4056:
1069:, using its pressure force to retain and drown his victims is not viable on land; these crocodylomorphs also had more elongated limbs, stiff back, ideal to hold the weight and give strength to run. Langston had already suggested that these teeth were indicative that this animal had more terrestrial habits than those of existing species and the Miocene contemporary crocodylians, and in the modern world only could be vaguely compared with the less aquatic habits of the caimans of the genus
2643:
1018:
142:
124:
3316:
482:
3888:
3335:
5016:
3881:
831:: The largest fragment dental crown reported of La Venta, locality V-4528. Reaches up to 20.5 mm. high, but its base is much wider than other teeth, reaching 23.8 mm. Apart from its size, three times that of the teeth of the holotype of the species, this tooth is characterized by a blunt, relatively straight edges and a more oval profile tranverasal fastened with small saws. It may not correspond to
3895:
4108:
4090:
3903:
5028:
3328:
4072:
4063:
4081:
4117:
4099:
4135:
3322:
4128:
1079:, pp. 134–135) Busbey suggested the fact that his skull was high implied that these animals seek not to make pressure during the bite indefinitely, but to produce cuts then back quickly so that the bleeding weakens their prey slowly up to devour, similar to what has been thought to be the case for the extant
920:
rib attached, with a wide, rectangular neural spine; it measured 10.7 cm high by 5.5 cm wide, while it is estimated that the sacral ribs should be 10 cm wide at its rear end. The other is an anterior caudal vertebrae, perhaps the first, with incomplete neural spine, with a broad neural
782:
specimen. Their teeth are quite compressed and has a first side facing forward tooth, the second is directed upwards and the third also slopes, while others are vertical; the first teeth have a wide separation between them (diastema). Symphysis extending beyond the sixth tooth, while the bone surface
1036:
with jaws and teeth even more flattened, although its general proportions are reminiscent of this species, which suggests that their skull would be higher and laterally flattened, in contrast to modern crocodilians, which generally have a horizontally flattened skull with conical teeth. Teeth are of
915:
in that fitted the fourth lower tooth that exceeded the 3.5 centimeters; has a narrow palate, lateral nasal aperture and without mild depression around. The dentary is high, with four sockets, two projecting forward and a third and fourth straight, the latter being the largest. The associated
1037:
the zyphodont kind, with very flattened sides, slightly curved back and with serrated edges with small denticles (between 5-6 denticles per millimeter in the teeth of this genus) and also without any grooves on its surface. This type of teeth appears in the close relatives of sebecids, the
2331:
Carvalho, Ismar de Souza; Antunes
Texeira, Vicente de Paula; Da Fonseca Ferraz, Mara Lúcia; Borges Ribeiro, Luiz Carlos; Martinelli, Agustín Guillermo; Macedo Neto, Francísco; Sertich, Joseph J. W.; Cardoso Cunha, Gabriel; Cardoso Cunha, Isabella; Fonseca Ferraz, Patrícia (2011).
1136:- indicate that the climate was warm and humid with heavy rainfall and with drought periods not extending beyond 3–4 months, consisting of watersheds, forests and certain adjacent areas of open grasslands where it could hunt down a large variety of species belonging to the
596:, it was also proportionally thinner, and had with more recurved teeth being laterally compressed. Additionally Langston referred a series of zyphodont teeth in the area found this species, with some teeth referred to as similar but classified generally to
2656:
854:: Includes a portion of the left anterior premaxillary, middle left maxillary fragment, a fragment of the left splenial bone, back of the left suspensory bone formed by parts of the quadratojugal, the
1157:
were likely apex predators in their environment, and as an effect they would be particularly susceptible to ecological changes that caused other lineages, particularly hoofed mammals (such as groups
809:: An elongated, slender and compressed laterally tooth, with a slight curvature, intermediate in size between 44561 and other teeth reported, with larger saws (3 per millimeter tooth; in
939:: A tooth of the Duke 106 locality. It is also incomplete, but its measurements (52 mm high by 30 mm wide) makes it the largest tooth reported for a sebecosuchian in Colombia.
778:, with six tooth sockets of which five have broken teeth, collected in 1945 in the locality V-4517. This element measures 189.0 millimeters in length, being 68% higher than the rate of
2542:
Prevosti, Francisco J.; Forasiepi, Analía; Zimicz, Natalia (2013). "The evolution of the
Cenozoic terrestrial mammalian predator guild in South America: competition or replacement?".
996:
in Brazil. Includes a full left premaxilla with a tooth preserved, and a piece of left maxilla with two alveoli. By having very flattened and spaced teeth looks remarkably similar to
4145:
955:
right tooth, measures 20 mm in height at the crown and 12 on the base, while its root has a length equivalent to 2.5 times the height of the crown. Unlike other teeth of
645:
Langston, along with the
Argentine paleontologist Zulma Gasparini, reported new remains of the species in a contribution on fossil crocodylomorphs of Colombia for the book
3967:
975:: A single tooth found in the area of Fitzcarrald Arch in the localities of Inuya and Mapuya in the Amazon region of Peru; its size and shape are indistinguishable from
895:, the front left tooth, part of the dorsal border of a rostrum and two vertebrae. The premaxillary measured 12.8 cm in length, is tall and thin compared to the
755:
Here is a list of specimens assigned to the species, with a brief description of these. They are grouped according to the institution in which they are preserved:
702:, in honor of its initial descriptor and his "(..)advances in knowledge of crocodylians of Tertiary Colombia in particular and South America in general", while
4372:
4127:
5113:
870:. These fragments were recovered in 1976 in the "Monkey Pod" locality of La Venta (Baraya Member of the Villavieja Formation) by Dr Lundelius Ernest Jr.
529:
named the "lechos de monos" (Monkey Beds) in 1945. Many of the remains discovered were then sent to the collections of the Museum of
Paleontology at the
5143:
1028:
Due to the fragmentary remains generally known for this family, it's only possible to describe some general aspects of the appearance and biology of
2601:
Salas-Gismondi, R.; Antoine, P.-O.; Baby, P.; Brusset, S.; Benammi, M.; Espurt, N.; de
Franceschi, D.; Pujos, F.; Tejada, J.; Urbina, M. (2007).
592:. Langston gave the taxon a new species considering that besides being larger, as the dentary fragment is 68% larger than the Argentine species,
5138:
3960:
2691:
933:: Tooth from the Duke 196 locality, is incomplete and worn, but is similar to the tooth UCMP 40186. It measures 39 mm high by 27 wide.
3930:
903:
in the premaxilla, from the edge of the second tooth to the fourth, which is particularly enlarged, being twice the third (reverse that in
4055:
622:
Arthur Busbey (1986) later brought new remains of the species also from the "Monkey Beds" of Huila in
Colombia, this time a fragment of
2617:
530:
5019:
3953:
541:, in which he made a detailed analysis of several fossils of crocodylomorphs in Colombia, including the remains of other species as
2400:
1065:. These forms were land animals that used their teeth to cut and tear their prey, because the traditional method of capture of the
1075:, which have a higher muzzle and a higher degree of ossification around the eye socket, so that they partially resemble sebecids.(
927:: A tooth found in the Duke 91 locality. The crown of this is twice as high as it is wide, reaching 36 mm. high and 17 wide.
5108:
649:(1997). In this case it would be new specimens of teeth, a somewhat fuller premaxilla, an anterior fragment of dentary, and two
533:(UCMP), where they are still housed. The description of the specimens did not come until 1965, when the American paleontologist
841:: The lower teeth reported by Langston (1965). Recovered in the locality V-4421, measuring just 6.1 mm. wide at the base.
657:, which also were unknown to this group. The specimens discovered on this occasion were deposited in the Geological Museum of
5133:
615:
the presence of a huge cranial portion (area of the snout), referring it to this genus, and although its age coincides with
4363:
4158:
5128:
4268:
2449:
Langston (Sebecia, Metasuchia) in the Middle
Paleocene of the Itaboraí Basin, Rio de Janeiro State, Southeastern Brazil"
2364:
Kay, R.; Madden, R. (1997). "Paleogeography and paleoecology". In Kay, R.; Madden, R.; Cifelli, R.; et al. (eds.).
735:(and where have been found fossils of other sebecids) has been classified as related to this species, under the name of
716:. Paolillo and Linares also clarified the status of the Peruvian fossil before assigned to the species, assigning it to
720:, although this does not rule out its presence in more southern areas; in the area of the Peruvian Amazon known as the
611:
Subsequently, Éric
Buffetaut and Robert Hoffstetter (1977) reported from the Ipururo Formation in the Amazon region of
4238:
2373:
Langston, W. (1965). "Fossil crocodilians from
Colombia and the Cenozoic History of the Crocodylia in South America".
2602:
1188:, a lineage of ziphodont terrestrial crocodylomorphs forming part of a separate lineage of the southern continent of
803:: Tooth of the Late Oligocene fauna found in Coyaima, locality V-4411. It measures 22.5 mm. height at the crown.
3887:
141:
4329:
4306:
3315:
825:: A fragment of the dental crown, found in the locality V-4528; it measured 16.3 mm at the base of the crown.
747:, suggesting that the latter is part of a lineage of sebecids with an extensive temporal and geographical record.
