467:
392:. Further exposure over the course of 20–60 minutes results in the flesh becoming grayish to blackish. The flesh has no distinctive door or taste. The pore surface is initially dull yellow, and sometimes ages to dingy olive-brown. Unlike many other boletes, it does not turn blue when bruised, although it may have natural blue-green stains. The pores are circular, measuring less than 1 mm, while the tubes extend to 8–14 mm deep. The stipe measures 3–10 cm (1.2–3.9 in) long by 1–3 cm (0.4–1.2 in) thick. It is nearly equal in length throughout or tapered from the top to base. Its color is pale yellow to brownish underneath the pale brown scabers that darken in age.
46:
556:
396:
33:
388:
shows through. The cap tends to undergo significant color changes throughout its development—first bright yellow, then dark brown, then finally pale tan—which may make it difficult to identify in the field. The flesh is white to pale yellow, and it stains reddish to burgundy when cut or bruised. This staining is most prominent at the junction of the cap and the
539:
mushrooms have been described variously as "great", and "of poor quality". They have a nutty flavor and firm texture; older specimens are less firm but retain the flavor. Drying the mushrooms enhances the flavor. The stipe tends to harbor insect larvae and should be cleaned before consumption. The
387:
The convex cap measures 5–15 cm (2.0–5.9 in) wide. Its color is orange-yellow, aging to yellow-brown. The cap margin has a narrow flap of sterile tissue. The surface of the cap is dry, with wrinkles and pits at maturity. It often becomes cracked in age, and the whitish flesh underneath
586:) is a frequent host. The bolete fruits singly or in groups in forests, shaded lawns, and often found in areas disturbed by human activity, such as pathsides and picnic areas. Fruiting typically occurs from July to September. A Chinese study evaluating the concentrations of
633:
trend. In Asia, the species has been reported from India, Korea, China, and Taiwan. Taiwanese specimens tend to have slightly smaller spores (10–16 by 4–5 μm) than those from mainland China or from
America.
497:, found in Belize, features a similar cap and scaber pigmentation on the stipe, and similar color changes in response to injury in the flesh of the cap and the apex of the stipe; unlike
513:. It generally has a darker cap, paler scabers, and somewhat wider spores, although these characteristics are variable. In his original species description, Charles Peck noted that
466:
1537:
258:
that stains initially pinkish to reddish and then to grayish or blackish when injured. The pore surface on the cap underside is yellowish. Fruitbodies are
1688:
439:
1617:
1485:
1222:
Sha JZ, Zhang D, Zeng XD, Wang JP (2011). "Heavy metal contents of five kinds in
Boletaceae collected in Xichang City and the safety evaluation".
1183:
1067:
1008:
968:
928:
896:
254:
is up to 10 cm (3.9 in) long and 3 cm (1.2 in) wide, with brown scabers on an underlying yellowish surface. It has firm
1673:
427:). The cap flesh is bilateral and inamyloid. The cystidia on the pores are present as conspicuous pleuro- and cheilocystidia. The
1668:
1678:
643:
450:
solution turns the cap cuticle a reddish color or is unreactive, and yellow or unreactive on the flesh. A drop of dilute
250:
up to 15 cm (5.9 in) in diameter. In age, the cap surface becomes wrinkled, often revealing white cracks. The
45:
1542:
721:
353:
519:
338:
290:
1693:
1300:
Halling RE, Osmundson TW, Neves MA (2008). "Pacific boletes: Implications for biogeographic relationships".
587:
505:
483:
1683:
1427:
1355:
1126:
750:
1663:
140:
458:
solution produces a gray color on the cap surface, and greenish-gray to olive coloration on the flesh.
