesoeucrocodylian from the Rio Loro Formation (Palaeocene) of north-western Argentina" 2477: 2311: 1412: 526: 317: 731:
Finally, a premaxilla and maxilla fragment found in Itaboraí Basin in Brazil dating from the Middle
506: 4652: 4441: 4246: 4211: 4196: 4062: 2648: 2338:
gen. et sp. nov., a new Late Cretaceous baurusuchid (Crocodyliformes) from the Bauru Basin, Brazil"
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has been found a characteristically flattened tooth from the middle Miocene therefore assigned to
1017: 4999: 4991: 4950: 4906: 4898: 4887: 4616: 4469: 4342: 4314: 4080: 4012: 3630: 2569: 2493: 1173:) to die out, thus leading to extinction to the last notosuchian crocodilomorphs of the world. 1120: 418: 414: 369: 360: 136: 5080: 4178: 4017: 2582: 3321: 5118: 5103: 4798: 4790: 4763: 4623: 4602: 4581: 4455: 4406: 4286: 4281: 4116: 4089: 3190: 2920: 2613: 534: 5027: 4166: 3774: 2736: 2559: 2551: 2528: 2518: 2485: 2390:, and the South American gavials". In Kay, R.; Madden, R.; Cifelli, R.; et al. (eds.). 2352: 2319: 2267: 1114:
in its ecosystem. The fossils found of other species - big fishes, manatees, large browsing
1058: 131:
Restoration showing known elements, with remaining material filled in after related species
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caused major ecological changes in South America in the mid-Miocene. The last sebecids,
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bone, the suspensory and retroarticular joint in the jaw, and a fragment of bone scute (
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The Tatacoa Desert, in which has been discovered numerous Miocene fossils, including to
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Riff, D.; de Souza, R.G.; Cidade, G.M.; Martinelli, A.G.; de Souza-Filho, J.P. (2012).
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as in the southern part of the continent, these sebecids were probably the terrestrial
1107: 561: 510: 481: 2237: 1938: 5097: 4943: 4880: 4866: 4779: 4539: 4399: 3842: 3815: 3781: 3742: 3727: 3700: 3693: 3639: 3605: 3598: 3589: 3551: 3516: 3428: 3398: 3391: 3260: 3176: 3169: 3153: 3079: 3051: 2523: 2506: 2272: 2255: 1487: 1467: 1341: 1158: 1132: 1111: 1087: 1080: 395: 74: 2573: 2497: 1086:
Since in the fauna of La Venta, although there were some large mammalian predators (
866:, the left retroarticular processes (partial) right to the jaw and a fragment of an 4966: 4958: 4859: 4817: 4702: 4546: 4504: 4497: 4427: 3822: 3718: 3672: 3463: 3406: 3232: 3204: 3162: 3121: 3114: 3098: 3071: 3064: 3026: 2963: 2761: 2425:"Nuevos cocodrilos Sebecosuchia del Cenozoico Suramericano (Mesosuchia:Crocodylia)" 2323: 1545: 1268: 1185: 1170: 1162: 1153: 1126: 1083:, thereby adopting a strategy of stalking in areas where prey transit frequently. 666: 466: 267: 209: 123: 2392:
Vertebrate Paleontology in the Neotropics. The Miocene Fauna of La Venta, Colombia
2366:
Vertebrate Paleontology in the Neotropics. The Miocene Fauna of La Venta, Colombia
2302:(Crocodylia: Sebecosuchidae) from the Miocene of La Venta Formation of Colombia". 2489: 2424: 2405:
and the iberoccitanian crocodile. A morphological and ultrastructural comparison"
959:, has a slight groove at the base of the tooth crown, on the inner side thereof. 4852: 4845: 4838: 4595: 4588: 4567: 4560: 4490: 4476: 4434: 4420: 4413: 4379: 4256: 3735: 3653: 3612: 3524: 3496: 3383: 3253: 3239: 3142: 3040: 2991: 2977: 2970: 2956: 2929: 2887: 2856: 2838: 2825: 2788: 2700: 1673: 1593: 1576: 1494: 1474: 1214: 1166: 1071: 1062: 1050: 1042: 1038: 708: 555: 549: 522: 49: 5066: 2464:
Pol, D.; Leardi, J.M.; Lecuona, A.; Krause, M. (2012). "Postcranial anatomy of
4980: 4973: 4695: 4574: 4553: 4511: 3788: 3750: 3470: 3449: 3421: 3365: 3352: 3283: 3246: 3218: 3211: 3135: 3019: 3012: 2909: 2745: 2730: 2638: 2555: 2356: 1521: 1046: 888: 876: 658: 623: 566: 501:
were discovered in the province of Huila in Colombia by the Spanish geologist
429: 251: 94: 59: 887:: Found in the Duke 57 locality in Villavieja Formation; consists of a right 4752: 4731: 4723: 4448: 4134: 3578: 3128: 2998: 2947: 2894: 1507: 1362: 1219: 1197: 1137: 1098: 1066: 867: 859: 797:, Mugrosa Formation). It measures 14.2 millimeters in diameter at the base. 775: 732: 687: 671: 635: 589: 518: 514: 436: 296: 196: 153: 99: 43: 4171: 3945: 2610:
4th European Meeting on the Palaeontology and Stratigraphy of Latin America
2333: 2623: 1200:
have consistently shown that these forms a clade within Sebecidae next to
5060: 4525: 2724: 1189: 1092: 1054: 912: 793:: Base of a tooth of the Upper Eocene, from the V-4620 town (Tama fauna, 650: 574: 448: 444: 173: 89: 84: 69: 64: 54: 34: 1102:), neither of them equated in size to the contemporary crocodylomorphs, 647:
Vertebrate Paleontology of the Neotropics: The Miocene Fauna of La Venta
4532: 3033: 2873: 2533: 1656: 952: 951:: A tooth of the Duke 41 locality, that probably correspond to a first 945:: Tooth from Duke 40 locality. It measures 30 mm high by 17 wide. 911:, p. 114) After the fourth premaxillary tooth there is a space or 892: 691: 579: 570: 460: 440: 422: 379: 283: 183: 104: 79: 30: 2564: 819:: A large tooth, locality V.4523, measures about 42.4 mm height. 505:, during the expeditions in the region by the American paleontologist 4661: 2718: 2603:"Middle Miocene crocodiles from the Fitzcarrald Arch, Amazonian Peru" 917: 855: 674:. In their article they considered that the traits of two species of 642:, based on its unique anatomy, less spaced teeth and a wider nose. 601: 585: 494: 163: 5037: 1032:. As mentioned above, the fossil type of the species is larger than 694:, and their age justified the creation of separate genres for both: 2583:"Crocodilomorfos. A maior diversidade de répteis fósseis do Brasil" 899:
being covered with fine grooves on its outer surface. It has three
2281:
Buffetaut, E.; Hoffstetter, R. (1977). "Découverte du Crocodilien
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Department of Vertebrate Paleontology, Museum of Natural History,
863: 631: 480: 425: 992:: consists of material of sebecid found in Itaboraí Basin in the 2159: 1951: 612: 5041: 3949: 3919: 3350: 2823: 2711: 2673: 1106:, p. 513), and given the absence of large predatory birds 712:, although some authors still maintain both species within 630:, and equally thin. He also described the first remains of the 2669: 774:: The front part of a right dentary preserving the mandibular 1192:. As noted earlier, this species formerly part of the genus 4133: 4126: 2468:(Crocodyliformes, Sebecidae) from the Eocene of Patagonia". 2399:
Legasa, O.; Buscalioni, Ángela D.; Gasparini, Zulma (1993).
2375:
University of California Publications of Geological Sciences
2211: 619:, distinguished it through it being larger and more robust. 2443:
Pinheiro, A.E.P.; Campos, D.A.; Bergqvist, L.P. (2011).
763:
Not all materials from this collection were assigned to
2224: 2172: 1964: 1927: 767:
but are included because of their similar morphology.