5123:
4188:
2684:
2256:"Evolution of South American mammalian predators (Borhyaenoidea): anatomical & palaeobiological implications"
965:
3334:
2444:
1429:
458:
was named in 2007 by
Alfredo Paolillo and Omar Linares for fossils originally described by Langston in 1965 as
3880:
3894:
993:
794:
605:
5043:
4831:
4716:
4038:
638:), which were the earliest known for this family. Busbey also ruled out the Peruvian fossil belonging to
3902:
664:
Moreover, Alfredo Paolillo and Omar Linares described a new genus of a large Venezuelan Miocene sebecid,
5075:
4462:
4390:
4276:
4206:
4201:
4002:
2776:
1143:
The disappearance of the system of large rivers of the Amazon lake system and the gradual uplift of the
584:; thus extending the time range of this genus and the family, hitherto known only from remains of the
543:
451:
340:
5023:
4672:
4251:
4071:
502:
721:
4924:
4916:
4772:
4319:
4098:
3327:
2677:
2507:"A new sebecid mesoeucrocodylian from the Rio Loro Formation (Palaeocene) of north-western Argentina"
2477:
2311:
1412:
526:
317:
731:
Finally, a premaxilla and maxilla fragment found in Itaboraí Basin in Brazil dating from the Middle
506:
4652:
4441:
4246:
4211:
4196:
4062:
2648:
2338:
gen. et sp. nov., a new Late Cretaceous baurusuchid (Crocodyliformes) from the Bauru Basin, Brazil"
724:
has been found a characteristically flattened tooth from the middle Miocene therefore assigned to
1017:
4999:
4991:
4950:
4906:
4898:
4887:
4616:
4469:
4342:
4314:
4080:
4012:
3630:
2569:
2493:
1173:) to die out, thus leading to extinction to the last notosuchian crocodilomorphs of the world.
1120:
418:
414:
369:
360:
136:
5080:
4178:
4017:
2582:
3321:
5118:
5103:
4798:
4790:
4763:
4623:
4602:
4581:
4455:
4406:
4286:
4281:
4116:
4089:
3190:
2920:
2613:
534:
5027:
4166:
3774:
2736:
2559:
2551:
2528:
2518:
2485:
2390:, and the South American gavials". In Kay, R.; Madden, R.; Cifelli, R.; et al. (eds.).
2352:
2319:
2267:
1114:
in its ecosystem. The fossils found of other species - big fishes, manatees, large browsing
1058:
131:
Restoration showing known elements, with remaining material filled in after related species
4934:
4824:
4709:
4681:
4609:
4337:
4223:
4218:
3833:
3808:
3646:
3619:
3564:
3539:
3532:
3486:
3439:
3413:
3225:
3197:
2936:
1379:
1328:
1304:
900:
654:
432:
235:
222:
2481:
2315:
1147:
caused major ecological changes in South America in the mid-Miocene. The last sebecids,
634:
bone, the suspensory and retroarticular joint in the jaw, and a fragment of bone scute (
485:
The Tatacoa Desert, in which has been discovered numerous Miocene fossils, including to
4873:
4808:
4743:
4688:
4641:
4483:
4296:
4291:
4228:
4107:
4033:
4007:
3997:
3849:
3799:
3761:
3707:
3685:
3571:
3509:
3456:
3183:
3107:
3086:
2984:
2901:
2581:
Riff, D.; de Souza, R.G.; Cidade, G.M.; Martinelli, A.G.; de Souza-Filho, J.P. (2012).
1251:
1115:
1110:
as in the southern part of the continent, these sebecids were probably the terrestrial
1107:
561:
510:
481:
2237:
1938:
5097:
4943:
4880:
4866:
4779:
4539:
4399:
3842:
3815:
3781:
3742:
3727:
3700:
3693:
3639:
3605:
3598:
3589:
3551:
3516:
3428:
3398:
3391:
3260:
3176:
3169:
3153:
3079:
3051:
2523:
2506:
2272:
2255:
1487:
1467:
1341:
1158:
1132:
1111:
1087:
1080:
395:
74:
2573:
2497:
1086:
Since in the fauna of La Venta, although there were some large mammalian predators (
866:, the left retroarticular processes (partial) right to the jaw and a fragment of an
4966:
4958:
4859:
4817:
4702:
4546:
4504:
4497:
4427:
3822:
3718:
3672:
3463:
3406:
3232:
3204:
3162:
3121:
3114:
3098:
3071:
3064:
3026:
2963:
2761:
2425:"Nuevos cocodrilos Sebecosuchia del Cenozoico Suramericano (Mesosuchia:Crocodylia)"
2323:
1545:
1268:
1185:
1170:
1162:
1153:
1126:
1083:, thereby adopting a strategy of stalking in areas where prey transit frequently.
666:
466:
267:
209:
123:
2392:
Vertebrate Paleontology in the Neotropics. The Miocene Fauna of La Venta, Colombia
2366:
Vertebrate Paleontology in the Neotropics. The Miocene Fauna of La Venta, Colombia
2302:(Crocodylia: Sebecosuchidae) from the Miocene of La Venta Formation of Colombia".
2489:
2424:
2405:
and the iberoccitanian crocodile. A morphological and ultrastructural comparison"
959:, has a slight groove at the base of the tooth crown, on the inner side thereof.
4852:
4845:
4838:
4595:
4588:
4567:
4560:
4490:
4476:
4434:
4420:
4413:
4379:
4256:
3735:
3653:
3612:
3524:
3496:
3383:
3253:
3239:
3142:
3040:
2991:
2977:
2970:
2956:
2929:
2887:
2856:
2838:
2825:
2788:
2700:
1673:
1593:
1576:
1494:
1474:
1214:
1166:
1071:
1062:
1050:
1042:
1038:
708:
555:
549:
522:
49:
5066:
2464:
Pol, D.; Leardi, J.M.; Lecuona, A.; Krause, M. (2012). "Postcranial anatomy of
4980:
4973:
4695:
4574:
4553:
4511:
3788:
3750:
3470:
3449:
3421:
3365:
3352:
3283:
3246:
3218:
3211:
3135:
3019:
3012:
2909:
2745:
2730:
2638:
2555:
2356:
1521:
1046:
888:
876:
658:
623:
566:
501:
were discovered in the province of Huila in Colombia by the Spanish geologist
429:
251:
94:
59:
887:: Found in the Duke 57 locality in Villavieja Formation; consists of a right
4752:
4731:
4723:
4448:
4134:
3578:
3128:
2998:
2947:
2894:
1507:
1362:
1219:
1197:
1137:
1098:
1066:
867:
859:
797:, Mugrosa Formation). It measures 14.2 millimeters in diameter at the base.
775:
732:
687:
671:
635:
589:
518:
514:
436:
296:
196:
153:
99:
43:
4171:
3945:
2610:
4th European Meeting on the Palaeontology and Stratigraphy of Latin America
2333:
2623:
1200:
have consistently shown that these forms a clade within Sebecidae next to
5060:
4525:
2724:
1189:
1092:
1054:
912:
793:: Base of a tooth of the Upper Eocene, from the V-4620 town (Tama fauna,
650:
574:
448:
444:
173:
89:
84:
69:
64:
54:
34:
1102:), neither of them equated in size to the contemporary crocodylomorphs,
647:
Vertebrate Paleontology of the Neotropics: The Miocene Fauna of La Venta
4532:
3033:
2873:
2533:
1656:
952:
951:: A tooth of the Duke 41 locality, that probably correspond to a first
945:: Tooth from Duke 40 locality. It measures 30 mm high by 17 wide.
911:, p. 114) After the fourth premaxillary tooth there is a space or
892:
691:
579:
570:
460:
440:
422:
379:
283:
183:
104:
79:
30:
2564:
819:: A large tooth, locality V.4523, measures about 42.4 mm height.
505:, during the expeditions in the region by the American paleontologist
4661:
2718:
2603:"Middle Miocene crocodiles from the Fitzcarrald Arch, Amazonian Peru"
917:
855:
674:. In their article they considered that the traits of two species of
642:, based on its unique anatomy, less spaced teeth and a wider nose.
601:
585:
494:
163:
5037:
1032:. As mentioned above, the fossil type of the species is larger than
694:, and their age justified the creation of separate genres for both:
2583:"Crocodilomorfos. A maior diversidade de répteis fósseis do Brasil"
899:
being covered with fine grooves on its outer surface. It has three
2281:
Buffetaut, E.; Hoffstetter, R. (1977). "Découverte du Crocodilien
1144:
1016:
964:
Department of Vertebrate Paleontology, Museum of Natural History,
863:
631:
480:
425:
992:: consists of material of sebecid found in Itaboraí Basin in the
2159:
1951:
612:
5041:
3949:
3919:
3350:
2823:
2711:
2673:
1106:, p. 513), and given the absence of large predatory birds
712:, although some authors still maintain both species within
630:, and equally thin. He also described the first remains of the
2669:
774:: The front part of a right dentary preserving the mandibular
1192:. As noted earlier, this species formerly part of the genus
4133:
4126:
2468:(Crocodyliformes, Sebecidae) from the Eocene of Patagonia".
2399:
Legasa, O.; Buscalioni, Ángela D.; Gasparini, Zulma (1993).
2375:
University of California Publications of Geological Sciences
2211:
619:, distinguished it through it being larger and more robust.
2443:
Pinheiro, A.E.P.; Campos, D.A.; Bergqvist, L.P. (2011).