1257:
Rodríguez-Ramírez EC, Moreno CE (2010). "Bolete diversity in two relict forests of the
Mexican beech (
1609:
1477:
493:
347:
234:
fungus. It is found in Asia, North
America, Central America, and South America, where it grows in an
1356:"Observations of the ectomycorrhizal roots collected from the bases of the basidiocarps in Chungbuk"
836:
811:
773:
666:
1529:
451:
424:
278:
271:
157:
1397:
1378:
1239:
1153:
1107:
793:
447:
304:
166:
40:
1658:
1591:
1550:
1459:
1317:
1282:
1179:
1063:
1004:
1000:
993:
964:
924:
892:
607:
372:
247:
1235:
1555:
1370:
1309:
1274:
1231:
1145:
1099:
785:
555:
455:
432:
395:
389:
255:
251:
32:
630:
532:
491:
by its dark brown to dark reddish-brown color, and flesh that does not stain with injury.
259:
1030:
523:, "from which it is easily separated by its dry pileus, smaller tubes and stouter stem."
1604:
1472:
571:
361:
286:
235:
87:
1652:
1243:
541:
454:(KOH) turns the cap surface red, and the flesh yellowish to orangish. Application of
77:
1382:
1335:
Adhikari MK. (2001). "New record of fleshy fungi from
Kathmandu valley, Nepal – 4".
1157:
501:, however, it also stains at the stipe base, and the cap is sticky rather than dry.
1464:
1374:
408:
1596:
1582:
1450:
1313:
1524:
1498:
625:
in the United States. The distribution extends south to Mexico, Costa Rica, and
618:
428:
404:
376:
312:
294:
243:
208:
117:
1149:
862:
692:
478:
412:
107:
416:
97:
57:
1321:
1286:
1178:. Morgantown, West Virginia: West Virginia University Press. p. 169.
1630:
1576:
1511:
1444:
626:
622:
544:
420:
342:
299:
275:
1278:
1622:
1490:
1111:
797:
614:
595:
579:
1503:
1635:
1516:
1199:
Asawa K, Yoshimoto T (1983). "Isolation of mannitol from a mushroom (
603:
231:
67:
1421:
1103:
789:
961:
North
American Boletes: A Color Guide to the Fleshy Pored Mushrooms
999:. Ann Arbor, Michigan: The University of Michigan Press. pp.
554:
465:
394:
368:
1205:
Bulletin of the
Yamaguchi Agricultural Experiment Station (Japan)
963:. Syracuse, New York: Syracuse University Press. pp. 213–4.
599:
1425:
1062:. Urbana, Illinois: University of Illinois Press. p. 254.
575:
239:
610:
exceeded that of national safety standards for edible fungi.
327:
are now obsolete genera that have since been subsumed into
629:. It is one of several boletes that have a north to south
341:
in a grouping of species that are associated with oak and
891:. Austin, Texas: University of Texas Press. p. 232.
1176:
Macrofungi
Associated with Oaks of Eastern North America
1024:
1022:
1020:
921:
Common
Mushrooms of the Talamanca Mountains, Costa Rica
487:
is a closely allied species. It is distinguished from
509:
is a lookalike that is difficult to distinguish from
411:
are spindle shaped, measuring 15–19 long by 5–6
371:
roots for "rough" and "head", refers to its wrinkled
693:"Nomenclatural problems concerning the generic name
1566:
1434:
923:. New York: New York Botanical Garden. p. 62.
262:, although opinions vary as to their desirability.
992:
613:The bolete is found from eastern Canada south to
954:
952:
950:
948:
946:
944:
942:
940:
442:can be used to help verify an identification of
1081:
1079:
914:
912:
910:
908:
1169:
1167:
686:
684:
660:
658:
8:
1354:Lee SS, Ching HC, Kim DH, Heyser W (2000).
1053:
1051:
986:
984:
982:
980:
744:
742:
1422:
959:Bessette AR, Bessette A, Roody WC (2000).
31:
20:
1203:) and its infrared absorption spectrum".
407:color ranges from brown to olive-brown.
1131:(Basidiomycota, Boletales) from Belize"
654:
1236:10.4028/www.scientific.net/AMR.422.555
415:. They have a smooth surface, and are
755:Bulletin of the New York State Museum
399:Spores are smooth and spindle shaped.