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Museum of Paleontology at the University of California
577:
of a new species a fragmentary taxon, which he called
1852: 5050: 4990: 4933: 4915: 4897: 4807: 4789: 4762: 4742: 4671: 4651: 4633: 4389: 4362: 4355: 4328: 4305: 4267: 4237: 4187: 4157: 4144: 4026: 3990: 3981: 3832: 3798: 3760: 3717: 3671: 3629: 3588: 3550: 3495: 3484: 3438: 3363: 3282: 3152: 3097: 3050: 2946: 2919: 2871: 2836: 2787: 2760: 2386:Langston, W.; Gasparini, Z. (1997). "Crocodilians, 921:canal, and measures 2.8 cm wide and 2.1 high. 2394:. Smithsonian Institution Press. pp. 113–154. 2368:. Smithsonian Institution Press. pp. 520–550. 2287:Comptes Rendus de l'Académie des Sciences de Paris 661:(previous name of Colombian Geological Service). 569:). As for the sebecid material he designated the 2146: 2133: 2120: 2107: 2094: 2081: 2068: 1891: 908: 600:sp., which do not come from the Miocene but the 1904: 1774: 907:), while the first two teeth should be small.( 513:at the locality V-4517, characterized by gray 447:land-mammal age), in the "Monkey Beds" of the 3961: 2685: 8: 2423:Paolillo, Alfredo; Linares, Omar J. (2007). 916:vertebrae are an amphycoelian sacral with a 5038: 4359: 4154: 3987: 3968: 3954: 3946: 3916: 3766: 3677: 3556: 3501: 3492: 3375: 3371: 3360: 3347: 3279: 3056: 2879: 2848: 2844: 2833: 2820: 2766: 2757: 2708: 2692: 2678: 2670: 2608:. In Díaz-Martínez, E; Rábano, I. (eds.). 984:Museu de Ciências da Terra, Rio de Janeiro 743:, whose main features are very similar to 122: 20: 2563: 2532: 2522: 2511:Zoological Journal of the Linnean Society 2271: 2260:Zoological Journal of the Linnean Society 2198: 1916: 1223: 509:. They were found in the area called the 2042: 2029: 2016: 2003: 1990: 1977: 1839: 1826: 1813: 1800: 1787: 1076: 1766: 2185: 2055: 1878: 1865: 2505:Pol, Diego; Powell, Jaime E. (2011). 2225:Prevosti, Forasiepi & Zimicz 2013 2173:Pinheiro, Campos & Bergqvist 2011 1965:Pinheiro, Campos & Bergqvist 2011 1103: 521:sediments. These have been named the 7: 2298:Busbey, A. (1986). "New material of 2285:dans le Miocène du Pérou oriental". 1000:, being provisionally classified as 1618: 1568: 1561: 1537: 1513: 1463: 1404: 1354: 1347: 1320: 1296: 1243: 1236: 1229: 2470:Journal of Vertebrate Paleontology 2304:Journal of Vertebrate Paleontology 531:University of California, Berkeley 14: 5114:Miocene reptiles of South America 539:Fossil Crocodylians from Colombia 5144:Prehistoric pseudosuchian genera 5026: 5015: 5014: 4115: 4106: 4097: 4088: 4079: 4070: 4061: 4054: 3901: 3893: 3886: 3879: 3333: 3326: 3320: 3314: 2641: 2524:10.1111/j.1096-3642.2011.00714.x 2273:10.1111/j.1096-3642.2004.00110.x 1853:Buffetaut & Hoffstetter 1977 393:Langston & Gasparini, 1997 ( 140: 2412:Studia Geologica Salmanticensia 2544:Journal of Mammalian Evolution 2324:10.1080/02724634.1986.10011595 813:typically are 4 to 6 mm). 417:", in honor of paleontologist 1: 5139:Fossil taxa described in 1965 2147:Langston & Gasparini 1997 2134:Langston & Gasparini 1997 2121:Langston & Gasparini 1997 2108:Langston & Gasparini 1997 2095:Langston & Gasparini 1997 2082:Langston & Gasparini 1997 2069:Langston & Gasparini 1997 1892:Langston & Gasparini 1997 909:Langston & Gasparini 1997 525:, of the La Venta fauna, the 2490:10.1080/02724634.2012.646833 1208:, not so closely related to 3931:Terrestrial crocodylomorphs 1905:Paolillo & Linares 2007 1775:Paolillo & Linares 2007 877:Colombian Geological Survey 846:Texas Memorial Museum (TMM) 5160: 2459:(4, Supplemento): R91–R92. 2160:Salas-Gismondi et al. 2007 1952:Salas-Gismondi et al. 2007 16:Extinct species of reptile 5012: 4049: 3926: 3915: 3877: 3769: 3680: 3559: 3504: 3378: 3374: 3359: 3346: 3312: 3278: 3059: 2882: 2851: 2847: 2832: 2819: 2769: 2756: 2707: 2556:10.1007/s10914-011-9175-9 2432:Paleobiologia Neotropical 2357:10.11646/zootaxa.2871.1.2 1670: 1653: 1636: 1623: 1616: 1590: 1573: 1566: 1559: 1542: 1535: 1518: 1511: 1491: 1471: 1461: 1426: 1409: 1402: 1376: 1359: 1352: 1345: 1325: 1318: 1301: 1294: 1265: 1248: 1241: 1234: 966:Universidad de San Marcos 439:. It lived in the middle 375: 368: 346: 339: 137:Scientific classification 135: 130: 121: 23: 3976:Paleontology in Colombia 1430:Baurusuchus salgadoensis 537:published his monograph 316:Paolillo & Linares, 5109:Miocene crocodylomorphs 2401:"The serrated teeth of 2289:. Série D (in French). 1917:Pol & Powell (2011) 1224:Pol & Powell (2011) 1021:Life reconstruction of 994:state of Rio de Janeiro 858:bone, opisthotic and a 795:department of Santander 606:department of Santander 556:Gryposuchus colombianus 4717:Montrichardia aquatica 4138: 4131: 1184:is considered part of 1130:, and some grazers as 1025: 562:Charactosuchus fieldsi 490: 390:Sebecosuchus huilensis 5134:Honda Group, Colombia 5076:Paleobiology Database 4463:Galbula hylochoreutes 4137: 4130: 2447:Sebecus cf. huilensis 2300:Sebecus cf. huilensis 2199:Kay & Madden 1997 1939:Carvalho et al. (2011 1198:phylogenetic analyzes 1093:Lycopsis longirostrus 1020: 765:Langstonia huilensis 544:Purussaurus neivensis 484: 443:(specifically in the 4773:Pachyrhizodus etayoi 2336:Campinasuchus dinizi 2238:Carvalho et al. 2011 1855:, Figure 1; p. 1664) 1413:Baurusuchus pachecoi 783:is smoother than in 698:was given the genus 527:geological formation 477:Discovery and naming 452:Villavieja Formation 353:Langstonia huilensis 5129:Fossils of Colombia 4442:Dracaena colombiana 4150:stratigraphic units 2649:Paleontology portal 2482:2012JVPal..32..328P 2466:Sebecus icaeorhinus 2316:1986JVPal...6...20B 2201:, pp. 534–535) 2136:, pp. 116–117) 2123:, pp. 116–117) 2110:, pp. 116–117) 1034:Sebecus icaeorhinus 875:Museum of Geology, 862:, a left part of a 780:Sebecus icaeorhinus 4888:Stenorhynchosaurus 4470:Granastrapotherium 4139: 4132: 3921:Related categories 3631:Pissarrachampsinae 2254:Argot, C. (2004). 