763:
Not all materials from this collection were assigned to
2224:
2172:
1964:
1927:
767:
but are included because of their similar morphology.
759:
Museum of Paleontology at the University of California
577:
of a new species a fragmentary taxon, which he called
1852:
5050:
4990:
4933:
4915:
4897:
4807:
4789:
4762:
4742:
4671:
4651:
4633:
4389:
4362:
4355:
4328:
4305:
4267:
4237:
4187:
4157:
4144:
4026:
3990:
3981:
3832:
3798:
3760:
3717:
3671:
3629:
3588:
3550:
3495:
3484:
3438:
3363:
3282:
3152:
3097:
3050:
2946:
2919:
2871:
2836:
2787:
2760:
2386:Langston, W.; Gasparini, Z. (1997). "Crocodilians,
921:canal, and measures 2.8 cm wide and 2.1 high.
2394:. Smithsonian Institution Press. pp. 113–154.
2368:. Smithsonian Institution Press. pp. 520–550.
2287:Comptes Rendus de l'Académie des Sciences de Paris
661:(previous name of Colombian Geological Service).
569:). As for the sebecid material he designated the
2146:
2133:
2120:
2107:
2094:
2081:
2068:
1891:
908:
600:sp., which do not come from the Miocene but the
1904:
1774:
907:), while the first two teeth should be small.(
513:at the locality V-4517, characterized by gray
447:land-mammal age), in the "Monkey Beds" of the
3961:
2685:
8:
2423:Paolillo, Alfredo; Linares, Omar J. (2007).
916:vertebrae are an amphycoelian sacral with a
5038:
4359:
4154:
3987:
3968:
3954:
3946:
3916:
3766:
3677:
3556:
3501:
3492:
3375:
3371:
3360:
3347:
3279:
3056:
2879:
2848:
2844:
2833:
2820:
2766:
2757:
2708:
2692:
2678:
2670:
2608:. In Díaz-Martínez, E; Rábano, I. (eds.).
984:Museu de Ciências da Terra, Rio de Janeiro
743:, whose main features are very similar to
122:
20:
2563:
2532:
2522:
2511:Zoological Journal of the Linnean Society
2271:
2260:Zoological Journal of the Linnean Society
2198:
1916:
1223:
509:. They were found in the area called the
2042:
2029:
2016:
2003:
1990:
1977:
1839:
1826:
1813:
1800:
1787:
1076:
1766:
2185:
2055:
1878:
1865:
2505:Pol, Diego; Powell, Jaime E. (2011).
2225:Prevosti, Forasiepi & Zimicz 2013
2173:Pinheiro, Campos & Bergqvist 2011
1965:Pinheiro, Campos & Bergqvist 2011
1103:
521:sediments. These have been named the
7:
2298:Busbey, A. (1986). "New material of
2285:dans le Miocène du Pérou oriental".
1000:, being provisionally classified as
1618:
1568:
1561:
1537:
1513:
1463:
1404:
1354:
1347:
1320:
1296:
1243:
1236:
1229:
2470:Journal of Vertebrate Paleontology
2304:Journal of Vertebrate Paleontology
531:University of California, Berkeley
14:
5114:Miocene reptiles of South America
539:Fossil Crocodylians from Colombia
5144:Prehistoric pseudosuchian genera
5026:
5015:
5014:
4115:
4106:
4097:
4088:
4079:
4070:
4061:
4054:
3901:
3893:
3886:
3879:
3333:
3326:
3320:
3314:
2641:
2524:10.1111/j.1096-3642.2011.00714.x
2273:10.1111/j.1096-3642.2004.00110.x
1853:Buffetaut & Hoffstetter 1977
393:Langston & Gasparini, 1997 (
140:
2412:Studia Geologica Salmanticensia
2544:Journal of Mammalian Evolution
2324:10.1080/02724634.1986.10011595
813:typically are 4 to 6 mm).
417:", in honor of paleontologist
1:
5139:Fossil taxa described in 1965
2147:Langston & Gasparini 1997
2134:Langston & Gasparini 1997
2121:Langston & Gasparini 1997
2108:Langston & Gasparini 1997
2095:Langston & Gasparini 1997
2082:Langston & Gasparini 1997
2069:Langston & Gasparini 1997
1892:Langston & Gasparini 1997
909:Langston & Gasparini 1997
525:, of the La Venta fauna, the
2490:10.1080/02724634.2012.646833
1208:, not so closely related to
3931:Terrestrial crocodylomorphs
1905:Paolillo & Linares 2007
1775:Paolillo & Linares 2007
877:Colombian Geological Survey
846:Texas Memorial Museum (TMM)
5160:
2459:(4, Supplemento): R91–R92.
2160:Salas-Gismondi et al. 2007
1952:Salas-Gismondi et al. 2007
16:Extinct species of reptile
5012:
4049:
3926:
3915:
3877:
3769:
3680:
3559:
3504:
3378:
3374:
3359:
3346:
3312:
3278:
3059:
2882:
2851:
2847:
2832:
2819:
2769:
2756:
2707:
2556:10.1007/s10914-011-9175-9
2432:Paleobiologia Neotropical
2357:10.11646/zootaxa.2871.1.2
1670:
1653:
1636:
1623:
1616:
1590:
1573:
1566:
1559:
1542:
1535:
1518:
1511:
1491:
1471:
1461:
1426:
1409:
1402:
1376:
1359:
1352:
1345:
1325:
1318:
1301:
1294:
1265:
1248:
1241:
1234:
966:Universidad de San Marcos
439:. It lived in the middle
375:
368:
346:
339:
137:Scientific classification
135:
130:
121:
23:
3976:Paleontology in Colombia
1430:Baurusuchus salgadoensis
537:published his monograph
316:Paolillo & Linares,
5109:Miocene crocodylomorphs
2401:"The serrated teeth of
2289:. Série D (in French).
1917:Pol & Powell (2011)
1224:Pol & Powell (2011)
1021:Life reconstruction of
994:state of Rio de Janeiro
858:bone, opisthotic and a
795:department of Santander
606:department of Santander
556:Gryposuchus colombianus
4717:Montrichardia aquatica
4138:
4131:
1184:is considered part of
1130:, and some grazers as
1025:
562:Charactosuchus fieldsi
490:
390:Sebecosuchus huilensis
5134:Honda Group, Colombia
5076:Paleobiology Database
4463:Galbula hylochoreutes
4137:
4130:
2447:Sebecus cf. huilensis
2300:Sebecus cf. huilensis
2199:Kay & Madden 1997
1939:Carvalho et al. (2011
1198:phylogenetic analyzes
1093:Lycopsis longirostrus
1020:
765:Langstonia huilensis
544:Purussaurus neivensis
484:
443:(specifically in the
4773:Pachyrhizodus etayoi
2336:Campinasuchus dinizi
2238:Carvalho et al. 2011
1855:, Figure 1; p. 1664)
1413:Baurusuchus pachecoi
783:is smoother than in
698:was given the genus
527:geological formation
477:Discovery and naming
452:Villavieja Formation
353:Langstonia huilensis
5129:Fossils of Colombia
4442:Dracaena colombiana
4150:stratigraphic units
2649:Paleontology portal
2482:2012JVPal..32..328P
2466:Sebecus icaeorhinus
2316:1986JVPal...6...20B
2201:, pp. 534–535)
2136:, pp. 116–117)
2123:, pp. 116–117)
2110:, pp. 116–117)
1034:Sebecus icaeorhinus
875:Museum of Geology,
862:, a left part of a
780:Sebecus icaeorhinus
4888:Stenorhynchosaurus
4470:Granastrapotherium
4139:
4132:
3921:Related categories
3631:Pissarrachampsinae
2254:Argot, C. (2004).
1121:Granastrapotherium
1026:
573:UCMP 37877 as the
550:Mourasuchus atopus
491:
419:Wann Langston, Jr.
5091:
5090:
5044:Taxon identifiers
5035:
5034:
5008:
5007:
4624:Xenastrapotherium
4603:Saimiri annectens
4456:Eunectes stirtoni
4407:Aramus paludigrus
4351:
4350:
4047:
4046:
3943:
3942:
3939:
3938:
3911:
3910:
3875:
3874:
3871:
3870:
3867:
3866:
3863:
3862:
3859:
3858:
3667:
3666:
3663:
3662:
3480:
3479:
3342:
3341:
3310:
3309:
3306:
3305:
3302:
3301:
3274:
3273:
3270:
3269:
3191:Colhuehuapisuchus
2921:Mahajangasuchidae
2867:
2866:
2815:
2814:
2811:
2810:
2807:
2806:
2659:Sebecus huilensis
2592:(in Portuguese).
2445:"The presence of
1980:, pp. 15–16)
1928:Pol et al. (2012)
1757:
1756:
1748:
1747:
1739:
1738:
1730:
1729:
1721:
1720:
1712:
1711:
1703:
1702:
1694:
1693:
1685:
1684:
1625:Form of Lumbrera
1605:
1604:
1450:
1449:
1441:
1440:
1391:
1390:
1280:
1279:
990:MCT 1795-R/1796-R
653:: a sacral and a
565:(plus a possible
535:Wann Langston Jr.
