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division, most notably CDC28, which encodes the yeast Cdk kinase. Other significant discoveries include introduction of the concept of cell cycle "checkpoints", which delay cell division when cellular insults are generated and also the identification and characterization of the mating signal transduction pathway.
381:
His earliest publications focused on the isolation of temperature sensitive yeast mutants disabled in basic biological processes, including DNA, RNA and protein synthesis. This led to the identification of the CDC (Cell
Division Cycle) genes, which function in promoting the progression through cell
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This award is given to scientists whose research in yeast has made the most impact in the broader areas of biology. Recipients of the award also give a lecture at the biennial Yeast
Genetics Meeting.
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256:
Working in yeast, Hartwell identified the fundamental role of checkpoints in cell cycle control, and CDC genes such as CDC28, which controls the start of the cycle—the progression through G1.
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394:, a non-profit organization dedicated to developing new technologies for the early detection of cancer. He is also a founding co-chair of the
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Arizona State
University and Virginia G. Piper Charitable Trust tap Nobel Prize winner Dr. Lee Hartwell to lead major health initiative
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1939:
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520:
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398:, and a member of its executive committee. In September 2009, it was announced that Hartwell would join the faculty of
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awarded the Medal of Merit, the state's highest honor, to
Hartwell. He is also a recipient of the
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1911:
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1325:
1309:
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406:'s Center for Sustainable Health with Dr. Michael Birt. He is also adjunct faculty at
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Lee
Hartwell, at 70, Tackles Personalized Medicine, Education in Latest Career Phase
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as the
Virginia G. Piper Chair of Personalized Medicine and co-director of the
292:
in 1968. In a series of experiments from 1970 to 1971, Hartwell discovered the
17:
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2464:
2030:
1289:
1237:
1184:
536:"Genetic control of the cell-division cycle in yeast. I. Detection of mutants"
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and in 1997 became its president and director until he retired in 2010.
234:
162:
633:
506:"The Nobel Prize in Physiology or Medicine 2001, Illustrated Lecture"
197:
191:
675:"Macromolecule synthesis in temperature-sensitive mutants of yeast"
390:
Hartwell is the
Chairman of the Scientific Advisory Board at the
2808:
Recipients of the Albert Lasker Award for Basic
Medical Research
229:(born October 30, 1939) is former president and director of the
2643:
2359:
816:
2788:
Massachusetts
Institute of Technology School of Science alumni
772:"Amrita News: Nobel-prize winner is adjunct faculty at Amrita"
471:
2002 – Leland
Hartwell, Fred Hutchinson Cancer Research Center
280:
in 1961. In 1964, he received his PhD in biology from the
198:
Studies on the induction of histidase in Bacillus subtilis
2803:
Members of the United States National Academy of Sciences
326:, Hartwell has received awards and honors including the
426:, Friedrich Miescher Institute for Biomedical Research
2724:
2677:
2536:
2505:
2474:
2450:
2426:
2395:
2013:
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1339:
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854:
314:in the genes are involved in some types of cancer.
208:
190:
168:
158:
87:
77:
63:
41:
34:
648:"The Official Site of Louisa Gross Horwitz Prize"
338:in 1987. In 1996, Hartwell joined the faculty of
660:Luke Timmerman for Xconomy. September 20, 2010
630:"Lee Hartwell's profile, Community of Science"
371:Komen Brinker Award for Scientific Distinction
347:Albert Lasker Award for Basic Medical Research
2823:Fred Hutchinson Cancer Research Center people
2655:
2371:
828:
722:Pacific Health Summit, About the Summit, 2011
489:"Leland H. Hartwell - A Superstar of Science"
8:
534:Hartwell LH, Culotti J, Reid B (June 1970).
310:). These genes regulate the cell cycle and
2662:
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2640:
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31:
2783:California Institute of Technology alumni
2778:Nobel laureates in Physiology or Medicine
698:
569:
559:
500:
498:
2742:Office of HIV/AIDS Network Coordination
521:"It's Now Dr. Hartwell, Nobel Laureate"
480:
2671:Fred Hutchinson Cancer Research Center
2443:(Trinidad & Tobago/United Kingdom)
340:Fred Hutchinson Cancer Research Center
284:. From 1965 to 1968, he worked at the
231:Fred Hutchinson Cancer Research Center
173:Fred Hutchinson Cancer Research Center
848:Nobel Prize in Physiology or Medicine
282:Massachusetts Institute of Technology
243:Nobel Prize in Physiology or Medicine
129:Nobel Prize in Physiology or Medicine
72:Massachusetts Institute of Technology
7:
447:, University of California, Berkeley
440:, University of Washington, Seattle
334:in 1995. He became a member of the
278:California Institute of Technology
68:California Institute of Technology
25:
359:University of Southern California
100:Genetics Society of America Medal
2798:Arizona State University faculty
2793:University of Washington faculty
519:Paulson, Tom (October 8, 2001).