319:, published by Josef Šutara in 1982.
7:
751:"Report of the state botanist, 1904"
289:collection was made in the woods of
1125:Ortiz-Santana B, Halling R (2009).
843:. Index Fungorum. CAB International
818:. Index Fungorum. CAB International
274:scientifically in 1904 by American
431:is present as a hymeniform layer.
379:known as the "wrinkled Leccinum".
345:. Others in this grouping include
14:
1689:Taxa named by Charles Horton Peck
44:
1396:Fu SZ, Wang QB, Yao YJ (2006).
919:Halling RE, Mueller GM (2005).
667:"Das System der Agaricales. II"
547:is present in the fruitbodies.
1375:10.1080/12298093.2000.12015725
889:Texas Mushrooms: A Field Guide
644:List of North American boletes
1:
887:Metzler V, Metzler S (2010).
774:"New Boletaceae from Florida"
1314:10.1016/j.mycres.2007.11.021
578:. In eastern North America,
574:fungus that associates with
367:, which is derived from the
1398:"An annotated checklist of
1224:Advanced Materials Research
594:fruitbodies, the levels of
1710:
1267:American Journal of Botany
1090:species from Costa Rica".
1150:10.1007/s12228-008-9067-3
1086:Halling RE. (1999). "New
1058:Kuo M, Methven A (2014).
590:in boletes found that in
520:Hemileccinum rubropunctum
317:Krombholziella rugosiceps
197:Krombholziella rugosiceps
172:
165:
146:
139:
41:Scientific classification
39:
30:
23:
1674:Fungi of Central America
1060:Mushrooms of the Midwest
551:Habitat and distribution
291:Port Jefferson, New York
226:, commonly known as the
1669:Fungi described in 1904
471:Leccinellum crocipodium
246:have convex, yellowish
1679:Fungi of North America
568:Leccinellum rugosiceps
564:
537:Leccinellum rugosiceps
484:Leccinum neotropicalis
474:
400:
335:Leccinellum rugosiceps
309:Krombholzia rugosiceps
270:The species was first
223:Leccinellum rugosiceps
189:Krombholzia rugosiceps
150:Leccinellum rugosiceps
25:Leccinellum rugosiceps
558:
469:
398:
354:L. pseudoscabrum
1302:Mycological Research
1037:. MushroomExpert.com
995:100 Edible Mushrooms
869:. MushroomExpert.com
200:(Peck) Šutara (1982)
192:(Peck) Singer (1942)
1436:Leccinum rugosiceps
1279:10.3732/ajb.0900284
1201:Leccinum rugosiceps
1033:Leccinum rugosiceps
861:Kuo M. (May 2007).
761:: 5–58 (see p. 20).
724:Leccinum rugosiceps
506:L. crocipodium
452:potassium hydroxide
279:Charles Horton Peck
205:Leccinum rugosiceps
1568:Boletus rugosiceps
1174:Binion D. (2008).
1127:"A new species of
772:Singer R. (1945).
691:Šutara J. (1982).
671:Annales Mycologici
665:Singer R. (1942).
592:L. rugosiceps
565:
561:L. rugosiceps
515:L. rugosiceps
511:L. rugosiceps
499:L. rugosiceps
489:L. rugosiceps
475:
448:ammonium hydroxide
444:L. rugosiceps
401:
297:transferred it to
283:Boletus rugosiceps
230:, is a species of
181:Boletus rugosiceps
132:L. rugosiceps
1646:
1645:
1551:Open Tree of Life
1428:Taxon identifiers
1259:Fagus grandifolia
1185:978-1-933202-36-5
1069:978-0-252-07976-4
1010:978-0-472-03126-9
970:978-0-8156-0588-1
930:978-0-89327-460-3
898:978-0-292-78626-4
837:"Record Details:
812:"Record Details:
749:Peck CH. (1905).