1121:Granastrapotherium 1026: 573:UCMP 37877 as the 550:Mourasuchus atopus 491: 419:Wann Langston, Jr. 5091: 5090: 5044:Taxon identifiers 5035: 5034: 5008: 5007: 4624:Xenastrapotherium 4603:Saimiri annectens 4456:Eunectes stirtoni 4407:Aramus paludigrus 4351: 4350: 4047: 4046: 3943: 3942: 3939: 3938: 3911: 3910: 3875: 3874: 3871: 3870: 3867: 3866: 3863: 3862: 3859: 3858: 3667: 3666: 3663: 3662: 3480: 3479: 3342: 3341: 3310: 3309: 3306: 3305: 3302: 3301: 3274: 3273: 3270: 3269: 3191:Colhuehuapisuchus 2921:Mahajangasuchidae 2867: 2866: 2815: 2814: 2811: 2810: 2807: 2806: 2659:Sebecus huilensis 2592:(in Portuguese). 2445:"The presence of 1980:, pp. 15–16) 1928:Pol et al. (2012) 1757: 1756: 1748: 1747: 1739: 1738: 1730: 1729: 1721: 1720: 1712: 1711: 1703: 1702: 1694: 1693: 1685: 1684: 1625:Form of Lumbrera 1605: 1604: 1450: 1449: 1441: 1440: 1391: 1390: 1280: 1279: 990:MCT 1795-R/1796-R 653:: a sacral and a 565:(plus a possible 535:Wann Langston Jr. 503:José Royo y Gómez 406: 405: 400: 386: 332:L. huilensis 320: 5151: 5124:Neogene Colombia 5084: 5083: 5071: 5070: 5069: 5039: 5030: 5018: 5017: 4360: 4155: 4119: 4110: 4101: 4092: 4083: 4074: 4065: 4058: 3988: 3970: 3963: 3956: 3947: 3917: 3906: 3905: 3898: 3897: 3891: 3890: 3883: 3775:Adamantinasuchus 3767: 3678: 3557: 3502: 3493: 3376: 3372: 3361: 3348: 3337: 3331: 3330: 3324: 3318: 3280: 3057: 2880: 2849: 2845: 2834: 2821: 2767: 2758: 2751: 2750: 2737:Mesoeucrocodylia 2709: 2694: 2687: 2680: 2671: 2651: 2646: 2645: 2644: 2630: 2628: 2622:. Archived from 2607: 2597: 2587: 2577: 2567: 2538: 2536: 2526: 2501: 2460: 2439: 2429: 2419: 2409: 2395: 2382: 2369: 2360: 2342: 2327: 2294: 2277: 2275: 2241: 2234: 2228: 2221: 2215: 2212:Riff et al. 2012 2208: 2202: 2195: 2189: 2182: 2176: 2169: 2163: 2156: 2150: 2143: 2137: 2130: 2124: 2117: 2111: 2104: 2098: 2091: 2085: 2078: 2072: 2065: 2059: 2052: 2046: 2039: 2033: 2026: 2020: 2013: 2007: 2000: 1994: 1987: 1981: 1974: 1968: 1961: 1955: 1948: 1942: 1936: 1930: 1925: 1919: 1914: 1908: 1901: 1895: 1888: 1882: 1875: 1869: 1862: 1856: 1849: 1843: 1836: 1830: 1823: 1817: 1810: 1804: 1797: 1791: 1784: 1778: 1771: 1619: 1569: 1562: 1538: 1514: 1464: 1405: 1355: 1348: 1321: 1297: 1244: 1237: 1230: 891:and part of the 722:Fitzcarrald Arch 392: 384: 355: 351: 315: 308: 295: 282: 266: 250: 145: 144: 126: 116: 46: 29:Temporal range: 21: 5159: 5158: 5154: 5153: 5152: 5150: 5149: 5148: 5094: 5093: 5092: 5087: 5079: 5074: 5065: 5064: 5059: 5046: 5036: 5031: 5004: 4986: 4929: 4911: 4907:Dinosaur tracks 4893: 4825:Callawayasaurus 4803: 4785: 4758: 4738: 4710:Cerrejonisuchus 4682:Acherontisuchus 4667: 4647: 4629: 4610:Saimiri fieldsi 4385: 4356:Notable fossils 4347: 4324: 4301: 4277:La Frontera Fm. 4263: 4233: 4202:Castilletes Fm. 4183: 4149: 4147: 4140: 4125: 4124: 4123: 4122: 4121: 4120: 4112: 4111: 4103: 4102: 4094: 4093: 4085: 4084: 4076: 4075: 4067: 4066: 4043: 4022: 3983: 3977: 3974: 3944: 3935: 3922: 3907: 3900: 3892: 3885: 3855: 3834:Caipirasuchinae 3828: 3809:Armadillosuchus 3794: 3756: 3713: 3659: 3647:Pissarrachampsa 3625: 3620:Stratiotosuchus 3584: 3565:Cynodontosuchus 3546: 3540:Razanandrongobe 3533:Pehuenchesuchus 3489: 3487:Xenodontosuchia 3476: 3440:Candidodontidae 3434: 3414:Razanandrongobe 3368: 3355: 3338: 3325: 3298: 3266: 3226:Montealtosuchus 3198:Gasparinisuchus 3148: 3093: 3046: 2942: 2937:Mahajangasuchus 2915: 2876: 2863: 2841: 2828: 2803: 2783: 2752: 2714: 2713: 2703: 2698: 2647: 2642: 2640: 2637: 2626: 2620: 2605: 2600: 2585: 2580: 2541: 2504: 2463: 2442: 2427: 2422: 2407: 2398: 2385: 2372: 2363: 2340: 2330: 2297: 2280: 2253: 2250: 2245: 2244: 2235: 2231: 2222: 2218: 2209: 2205: 2196: 2192: 2183: 2179: 2170: 2166: 2157: 2153: 2144: 2140: 2131: 2127: 2118: 2114: 2105: 2101: 2092: 2088: 2079: 2075: 2066: 2062: 2053: 2049: 2040: 2036: 2027: 2023: 2014: 2010: 2001: 1997: 1988: 1984: 1975: 1971: 1962: 1958: 1949: 1945: 1937: 1933: 1926: 1922: 1915: 1911: 1902: 1898: 1889: 1885: 1876: 1872: 1863: 1859: 1850: 1846: 1837: 1833: 1824: 1820: 1811: 1807: 1798: 1794: 1785: 1781: 1772: 1768: 1763: 1758: 1749: 1740: 1731: 1722: 1713: 1704: 1695: 1686: 1606: 1451: 1442: 1392: 1380:Stratiotosuchus 1329:Cynodontosuchus 1305:Pehuenchesuchus 1281: 1179: 1116:meridiungulates 1059:pristichampsids 1015: 986: 969: 881: 848: 761: 753: 751:Specimens known 626:, smaller than 604:, found in the 479: 364: 357: 349: 348: 335: 314: 306: 293: 280: 264: 248: 236:Crocodyliformes 223:Crocodylomorpha 139: 117: 115: 114: 113: 112: 107: 102: 97: 92: 87: 82: 77: 72: 67: 62: 57: 52: 41: 40: 38: 27: 17: 12: 11: 5: 5157: 5155: 5147: 5146: 5141: 5136: 5131: 5126: 5121: 5116: 5111: 5106: 5096: 