288:as a professor. He moved to the
286:University of California, Irvine
691:10.1128/JB.93.5.1662-1670.1967
600:. Nobel Foundation: Stockholm.
227:Leland Harrison (Lee) Hartwell
27:American biologist (born 1939)
1:
593:Tore Frängsmyr., ed. (2002).
523:. Seattle Post-Intelligencer.
58:Los Angeles, California, U.S.
740:Hartwell to join ASU faculty
540:Proc. Natl. Acad. Sci. U.S.A
336:National Academy of Sciences
2685:William B. Hutchinson, M.D.
1940:Christiane NĂĽsslein-Volhard
757:September 29, 2009, at the
361:in 2000. On July 9, 2003,
2839:
2732:HIV Vaccine Trials Network
619:, accessed 11 October 2020
328:Louisa Gross Horwitz Prize
185:Amrita Vishwa Vidyapeetham
2567:
1137:Charles Scott Sherrington
1113:Frederick Gowland Hopkins
220:
151:
2773:American Nobel laureates
2405:William Standish Knowles
2127:Françoise Barré-Sinoussi
761:Biodesign Institute News
433:, Harvard Medical School
400:Arizona State University
345:In 1998 he received the
307:Saccharomyces cerevisiae
290:University of Washington
272:, and then received his
177:Arizona State University
2813:Massry Prize recipients
2725:Organizations on campus
786:"Lee Hartwell Lectures"
679:Journal of Bacteriology
355:Keck School of Medicine
2573:Nobel Prize recipients
2507:Physiology or Medicine
908:Santiago RamĂłn y Cajal
727:March 4, 2016, at the
1409:Dickinson W. Richards
1093:Julius Wagner-Jauregg
598:The Nobel Prizes 2001
561:10.1073/pnas.66.2.352
468:, Whitehead Institute
396:Pacific Health Summit
241:. He shared the 2001
82:Cell cycle regulation
2417:Karl Barry Sharpless
1832:Rita Levi-Montalcini
1193:Albert Szent-Györgyi
673:Hartwell LH (1967).
270:Glendale, California
266:Glendale High School
2818:American scientists
2224:William C. Campbell
2135:Elizabeth Blackburn
1972:Robert F. Furchgott
1964:Stanley B. Prusiner
1956:Rolf M. Zinkernagel
1856:George H. Hitchings
1820:Joseph L. Goldstein
1780:Bengt I. Samuelsson
1609:Earl Sutherland Jr.
552:1970PNAS...66..352H
461:, Oxford University
404:Biodesign Institute
332:Columbia University
322:In addition to the
294:cell division cycle
274:Bachelor of Science
181:Biodesign Institute
115:Albert Lasker Award
107:Komen Brinker Award
2706:Leland H. Hartwell
2515:Leland H. Hartwell
2484:Eric Allin Cornell
2284:William Kaelin Jr.
2280:Peter J. Ratcliffe
2023:Leland H. Hartwell
1912:Richard J. Roberts
1792:Barbara McClintock
1736:Godfrey Hounsfield
1637:Nikolaas Tinbergen
1569:Marshall Nirenberg
1537:Charles B. Huggins
1326:Tadeusz Reichstein
1314:AntĂłnio Egas Moniz
1109:Christiaan Eijkman
804:on Nobelprize.org
802:Leland H. Hartwell
774:. October 6, 2011.
636:on April 27, 2006.