584:Quercus palustris
433:Clamp connections
238:association with
228:wrinkled Leccinum
219:
218:
201:
193:
185:
16:Species of fungus
1701:
1639:
1638:
1626:
1625:
1613:
1612:
1600:
1599:
1587:
1586:
1585:
1559:
1558:
1546:
1545:
1533:
1532:
1520:
1519:
1507:
1506:
1494:
1493:
1481:
1480:
1468:
1467:
1455:
1454:
1453:
1423:
1417:
1416:
1406:
1393:
1387:
1386:
1360:
1351:
1345:
1344:
1332:
1326:
1325:
1297:
1291:
1290:
1254:
1248:
1247:
1219:
1213:
1212:
1196:
1190:
1189:
1171:
1162:
1161:
1135:
1122:
1116:
1115:
1083:
1074:
1073:
1055:
1046:
1045:
1043:
1042:
1026:
1015:
1014:
998:
988:
975:
974:
956:
935:
934:
916:
903:
902:
884:
878:
877:
875:
874:
858:
852:
851:
849:
848:
833:
827:
826:
824:
823:
808:
802:
801:
769:
763:
762:
746:
737:
736:
734:
733:
718:
712:
711:
701:
700:(PDF (abstract))
688:
679:
678:
662:
559:A collection of
494:L. viscosum
456:iron(II) sulfate
425:Melzer's reagent
362:specific epithet
348:L. albellum
199:
191:
183:
152:
49:
48:
35:
21:
1709:
1708:
1704:
1703:
1702:
1700:
1699:
1698:
1649:
1648:
1647:
1642:
1634:
1629:
1621:
1616:
1608:
1603:
1595:
1590:
1581:
1580:
1575:
1562:
1554:
1549:
1541:
1536:
1528:
1523:
1515:
1510:
1502:
1497:
1489:
1484:
1476:
1471:
1463:
1458:
1449:
1448:
1443:
1430:
1420:
1404:
1395:
1394:
1390:
1358:
1353:
1352:
1348:
1334:
1333:
1329:
1299:
1298:
1294:
1256:
1255:
1251:
1221:
1220:
1216:
1198:
1197:
1193:
1186:
1173:
1172:
1165:
1133:
1124:
1123:
1119:
1104:10.2307/4110871
1085:
1084:
1077:
1070:
1057:
1056:
1049:
1040:
1038:
1028:
1027:
1018:
1011:
991:Kuo M. (2007).
990:
989:
978:
971:
958:
957:
938:
931:
918:
917:
906:
899:
886:
885:
881:
872:
870:
860:
859:
855:
846:
844:
835:
834:
830:
821:
819:
810:
809:
805:
790:10.2307/3755143
771:
770:
766:
748:
747:
740:
731:
729:
720:
719:
715:
704:Ceská Mykologie
699:
690:
689:
682:
664:
663:
656:
652:
640:
572:ectomycorrhizal
553:
529:
464:
462:Similar species
385:
311:, published by
268:
236:ectomycorrhizal
215:
214:
176:
161:
154:
148:
135:
43:
17:
12:
11:
5:
1707:
1705:
1697:
1696:
1694:Fungus species
1691:
1686:
1681:
1676:
1671:
1666:
1661:
1651:
1650:
1644:
1643:
1641:
1640:
1627:
1614:
1601:
1588:
1572:
1570:
1564:
1563:
1561:
1560:
1547:
1534:
1521:
1508:
1495:
1482:
1469:
1456:
1440:
1438:
1432:
1431:
1426:
1419:
1418:
1388:
1346:
1327:
1292:
1265:; Fagaceae)".