5095: 5089: 5088: 5086: 5085: 5072: 5056: 5054: 5048: 5047: 5042: 5033: 5032: 5013: 5010: 5009: 5006: 5005: 5003: 5002: 4996: 4994: 4988: 4987: 4985: 4984: 4977: 4970: 4963: 4954: 4947: 4939: 4937: 4931: 4930: 4928: 4927: 4921: 4919: 4917:Valle Alto Fm. 4913: 4912: 4910: 4909: 4903: 4901: 4895: 4894: 4892: 4891: 4884: 4877: 4874:Platypterygius 4870: 4863: 4856: 4849: 4842: 4835: 4828: 4821: 4813: 4811: 4805: 4804: 4802: 4801: 4795: 4793: 4787: 4786: 4784: 4783: 4776: 4768: 4766: 4760: 4759: 4757: 4756: 4748: 4746: 4740: 4739: 4737: 4736: 4727: 4720: 4713: 4706: 4699: 4692: 4689:Anthracosuchus 4685: 4677: 4675: 4669: 4668: 4666: 4665: 4657: 4655: 4649: 4648: 4646: 4645: 4642:Colombitherium 4637: 4635: 4631: 4630: 4628: 4627: 4620: 4613: 4606: 4599: 4592: 4585: 4578: 4571: 4564: 4557: 4550: 4543: 4536: 4529: 4522: 4515: 4508: 4501: 4494: 4487: 4484:Hilarcotherium 4480: 4473: 4466: 4459: 4452: 4445: 4438: 4431: 4424: 4417: 4410: 4403: 4395: 4393: 4387: 4386: 4384: 4383: 4376: 4368: 4366: 4357: 4353: 4352: 4349: 4348: 4346: 4345: 4340: 4334: 4332: 4326: 4325: 4323: 4322: 4320:Valle Alto Fm. 4317: 4311: 4309: 4303: 4302: 4300: 4299: 4294: 4289: 4284: 4279: 4273: 4271: 4265: 4264: 4262: 4261: 4260: 4259: 4249: 4243: 4241: 4235: 4234: 4232: 4231: 4226: 4221: 4216: 4215: 4214: 4204: 4199: 4193: 4191: 4185: 4184: 4182: 4181: 4176: 4175: 4174: 4163: 4161: 4152: 4142: 4141: 4114: 4113: 4105: 4104: 4096: 4095: 4087: 4086: 4078: 4077: 4069: 4068: 4060: 4059: 4053: 4052: 4051: 4050: 4048: 4045: 4044: 4042: 4041: 4036: 4030: 4028: 4024: 4023: 4021: 4020: 4015: 4010: 4005: 4000: 3994: 3992: 3985: 3979: 3978: 3975: 3973: 3972: 3965: 3958: 3950: 3941: 3940: 3937: 3936: 3934: 3933: 3927: 3924: 3923: 3920: 3913: 3912: 3909: 3908: 3878: 3876: 3873: 3872: 3869: 3868: 3865: 3864: 3861: 3860: 3857: 3856: 3854: 3853: 3850:Morrinhosuchus 3846: 3838: 3836: 3830: 3829: 3827: 3826: 3819: 3812: 3804: 3802: 3800:Sphagesaurinae 3796: 3795: 3793: 3792: 3785: 3778: 3770: 3764: 3762:Sphagesauridae 3758: 3757: 3755: 3754: 3747: 3739: 3732: 3723: 3721: 3715: 3714: 3712: 3711: 3708:Eptalofosuchus 3704: 3697: 3690: 3686:Brasileosaurus 3681: 3675: 3669: 3668: 3665: 3664: 3661: 3660: 3658: 3657: 3650: 3643: 3635: 3633: 3627: 3626: 3624: 3623: 3616: 3609: 3602: 3594: 3592: 3586: 3585: 3583: 3582: 3575: 3572:Gondwanasuchus 3568: 3560: 3554: 3548: 3547: 3545: 3544: 3536: 3529: 3521: 3513: 3510:Chimaerasuchus 3505: 3499: 3490: 3485: 3482: 3481: 3478: 3477: 3475: 3474: 3467: 3460: 3457:Lavocatchampsa 3453: 3445: 3443: 3436: 3435: 3433: 3432: 3425: 3418: 3410: 3403: 3395: 3388: 3379: 3369: 3364: 3357: 3356: 3351: 3344: 3343: 3340: 3339: 3313: 3311: 3308: 3307: 3304: 3303: 3300: 3299: 3297: 3296: 3295: 3294: 3288: 3286: 3276: 3275: 3272: 3271: 3268: 3267: 3265: 3264: 3257: 3250: 3243: 3236: 3229: 3222: 3215: 3208: 3201: 3194: 3187: 3184:Bayomesasuchus 3180: 3173: 3166: 3158: 3156: 3150: 3149: 3147: 3146: 3139: 3132: 3125: 3118: 3111: 3108:Barreirosuchus 3103: 3101: 3095: 3094: 3092: 3091: 3087:Stolokrosuchus 3083: 3076: 3068: 3060: 3054: 3048: 3047: 3045: 3044: 3037: 3030: 3023: 3016: 3009: 3002: 2995: 2988: 2985:Dentaneosuchus 2981: 2974: 2967: 2960: 2952: 2950: 2944: 2943: 2941: 2940: 2933: 2925: 2923: 2917: 2916: 2914: 2913: 2906: 2902:Trematochampsa 2898: 2891: 2883: 2877: 2872: 2869: 2868: 2865: 2864: 2862: 2861: 2852: 2842: 2837: 2830: 2829: 2824: 2817: 2816: 2813: 2812: 2809: 2808: 2805: 2804: 2802: 2801: 2800: 2799: 2793: 2791: 2785: 2784: 2782: 2781: 2780: 2779: 2770: 2764: 2754: 2753: 2749: 2748: 2739: 2733: 2727: 2721: 2712: 2705: 2704: 2699: 2697: 2696: 2689: 2682: 2674: 2668: 2667: 2653: 2652: 2636: 2635:External links 2633: 2632: 2631: 2629:on 2009-07-04. 2619:978-8478407071 2618: 2598: 2578: 2539: 2502: 2476:(2): 328–354. 2461: 2440: 2434:(in Spanish). 2420: 2396: 2383: 2370: 2361: 2328: 2295: 2278: 2266:(4): 487–521. 2249: 2246: 2243: 2242: 2229: 2216: 2203: 2190: 2177: 2175:, p. R92) 2164: 2162:, p. 356) 2151: 2149:, p. 117) 2138: 2125: 2112: 2099: 2097:, p. 116) 2086: 2084:, p. 116) 2073: 2071:, p. 115) 2060: 2047: 2034: 2021: 2008: 1995: 1982: 1969: 1967:, p. R92) 1956: 1954:, p. 356) 1943: 1931: 1920: 1909: 1896: 1894:, p. 114) 1883: 1870: 1857: 1844: 1831: 1818: 1805: 1792: 1779: 1765: 1764: 1762: 1759: 1755: 1754: 1751: 1750: 1746: 1745: 1742: 1741: 1737: 1736: 1733: 1732: 1728: 1727: 1724: 1723: 1719: 1718: 1715: 1714: 1710: 1709: 1706: 1705: 1701: 1700: 1697: 1696: 1692: 1691: 1688: 1687: 1683: 1682: 1679: 1678: 1669: 1666: 1665: 1662: 1661: 1652: 1649: 1648: 1645: 1644: 1635: 1632: 1631: 1628: 1627: 1622: 1617: 1615: 1612: 1611: 1608: 1607: 1603: 1602: 1599: 1598: 1589: 1586: 1585: 1582: 1581: 1572: 1567: 1565: 1560: 1558: 1555: 1554: 1551: 1550: 1541: 1536: 1534: 1531: 1530: 1527: 1526: 1517: 1512: 1510: 1504: 1503: 1500: 1499: 1490: 1484: 1483: 1480: 1479: 1470: 1462: 1460: 1457: 1456: 1453: 1452: 1448: 1447: 1444: 1443: 1439: 1438: 1435: 1434: 1425: 1422: 1421: 1418: 1417: 1408: 1403: 1401: 1398: 1397: 1394: 1393: 1389: 1388: 1385: 1384: 1375: 1372: 1371: 1368: 1367: 1358: 1353: 1351: 1346: 1344: 1338: 1337: 1334: 1333: 1324: 1319: 1317: 1314: 1313: 1310: 1309: 1300: 1295: 1293: 1287: 1286: 1283: 1282: 1278: 1277: 1274: 1273: 1264: 1261: 1260: 1257: 1256: 1252:Chimaerasuchus 1247: 1242: 1240: 1235: 1233: 1228: 1202:S. icaeorhinus 1178: 1175: 1112:apex predators 1014: 1011: 1010: 1009: 985: 982: 981: 980: 968: 962: 961: 960: 946: 940: 934: 928: 922: 905:S. icaeorhinus 901:dental alveoli 897:S. icaeorhinus 880: 873: 872: 871: 847: 844: 843: 842: 836: 826: 820: 814: 804: 798: 788: 785:S. icaeorhinus 760: 757: 752: 749: 628:S. icaeorhinus 608:in Colombia. 