613:on Nobelprize.org
611:Leland H. Hartwell
454:, Kyoto University
452:Mitsuhiro Yanagida
414:Lee Hartwell Award
264:Hartwell attended
36:Leland H. Hartwell
2750:
2749:
2711:E. Donnall Thomas
2637:
2636:
2538:Economic Sciences
2490:Wolfgang Ketterle
2353:
2352:
2318:Ardem Patapoutian
2167:Ralph M. Steinman
2163:Jules A. Hoffmann
2151:Robert G. Edwards
2119:Harald zur Hausen
2043:H. Robert Horvitz
1944:Eric F. Wieschaus
1880:E. Donnall Thomas
1864:J. Michael Bishop
1852:Gertrude B. Elion
1724:Hamilton O. Smith
1649:Christian de Duve
1385:Frederick Robbins
1262:Alexander Fleming
1201:Corneille Heymans
1047:Frederick Banting
952:Allvar Gullstrand
408:Amrita University
392:Canary Foundation
318:Awards and honors
224:
223:
153:Scientific career
147:
16:(Redirected from
2830:
2737:Sage Bionetworks
2664:
2657:
2650:
2641:
2575:
2529:(United Kingdom)
2523:(United Kingdom)
2380:
2373:
2366:
2357:
2346:
2331:
2320:
2305:
2299:Michael Houghton
2286:
2276:Gregg L. Semenza
2270:
2264:James P. Allison
2258:
2256:Michael W. Young
2242:
2240:Yoshinori Ohsumi
2234:
2218:
2202:
2200:Thomas C. SĂĽdhof
2186:
2174:
2153:
2145:
2139:Carol W. Greider
2129:
2113:
2097:
2085:
2073:
2061:
2049:
2033:
2006:
1990:
1982:
1966:
1958:
1952:Peter C. Doherty
1946:
1930:
1924:Alfred G. Gilman
1918:
1906:
1894:
1882:
1870:
1868:Harold E. Varmus
1858:
1842:
1834:
1822:
1810:
1794:
1786:
1770:
1754:
1744:Baruj Benacerraf
1738:
1726:
1710:
1694:
1671:
1655:
1639:
1623:
1611:
1603:
1587:
1571:
1561:Robert W. Holley
1555:
1539:
1527:
1511:
1499:
1483:
1467:
1465:Georg von Békésy
1459:
1447:
1435:
1433:Joshua Lederberg
1419:
1411:
1405:Werner Forssmann
1395:
1387:
1371:
1359:
1351:
1332:
1316:
1304:
1296:
1294:Bernardo Houssay
1280:
1272:
1256:
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1232:
1231:
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1224:
1218:
1217:
1211:
1203:
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1151:
1143:
1131:
1123:
1121:Karl Landsteiner
1115:
1103:
1095:
1087:
1085:Johannes Fibiger
1068:
1067:
1061:
1059:Willem Einthoven
1053:
1041:
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924:Élie Metchnikoff
918:
916:Alphonse Laveran
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650:. June 14, 2018.
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632:. Archived from
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596:Les Prix Nobel.
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210:Doctoral advisor
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93:Rosenstiel Award
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52:October 30, 1939
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2690:Fred Hutchinson
2673:
2668:
2638:
2633:
2571:
2563:
2560:(United States)
2558:Joseph Stiglitz
2554:(United States)
2548:(United States)
2532:
2517:(United States)
2501:
2498:(United States)
2486:(United States)
2470:
2446:
2422:
2419:(United States)
2407:(United States)
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2384:
2354:
2349:
2334:
2323:
2308:
2303:Charles M. Rice
2295:Harvey J. Alter
2289:
2273:
2261:
2252:Michael Rosbash
2248:Jeffrey C. Hall
2245:
2237:
2221:
2212:May-Britt Moser
2205:
2189:
2184:Shinya Yamanaka
2177:
2156:
2148:
2143:Jack W. Szostak
2132:
2116:
2111:Oliver Smithies
2100:
2088:
2076:
2064:
2059:Peter Mansfield
2052:
2047:John E. Sulston
2036:
2020:
2009:
1993:
1985:
1969:
1961:
1949:
1936:Edward B. Lewis
1933:
1921:
1909:
1897:
1885:
1873:
1861:
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1840:Susumu Tonegawa
1837:
1825:
1813:
1797:
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1729:
1713:
1700:Roger Guillemin
1697:
1692:Daniel Gajdusek
1688:Baruch Blumberg
1685:
1674:
1665:Renato Dulbecco
1661:David Baltimore
1658:
1642:
1629:Karl von Frisch
1626:
1614:
1606:
1590:
1574:
1558:
1549:Haldan Hartline
1542:
1530:
1514:
1502:
1486:
1481:Maurice Wilkins
1470:
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1445:Arthur Kornberg
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1250:Joseph Erlanger
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1214:
1206:
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1101:Charles Nicolle
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944:Albrecht Kossel
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466:Susan Lindquist
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386:Other positions
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215:Boris Magasanik
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64:Alma mater
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28:
23:
22:
18:Leland Hartwell
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12:
11:
5:
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2695:Lawrence Corey
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2659:
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2621:
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2611:
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2601:
2596:
2591:
2586:
2581:
2576:
2568:
2565:
2564:
2562:
2561:
2555:
2552:Michael Spence
2549:
2546:George Akerlof
2542:
2540:
2534:
2533:
2531:
2530:
2524:
2518:
2511:
2509:
2503:
2502:
2500:
2499:
2493:
2487:
2480:
2478:
2472:
2471:
2469:
2468:
2462:
2460:United Nations
2456:
2454:
2448:
2447:
2445:
2444:
2437:
2435:
2424:
2423:
2421:
2420:
2414:
2408:
2401:
2399:
2393:
2392:
2385:
2383:
2382:
2375:
2368:
2360:
2351:
2350:
2348:
2347:
2340:Katalin KarikĂł
2332:
2321:
2306:
2287:
2271:
2259:
2243:
2235:
2219:
2203:
2196:Randy Schekman
2187:
2175:
2154:
2146:
2130:
2123:Luc Montagnier
2114:
2103:Mario Capecchi
2098:
2086:
2079:Barry Marshall
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44:
40:
33:
30:
19:
2705:
2603:
2514:
2411:RyĹŤji Noyori
2329:Svante Pääbo
2314:David Julius
2268:Tasuku Honjo
2216:Edvard Moser
2208:John O'Keefe
2171:posthumously
2170:
2107:Martin Evans
2083:Robin Warren
2067:Richard Axel
2022:
2014:2001–present
1892:Bert Sakmann
1760:Roger Sperry
1752:George Snell
1748:Jean Dausset
1716:Werner Arber
1669:Howard Temin
1593:Bernard Katz
1577:Max DelbrĂĽck
1533:Francis Rous
1509:Feodor Lynen
1505:Konrad Bloch
1493:Alan Hodgkin
1477:James Watson
1453:Frank Burnet
1441:Severo Ochoa
1429:Edward Tatum
1417:Daniel Bovet
1330:Philip Hench
1242:Edward Doisy
1173:Hans Spemann
1161:George Minot
1141:Edgar Adrian
1129:Otto Warburg
1051:John Macleod
1020:August Krogh
1012:Jules Bordet
928:Paul Ehrlich
880:Niels Finsen
864:Emil Behring
780:
766:
747:
735:
717:
682:
678:
668:
656:
642:
634:the original
624:
606:
597:
594:
588:
546:(2): 352–9.
543:
539:
529:
514:
483:
424:Susan Gasser
417:
389:
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351:Massry Prize
344:
321:
305:
297:
263:
255:
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196:
169:Institutions
152:
136:Mendel Medal
122:Massry Prize
29:
2763:1939 births
2697:, President
2496:Carl Wieman
2388:Nobel Prize
2180:John Gurdon
2095:Craig Mello
2091:Andrew Fire
2004:Eric Kandel
1980:Ferid Murad
1888:Erwin Neher
1800:Niels Jerne
1565:Har Khorana
1553:George Wald
1521:André Lwoff
1489:John Eccles
1377:John Enders
1349:Max Theiler
1310:Walter Hess
1302:Paul MĂĽller
1266:Ernst Chain
936:Emil Kocher
896:Robert Koch
888:Ivan Pavlov
872:Ronald Ross
459:Kim Nasmyth
438:Stan Fields
324:Nobel Prize
300:) genes in
2757:Categories
2678:Key people
2527:Paul Nurse
2465:Kofi Annan
2428:Literature
2031:Paul Nurse
1365:Hans Krebs
1290:Gerty Cori
1238:Henrik Dam
1185:Otto Loewi
1181:Henry Dale
476:References
410:in India.