503:José Royo y Gómez
406:
405:
400:
386:
332:L. huilensis
320:
5151:
5124:Neogene Colombia
5084:
5083:
5071:
5070:
5069:
5039:
5030:
5018:
5017:
4360:
4155:
4119:
4110:
4101:
4092:
4083:
4074:
4065:
4058:
3988:
3970:
3963:
3956:
3947:
3917:
3906:
3905:
3898:
3897:
3891:
3890:
3883:
3775:Adamantinasuchus
3767:
3678:
3557:
3502:
3493:
3376:
3372:
3361:
3348:
3337:
3331:
3330:
3324:
3318:
3280:
3057:
2880:
2849:
2845:
2834:
2821:
2767:
2758:
2751:
2750:
2737:Mesoeucrocodylia
2709:
2694:
2687:
2680:
2671:
2651:
2646:
2645:
2644:
2630:
2628:
2622:. Archived from
2607:
2597:
2587:
2577:
2567:
2538:
2536:
2526:
2501:
2460:
2439:
2429:
2419:
2409:
2395:
2382:
2369:
2360:
2342:
2327:
2294:
2277:
2275:
2241:
2234:
2228:
2221:
2215:
2212:Riff et al. 2012
2208:
2202:
2195:
2189:
2182:
2176:
2169:
2163:
2156:
2150:
2143:
2137:
2130:
2124:
2117:
2111:
2104:
2098:
2091:
2085:
2078:
2072:
2065:
2059:
2052:
2046:
2039:
2033:
2026:
2020:
2013:
2007:
2000:
1994:
1987:
1981:
1974:
1968:
1961:
1955:
1948:
1942:
1936:
1930:
1925:
1919:
1914:
1908:
1901:
1895:
1888:
1882:
1875:
1869:
1862:
1856:
1849:
1843:
1836:
1830:
1823:
1817:
1810:
1804:
1797:
1791:
1784:
1778:
1771:
1619:
1569:
1562:
1538:
1514:
1464:
1405:
1355:
1348:
1321:
1297:
1244:
1237:
1230:
891:and part of the
722:Fitzcarrald Arch
392:
384:
355:
351:
315:
308:
295:
282:
266:
250:
145:
144:
126:
116:
46:
29:Temporal range:
21:
5159:
5158:
5154:
5153:
5152:
5150:
5149:
5148:
5094:
5093:
5092:
5087:
5079:
5074:
5065:
5064:
5059:
5046:
5036:
5031:
5004:
4986:
4929:
4911:
4907:Dinosaur tracks
4893:
4825:Callawayasaurus
4803:
4785:
4758:
4738:
4710:Cerrejonisuchus
4682:Acherontisuchus
4667:
4647:
4629:
4610:Saimiri fieldsi
4385:
4356:Notable fossils
4347:
4324:
4301:
4277:La Frontera Fm.
4263:
4233:
4202:Castilletes Fm.
4183:
4149:
4147:
4140:
4125:
4124:
4123:
4122:
4121:
4120:
4112:
4111:
4103:
4102:
4094:
4093:
4085:
4084:
4076:
4075:
4067:
4066:
4043:
4022:
3983:
3977:
3974:
3944:
3935:
3922:
3907:
3900:
3892:
3885:
3855:
3834:Caipirasuchinae
3828:
3809:Armadillosuchus
3794:
3756:
3713:
3659:
3647:Pissarrachampsa
3625:
3620:Stratiotosuchus
3584:
3565:Cynodontosuchus
3546:
3540:Razanandrongobe
3533:Pehuenchesuchus
3489:
3487:Xenodontosuchia
3476:
3440:Candidodontidae
3434:
3414:Razanandrongobe
3368:
3355:
3338:
3325:
3298:
3266:
3226:Montealtosuchus
3198:Gasparinisuchus
3148:
3093:
3046:
2942:
2937:Mahajangasuchus
2915:
2876:
2863:
2841:
2828:
2803:
2783:
2752:
2714:
2713:
2703:
2698:
2647:
2642:
2640:
2637:
2626:
2620:
2605:
2600:
2585:
2580:
2541:
2504:
2463:
2442:
2427:
2422:
2407:
2398:
2385:
2372:
2363:
2340:
2330:
2297:
2280:
2253:
2250:
2245:
2244:
2235:
2231:
2222:
2218:
2209:
2205:
2196:
2192:
2183:
2179:
2170:
2166:
2157:
2153:
2144:
2140:
2131:
2127:
2118:
2114:
2105:
2101:
2092:
2088:
2079:
2075:
2066:
2062:
2053:
2049:
2040:
2036:
2027:
2023:
2014:
2010:
2001:
1997:
1988:
1984:
1975:
1971:
1962:
1958:
1949:
1945:
1937:
1933:
1926:
1922:
1915:
1911:
1902:
1898:
1889:
1885:
1876:
1872:
1863:
1859:
1850:
1846:
1837:
1833:
1824:
1820:
1811:
1807:
1798:
1794:
1785:
1781:
1772:
1768:
1763:
1758:
1749:
1740:
1731:
1722:
1713:
1704:
1695:
1686:
1606:
1451:
1442:
1392:
1380:Stratiotosuchus
1329:Cynodontosuchus
1305:Pehuenchesuchus
1281:
1179:
1116:meridiungulates
1059:pristichampsids
1015:
986:
969:
881:
848:
761:
753:
751:Specimens known
626:, smaller than
604:, found in the
479:
364:
357:
349:
348:
335:
314:
306:
293:
280:
264:
248:
236:Crocodyliformes
223:Crocodylomorpha
139:
117:
115:
114:
113:
112:
107:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
41:
40:
38:
27:
17:
12:
11:
5:
5157:
5155:
5147:
5146:
5141:
5136:
5131:
5126:
5121:
5116:
5111:
5106:
5096:
5095:
5089:
5088:
5086:
5085:
5072:
5056:
5054:
5048:
5047:
5042:
5033:
5032:
5013:
5010:
5009:
5006:
5005:
5003:
5002:
4996:
4994:
4988:
4987:
4985:
4984:
4977:
4970:
4963:
4954:
4947:
4939:
4937:
4931:
4930:
4928:
4927:
4921:
4919:
4917:Valle Alto Fm.
4913:
4912:
4910:
4909:
4903:
4901:
4895:
4894:
4892:
4891:
4884:
4877:
4874:Platypterygius
4870:
4863:
4856:
4849:
4842:
4835:
4828:
4821:
4813:
4811:
4805:
4804:
4802:
4801:
4795:
4793:
4787:
4786:
4784:
4783:
4776:
4768:
4766:
4760:
4759:
4757:
4756:
4748:
4746:
4740:
4739:
4737:
4736:
4727:
4720:
4713:
4706:
4699:
4692:
4689:Anthracosuchus
4685:
4677:
4675:
4669:
4668:
4666:
4665:
4657:
4655:
4649:
4648:
4646:
4645:
4642:Colombitherium
4637:
4635:
4631:
4630:
4628:
4627:
4620:
4613:
4606:
4599:
4592:
4585:
4578:
4571:
4564:
4557:
4550:
4543:
4536:
4529:
4522:
4515:
4508:
4501:
4494:
4487:
4484:Hilarcotherium
4480:
4473:
4466:
4459:
4452:
4445:
4438:
4431:
4424:
4417:
4410:
4403:
4395:
4393:
4387:
4386:
4384:
4383:
4376:
4368:
4366:
4357:
4353:
4352:
4349:
4348:
4346:
4345:
4340:
4334:
4332:
4326:
4325:
4323:
4322:
4320:Valle Alto Fm.