594:S. icaeorhinus 511:Tatacoa Desert 507:Robert Stirton 478: 475: 435:of the family 433:crocodylomorph 413:(meaning " of 404: 403: 402: 401: 387: 385:Langston, 1965 373: 372: 366: 365: 358: 344: 343: 337: 336: 328: 326: 322: 321: 304: 300: 299: 291: 287: 286: 278: 271: 270: 262: 255: 254: 246: 239: 238: 233: 226: 225: 220: 213: 212: 207: 200: 199: 194: 187: 186: 181: 177: 176: 171: 167: 166: 161: 157: 156: 151: 147: 146: 133: 132: 128: 127: 119: 118: 110: 109: 108: 103: 98: 93: 88: 83: 78: 73: 68: 63: 58: 53: 48: 47: 28: 15: 13: 10: 9: 6: 4: 3: 2: 5156: 5145: 5142: 5140: 5137: 5135: 5132: 5130: 5127: 5125: 5122: 5120: 5117: 5115: 5112: 5110: 5107: 5105: 5102: 5101: 5099: 5082: 5077: 5073: 5068: 5062: 5058: 5057: 5055: 5053: 5049: 5045: 5040: 5029: 5025: 5021: 5011: 5001: 4998: 4997: 4995: 4993: 4989: 4983: 4982: 4978: 4976: 4975: 4971: 4969: 4968: 4964: 4962: 4961: 4960: 4955: 4953: 4952: 4948: 4946: 4945: 4944:Antarctilamna 4941: 4940: 4938: 4936: 4932: 4926: 4923: 4922: 4920: 4918: 4914: 4908: 4905: 4904: 4902: 4900: 4896: 4890: 4889: 4885: 4883: 4882: 4881:Sachicasaurus 4878: 4876: 4875: 4871: 4869: 4868: 4867:Padillasaurus 4864: 4862: 4861: 4857: 4855: 4854: 4850: 4848: 4847: 4843: 4841: 4840: 4836: 4834: 4833: 4832:Desmatochelys 4829: 4827: 4826: 4822: 4820: 4819: 4815: 4814: 4812: 4810: 4806: 4800: 4797: 4796: 4794: 4792: 4788: 4782: 4781: 4780:Yaguarasaurus 4777: 4775: 4774: 4770: 4769: 4767: 4765: 4761: 4755: 4754: 4750: 4749: 4747: 4745: 4741: 4735: 4734: 4733: 4728: 4726: 4725: 4721: 4719: 4718: 4714: 4712: 4711: 4707: 4705: 4704: 4700: 4698: 4697: 4693: 4691: 4690: 4686: 4684: 4683: 4679: 4678: 4676: 4674: 4670: 4664: 4663: 4659: 4658: 4656: 4654: 4650: 4644: 4643: 4639: 4638: 4636: 4634:Gualanday Gp. 4632: 4626: 4625: 4621: 4619: 4618: 4614: 4612: 4611: 4607: 4605: 4604: 4600: 4598: 4597: 4593: 4591: 4590: 4586: 4584: 4583: 4579: 4577: 4576: 4572: 4570: 4569: 4565: 4563: 4562: 4558: 4556: 4555: 4551: 4549: 4548: 4544: 4542: 4541: 4540:Miocallicebus 4537: 4535: 4534: 4530: 4528: 4527: 4523: 4521: 4520: 4516: 4514: 4513: 4509: 4507: 4506: 4502: 4500: 4499: 4495: 4493: 4492: 4488: 4486: 4485: 4481: 4479: 4478: 4474: 4472: 4471: 4467: 4465: 4464: 4460: 4458: 4457: 4453: 4451: 4450: 4446: 4444: 4443: 4439: 4437: 4436: 4432: 4430: 4429: 4425: 4423: 4422: 4418: 4416: 4415: 4411: 4409: 4408: 4404: 4402: 4401: 4400:Anachlysictis 4397: 4396: 4394: 4392: 4388: 4382: 4381: 4377: 4375: 4374: 4370: 4369: 4367: 4365: 4361: 4358: 4354: 4344: 4341: 4339: 4336: 4335: 4333: 4331: 4327: 4321: 4318: 4316: 4313: 4312: 4310: 4308: 4304: 4298: 4295: 4293: 4290: 4288: 4285: 4283: 4280: 4278: 4275: 4274: 4272: 4270: 4266: 4258: 4255: 4254: 4253: 4250: 4248: 4245: 4244: 4242: 4240: 4236: 4230: 4227: 4225: 4222: 4220: 4217: 4213: 4210: 4209: 4208: 4205: 4203: 4200: 4198: 4197:Barzalosa Fm. 4195: 4194: 4192: 4190: 4186: 4180: 4177: 4173: 4170: 4169: 4168: 4165: 4164: 4162: 4160: 4156: 4153: 4151: 4148:fossiliferous 4143: 4136: 4129: 4118: 4109: 4100: 4091: 4082: 4073: 4064: 4057: 4040: 4037: 4035: 4032: 4031: 4029: 4025: 4019: 4016: 4014: 4011: 4009: 4006: 4004: 4001: 3999: 3996: 3995: 3993: 3989: 3986: 3980: 3971: 3966: 3964: 3959: 3957: 3952: 3951: 3948: 3932: 3929: 3928: 3925: 3918: 3914: 3904: 3899: 3896: 3889: 3882: 3852: 3851: 3847: 3845: 3844: 3843:Caipirasuchus 3840: 3839: 3837: 3835: 3831: 3825: 3824: 3820: 3818: 3817: 3816:Caryonosuchus 3813: 3811: 3810: 3806: 3805: 3803: 3801: 3797: 3791: 3790: 3786: 3784: 3783: 3782:Labidiosuchus 3779: 3777: 3776: 3772: 3771: 3768: 3765: 3763: 3759: 3753: 3752: 3748: 3745: 3744: 3743:Mariliasuchus 3740: 3738: 3737: 3733: 3730: 3729: 3728:Coronelsuchus 3725: 3724: 3722: 3720: 3716: 3710: 3709: 3705: 3703: 3702: 3701:Coronelsuchus 3698: 3696: 3695: 3694:Coringasuchus 3691: 3688: 3687: 3683: 3682: 3679: 3676: 3674: 3670: 3656: 3655: 3651: 3649: 3648: 3644: 3642: 3641: 3640:Campinasuchus 3637: 3636: 3634: 3632: 3628: 3622: 3621: 3617: 3615: 3614: 3610: 3608: 3607: 3606:Aplestosuchus 3603: 3601: 3600: 3599:Aphaurosuchus 3596: 3595: 3593: 3591: 3590:Baurusuchinae 3587: 3581: 3580: 3576: 3574: 3573: 3569: 3567: 3566: 3562: 3561: 3558: 3555: 3553: 3552:Baurusuchidae 3549: 3542: 3541: 3537: 3535: 3534: 3530: 3527: 3526: 3522: 3519: 3518: 3517:Comahuesuchus 3514: 3512: 3511: 3507: 3506: 3503: 3500: 3498: 3494: 3491: 3488: 3483: 3473: 3472: 3468: 3466: 3465: 3461: 3459: 3458: 3454: 3452: 3451: 3447: 3446: 3444: 3441: 3437: 3431: 3430: 3429:Uruguaysuchus 3426: 3424: 3423: 3419: 3416: 3415: 3411: 3409: 3408: 3404: 3401: 3400: 3399:Comahuesuchus 3396: 3394: 3393: 3392:Araripesuchus 3389: 3386: 3385: 3381: 3380: 3377: 3373: 3370: 3367: 3362: 3358: 3354: 3349: 3345: 3336: 3332: 3329: 3323: 3317: 3292: 3291: 3290: 3289: 3287: 3285: 3281: 3277: 3263: 3262: 3261:Uberabasuchus 3258: 3256: 3255: 3251: 3249: 3248: 3244: 3242: 