367:Gary Locke
363:Washington
349:, and the
247:Paul Nurse
239:Washington
48:1939-10-30
2716:Mark Roth
2492:(Germany)
2397:Chemistry
2390:laureates
2232:Tu Youyou
1784:John Vane
1679:1976–2000
1340:1951–1975
1286:Carl Cori
1076:1926–1950
855:1901–1925
844:Laureates
365:Governor
353:from the
312:mutations
276:from the
260:Education
2521:Tim Hunt
2027:Tim Hunt
755:Archived
742:ASU News
725:Archived
377:Research
251:Tim Hunt
2476:Physics
2467:(Ghana)
2413:(Japan)
846:of the
709:5337848
580:5271168
548:Bibcode
464:2004 –
457:2006 –
450:2008 –
443:2010 –
436:2012 –
429:2014 –
422:2016 –
235:Seattle
163:Biology
2274:2019:
2262:2018:
2246:2017:
2238:2016:
2222:2015:
2206:2014:
2190:2013:
2178:2012:
2157:2011:
2149:2010:
2133:2009:
2117:2008:
2101:2007:
2089:2006:
2077:2005:
2065:2004:
2053:2003:
2037:2002:
2021:2001:
1994:2000:
1986:1999:
1970:1998:
1962:1997:
1950:1996:
1934:1995:
1922:1994:
1910:1993:
1898:1992:
1886:1991:
1874:1990:
1862:1989:
1846:1988:
1838:1987:
1826:1986:
1814:1985:
1798:1984:
1790:1983:
1774:1982:
1758:1981:
1742:1980:
1730:1979:
1714:1978:
1698:1977:
1686:1976:
1659:1975:
1643:1974:
1627:1973:
1615:1972:
1607:1971:
1591:1970:
1575:1969:
1559:1968:
1543:1967:
1531:1966:
1515:1965:
1503:1964:
1487:1963:
1471:1962:
1463:1961:
1451:1960:
1439:1959:
1423:1958:
1415:1957:
1399:1956:
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1375:1954:
1363:1953:
1355:1952:
1347:1951:
1320:1950:
1308:1949:
1300:1948:
1284:1947:
1276:1946:
1260:1945:
1248:1944:
1236:1943:
1207:1939:
1199:1938:
1191:1937:
1179:1936:
1171:1935:
1155:1934:
1147:1933:
1135:1932:
1127:1931:
1119:1930:
1107:1929:
1099:1928:
1091:1927:
1083:1926:
1057:1924:
1045:1923:
1033:1922:
1018:1920:
1010:1919:
974:1914:
966:1913:
958:1912:
950:1911:
942:1910:
934:1909:
922:1908:
914:1907:
902:1906:
894:1905:
886:1904:
878:1903:
870:1902:
862:1901:
707:
700:276664
697:
578:
571:283051
568:
203:(1964)
201:
192:Thesis
159:Fields
146:(2003)
139:(2001)
132:(2001)
125:(2000)
118:(1998)
110:(1998)
103:(1994)
96:(1992)
88:Awards
2604:2001
2452:Peace
2386:2001
330:from
245:with
2629:2006
2624:2005
2619:2004
2614:2003
2609:2002
2599:2000
2594:1999
2589:1998
2584:1997
2579:1996
2432:2001
2336:2023
2325:2022
2310:2021
2291:2020
1230:1942
1223:1941
1216:1940
1066:1925
1027:1921
1004:1918
997:1917
990:1916
983:1915
705:PMID
576:PMID
249:and
42:Born
695:PMC
687:doi
566:PMC
556:doi
298:CDC
268:in
233:in
2759::
2342:/
2338::
2327::
2316:/
2312::
2301:/
2297:/
2293::
2282:/
2278:/
2266:/
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2250:/
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2214:/
2210:/
2198:/
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2182:/
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2161:/
2141:/
2137:/
2125:/
2121:/
2109:/
2105:/
2093:/
2081:/
2069:/
2057:/
2045:/
2041:/
2029:/
2025:/
2002:/
1998:/
1978:/
1974:/
1954:/
1942:/
1938:/
1926:/
1914:/
1902:/
1890:/
1878:/
1866:/
1854:/
1850:/
1830:/
1818:/
1806:/
1802:/
1782:/
1778:/
1766:/
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1750:/
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1734:/
1722:/
1718:/
1706:/
1702:/
1690:/
1667:/
1663:/
1651:/
1647:/
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1631:/
1619:/
1599:/
1595:/
1583:/
1579:/
1567:/
1563:/
1551:/
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1523:/
1519:/
1507:/
1495:/
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1455:/
1443:/
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1427:/
1407:/
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1383:/
1379:/
1367:/
1328:/
1324:/
1312:/
1292:/
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1268:/
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1240:/
1183:/
1163:/
1159:/
1139:/
1111:/
1049:/
1037:/
926:/
906:/
703:.
693:.
683:93
681:.
677:.
574:.
564:.
554:.
544:66
542:.
538:.
497:^
373:.
357:,
237:,
70:,
50:)
2663:e
2656:t
2649:v
2434:)
2430:(
2379:e
2372:t
2365:v
2173:)
2169:(
836:e
829:t
822:v
788:.
711:.
689::
582:.
558::
550::
508:.
491:.
304:(
296:(
46:(
20:)
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