1249:
1214:
1191:
1184:
1163:
1117:
1075:
1068:
1047:
1016:
1009:
976:
969:
936:
929:
904:
897:
879:
853:
839:Krombholziella
828:
803:
764:
738:
726:(Peck) Singer"
713:
695:Krombholziella
680:
653:
651:
648:
647:
646:
639:
636:
552:
549:
528:
525:
473:is a lookalike
463:
460:
440:chemical tests
384:
381:
325:Krombholziella
267:
264:
217:
216:
213:
212:
202:
194:
186:
177:
174:
173:
170:
169:
163:
162:
160:) C. Hahn 2020
155:
144:
143:
137:
136:
129:
127:
123:
122:
115:
111:
110:
105:
101:
100:
95:
91:
90:
88:Agaricomycetes
85:
81:
80:
75:
71:
70:
65:
61:
60:
55:
51:
50:
37:
36:
28:
27:
15:
13:
10:
9:
6:
4:
3:
2:
1706:
1695:
1692:
1690:
1687:
1685:
1684:Fungi of Asia
1682:
1680:
1677:
1675:
1672:
1670:
1667:
1665:
1662:
1660:
1657:
1656:
1654:
1637:
1632:
1628:
1624:
1619:
1615:
1611:
1606:
1602:
1598:
1593:
1589:
1584:
1578:
1574:
1573:
1571:
1569:
1565:
1557:
1552:
1548:
1544:
1539:
1535:
1531:
1526:
1522:
1518:
1513:
1509:
1505:
1500:
1496:
1492:
1487:
1483:
1479:
1474:
1470:
1466:
1461:
1457:
1452:
1446:
1442:
1441:
1439:
1437:
1433:
1429:
1424:
1414:
1410:
1403:
1401:
1392:
1389:
1384:
1380:
1376:
1372:
1368:
1364:
1357:
1350:
1347:
1342:
1338:
1331:
1328:
1323:
1319:
1315:
1311:
1308:(4): 437–47.
1307:
1303:
1296:
1293:
1288:
1284:
1280:
1276:
1272:
1268:
1264:
1260:
1253:
1250:
1245:
1241:
1237:
1233:
1229:
1225:
1218:
1215:
1211:(11): 810–14.
1210:
1206:
1202:
1195:
1192:
1187:
1181:
1177:
1170:
1168:
1164:
1159:
1155:
1151:
1147:
1143:
1139:
1132:
1130:
1121:
1118:
1113:
1109:
1105:
1101:
1098:(3): 747–53.
1097:
1093:
1089:
1082:
1080:
1076:
1071:
1065:
1061:
1054:
1052:
1048:
1036:
1034:
1025:
1023:
1021:
1017:
1012:
1006:
1002:
997:
996:
987:
985:
983:
981:
977:
972:
966:
962:
955:
953:
951:
949:
947:
945:
943:
941:
937:
932:
926:
922:
915:
913:
911:
909:
905:
900:
894:
890:
883:
880:
868:
866:
857:
854:
842:
840:
832:
829:
817:
815:
807:
804:
799:
795:
791:
787:
783:
779:
775:
768:
765:
760:
756:
752:
745:
743:
739:
727:
725:
717:
714:
709:
705:
698:
696:
687:
685:
681:
676:
673:(in German).
672:
668:
661:
659:
655:
649:
645:
642:
641:
637:
635:
632:
628:
624:
620:
616:
611:
609:
605:
601:
597:
593:
589:
585:
581:
577:
573:
569:
562:
557:
550:
548:
546:
543:
542:sugar alcohol
538:
534:
526:
524:
522:
521:
516:
512:
508:
507:
502:
500:
496:
495:
490:
486:
485:
480:
472:
468:
461:
459:
457:
453:
449:
445:
441:
436:
434:
430:
426:
422:
418:
414:
410:
406:
397:
393:
391:
382:
380:
378:
374:
370:
366:
363:
358:
356:
355:
350:
349:
344:
340:
336:
332:
330:
326:
322:
318:
315:in 1942, and
314:
310:
306:
302:
301:
296:
292:
288:
284:
280:
277:
273:
265:
263:
261:
257:
253:
249:
245:
241:
237:
233:
229:
225:
224:
210:
206:
203:
198:
195:
190:
187:
182:
179:
178:
171:
168:
164:
159:
153:
151:
145:
142:
141:Binomial name
138:
134:
133:
128:
125:
124:
121:
120:
116:
113:
112:
109:
106:
103:
102:
99:
96:
93:
92:
89:
86:
83:
82:
79:
78:Basidiomycota
76:
73:
72:
69:
66:
63:
62:
59:
56:
53:
52:
47:
42:
38:
34:
29:
26:
22:
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1664:Edible fungi
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1273:(5): 893–8.