4317:
4311:
4309:
4303:
4302:
4300:
4299:
4294:
4289:
4284:
4279:
4273:
4271:
4265:
4264:
4262:
4261:
4260:
4259:
4249:
4243:
4241:
4235:
4234:
4232:
4231:
4226:
4221:
4216:
4215:
4214:
4204:
4199:
4193:
4191:
4185:
4184:
4182:
4181:
4176:
4175:
4174:
4163:
4161:
4152:
4142:
4141:
4114:
4113:
4105:
4104:
4096:
4095:
4087:
4086:
4078:
4077:
4069:
4068:
4060:
4059:
4053:
4052:
4051:
4050:
4048:
4045:
4044:
4042:
4041:
4036:
4030:
4028:
4024:
4023:
4021:
4020:
4015:
4010:
4005:
4000:
3994:
3992:
3985:
3979:
3978:
3975:
3973:
3972:
3965:
3958:
3950:
3941:
3940:
3937:
3936:
3934:
3933:
3927:
3924:
3923:
3920:
3913:
3912:
3909:
3908:
3878:
3876:
3873:
3872:
3869:
3868:
3865:
3864:
3861:
3860:
3857:
3856:
3854:
3853:
3850:Morrinhosuchus
3846:
3838:
3836:
3830:
3829:
3827:
3826:
3819:
3812:
3804:
3802:
3800:Sphagesaurinae
3796:
3795:
3793:
3792:
3785:
3778:
3770:
3764:
3762:Sphagesauridae
3758:
3757:
3755:
3754:
3747:
3739:
3732:
3723:
3721:
3715:
3714:
3712:
3711:
3708:Eptalofosuchus
3704:
3697:
3690:
3686:Brasileosaurus
3681:
3675:
3669:
3668:
3665:
3664:
3661:
3660:
3658:
3657:
3650:
3643:
3635:
3633:
3627:
3626:
3624:
3623:
3616:
3609:
3602:
3594:
3592:
3586:
3585:
3583:
3582:
3575:
3572:Gondwanasuchus
3568:
3560:
3554:
3548:
3547:
3545:
3544:
3536:
3529:
3521:
3513:
3510:Chimaerasuchus
3505:
3499:
3490:
3485:
3482:
3481:
3478:
3477:
3475:
3474:
3467:
3460:
3457:Lavocatchampsa
3453:
3445:
3443:
3436:
3435:
3433:
3432:
3425:
3418:
3410:
3403:
3395:
3388:
3379:
3369:
3364:
3357:
3356:
3351:
3344:
3343:
3340:
3339:
3313:
3311:
3308:
3307:
3304:
3303:
3300:
3299:
3297:
3296:
3295:
3294:
3288:
3286:
3276:
3275:
3272:
3271:
3268:
3267:
3265:
3264:
3257:
3250:
3243:
3236:
3229:
3222:
3215:
3208:
3201:
3194:
3187:
3184:Bayomesasuchus
3180:
3173:
3166:
3158:
3156:
3150:
3149:
3147:
3146:
3139:
3132:
3125:
3118:
3111:
3108:Barreirosuchus
3103:
3101:
3095:
3094:
3092:
3091:
3087:Stolokrosuchus
3083:
3076:
3068:
3060:
3054:
3048:
3047:
3045:
3044:
3037:
3030:
3023:
3016:
3009:
3002:
2995:
2988:
2985:Dentaneosuchus
2981:
2974:
2967:
2960:
2952:
2950:
2944:
2943:
2941:
2940:
2933:
2925:
2923:
2917:
2916:
2914:
2913:
2906:
2902:Trematochampsa
2898:
2891:
2883:
2877:
2872:
2869:
2868:
2865:
2864:
2862:
2861:
2852:
2842:
2837:
2830:
2829:
2824:
2817:
2816:
2813:
2812:
2809:
2808:
2805:
2804:
2802:
2801:
2800:
2799:
2793:
2791:
2785:
2784:
2782:
2781:
2780:
2779:
2770:
2764:
2754:
2753:
2749:
2748:
2739:
2733:
2727:
2721:
2712:
2705:
2704:
2699:
2697:
2696:
2689:
2682:
2674:
2668:
2667:
2653:
2652:
2636:
2635:External links
2633:
2632:
2631:
2629:on 2009-07-04.
2619:978-8478407071
2618:
2598:
2578:
2539:
2502:
2476:(2): 328–354.
2461:
2440:
2434:(in Spanish).
2420:
2396:
2383:
2370:
2361:
2328:
2295:
2278:
2266:(4): 487–521.
2249:
2246:
2243:
2242:
2229:
2216:
2203:
2190:
2177:
2175:, p. R92)
2164:
2162:, p. 356)
2151:
2149:, p. 117)
2138:
2125:
2112:
2099:
2097:, p. 116)
2086:
2084:, p. 116)
2073:
2071:, p. 115)
2060:
2047:
2034:
2021:
2008:
1995:
1982:
1969:
1967:, p. R92)
1956:
1954:, p. 356)
1943:
1931:
1920:
1909:
1896:
1894:, p. 114)
1883:
1870:
1857:
1844:
1831:
1818:
1805:
1792:
1779:
1765:
1764:
1762:
1759:
1755:
1754:
1751:
1750:
1746:
1745:
1742:
1741:
1737:
1736:
1733:
1732:
1728:
1727:
1724:
1723:
1719:
1718:
1715:
1714:
1710:
1709:
1706:
1705:
1701:
1700:
1697:
1696:
1692:
1691:
1688:
1687:
1683:
1682:
1679:
1678:
1669:
1666:
1665:
1662:
1661:
1652:
1649:
1648:
1645:
1644:
1635:
1632:
1631:
1628:
1627:
1622:
1617:
1615:
1612:
1611:
1608:
1607:
1603:
1602:
1599:
1598:
1589:
1586:
1585:
1582:
1581:
1572:
1567:
1565:
1560:
1558:
1555:
1554:
1551:
1550:
1541:
1536:
1534:
1531:
1530:
1527:
1526:
1517:
1512:
1510:
1504:
1503:
1500:
1499:
1490:
1484:
1483:
1480:
1479:
1470:
1462:
1460:
1457:
1456:
1453:
1452:
1448:
1447:
1444:
1443:
1439:
1438:
1435:
1434:
1425:
1422:
1421:
1418:
1417:
1408:
1403:
1401:
1398:
1397:
1394:
1393:
1389:
1388:
1385:
1384:
1375:
1372:
1371:
1368:
1367:
1358:
1353:
1351:
1346:
1344:
1338:
1337:
1334:
1333:
1324:
1319:
1317:
1314:
1313:
1310:
1309:
1300:
1295:
1293:
1287:
1286:
1283:
1282:
1278:
1277:
1274:
1273:
1264:
1261:
1260:
1257:
1256:
1252:Chimaerasuchus
1247:
1242:
1240:
1235:
1233:
1228:
1202:S. icaeorhinus
1178:
1175:
1112:apex predators
1014:
1011:
1010:
1009:
985:
982:
981:
980:
968:
962:
961:
960:
946:
940:
934:
928:
922:
905:S. icaeorhinus
901:dental alveoli
897:S. icaeorhinus
880:
873:
872:
871:
847:
844:
843:
842:
836:
826:
820:
814:
804:
798:
788:
785:S. icaeorhinus
760:
757:
752:
749:
628:S. icaeorhinus
608:in Colombia.
594:S. icaeorhinus
511:Tatacoa Desert
507:Robert Stirton
478:
475:
435:of the family
433:crocodylomorph
413:(meaning " of
404:
403:
402:
401:
387:
385:Langston, 1965
373:
372:
366:
365:
358:
344:
343:
337:
336:
328:
326:
322:
321:
304:
300:
299:
291:
287:
286:
278:
271:
270:
262:
255:
254:
246:
239:
238:
233:
226:
225:
220:
213:
212:
207:
200:
199:
194:
187:
186:
181:
177:
176:
171:
167:
166:
161:
157:
156:
151:
147:
146:
133:
132:
128:
127:
119:
118:
110:
109:
108:
103:
98:
93:
88:
83:
78:
73:
68:
63:
58:
53:
48:
47:
28:
15:
13:
10:
9:
6:
4:
3:
2:
5156:
5145:
5142:
5140:
5137:
5135:
5132:
5130:
5127:
5125:
5122:
5120:
5117:
5115:
5112:
5110:
5107:
5105:
5102:
5101:
5099:
5082:
5077:
5073:
5068:
5062:
5058:
5057:
5055:
5053:
5049:
5045:
5040:
5029:
5025:
5021:
5011:
5001:
4998:
4997:
4995:
4993:
4989:
4983:
4982:
4978:
4976:
4975:
4971:
4969:
4968:
4964:
4962:
4961:
4960:
4955:
4953:
4952:
4948:
4946:
4945:
4944:Antarctilamna
4941:
4940:
4938:
4936:
4932:
4926:
4923:
4922:
4920:
4918:
4914:
4908:
4905:
4904:
4902:
4900:
4896:
4890:
4889:
4885:
4883:
4882:
4881:Sachicasaurus
4878:
4876:
4875:
4871:
4869:
4868:
4867:Padillasaurus
4864:
4862:
4861:
4857:
4855:
4854:
4850:
4848:
4847:
4843:
4841:
4840:
4836:
4834:
4833:
4832:Desmatochelys
4829:
4827:
4826:
4822:
4820:
4819:
4815:
4814:
4812:
4810:
4806:
4800:
4797:
4796:
4794:
4792:
4788:
4782:
4781:
4780:Yaguarasaurus
4777:
4775:
4774:
4770:
4769:
4767:
4765:
4761:
4755:
4754:
4750:
4749:
4747:
4745:
4741:
4735:
4734:
4733:
4728:
4726:
4725:
4721:
4719:
4718:
4714:
4712:
4711:
4707:
4705:
4704:
4700:
4698:
4697:
4693:
4691:
4690:
4686:
4684:
4683:
4679:
4678:
4676:
4674:
4670:
4664:
4663:
4659:
4658:
4656:
4654:
4650:
4644:
4643:
4639:
4638:
4636:
4634:Gualanday Gp.
4632:
4626:
4625:
4621:
4619:
4618:
4614:
4612:
4611:
4607:
4605:
4604:
4600:
4598:
4597:
4593:
4591:
4590:
4586:
4584:
4583:
4579:
4577:
4576:
4572:
4570:
4569:
4565:
4563:
4562:
4558:
4556:
4555:
4551:
4549:
4548:
4544:
4542:
4541:
4540:Miocallicebus
4537:
4535:
4534:
4530:
4528:
4527:
4523:
4521:
4520:
4516:
4514:
4513:
4509:
4507:
4506:
4502:
4500:
4499:
4495:
4493:
4492:
4488:
4486:
4485:
4481:
4479:
4478:
4474:
4472:
4471:
4467:
4465:
4464:
4460:
4458:
4457:
4453:
4451:
4450:
4446:
4444:
4443:
4439:
4437:
4436:
4432:
4430:
4429:
4425:
4423:
4422:
4418:
4416:
4415:
4411:
4409:
4408:
4404:
4402:
4401:
4400:Anachlysictis
4397:
4396:
4394:
4392:
4388:
4382:
4381:
4377:
4375:
4374:
4370:
4369:
4367:
4365:
4361:
4358:
4354:
4344:
4341:
4339:
4336:
4335:
4333:
4331:
4327:
4321:
4318:
4316:
4313:
4312:
4310:
4308:
4304:
4298:
4295:
4293:
4290:
4288:
4285:
4283:
4280:
4278:
4275:
4274:
4272:
4270:
4266:
4258:
4255:
4254:
4253:
4250:
4248:
4245:
4244:
4242:
4240:
4236:
4230:
4227:
4225:
4222:
4220:
4217:
4213:
4210:
4209:
4208:
4205:
4203:
4200:
4198:
4197:Barzalosa Fm.
