102:
124:
2799:
553:
304:
340:
Members of the family are generally detritivorous or scavengers feeding on carrion or decaying organic matter like dung, or are specialised on feeding on specific types of fungus. Many species have reduced wings, with about half of all described species being flightless.
309:
308:
305:
332:
due to the globular shape of many species, although some are more elongated in shape. They are generally small or very small beetles (less than 10 mm in length) and many (but not all) species have clubbed
310:
307:
2660:
2699:
2471:
306:
2840:
2634:
2673:
584:
2771:
2678:
101:
2686:
2833:
2556:
2533:
2613:
123:
2704:
2626:
220:
2874:
2869:
577:
379:
295:
2864:
2826:
540:
Web-based taxonomy pages ("scratchpad") for the
Cholevinae of the world (here treated as a separate family)
2486:
351:
2735:
2561:
463:
399:
Bouchard, Patrice; Bousquet, Yves; Davies, Anthony E.; Alonso-Zarazaga, Miguel A.; et al. (2011).
2753:
2776:
2714:
2595:
1723:
501:
408:
1942:
570:
346:
244:
110:
2859:
517:
489:
118:
2740:
236:
328:
with around 3800 described species found worldwide. Members of this family are commonly called
260:
2722:
2582:
2211:
1628:
490:"Two new polyphagan beetles (Tenebrionidae, Leiodidae) from lower Cenomanian amber of Myanmar"
444:
426:
334:
2810:
2763:
2665:
2727:
2308:
2087:
2082:
2005:
1763:
779:
509:
434:
416:
195:
1997:
1950:
1795:
1779:
1455:
1215:
1184:
1099:
1017:
908:
876:
2524:
557:
505:
412:
2621:
2608:
2400:
2351:
2206:
2013:
1681:
1665:
1604:
1499:
1470:
1398:
1362:
1346:
1262:
1207:
1112:
1009:
858:
820:
800:
696:
439:
276:
205:
534:
2853:
2382:
2186:
2177:
1918:
1889:
1881:
1873:
1847:
1822:
1718:
1705:
1644:
1562:
1465:
1460:
1341:
1275:
1246:
1230:
1194:
1164:
1117:
1071:
1038:
954:
946:
917:
755:
521:
367:
59:
2798:
2587:
2359:
2283:
2029:
2021:
1976:
1960:
1910:
1831:
1787:
1739:
1657:
1570:
1439:
1426:
1406:
1367:
1267:
1254:
1225:
1159:
1147:
1043:
1000:
983:
975:
925:
763:
733:
664:
653:
637:
400:
288:
513:
2538:
2691:
2647:
2518:
2445:
2416:
2390:
2377:
2317:
2275:
2259:
2251:
2243:
2227:
2219:
2155:
2113:
2105:
2074:
1989:
1968:
1934:
1926:
1839:
1771:
1755:
1747:
1710:
1697:
1673:
1591:
1583:
1533:
1525:
1483:
1447:
1418:
1390:
1385:
1301:
1296:
1288:
1238:
1174:
1169:
1083:
1048:
1033:
1025:
941:
933:
750:
34:
552:
2372:
2333:
2325:
2267:
2130:
2092:
2045:
2037:
1857:
1805:
1689:
1620:
1491:
1475:
1431:
1372:
1336:
1316:
1280:
1189:
1179:
1053:
787:
771:
363:
252:
79:
44:
2509:
562:
430:
2437:
2429:
2408:
2364:
2163:
2138:
2066:
1894:
1865:
1549:
1380:
1354:
1306:
1220:
1122:
1091:
1063:
896:
871:
866:
847:
812:
701:
684:
676:
625:
613:
359:
268:
185:
155:
135:
84:
448:
421:
2569:
2503:
2341:
2291:
2235:
2194:
2143:
1800:
1731:
1612:
1324:
1130:
1104:
962:
795:
742:
721:
631:
355:
74:
69:
54:
49:
39:
2548:
2574:
2058:
2050:
1902:
1636:
1557:
1541:
89:
64:
2652:
1649:
1509:
1142:
619:
607:
594:
325:
175:
165:
145:
2639:
2480:
539:
366:, with Cretaceous members of the family being primarily known from
2748:
302:
2484:
893:
844:
718:
650:
566:
2600:
965:(furniture beetles, death watch beetles, spider beetles)
2814:
2493:
2399:
2350:
2316:
2307:
2185:
2176:
2129:
2104:
1988:
1959:
1856:
1830:
1821:
1603:
1582:
1508:
1315:
1206:
1062:
1008:
999:
974:
916:
907:
732:
488:Bao, Tong; Antunes-Carvalho, Caio (December 2020).