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1144:(2): 172–4.
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1092:Kew Bulletin
1091:
1087:
1059:
1039:. Retrieved
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920:
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871:. Retrieved
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784:(6): 797–9.
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588:heavy metals
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24:
18:
1525:NatureServe
1499:iNaturalist
1369:(2): 62–9.
1363:Mycobiology
1337:Green World
863:"The genus
814:Krombholzia
722:"Mycobank:
710:(2): 77–84.
619:Mississippi
563:fruitbodies
479:Costa Rican
429:cap cuticle
419:(i.e., not
405:spore print
383:Description
321:Krombholzia
313:Rolf Singer
295:Rolf Singer
244:Fruitbodies
184:Peck (1904)
119:Leccinellum
1653:Categories
1230:: 555–60.
1041:2015-02-20
873:2015-02-20
847:2015-02-06
822:2015-02-06
732:2024-08-05
728:. Mycobank
650:References
621:, west to
517:grew with
365:rugosiceps
339:classified
276:mycologist
108:Boletaceae
74:Division:
1583:Q59537266
1530:2.1072300
1451:Q20720954
1409:Mycotaxon
1402:in China"
1343:(1): 3–7.
1244:137366362
1138:Brittonia
816:P. Karst"
778:Mycologia
697:R. Maire"
535:species,
527:Edibility
417:inamyloid
303:in 1945.
272:described
126:Species:
98:Boletales
64:Kingdom:
58:Eukaryota
1659:Leccinum
1631:MycoBank
1605:Fungorum
1577:Wikidata
1512:MycoBank
1473:Fungorum
1445:Wikidata
1415:: 47–50.
1400:Leccinum
1383:86246335
1322:18316181
1287:21622453
1263:mexicana
1158:41438436
1129:Leccinum
1088:Leccinum
865:Leccinum
638:See also
627:Colombia
623:Michigan
545:mannitol
438:Several
421:staining
377:commonly
375:. It is
343:hornbeam
329:Leccinum
307:include
305:Synonyms
300:Leccinum
266:Taxonomy
167:Synonyms
104:Family:
54:Domain:
1623:6015799
1491:3354356
1112:4110871
1029:Kuo M.
798:3755143
615:Florida
608:mercury
596:cadmium
580:pin oak
481:bolete
207:(Peck)
114:Genus:
94:Order:
84:Class:
1636:195004
1610:195004
1556:504054
1543:279511
1517:287524
1504:334866
1478:287524
1381:
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1007:
967:
927:
895:
841:Maire"
796:
631:clinal
606:, and
604:copper
570:is an
533:edible
409:Spores
285:. The
260:edible
232:bolete
211:(1945)
209:Singer
1465:3STF8
1405:(PDF)
1379:S2CID
1359:(PDF)
1261:var.
1240:S2CID
1154:S2CID
1134:(PDF)
1108:JSTOR
1001:186–9
794:JSTOR
677:: 34.
423:with
390:stipe
369:Latin
256:flesh
252:stipe
68:Fungi
1618:GBIF
1597:MD3W
1538:NCBI
1486:GBIF
1318:PMID
1283:PMID
1180:ISBN
1064:ISBN
1005:ISBN
965:ISBN
925:ISBN
893:ISBN
617:and
600:zinc
477:The
403:The
360:The
351:and
323:and
287:type
248:caps
175:List
158:Peck
1592:CoL
1460:CoL
1371:doi
1310:doi
1306:112
1275:doi
1232:doi
1228:422
1146:doi
1100:doi
786:doi
576:oak
531:An
373:cap
337:is
281:as
240:oak
1655::
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