3241: 3237: 3235: 3234: 3230: 3228: 3227: 3223: 3221: 3220: 3216: 3214: 3213: 3209: 3207: 3206: 3202: 3200: 3199: 3195: 3193: 3192: 3188: 3186: 3185: 3181: 3179: 3178: 3177:Barrosasuchus 3174: 3172: 3171: 3170:Barcinosuchus 3167: 3165: 3164: 3160: 3159: 3157: 3155: 3154:Peirosaurinae 3151: 3145: 3144: 3140: 3138: 3137: 3133: 3131: 3130: 3126: 3124: 3123: 3119: 3117: 3116: 3112: 3110: 3109: 3105: 3104: 3102: 3100: 3096: 3089: 3088: 3084: 3082: 3081: 3080:Miadanasuchus 3077: 3074: 3073: 3069: 3067: 3066: 3062: 3061: 3058: 3055: 3053: 3052:Peirosauridae 3049: 3043: 3042: 3038: 3036: 3035: 3031: 3029: 3028: 3024: 3022: 3021: 3017: 3015: 3014: 3010: 3008: 3007: 3003: 3001: 3000: 2996: 2994: 2993: 2989: 2987: 2986: 2982: 2980: 2979: 2975: 2973: 2972: 2968: 2966: 2965: 2961: 2959: 2958: 2954: 2953: 2951: 2949: 2945: 2939: 2938: 2934: 2932: 2931: 2927: 2926: 2924: 2922: 2918: 2912: 2911: 2907: 2904: 2903: 2899: 2897: 2896: 2892: 2890: 2889: 2885: 2884: 2881: 2878: 2875: 2870: 2859: 2858: 2854: 2853: 2850: 2846: 2843: 2840: 2835: 2831: 2827: 2822: 2818: 2797: 2796: 2795: 2794: 2792: 2790: 2786: 2778: 2774: 2773: 2772: 2771: 2768: 2765: 2763: 2759: 2755: 2747: 2743: 2740: 2738: 2734: 2732: 2728: 2726: 2722: 2720: 2716: 2715: 2710: 2706: 2702: 2695: 2690: 2688: 2683: 2681: 2676: 2675: 2672: 2666: 2664: 2660: 2655: 2654: 2650: 2639: 2634: 2625: 2621: 2615: 2611: 2604: 2599: 2596:(1–2): 12–40. 2595: 2591: 2584: 2579: 2575: 2571: 2566: 2561: 2557: 2553: 2549: 2545: 2540: 2535: 2530: 2525: 2520: 2516: 2512: 2508: 2503: 2499: 2495: 2491: 2487: 2483: 2479: 2475: 2471: 2467: 2462: 2458: 2454: 2450: 2448: 2441: 2437: 2433: 2426: 2421: 2417: 2413: 2406: 2404: 2397: 2393: 2389: 2384: 2380: 2376: 2371: 2367: 2362: 2358: 2354: 2350: 2346: 2339: 2337: 2329: 2325: 2321: 2317: 2313: 2309: 2305: 2301: 2296: 2292: 2288: 2284: 2279: 2274: 2269: 2265: 2261: 2257: 2252: 2251: 2247: 2240:, p. 38) 2239: 2233: 2230: 2227:, p. 20) 2226: 2220: 2217: 2214:, p. 29) 2213: 2207: 2204: 2200: 2194: 2191: 2188:, p. 27) 2187: 2181: 2178: 2174: 2168: 2165: 2161: 2155: 2152: 2148: 2142: 2139: 2135: 2129: 2126: 2122: 2116: 2113: 2109: 2103: 2100: 2096: 2090: 2087: 2083: 2077: 2074: 2070: 2064: 2061: 2058:, p. 20) 2057: 2051: 2048: 2045:, p. 17) 2044: 2043:Langston 1965 2038: 2035: 2032:, p. 16) 2031: 2030:Langston 1965 2025: 2022: 2019:, p. 17) 2018: 2017:Langston 1965 2012: 2009: 2006:, p. 17) 2005: 2004:Langston 1965 1999: 1996: 1993:, p. 17) 1992: 1991:Langston 1965 1986: 1983: 1979: 1978:Langston 1965 1973: 1970: 1966: 1960: 1957: 1953: 1947: 1944: 1941:, p. 38) 1940: 1935: 1932: 1929: 1924: 1921: 1918: 1913: 1910: 1907:, p. 11) 1906: 1900: 1897: 1893: 1887: 1884: 1881:, p. 26) 1880: 1874: 1871: 1868:, p. 20) 1867: 1861: 1858: 1854: 1848: 1845: 1842:, p. 16) 1841: 1840:Langston 1965 1835: 1832: 1829:, p. 14) 1828: 1827:Langston 1965 1822: 1819: 1816:, p. 14) 1815: 1814:Langston 1965 1809: 1806: 1802: 1801:Langston 1965 1796: 1793: 1789: 1788:Langston 1965 1783: 1780: 1776: 1770: 1767: 1760: 1753: 1752: 1744: 1743: 1735: 1734: 1726: 1725: 1717: 1716: 1708: 1707: 1699: 1698: 1690: 1689: 1681: 1680: 1677: 1676: 1675: 1668: 1667: 1664: 1663: 1660: 1659: 1658: 1651: 1650: 1647: 1646: 1643: 1642: 1641: 1634: 1633: 1630: 1629: 1626: 1621: 1620: 1614: 1613: 1610: 1609: 1601: 1600: 1597: 1596: 1595: 1588: 1587: 1584: 1583: 1580: 1579: 1578: 1571: 1570: 1564: 1563: 1557: 1556: 1553: 1552: 1549: 1548: 1547: 1540: 1539: 1533: 1532: 1529: 1528: 1525: 1524: 1523: 1516: 1515: 1509: 1506: 1505: 1502: 1501: 1498: 1497: 1496: 1489: 1488:Iberosuchidae 1486: 1485: 1482: 1481: 1478: 1477: 1476: 1469: 1468:Bergisuchidae 1466: 1465: 1459: 1458: 1455: 1454: 1446: 1445: 1437: 1436: 1433: 1432: 1431: 1424: 1423: 1420: 1419: 1416: 1415: 1414: 1407: 1406: 1400: 1399: 1396: 1395: 1387: 1386: 1383: 1382: 1381: 1374: 1373: 1370: 1369: 1366: 1365: 1364: 1357: 1356: 1350: 1349: 1343: 1342:Baurusuchidae 1340: 1339: 1336: 1335: 1332: 1331: 1330: 1323: 1322: 1316: 1315: 1312: 1311: 1308: 1307: 1306: 1299: 1298: 1292: 1289: 1288: 1285: 1284: 1276: 1275: 1272: 1271: 1270: 1263: 1262: 1259: 1258: 1255: 1254: 1253: 1246: 1245: 1239: 1238: 1232: 1231: 1227: 1225: 1221: 1217: 1216: 1211: 1207: 1203: 1199: 1195: 1191: 1187: 1183: 1176: 1174: 1172: 1168: 1164: 1160: 1159:Astrapotheria 1156: 1155: 1150: 1146: 1141: 1139: 1135: 1134: 1133:Pericotoxodon 1129: 1128: 1123: 1122: 1117: 1113: 1109: 1108:phorusrhacids 1105: 1101: 1100: 1095: 1094: 1089: 1088:sparassodonts 1084: 1082: 1081:Komodo dragon 1078: 1077:Langston 1965 1074: 1073: 1068: 1064: 1060: 1056: 1052: 1048: 1044: 1040: 1035: 1031: 1024: 1019: 1012: 1007: 1003: 999: 995: 991: 988: 987: 983: 978: 974: 971: 970: 967: 963: 958: 954: 950: 947: 944: 941: 938: 935: 932: 929: 926: 923: 919: 914: 910: 906: 902: 898: 894: 890: 886: 883: 882: 878: 874: 869: 865: 861: 857: 853: 850: 849: 845: 840: 837: 834: 830: 827: 824: 821: 818: 815: 812: 808: 805: 802: 799: 796: 792: 789: 786: 781: 777: 773: 770: 769: 768: 766: 758: 756: 750: 748: 746: 742: 738: 734: 729: 727: 723: 719: 715: 711: 710: 705: 704:S. querejazus 701: 697: 693: 689: 685: 684:S. querejazus 681: 677: 673: 669: 668: 662: 660: 656: 652: 648: 643: 641: 637: 633: 629: 625: 620: 618: 614: 609: 607: 603: 599: 595: 591: 