4195:
4194:
4192:
4190:
4186:
4180:
4177:
4173:
4170:
4169:
4168:
4165:
4164:
4162:
4160:
4156:
4153:
4151:
4148:fossiliferous
4143:
4136:
4129:
4118:
4109:
4100:
4091:
4082:
4073:
4064:
4057:
4040:
4037:
4035:
4032:
4031:
4029:
4025:
4019:
4016:
4014:
4011:
4009:
4006:
4004:
4001:
3999:
3996:
3995:
3993:
3989:
3986:
3980:
3971:
3966:
3964:
3959:
3957:
3952:
3951:
3948:
3932:
3929:
3928:
3925:
3918:
3914:
3904:
3899:
3896:
3889:
3882:
3852:
3851:
3847:
3845:
3844:
3843:Caipirasuchus
3840:
3839:
3837:
3835:
3831:
3825:
3824:
3820:
3818:
3817:
3816:Caryonosuchus
3813:
3811:
3810:
3806:
3805:
3803:
3801:
3797:
3791:
3790:
3786:
3784:
3783:
3782:Labidiosuchus
3779:
3777:
3776:
3772:
3771:
3768:
3765:
3763:
3759:
3753:
3752:
3748:
3745:
3744:
3743:Mariliasuchus
3740:
3738:
3737:
3733:
3730:
3729:
3728:Coronelsuchus
3725:
3724:
3722:
3720:
3716:
3710:
3709:
3705:
3703:
3702:
3701:Coronelsuchus
3698:
3696:
3695:
3694:Coringasuchus
3691:
3688:
3687:
3683:
3682:
3679:
3676:
3674:
3670:
3656:
3655:
3651:
3649:
3648:
3644:
3642:
3641:
3640:Campinasuchus
3637:
3636:
3634:
3632:
3628:
3622:
3621:
3617:
3615:
3614:
3610:
3608:
3607:
3606:Aplestosuchus
3603:
3601:
3600:
3599:Aphaurosuchus
3596:
3595:
3593:
3591:
3590:Baurusuchinae
3587:
3581:
3580:
3576:
3574:
3573:
3569:
3567:
3566:
3562:
3561:
3558:
3555:
3553:
3552:Baurusuchidae
3549:
3542:
3541:
3537:
3535:
3534:
3530:
3527:
3526:
3522:
3519:
3518:
3517:Comahuesuchus
3514:
3512:
3511:
3507:
3506:
3503:
3500:
3498:
3494:
3491:
3488:
3483:
3473:
3472:
3468:
3466:
3465:
3461:
3459:
3458:
3454:
3452:
3451:
3447:
3446:
3444:
3441:
3437:
3431:
3430:
3429:Uruguaysuchus
3426:
3424:
3423:
3419:
3416:
3415:
3411:
3409:
3408:
3404:
3401:
3400:
3399:Comahuesuchus
3396:
3394:
3393:
3392:Araripesuchus
3389:
3386:
3385:
3381:
3380:
3377:
3373:
3370:
3367:
3362:
3358:
3354:
3349:
3345:
3336:
3332:
3329:
3323:
3317:
3292:
3291:
3290:
3289:
3287:
3285:
3281:
3277:
3263:
3262:
3261:Uberabasuchus
3258:
3256:
3255:
3251:
3249:
3248:
3244:
3242:
3241:
3237:
3235:
3234:
3230:
3228:
3227:
3223:
3221:
3220:
3216:
3214:
3213:
3209:
3207:
3206:
3202:
3200:
3199:
3195:
3193:
3192:
3188:
3186:
3185:
3181:
3179:
3178:
3177:Barrosasuchus
3174:
3172:
3171:
3170:Barcinosuchus
3167:
3165:
3164:
3160:
3159:
3157:
3155:
3154:Peirosaurinae
3151:
3145:
3144:
3140:
3138:
3137:
3133:
3131:
3130:
3126:
3124:
3123:
3119:
3117:
3116:
3112:
3110:
3109:
3105:
3104:
3102:
3100:
3096:
3089:
3088:
3084:
3082:
3081:
3080:Miadanasuchus
3077:
3074:
3073:
3069:
3067:
3066:
3062:
3061:
3058:
3055:
3053:
3052:Peirosauridae
3049:
3043:
3042:
3038:
3036:
3035:
3031:
3029:
3028:
3024:
3022:
3021:
3017:
3015:
3014:
3010:
3008:
3007:
3003:
3001:
3000:
2996:
2994:
2993:
2989:
2987:
2986:
2982:
2980:
2979:
2975:
2973:
2972:
2968:
2966:
2965:
2961:
2959:
2958:
2954:
2953:
2951:
2949:
2945:
2939:
2938:
2934:
2932:
2931:
2927:
2926:
2924:
2922:
2918:
2912:
2911:
2907:
2904:
2903:
2899:
2897:
2896:
2892:
2890:
2889:
2885:
2884:
2881:
2878:
2875:
2870:
2859:
2858:
2854:
2853:
2850:
2846:
2843:
2840:
2835:
2831:
2827:
2822:
2818:
2797:
2796:
2795:
2794:
2792:
2790:
2786:
2778:
2774:
2773:
2772:
2771:
2768:
2765:
2763:
2759:
2755:
2747:
2743:
2740:
2738:
2734:
2732:
2728:
2726:
2722:
2720:
2716:
2715:
2710:
2706:
2702:
2695:
2690:
2688:
2683:
2681:
2676:
2675:
2672:
2666:
2664:
2660:
2655:
2654:
2650:
2639:
2634:
2625:
2621:
2615:
2611:
2604:
2599:
2596:(1–2): 12–40.
2595:
2591:
2584:
2579:
2575:
2571:
2566:
2561:
2557:
2553:
2549:
2545:
2540:
2535:
2530:
2525:
2520:
2516:
2512:
2508:
2503:
2499:
2495:
2491:
2487:
2483:
2479:
2475:
2471:
2467:
2462:
2458:
2454:
2450:
2448:
2441:
2437:
2433:
2426:
2421:
2417:
2413:
2406:
2404:
2397:
2393:
2389:
2384:
2380:
2376:
2371:
2367:
2362:
2358:
2354:
2350:
2346:
2339:
2337:
2329:
2325:
2321:
2317:
2313:
2309:
2305:
2301:
2296:
2292:
2288:
2284:
2279:
2274:
2269:
2265:
2261:
2257:
2252:
2251:
2247:
2240:, p. 38)
2239:
2233:
2230:
2227:, p. 20)
2226:
2220:
2217:
2214:, p. 29)
2213:
2207:
2204:
2200:
2194:
2191:
2188:, p. 27)
2187:
2181:
2178:
2174:
2168:
2165:
2161:
2155:
2152:
2148:
2142:
2139:
2135:
2129:
2126:
2122:
2116:
2113:
2109:
2103:
2100:
2096:
2090:
2087:
2083:
2077:
2074:
2070:
2064:
2061:
2058:, p. 20)
2057:
2051:
2048:
2045:, p. 17)
2044:
2043:Langston 1965
2038:
2035:
2032:, p. 16)
2031:
2030:Langston 1965
2025:
2022:
2019:, p. 17)
2018:
2017:Langston 1965
2012:
2009:
2006:, p. 17)
2005:
2004:Langston 1965
1999:
1996:
1993:, p. 17)
1992:
1991:Langston 1965
1986:
1983:
1979:
1978:Langston 1965
1973:
1970:
1966:
1960:
1957:
1953:
1947:
1944:
1941:, p. 38)
1940:
1935:
1932:
1929:
1924:
1921:
1918:
1913:
1910:
1907:, p. 11)
1906:
1900:
1897:
1893:
1887:
1884:
1881:, p. 26)
1880:
1874:
1871:
1868:, p. 20)
1867:
1861:
1858:
1854:
1848:
1845:
1842:, p. 16)
1841:
1840:Langston 1965
1835:
1832:
1829:, p. 14)
1828:
1827:Langston 1965
1822:
1819:
1816:, p. 14)
1815:
1814:Langston 1965
1809:
1806:
1802:
1801:Langston 1965
1796:
1793:
1789:
1788:Langston 1965
1783:
1780:
1776:
1770:
1767:
1760:
1753:
1752:
1744:
1743:
1735:
1734:
1726:
1725:
1717:
1716:
1708:
1707:
1699:
1698:
1690:
1689:
1681:
1680:
1677:
1676:
1675:
1668:
1667:
1664:
1663:
1660:
1659:
1658:
1651:
1650:
1647:
1646:
1643:
1642:
1641:
1634:
1633:
1630:
1629:
1626:
1621:
1620:
1614:
1613:
1610:
1609:
1601:
1600:
1597:
1596:
1595:
1588:
1587:
1584:
1583:
1580:
1579:
1578:
1571:
1570:
1564:
1563:
1557:
1556:
1553:
1552:
1549:
1548:
1547:
1540:
1539:
1533:
1532:
1529:
1528:
1525:
1524:
1523:
1516:
1515:
1509:
1506:
1505:
1502:
1501:
1498:
1497:
1496:
1489:
1488:Iberosuchidae
1486:
1485:
1482:
1481:
1478:
1477:
1476:
1469:
1468:Bergisuchidae
1466:
1465:
1459:
1458:
1455:
1454:
1446:
1445:
1437:
1436:
1433:
1432:
1431:
1424:
1423:
1420:
1419:
1416:
1415:
1414:
1407:
1406:
1400:
1399:
1396:
1395:
1387:
1386:
1383:
1382:
1381:
1374:
1373:
1370:
1369:
1366:
1365:
1364:
1357:
1356:
1350:
1349:
1343:
1342:Baurusuchidae
1340:
1339:
1336:
1335:
1332:
1331:
1330:
1323:
1322:
1316:
1315:
1312:
1311:
1308:
1307:
1306:
1299:
1298:
1292:
1289:
1288:
1285:
1284:
1276:
1275:
1272:
1271:
1270:
1263:
1262:
1259:
1258:
1255:
1254:
1253:
1246:
1245:
1239:
1238:
1232:
1231:
1227:
1225:
1221:
1217:
1216:
1211:
1207:
1203:
1199:
1195:
1191:
1187:
1183:
1176:
1174:
1172:
1168:
1164:
1160:
1159:Astrapotheria
1156:
1155:
1150:
1146:
1141:
1139:
1135:
1134:
1133:Pericotoxodon
1129:
1128:
1123:
1122:
1117:
1113:
1109:
1108:phorusrhacids
1105:
1101:
1100:
1095:
1094:
1089:
1088:sparassodonts
1084:
1082:
1081:Komodo dragon
1078:
1077:Langston 1965
1074:
1073:
1068:
1064:
1060:
1056:
1052:
1048:
1044:
1040:
1035:
1031:
1024:
1019:
1012:
1007:
1003:
999:
995:
991:
988:
987:
983:
978:
974:
971:
970:
967:
963:
958:
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950:
947:
944:
941:
938:
935:
932:
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926:
923:
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898:
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890:
886:
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874:
869:
865:
861:
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850:
849:
845:
840:
837:
834:
830:
827:
824:
821:
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815:
812:
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805:
802:
799:
796:
792:
789:
786:
781:
777:
773:
770:
769:
768:
766:
758:
756:
750:
748:
746:
742:
738:
734:
729:
727:
723:
719:
715:
711:
710:
705:
704:S. querejazus
701:
697:
693:
689:
685:
684:S. querejazus
681:
677:
673:
669:
668:
662:
660:
656:
652:
648:
643:
641:
637:
633:
629:
625:
620:
618:
614:
609:
607:
603:
599:
595:
591:
587:
583:
581:
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572:
568:
564:
563:
558:
557:
552:
551:
546:
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540:
536:
532:
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516:
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508:
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488:
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476:
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468:
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457:
453:
450:
446:
442:
438:
434:
431:
427:
424:
420:
416:
412:
411:
398:
397:
396:lapsus calami
391:
388:
383:
381:
377:
376:
374:
371:
367:
362:
356:
354:
345:
342:
341:Binomial name
338:
334:
333:
327:
324:
323:
319:
313:
312:
305:
302:
301:
298:
292:
289:
288:
285:
279:
276:
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269:
263:
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257:
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198:
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192:
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188:
185:
182:
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169:
168:
165:
162:
159:
158:
155:
152:
149:
148:
143:
138:
134:
129:
125:
120:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
45:
36:
32:
26:
22:
19:
5051:
4992:Floresta Fm.
4979:
4972:
4967:Holoptychius
4965:
4959:Bothriolepis
4957:
4956:
4949:
4942:
4899:Arcabuco Fm.
4886:
4879:
4872:
4865:
4860:Muiscasaurus
4858:
4851:
4844:
4837:
4830:
4823:
4818:Acostasaurus
4816:
4778:
4771:
4751:
4730:
4729:
4722:
4715:
4708:
4703:Cerrejonemys
4701:
4694:
4687:
4680:
4673:Cerrejón Fm.
4660:
4640:
4622:
4615:
4608:
4601:
4594:
4587:
4580:
4573:
4566:
4559:
4552:
4547:Miocochilius
4545:
4538:
4531:
4524:
4518:
4517:
4510:
4505:Huilatherium
4503:
4498:Hondadelphys
4496:
4489:
4482:
4475:
4468:
4461:
4454:
4447:
4440:
4433:
4428:Cebupithecia
4426:
4419:
4412:
4405:
4398:
4378:
4373:Gomphotheres
4371:
4343:Floresta Fm.
4315:Arcabuco Fm.
4252:Cerrejón Fm.
4027:19th century
3884:
3848:
3841:
3823:Sphagesaurus
3821:
3814:
3807:
3787:
3780:
3773:
3749:
3741:
3734:
3726:
3719:Notosuchidae
3706:
3699:
3692:
3684:
3673:Sphagesauria
3652:
3645:
3638:
3618:
3611:
3604:
3597:
3577:
3570:
3563:
3538:
3531:
3523:
3515:
3508:
3469:
3464:Malawisuchus
3462:
3455:
3448:
3427:
3420:
3412:
3407:Libycosuchus
3405:
3397:
3390:
3382:
3319:
3259:
3252:
3245:
3238:
3233:Patagosuchus
3231:
3224:
3217:
3210:
3205:Hamadasuchus
3203:
3196:
3189:
3182:
3175:
3168:
3163:Antaeusuchus
3161:
3141:
3134:
3127:
3122:Epoidesuchus
3120:
3115:Caririsuchus
3113:
3106:
3099:Pepesuchinae
3085:
3078:
3072:Fortignathus
3070:
3065:Amargasuchus
3063:
3039:
3032:
3027:Sahitisuchus
3025:
3018:
3011:
3005:
3004:
2997:
2990:
2983:
2976:
2969:
2964:Barinasuchus
2962:
2955:
2935:
2928:
2908:
2900:
2893:
2886:
2855:
2777:Pseudosuchia
2762:Pseudosuchia
2741:
2662:
2658:
2624:the original
2609:
2593:
2589:
2547:
2543:
2514:
2510:
2473:
2469:
2465:
2456:
2452:
2446:
2435:
2431:
2415:
2411:
2402:
2391:
2387:
2378:
2374:
2365:
2348:
2344:
2335:
2310:(1): 20–27.
2307:
2303:
2299:
2293:: 1663–1666.
2290:
2286:
2282:
2263:
2259:
2248:Bibliography
2232:
2219:
2206:
2193:
2180:
2167:
2154:
2141:
2128:
2115:
2102:
2089:
2076:
2063:
2050:
2037:
2024:
2011:
1998:
1985:
1972:
1959:
1946:
1934:
1923:
1912:
1899:
1886:
1873:
1860:
1847:
1834:
1821:
1808:
1803:, p. 6)
1795:
1782:
1769:
1672:
1671:
1655:
1654:
1639:
1638:
1637:
1624:
1592:
1591:
1575:
1574:
1546:Barinasuchus
1544:
1543:
1520:
1519:
1493:
1492:
1473:
1472:
1428:
1427:
1411:
1410:
1378:
1377:
1361:
1360:
1327:
1326:
1303:
1302:
1291:Sebecosuchia
1290:
1269:Sphagesaurus
1267:
1266:
1250:
1249:
1213:
1210:Barinasuchus
1209:
1205:
1201:
1193:
1186:Sebecosuchia
1181:
1180:
1171:Notohippidae
1163:Leontiniidae
1154:Barinasuchus
1152:
1148:
1142:
1131:
1127:Huilatherium
1125:
1119:
1097:
1091:
1085:
1070:
1063:mekosuchines
1051:crocodilians
1043:baurusuchids
1039:peirosaurids
1033:
1029:
1027:
1022:
1013:Paleobiology
1005:
1001:
997:
989:
977:L. huilensis
976:
972:
957:L. huilensis
956:
948:
942:
936:
930:
924:
904:
896:
884:
851:
838:
832:
828:
822:
816:
810:
806:
800:
790:
784:
779:
771:
764:
762:
754:
744:
740:
736:
730:
725:
718:Barinasuchus
717:
713:
707:
706:was renamed
703:
699:
696:S. huilensis
695:
683:
680:S. huilensis
679:
675:
667:Barinasuchus
665:
663:
646:
644:
640:S. huilensis
639:
627:
621:
617:S. huilensis
616:
610:
597:
593:
578:
560:
554:
548:
542:
538:
498:
492:
486:
471:L. huilensis
470:
467:type species
465:. Thus, the
459:
455:
409:
408:
407:
394:
389:
378:
352:
347:
331:
330:
310:
309:
274:
268:Sebecosuchia
258:
242:
229:
216:
210:Pseudosuchia
203:
190:
24:
18:
4951:Asterolepis
4853:Leyvachelys
4846:Leivanectes
4839:Kronosaurus
4764:Hondita Fm.