362:). Members of modern subfamilies appear during the
1842:(jewel beetles, or metallic wood-boring beetles)
1565:(true weevils, bark beetles, ambrosia beetles)
464:"Leiodidae Fleming, 1821, Coleoptera, Beetles"
2834:
578:
8:
1808:(ironclad beetles, cylindrical bark beetles)
401:"Family-group names in Coleoptera (Insecta)"
2841:
2827:
2481:
2313:
2182:
1827:
1005:
913:
904:
890:
841:
729:
715:
647:
585:
571:
563:
100:
20:
2472:List of subgroups of the order Coleoptera
1552:(straight snout weevils, New York weevil)
438:
420:
1979:(cicada beetle, cicada parasite beetles)
391:
2069:(false fireflies, long-lipped beetles)
7:
2795:
2793:
2754:D85C87B3-461C-EA06-C4DD-ECB71D2EE065
2715:7cddb66d-0c2b-4967-93c1-49f7ac79a921
2627:b38f6aa3-0aee-4521-beb1-61769e8d2968
2095:(false metallic wood-boring beetles)
466:. In Schmidt-Rhaesa, Andreas (ed.).
344:The oldest fossil of the family is
2813:. You can help Knowledge (XXG) by
14:
2797:
1706:Perimylopidae, or Promecheilidae
1291:(minute brown scavenger beetles)
551:
122:
1921:(variegated mud-loving beetles)
1:
1750:(narrow-waisted bark beetles)
1442:(parasitic flat bark beetles)
1393:(short-winged flower beetles)
1249:(lady beetles, or God's cows)
986:(tooth-necked fungus beetles)
514:10.1016/j.cretres.2020.104599
2262:(sand-loving scarab beetles)
1652:(minute tree-fungus beetles)
1478:(silvanid flat bark beetles)
470:. Berlin, Boston: De Gruyter
354:, Mongolia, dating the Late
2411:(primitive carrion beetles)
2367:(minute mud-loving beetles)
2230:(bumble bee scarab beetles)
2000:(soft-bodied plant beetles)
1971:(soft bodied plant beetles)
774:(predaceous diving beetles)
2891:
2792:
2246:(scavenger scarab beetles)
2238:(enigmatic scarab beetles)
1411:Lamingtonium binnaberrense
1125:(soft-wing flower beetles)
468:Handbook of Zoology Online
462:Newton, Alfred F. (2016).
2469:
2385:(water scavenger beetles)
2118:Rhinorhipus tamborinensis
1897:(long-toed water beetles)
1790:(polypore fungus beetles)
1692:(palm and flower beetles)
1676:(tumbling flower beetles)
1623:(ant-like flower beetles)
1401:(lined flat bark beetles)
1375:(pleasing fungus beetles)
1270:(handsome fungus beetles)
903:
889:
854:
840:
815:(burrowing water beetles)
728:
714:
660:
646:
602:
294:
287:
233:
228:
119:Scientific classification
117:
108:
99:
23:
2432:(feather-winged beetles)
1766:(false longhorn beetles)
1713:(jugular-horned beetles)
1660:(false darkling beetles)
1631:(cryptic fungus beetles)
1450:(shining flower beetles)
790:(crawling water beetles)
689:Sikhotealinia zhiltzovae
380:List of Leiodidae genera
1913:(forest stream beetles)
1774:(synchroa bark beetles)
1734:(dead log bark beetles)
1726:(fire-coloured beetles)
1615:(ant-like leaf beetles)
1349:(silken fungus beetles)
1283:(well polished beetles)
1257:(minute fungus beetles)
1076:Acanthocnemus nigricans
2809:-related article is a
2424:(round fungus beetles)
1758:(false flower beetles)
1742:(wedge-shaped beetles)
1700:(false blister beetle)
1684:(hairy fungus beetles)
1639:(conifer bark beetles)
1536:(leaf-rolling weevils)
1197:(bark-gnawing beetles)