587: 583: 581: 576: 572: 568: 564: 563: 558: 557: 552: 551: 546: 545: 540: 536: 532: 528: 524: 520: 516: 512: 508: 504: 500: 496: 488: 483: 476: 474: 472: 468: 464: 462: 457: 453: 450: 446: 442: 438: 434: 431: 427: 424: 420: 416: 412: 411: 398: 397: 396:lapsus calami 391: 388: 383: 381: 377: 376: 374: 371: 367: 362: 356: 354: 345: 342: 341:Binomial name 338: 334: 333: 327: 324: 323: 319: 313: 312: 305: 302: 301: 298: 292: 289: 288: 285: 279: 276: 273: 272: 269: 263: 260: 257: 256: 253: 247: 244: 241: 240: 237: 234: 231: 228: 227: 224: 221: 218: 215: 214: 211: 208: 205: 202: 201: 198: 195: 192: 189: 188: 185: 182: 179: 178: 175: 172: 169: 168: 165: 162: 159: 158: 155: 152: 149: 148: 143: 138: 134: 129: 125: 120: 106: 101: 96: 91: 86: 81: 76: 71: 66: 61: 56: 51: 45: 36: 32: 26: 22: 19: 5051: 4992:Floresta Fm. 4979: 4972: 4967:Holoptychius 4965: 4959:Bothriolepis 4957: 4956: 4949: 4942: 4899:Arcabuco Fm. 4886: 4879: 4872: 4865: 4860:Muiscasaurus 4858: 4851: 4844: 4837: 4830: 4823: 4818:Acostasaurus 4816: 4778: 4771: 4751: 4730: 4729: 4722: 4715: 4708: 4703:Cerrejonemys 4701: 4694: 4687: 4680: 4673:Cerrejón Fm. 4660: 4640: 4622: 4615: 4608: 4601: 4594: 4587: 4580: 4573: 4566: 4559: 4552: 4547:Miocochilius 4545: 4538: 4531: 4524: 4518: 4517: 4510: 4505:Huilatherium 4503: 4498:Hondadelphys 4496: 4489: 4482: 4475: 4468: 4461: 4454: 4447: 4440: 4433: 4428:Cebupithecia 4426: 4419: 4412: 4405: 4398: 4378: 4373:Gomphotheres 4371: 4343:Floresta Fm. 4315:Arcabuco Fm. 4252:Cerrejón Fm. 4027:19th century 3884: 3848: 3841: 3823:Sphagesaurus 3821: 3814: 3807: 3787: 3780: 3773: 3749: 3741: 3734: 3726: 3719:Notosuchidae 3706: 3699: 3692: 3684: 3673:Sphagesauria 3652: 3645: 3638: 3618: 3611: 3604: 3597: 3577: 3570: 3563: 3538: 3531: 3523: 3515: 3508: 3469: 3464:Malawisuchus 3462: 3455: 3448: 3427: 3420: 3412: 3407:Libycosuchus 3405: 3397: 3390: 3382: 3319: 3259: 3252: 3245: 3238: 3233:Patagosuchus 3231: 3224: 3217: 3210: 3205:Hamadasuchus 3203: 3196: 3189: 3182: 3175: 3168: 3163:Antaeusuchus 3161: 3141: 3134: 3127: 3122:Epoidesuchus 3120: 3115:Caririsuchus 3113: 3106: 3099:Pepesuchinae 3085: 3078: 3072:Fortignathus 3070: 3065:Amargasuchus 3063: 3039: 3032: 3027:Sahitisuchus 3025: 3018: 3011: 3005: 3004: 2997: 2990: 2983: 2976: 2969: 2964:Barinasuchus 2962: 2955: 2935: 2928: 2908: 2900: 2893: 2886: 2855: 2777:Pseudosuchia 2762:Pseudosuchia 2741: 2662: 2658: 2624:the original 2609: 2593: 2589: 2547: 2543: 2514: 2510: 2473: 2469: 2465: 2456: 2452: 2446: 2435: 2431: 2415: 2411: 2402: 2391: 2387: 2378: 2374: 2365: 2348: 2344: 2335: 2310:(1): 20–27. 2307: 2303: 2299: 2293:: 1663–1666. 2290: 2286: 2282: 2263: 2259: 2248:Bibliography 2232: 2219: 2206: 2193: 2180: 2167: 2154: 2141: 2128: 2115: 2102: 2089: 2076: 2063: 2050: 2037: 2024: 2011: 1998: 1985: 1972: 1959: 1946: 1934: 1923: 1912: 1899: 1886: 1873: 1860: 1847: 1834: 1821: 1808: 1803:, p. 6) 1795: 1782: 1769: 1672: 1671: 1655: 1654: 1639: 1638: 1637: 1624: 1592: 1591: 1575: 1574: 1546:Barinasuchus 1544: 1543: 1520: 1519: 1493: 1492: 1473: 1472: 1428: 1427: 1411: 1410: 1378: 1377: 1361: 1360: 1327: 1326: 1303: 1302: 1291:Sebecosuchia 1290: 1269:Sphagesaurus 1267: 1266: 1250: 1249: 1213: 1210:Barinasuchus 1209: 1205: 1201: 1193: 1186:Sebecosuchia 1181: 1180: 1171:Notohippidae 1163:Leontiniidae 1154:Barinasuchus 1152: 1148: 1142: 1131: 1127:Huilatherium 1125: 1119: 1097: 1091: 1085: 1070: 1063:mekosuchines 1051:crocodilians 1043:baurusuchids 1039:peirosaurids 1033: 1029: 1027: 1022: 1013:Paleobiology 1005: 1001: 997: 989: 977:L. huilensis 976: 972: 957:L. huilensis 956: 948: 942: 936: 930: 924: 904: 896: 884: 851: 838: 832: 828: 822: 816: 810: 806: 800: 790: 784: 779: 771: 764: 762: 754: 744: 740: 736: 730: 725: 718:Barinasuchus 717: 713: 707: 706:was renamed 703: 699: 696:S. huilensis 695: 683: 680:S. huilensis 679: 675: 667:Barinasuchus 665: 663: 646: 644: 640:S. huilensis 639: 627: 621: 617:S. huilensis 616: 610: 597: 593: 578: 560: 554: 548: 542: 538: 498: 492: 486: 471:L. huilensis 470: 467:type species 465:. Thus, the 459: 455: 409: 408: 407: 394: 389: 378: 352: 347: 331: 330: 310: 309: 274: 268:Sebecosuchia 258: 242: 229: 216: 210:Pseudosuchia 203: 190: 24: 18: 4951:Asterolepis 4853:Leyvachelys 4846:Leivanectes 4839:Kronosaurus 4764:Hondita Fm. 4596:Purussaurus 4589:Potamosiren 4568:Neotamandua 4561:Mourasuchus 4491:Hoazinoides 4477:Gryposuchus 4435:Colombophis 4421:Boreostemma 4414:Balanerodus 4380:Megatherium 4364:Pleistocene 4287:Hondita Fm. 4207:Honda Group 4159:Pleistocene 4013:Hoffstetter 3984:researchers 3736:Llanosuchus 3654:Wargosuchus 3613:Baurusuchus 3525:Eremosuchus 3497:Baurusuchia 3384:Anatosuchus 3254:Rukwasuchus 3240:Peirosaurus 3143:Roxochampsa 3041:Zulmasuchus 2992:Iberosuchus 2978:Bretesuchus 2971:Bergisuchus 2957:Ayllusuchus 2930:Kaprosuchus 2888:Baharijodon 2857:Microsuchus 2839:Ziphosuchia 2826:Ziphosuchia 2789:Ziphosuchia 2701:Ziphosuchia 2657:Photo of a 2550:(1): 3–21. 