4596:Purussaurus
4589:Potamosiren
4568:Neotamandua
4561:Mourasuchus
4491:Hoazinoides
4477:Gryposuchus
4435:Colombophis
4421:Boreostemma
4414:Balanerodus
4380:Megatherium
4364:Pleistocene
4287:Hondita Fm.
4207:Honda Group
4159:Pleistocene
4013:Hoffstetter
3984:researchers
3736:Llanosuchus
3654:Wargosuchus
3613:Baurusuchus
3525:Eremosuchus
3497:Baurusuchia
3384:Anatosuchus
3254:Rukwasuchus
3240:Peirosaurus
3143:Roxochampsa
3041:Zulmasuchus
2992:Iberosuchus
2978:Bretesuchus
2971:Bergisuchus
2957:Ayllusuchus
2930:Kaprosuchus
2888:Baharijodon
2857:Microsuchus
2839:Ziphosuchia
2826:Ziphosuchia
2789:Ziphosuchia
2701:Ziphosuchia
2657:Photo of a
2550:(1): 3–21.
2534:11336/69518
2453:Ameghiniana
2388:Gryposuchus
2186:Busbey 1986
2056:Busbey 1986
1879:Busbey 1986
1866:Busbey 1986
1674:Zulmasuchus
1594:Bretesuchus
1577:Ayllusuchus
1495:Iberosuchus
1475:Bergisuchus
1215:Bretesuchus
1206:Zulmasuchus
1167:Adianthidae
1104:Argot (2004
1072:Paleosuchus
949:IGM 184 165
937:IGM 250 427
931:IGM 250 541
925:IGM 184 427
852:TMM 41658-8
709:Zulmasuchus
523:Honda Group
430:notosuchian
197:Archosauria
42:13–12
31:Mid Miocene
5098:Categories
5052:Langstonia
5000:Trilobites
4981:Strepsodus
4974:Pterygotus
4696:Carbonemys
4653:Bogotá Fm.
4575:Nuciruptor
4554:Mohanamico
4519:Langstonia
4512:Lagonimico
4269:Cretaceous
4247:Bogotá Fm.
4167:Sabana Fm.
3991:After 1900
3789:Yacarerani
3751:Notosuchus
3471:Pakasuchus
3450:Candidodon
3422:Simosuchus
3366:Notosuchia
3353:Notosuchia
3293:see below↓
3284:Notosuchia
3247:Pepesuchus
3219:Lomasuchus
3212:Kinesuchus
3136:Pepesuchus
3020:Ogresuchus
3013:Lorosuchus
3006:Langstonia
2910:Wanosuchus
2798:see below↓
2746:Metasuchia
2731:Sauropsida
2663:Langstonia
2565:11336/2663
2517:: S7–S36.
2418:: 127–144.
1761:References
1640:Langstonia
1522:Lorosuchus
1182:Langstonia
1149:Langstonia
1067:crocodiles
1047:Cretaceous
1030:Langstonia
1023:Langstonia
998:Langstonia
943:IGM 184378
889:premaxilla
885:IGM 250816
839:UCMP 44566
833:Langstonia
829:UCMP 44561
823:UCMP 44563
817:UCMP 40220
811:Langstonia
807:UCMP 40186
801:UCMP 44562
791:UCMP 41308
772:UCMP 37877
745:Langstonia
726:Langstonia
700:Langstonia
659:Ingeominas
624:premaxilla
567:dyrosaurid
517:overlying
499:Langstonia
493:The first
487:Langstonia
456:Langstonia
410:Langstonia
311:Langstonia
252:Notosuchia
25:Langstonia
4935:Cuche Fm.
4799:Ammonites
4753:Eonatator
4744:Oliní Gp.
4732:Titanoboa
4724:Puentemys
4617:Stirtonia
4449:Dukecynus
4391:Honda Gp.
4338:Cuche Fm.
4292:Oliní Gp.
4239:Paleogene
4224:Uitpa Fm.
4219:Jimol Fm.
4179:Soatá Fm.
3579:Pabwehshi
3129:Itasuchus
2999:Ilchunaia
2948:Sebecidae
2895:Doratodon
2717:Kingdom:
2351:: 19–42.
1508:Sebecidae
1363:Pabwehshi
1222:based on
1220:Cladogram
1177:Phylogeny
1138:megafauna
1099:Dukecynus
1061:and some
1049:and some
1006:huilensis
868:osteoderm
860:squamosal
776:symphysis
741:huilensis
733:Paleocene
688:Paleocene
672:Venezuela
651:vertebrae
636:osteoderm
590:Argentina
582:huilensis
519:sandstone
515:claystone
463:huilensis
449:Colombian
437:Sebecidae
382:huilensis
325:Species:
297:Sebecidae
160:Kingdom:
154:Eukaryota
5119:Laventan
5104:Sebecids
5067:Q5236876
5061:Wikidata
5020:Category
4809:Paja Fm.
4791:Hiló Fm.
4582:Patasola
4526:Lycopsis
4330:Devonian
4307:Jurassic
4297:Paja Fm.
4282:Hiló Fm.
4257:Cerrejón
4229:Ware Fm.
4212:La Venta
4039:Humboldt
2725:Chordata
2723:Phylum:
2719:Animalia
2574:15751319
2498:86565764
2381:: 1–127.
1190:Gondwana
1055:Cenozoic
973:MUSM 912
953:maxillar
913:diastema
575:holotype
445:Laventan
421:) is an
415:Langston
370:Synonyms
361:Langston
290:Family:
184:Reptilia
174:Chordata
170:Phylum:
164:Animalia
150:Domain:
35:Laventan
4533:Micodon
4189:Neogene
4008:Hettner
3982:Notable
3034:Sebecus
2874:Sebecia
2735:Order:
2729:Class:
2665:) tooth
2478:Bibcode
2438:: 1–25.
2403:Sebecus
2345:Zootaxa
2312:Bibcode
2283:Sebecus
1657:Sebecus
1194:Sebecus
1057:as the
1053:of the
1045:of the
1002:Sebecus
893:maxilla
737:Sebecus
714:Sebecus
692:Bolivia
676:Sebecus
598:Sebecus
580:Sebecus
571:dentary
495:fossils
461:Sebecus
441:Miocene
423:extinct
380:Sebecus
363:, 1965)
350:†
329:†
303:Genus:
284:Sebecia
180:Class:
111:↓
5081:192480
5024:Images
4662:Etayoa
4172:Tibitó
4034:Cuvier
4018:Páramo
4003:Hammen
2616:
2590:Terrae
2572:
2496:
918:sacral
856:square
655:caudal
602:Eocene
586:Eocene
4925:Flora
4146:Major
3998:Etayo
2742:Clade
2627:(PDF)
2606:(PDF)
2586:(PDF)
2570:S2CID
2494:S2CID
2428:(PDF)
2408:(PDF)
2341:(PDF)
1145:Andes
879:(IGM)
864:jugal
632:jugal
426:genus
275:Clade
259:Clade
243:Clade
230:Clade
217:Clade
204:Clade
191:Clade
2775:see
2614:ISBN
2349:2871
1212:and
1204:and
1196:and
1169:and
1151:and
1124:and
1096:and
1041:and
1004:cf.
739:cf.
682:and
613:Peru
559:and
473:.
318:2007
50:PreꞒ
2560:hdl
2552:doi
2529:hdl
2519:doi
2515:163
2486:doi
2353:doi
2320:doi
2291:284
2268:doi
2264:140
1140:.
1118:as
728:.
690:of
686:of
670:of
588:of
497:of
469:is
428:of
5100::
5078::
5063::
5022:•
2744::
2661:(=
2612:.
2588:.
2568:.
2558:.
2548:20
2546:.
2527:.
2513:.
2509:.
2492:.
2484:.
2474:32
2472:.
2457:48
2455:.
2451:.
2430:.
2416:29
2414:.
2410:.
2379:52
2377:.
2347:.
2343:.
2318:.
2306:.
2262:.
2258:.
1226:.
1218:.
1165:,
1161:,
1090:,
1008:.
979:.
835:.
787:.
678:,
553:,
547:,
454:.
277::
261::
245::
232::
219::
206::
193::
100:Pg
44:Ma
3969:e
3962:t
3955:v
3746:?
3731:?
3689:?
3543:?
3528:?
3520:?
3442:?
3417:?
3402:?
3387:?
3090:?
3075:?
2905:?
2860:?
2693:e
2686:t
2679:v
2594:9
2576:.
2562::
2554::
2537:.
2531::
2521::
2500:.
2488::
2480::
2436:3
2359:.
2355::
2334:"
2326:.
2322::
2314::
2308:6
2276:.
2270::
2236:(
2223:(
2210:(
2197:(
2184:(
2171:(
2158:(
2145:(
2132:(
2119:(
2106:(
2093:(
2080:(
2067:(
2054:(
2041:(
2028:(
2015:(
2002:(
1989:(
1976:(
1963:(
1950:(
1903:(
1890:(
1877:(
1864:(
1851:(
1838:(
1825:(
1812:(
1799:(
1790:)
1786:(
1777:)
1773:(
489:.
399:)
359:(
307:†
294:†
281:†
265:†
249:†
105:N
95:K
90:J
85:T
80:P
75:C
70:D
65:S
60:O
55:Ꞓ
39:~
37:)
33:(
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