957:(wounded-tree beetles)
823:(false ground beetles)
745:(trout-stream beetles)
422:10.3897/zookeys.88.807
318:
2736:Paleobiology Database
2336:(false clown beetles)
2158:(plate-thigh beetles)
2040:(false click beetles)
1945:(water-penny beetles)
1594:(ship-timber beetles)
1573:(pine flower weevils)
1421:(root-eating beetles)
1241:(minute bark beetles)
1152:Phloiophilus edwardsi
679:(reticulated beetles)
313:
2622:Fauna Europaea (new)
2214:(false stag beetles)
2199:Belohina inexpectata
2061:(net-winged beetles)
2024:(rare click beetles)
1937:(travertine beetles)
1929:(minute mud beetles)
1494:(dry-fungus beetles)
1329:Agapytho foveicollis
1086:(false skin beetles)
949:(Jacobson's beetles)
669:Crowsoniella relicta
330:round fungus beetles
2212:Diphyllostomatidae
1544:(primitive weevils)
1357:(flat bark beetles)
1107:(checkered beetles)
1094:(fruitworm beetles)
782:(whirligig beetles)
506:2020CrRes.11604599B
494:Cretaceous Research
413:2011ZooK...88....1B
296:at least 380 genera
111:Anisotoma humeralis
2077:(glowworm beetles)
1782:(darkling beetles)
1629:Archeocrypticidae
1486:(palmetto beetles)
1233:(dry bark beetles)
1020:(longhorn beetles)
535:Family description
319:
2822:
2821:
2787:
2786:
2723:Open Tree of Life
2487:Taxon identifiers
2478:
2477:
2465:
2464:
2461:
2460:
2457:
2456:
2449:
2441:
2440:(carrion beetles)
2433:
2425:
2412:
2386:
2368:
2337:
2329:
2303:
2302:
2295:
2287:
2279:
2271:
2263:
2255:
2247:
2239:
2231:
2223:
2215:
2202:
2172:
2171:
2159:
2151:
2121:
2096:
2078:
2070:
2062:
2054:
2041:
2033:
2025:
2017:
2016:(soldier beetles)
2009:
2006:Brachypsectridae
2001:
1980:
1972:
1946:
1938:
1930:
1922:
1914:
1906:
1898:
1885:
1877:
1869:
1843:
1817:
1816:
1809:
1791:
1783:
1775:
1767:
1764:Stenotrachelidae
1759:
1751:
1743:
1735:
1727:
1714:
1701:
1693:
1685:
1677:
1669:
1668:(blister beetles)
1661:
1653:
1640:
1632:
1624:
1616:
1595:
1574:
1566:
1553:
1545:
1537:
1529:
1519:
1495:
1487:
1479:
1451:
1443:
1435:
1422:
1414:
1402:
1394:
1376:
1358:
1350:
1332:
1292:
1284:
1271:
1258:
1250:
1242:
1234:
1216:Akalyptoischiidae
1198:
1155:
1138:
1126:
1108:
1095:
1087:
1079:
1029:
1021:
995:
994:
987:
966:
958:
950:
937:
929:
885:
884:
862:
836:
835:
832:
831:
824:
816:
808:
791:
783:
775:
767:
759:
746:
710:
709:
692:
680:
672:
311:
301:
300:
281:
273:
265:
257:
249:
241:
224:
16:Family of beetles
2882:
2843:
2836:
2829:
2801:
2794:
2780:
2779:
2767:
2766:
2757:
2756:
2744:
2743:
2731:
2730:
2718:
2717:
2708:
2707:
2695:
2694:
2692:NHMSYS0001718790
2682:
2681:
2669:
2668:
2656:
2655:
2643:
2642:
2630:
2629:
2617:
2616:
2604:
2603:
2591:
2590:
2578:
2577:
2565:
2564:
2552:
2551:
2542:
2541:
2529:
2528:
2527:
2514:
2513:
2512:
2482:
2447:
2439:
2431:
2423:
2410:
2384:
2366:
2335:
2327:
2314:
2309:Staphyliniformia
2293:
2285:
2277:
2269:
2261:
2253:
2245:
2237:
2229:
2221:
2213:
2196:
2183:
2157:
2148:Declinia relicta
2145:
2115:
2094:
2088:Sinopyrophoridae
2083:Rhagophthalmidae
2076:
2068:
2060:
2052:
2039:
2031:
2023:
2015:
2007:
1999:
1998:Artematopodidae
1978:
1970:
1944:
1936:
1928:
1920:
1912:
1905:(riffle beetles)