2534:11336/69518 2453:Ameghiniana 2388:Gryposuchus 2186:Busbey 1986 2056:Busbey 1986 1879:Busbey 1986 1866:Busbey 1986 1674:Zulmasuchus 1594:Bretesuchus 1577:Ayllusuchus 1495:Iberosuchus 1475:Bergisuchus 1215:Bretesuchus 1206:Zulmasuchus 1167:Adianthidae 1104:Argot (2004 1072:Paleosuchus 949:IGM 184 165 937:IGM 250 427 931:IGM 250 541 925:IGM 184 427 852:TMM 41658-8 709:Zulmasuchus 523:Honda Group 430:notosuchian 197:Archosauria 42:13–12  31:Mid Miocene 5098:Categories 5052:Langstonia 5000:Trilobites 4981:Strepsodus 4974:Pterygotus 4696:Carbonemys 4653:Bogotá Fm. 4575:Nuciruptor 4554:Mohanamico 4519:Langstonia 4512:Lagonimico 4269:Cretaceous 4247:Bogotá Fm. 4167:Sabana Fm. 3991:After 1900 3789:Yacarerani 3751:Notosuchus 3471:Pakasuchus 3450:Candidodon 3422:Simosuchus 3366:Notosuchia 3353:Notosuchia 3293:see below↓ 3284:Notosuchia 3247:Pepesuchus 3219:Lomasuchus 3212:Kinesuchus 3136:Pepesuchus 3020:Ogresuchus 3013:Lorosuchus 3006:Langstonia 2910:Wanosuchus 2798:see below↓ 2746:Metasuchia 2731:Sauropsida 2663:Langstonia 2565:11336/2663 2517:: S7–S36. 2418:: 127–144. 1761:References 1640:Langstonia 1522:Lorosuchus 1182:Langstonia 1149:Langstonia 1067:crocodiles 1047:Cretaceous 1030:Langstonia 1023:Langstonia 998:Langstonia 943:IGM 184378 889:premaxilla 885:IGM 250816 839:UCMP 44566 833:Langstonia 829:UCMP 44561 823:UCMP 44563 817:UCMP 40220 811:Langstonia 807:UCMP 40186 801:UCMP 44562 791:UCMP 41308 772:UCMP 37877 745:Langstonia 726:Langstonia 700:Langstonia 659:Ingeominas 624:premaxilla 567:dyrosaurid 517:overlying 499:Langstonia 493:The first 487:Langstonia 456:Langstonia 410:Langstonia 311:Langstonia 252:Notosuchia 25:Langstonia 4935:Cuche Fm. 4799:Ammonites 4753:Eonatator 4744:Oliní Gp. 4732:Titanoboa 4724:Puentemys 4617:Stirtonia 4449:Dukecynus 4391:Honda Gp. 4338:Cuche Fm. 4292:Oliní Gp. 4239:Paleogene 4224:Uitpa Fm. 4219:Jimol Fm. 4179:Soatá Fm. 3579:Pabwehshi 3129:Itasuchus 2999:Ilchunaia 2948:Sebecidae 2895:Doratodon 2717:Kingdom: 2351:: 19–42. 1508:Sebecidae 1363:Pabwehshi 1222:based on 1220:Cladogram 1177:Phylogeny 1138:megafauna 1099:Dukecynus 1061:and some 1049:and some 1006:huilensis 868:osteoderm 860:squamosal 776:symphysis 741:huilensis 733:Paleocene 688:Paleocene 672:Venezuela 651:vertebrae 636:osteoderm 590:Argentina 582:huilensis 519:sandstone 515:claystone 463:huilensis 449:Colombian 437:Sebecidae 382:huilensis 325:Species: 297:Sebecidae 160:Kingdom: 154:Eukaryota 5119:Laventan 5104:Sebecids 5067:Q5236876 5061:Wikidata 5020:Category 4809:Paja Fm. 4791:Hiló Fm. 4582:Patasola 4526:Lycopsis 4330:Devonian 4307:Jurassic 4297:Paja Fm. 4282:Hiló Fm. 4257:Cerrejón 4229:Ware Fm. 4212:La Venta 4039:Humboldt 2725:Chordata 2723:Phylum: 2719:Animalia 2574:15751319 2498:86565764 2381:: 1–127. 1190:Gondwana 1055:Cenozoic 973:MUSM 912 953:maxillar 913:diastema 575:holotype 445:Laventan 421:) is an 415:Langston 370:Synonyms 361:Langston 290:Family: 184:Reptilia 174:Chordata 170:Phylum: 164:Animalia 150:Domain: 35:Laventan 4533:Micodon 4189:Neogene 4008:Hettner 3982:Notable 3034:Sebecus 2874:Sebecia 2735:Order: 2729:Class: 2665:) tooth 2478:Bibcode 2438:: 1–25. 2403:Sebecus 2345:Zootaxa 2312:Bibcode 2283:Sebecus 1657:Sebecus 1194:Sebecus 1057:as the 1053:of the 1045:of the 1002:Sebecus 893:maxilla 737:Sebecus 714:Sebecus 692:Bolivia 676:Sebecus 598:Sebecus 580:Sebecus 571:dentary 495:fossils 461:Sebecus 441:Miocene 423:extinct 380:Sebecus 363:, 1965) 350:† 329:† 303:Genus: 284:Sebecia 180:Class: 111:↓ 5081:192480 5024:Images 4662:Etayoa 4172:Tibitó 4034:Cuvier 4018:Páramo 4003:Hammen 2616:  2590:Terrae 2572:  2496:  918:sacral 856:square 655:caudal 602:Eocene 586:Eocene 4925:Flora 4146:Major 3998:Etayo 2742:Clade 2627:(PDF) 2606:(PDF) 2586:(PDF) 2570:S2CID 2494:S2CID 2428:(PDF) 2408:(PDF) 2341:(PDF) 1145:Andes 879:(IGM) 864:jugal 632:jugal 426:genus 275:Clade 259:Clade 243:Clade 230:Clade 217:Clade 204:Clade 191:Clade 2775:see 2614:ISBN 2349:2871 1212:and 1204:and 1196:and 1169:and 1151:and 1124:and 1096:and 1041:and 1004:cf. 739:cf. 682:and 613:Peru 559:and 473:. 318:2007 50:PreꞒ 2560:hdl 2552:doi 2529:hdl 2519:doi 2515:163 2486:doi 2353:doi 2320:doi 2291:284 2268:doi 2264:140 1140:. 1118:as 728:. 690:of 686:of 670:of 588:of 497:of 469:is 428:of 5100:: 5078:: 5063:: 5022:• 2744:: 2661:(= 2612:. 2588:. 2568:. 2558:. 2548:20 2546:. 2527:. 2513:. 2509:. 2492:. 2484:. 2474:32 2472:. 2457:48 2455:. 2451:. 2430:. 2416:29 2414:. 2410:. 2379:52 2377:. 2347:. 2343:. 2318:. 2306:. 2262:. 2258:. 1226:. 1218:. 1165:, 1161:, 1090:, 1008:. 979:. 835:. 787:. 678:, 553:, 547:, 454:. 277:: 261:: 245:: 232:: 219:: 206:: 193:: 100:Pg 44:Ma 3969:e 3962:t 3955:v 3746:? 3731:? 3689:? 3543:? 3528:? 3520:? 3442:? 3417:? 3402:? 3387:? 3090:? 3075:? 2905:? 2860:? 2693:e 2686:t 2679:v 2594:9 2576:. 2562:: 2554:: 2537:. 2531:: 2521:: 2500:. 2488:: 2480:: 2436:3 2359:. 2355:: 2334:" 2326:. 2322:: 2314:: 2308:6 2276:. 2270:: 2236:( 2223:( 2210:( 2197:( 2184:( 2171:( 2158:( 2145:( 2132:( 2119:( 2106:( 2093:( 2080:( 2067:( 2054:( 2041:( 2028:( 2015:( 2002:( 1989:( 1976:( 1963:( 1950:( 1903:( 1890:( 1877:( 1864:( 1851:( 1838:( 1825:( 1812:( 1799:( 1790:) 1786:( 1777:) 1773:( 489:. 399:) 359:( 307:† 294:† 281:† 265:† 249:† 105:N 95:K 90:J 85:T 80:P 75:C 70:D 65:S 60:O 55:Ꞓ 39:~ 37:) 33:(

Index

Mid Miocene
Laventan
Ma
PreꞒ

O
S
D
C
P
T
J
K
Pg
N
Skeleton restoration
Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Reptilia
Archosauria
Pseudosuchia
Crocodylomorpha
Crocodyliformes
Notosuchia
Sebecosuchia
Sebecia
Sebecidae

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