1904:
1896:
1884:(turtle beetles)
1883:
1875:
1867:
1841:
1828:
1807:
1789:
1781:
1773:
1765:
1757:
1749:
1741:
1733:
1725:
1712:
1699:
1691:
1683:
1675:
1667:
1659:
1651:
1638:
1630:
1622:
1614:
1593:
1572:
1564:
1551:
1543:
1535:
1528:(fungus weevils)
1527:
1518:
1517:
1513:
1500:Tasmosalpingidae
1493:
1485:
1477:
1449:
1441:
1433:
1420:
1408:
1400:
1392:
1374:
1356:
1348:
1326:
1290:
1282:
1269:
1256:
1248:
1240:
1232:
1196:
1149:
1132:
1124:
1106:
1093:
1085:
1073:
1072:Acanthocnemidae
1027:
1019:
1006:
985:
964:
956:
948:
935:
927:
914:
905:
891:
860:
842:
822:
814:
802:
789:
781:
773:
765:
758:(ground beetles)
757:
744:
730:
716:
686:
678:
666:
665:Crowsoniellidae
648:
587:
580:
573:
564:
556:Data related to
555:
526:
525:
485:
479:
478:
476:
475:
459:
453:
452:
442:
424:
396:
312:
279:
271:
263:
255:
247:
239:
219:
196:Staphyliniformia
127:
126:
104:
94:
31:
30:Tithonian–Recent
27:Temporal range:
21:
2890:
2889:
2885:
2884:
2883:
2881:
2880:
2879:
2875:Leiodidae stubs
2870:Beetle families
2850:
2849:
2848:
2847:
2790:
2788:
2783:
2775:
2770:
2762:
2760:
2752:
2747:
2739:
2734:
2726:
2721:
2713:
2711:
2703:
2698:
2690:
2685:
2677:
2672:
2664:
2659:
2651:
2646:
2638:
2633:
2625:
2620:
2612:
2607:
2599:
2594:
2586:
2581:
2573:
2568:
2560:
2555:
2547:
2545:
2537:
2532:
2523:
2522:
2517:
2508:
2507:
2502:
2489:
2479:
2474:
2453:
2395:
2346:
2328:(clown beetles)
2299:
2270:(betsy beetles)
2178:Scarabaeiformia
2168:
2125:
2100:
2032:(click beetles)
2008:(Texas beetles)
1984:
1955:
1951:Ptilodactylidae
1876:(cedar beetles)
1852:
1813:
1796:Trictenotomidae
1682:Mycetophagidae
1599:
1578:
1515:
1514:
1512:
1504:
1456:Phloeostichidae
1407:Lamingtoniidae
1399:Laemophloeidae
1347:Cryptophagidae
1311:
1202:
1185:Thanerocleridae
1148:Phloiophilidae
1135:Metaxina ornata
1100:Chaetosomatidae
1058:
991:
970:
928:(auger beetles)
909:Bostrichiformia
899:
881:
877:Torridincolidae
861:(skiff beetles)
859:Hydroscaphidae
850:
828:
821:Trachypachidae
766:(tiger beetles)
724:
706:
656:
642:
598:
591:
560:at Wikispecies
549:
544:
530:
529:
487:
486:
482:
473:
471:
461:
460:
456:
398:
397:
393:
388:
376:
324:is a family of
303:
218:
121:
95:
93:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
37:
29:
28:
25:
17:
12:
11:
5:
2888:
2886:
2878:
2877:
2872:
2867:
2865:Staphylinoidea
2862:
2852:
2851:
2846:
2845:
2838:
2831:
2823:
2820:
2819:
2802:
2785:
2784:
2782:
2781:
2768:
2758:
2745:
2732:
2719:
2709:
2696:
2683:
2670:
2657:
2644:
2631:
2618:
2609:Fauna Europaea
2605:
2592:
2579:
2566:
2553:
2543:
2530:
2515:
2499:
2497:
2491:
2490:
2485:
2476:
2475:
2470:
2467:
2466:
2463:
2462:
2459:
2458:
2455:
2454:
2452:
2451:
2448:(rove beetles)
2446:Staphylinidae
2443:
2435:
2427:
2419:
2414:
2405:
2403:
2401:Staphylinoidea
2397:
2396:
2394:
2393:
2388:
2383:Hydrophilidae
2380:
2375:
2370:
2362:
2356:
2354:
2352:Hydrophiloidea
2348:
2347:
2345:
2344:
2339:
2331:
2322:
2320:
2311:
2305:
2304:
2301:
2300:
2298:
2297:
2294:(hide beetles)
2289:
2281:
2278:(rain beetles)
2273:
2265:
2257:
2254:(stag beetles)
2249:
2241:
2233:
2225:
2217:
2209:
2207:Bolboceratidae
2204:
2191:
2189:
2180:
2174:
2173:
2170:
2169:
2167:
2166:
2161:
2153:
2141:
2135:
2133:
2127:
2126:
2124:
2123:
2114:Rhinorhipidae
2110:
2108:
2106:Rhinorhipoidea
2102:
2101:
2099:
2098:
2090:
2085:
2080:
2072:
2064:
2056:
2048:
2043:
2035:
2027:
2019:
2011:
2003:
1994:
1992:
1986:
1985:
1983:
1982:
1974:
1965:
1963:
1957:
1956:
1954:
1953:
1948:
1940:
1932:
1924:
1919:Heteroceridae
1916:
1908:
1900:
1892:
1887:
1882:Chelonariidae
1879:
1874:Callirhipidae
1871:
1868:(pill beetles)
1862:
1860:
1854:
1853:
1851:
1850:
1845:
1836:
1834:
1825:
1819:
1818:
1815:
1814:
1812:
1811:
1803:
1798:
1793:
1785:
1780:Tenebrionidae
1777:
1769:
1761:
1753:
1745:
1737:
1729:
1721:
1716:
1708:
1703:
1695:
1687:
1679:
1671:
1663:
1655:
1647:
1642:
1634:
1626:
1618:
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2815:expanding it
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2804:
2789:
2494:
2421:
2373:Helophoridae
2365:Georissidae
2360:Epimetopidae
2276:Pleocomidae
2260:Ochodaeidae
2244:Hybosoridae
2228:Glaphyridae
2220:Geotrupidae
2198:
2195:Belohinidae
2156:Eucinetidae
2147:
2144:Decliniidae
2117:
2075:Phengodidae
2014:Cantharidae
1969:Dascillidae
1961:Dascilloidea
1943:Psephenidae
1935:Lutrochidae
1927:Limnichidae
1840:Buprestidae
1832:Buprestoidea
1772:Synchroidae
1756:Scraptiidae
1748:Salpingidae
1711:Prostomidae
1698:Oedemeridae
1674:Mordellidae
1592:Lymexylidae
1584:Lymexyloidea
1534:Attelabidae
1526:Anthribidae
1484:Smicripidae
1448:Phalacridae
1432:Nitidulidae
1427:Myraboliidae
1419:Monotomidae
1410:
1391:Kateretidae
1368:Cyclaxyridae
1328:
1325:Agapythidae
1289:Latridiidae
1239:Cerylonidae
1226:Anamorphidae
1160:Phycosecidae
1151:
1134:
1131:Metaxinidae
1084:Biphyllidae
1075:
1044:Orsodacnidae
1001:Cucujiformia
934:Dermestidae
804:
743:Amphizoidae
688:
668:
654:Archostemata
638:Holometabola
636:Superorder:
630:Infraclass:
550:
497:
493:
483:
472:. Retrieved
467:
457:
404:
394:
345:
343:
339:
329:
321:
320:
314:
229:Subfamilies
215:
192:Infraorder:
109:
18:
2648:iNaturalist
2519:Wikispecies
2417:Hydraenidae
2391:Spercheidae
2378:Hydrochidae
2342:Synteliidae
2326:Histeridae
2318:Histeroidea
2268:Passalidae
2236:Glaresidae
2093:Throscidae
2053:(fireflies)
2051:Lampyridae
2038:Eucnemidae
2030:Elateridae
1990:Elateroidea
1806:Zopheridae
1690:Mycteridae
1621:Anthicidae
1492:Sphindidae
1476:Silvanidae
1386:Hobartiidae
1373:Erotylidae
1302:Mycetaeidae
1297:Murmidiidae
1281:Euxestidae
1175:Rentoniidae
1049:Oxypeltidae
1034:Disteniidae
867:Lepiceridae
796:Hygrobiidae
788:Haliplidae
772:Dytiscidae
751:Aspidytidae
256:Kirby, 1837
248:Hatch, 1927
2854:Categories
2438:Silphidae
2430:Ptiliidae
2422:Leiodidae
2409:Agyrtidae
2252:Lucanidae
2131:Scirtoidea
2067:Omethidae
2046:Jurasaidae
1895:Dryopidae
1866:Byrrhidae
1858:Byrrhoidea
1550:Brentidae
1355:Cucujidae
1337:Boganiidae
1317:Cucujoidea
1190:Thymalidae
1180:Rhadalidae
1123:Melyridae
1092:Byturidae
1054:Vesperidae
963:Ptiniidae
813:Noteridae
780:Gyrinidae
756:Carabidae
685:Jurodidae
677:Cupedidae
624:Subclass:
614:Arthropoda
595:Coleoptera
500:: 104599.
474:2022-10-02
386:References
364:Cretaceous
264:Horn, 1880
253:Cholevinae
237:Camiarinae
182:Suborder:
176:Coleoptera
156:Arthropoda
2860:Leiodidae
2807:Leiodidae
2539:Leiodidae
2525:Leiodidae
2495:Leiodidae
2292:Trogidae
2286:(scarabs)
2164:Scirtidae
2139:Clambidae
1732:Pythidae
1666:Meloidae
1613:Aderidae
1516:(weevils)
1381:Helotidae
1307:Teredidae
1221:Alexiidae
1105:Cleridae
1064:Cleroidea
897:Polyphaga
895:Suborder
848:Myxophaga
846:Suborder
801:Meruidae
720:Suborder
702:Ommatidae
652:Suborder
626:Pterygota
606:Kingdom:
558:Leiodidae
522:224930238
431:1313-2989
360:Tithonian
322:Leiodidae
289:Diversity
269:Leiodinae
261:Coloninae
216:Leiodidae
186:Polyphaga
142:Kingdom:
136:Eukaryota
24:Leiodidae
2570:BugGuide
2546:BioLib:
2504:Wikidata
2059:Lycidae
1903:Elmidae
1801:Ulodidae
1637:Boridae
1542:Belidae
1143:Peltidae
736:families
722:Adephaga
632:Neoptera
612:Phylum:
608:Animalia
597:families
449:21594053
374:See also
356:Jurassic
352:Shar-Teg
335:antennae
212:Family:
152:Phylum:
146:Animalia
132:Domain:
2764:4779668
2510:Q132279
1650:Ciidae
1558:Caridae
620:Insecta
618:Class:
593:Extant
502:Bibcode
440:3088472
409:Bibcode
405:ZooKeys
326:beetles
221:Fleming
172:Order:
166:Insecta
162:Class:
2777:150757
2761:uBio:
2741:346356
2728:215452
2712:NZOR:
2679:113817
2666:100848
734:Extant
520:
447:
437:
429:
223:, 1821
2805:This
2772:WoRMS
2749:Plazi
2705:66535
2661:IRMNG
2653:60571
2614:11109
2588:8VR3J
2575:13865
2562:75123
518:S2CID
350:from
2811:stub
2700:NCBI
2674:ITIS
2640:4738
2635:GBIF
2557:BOLD
2549:5363
445:PMID
427:ISSN
35:PreꞒ
2687:NBN
2601:378
2596:EoL
2583:CoL
2534:AFD
510:doi
498:116
435:PMC
417:doi
2856::
2774::
2751::
2738::
2725::
2702::
2689::
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516:.
508:.
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492:.
443:.
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415:.
403:.
370:.
337:.
85:Pg
2842:e
2835:t
2828:v
2817:.
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2197:(
2150:)
2146:(
2120:)
2116:(
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1327:(
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1150:(
1137:)
1133:(
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803:(
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687:(
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667:(
586:e
579:t
572:v
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512::
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477:.
451:.
419::
411::
358:(
90:N
80:K
75:J
70:T
65:P
60:C
55:D
50:S
45:O
40:Ꞓ
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