514:
41:
417:
458:
1708:
66:
535:
396:
1725:
300:, and the naked head and neck are adaptations to this, as are those of the vultures with which they often feed. A feathered head would become rapidly clotted with blood and other substances when a scavenging bird's head was inside a large corpse, and the bare head is easier to keep clean.
262:, typically 110–150 cm tall with a 210–250 cm wingspan. The three species each have a black upper body and wings, and white belly and undertail. The head and neck are bare like those of a
1883:
1922:
1857:
1896:
1465:
1991:
903:
893:
876:
585:
of this genus. Many fossils members of the genus were much larger than living species, standing as tall as a man, with the earliest being
513:
1111:
1986:
1707:
1901:
821:
995:"More bones of Leptoptilos robustus from Flores reveal new insights into giant marabou stork paleobiology and biogeography"
1909:
40:
1805:
1458:
1267:
52:
365:
144:
65:
1836:
959:
1927:
1062:"A new species of giant marabou stork (Aves: Ciconiiformes) from the Pleistocene of Liang Bua, Flores (Indonesia)"
1849:
1257:
416:
790:
695:
678:
558:
496:
1745:
1728:
1658:
1451:
457:
403:
798:
deposits (Late
Miocene? to Late Pliocene) may belong to this genus or to a closely related one (Louchart
636:
1948:
1810:
868:
Priority! The Dating of
Scientific Names in Ornithology: a Directory to the literature and its reviewers
727:
627:
587:
465:
440:
757:
1981:
1976:
1792:
1104:
1006:
816:
744:
732:
612:
1352:
1167:
948:. Vol. 1 (2nd ed.). Cambridge, Massachusetts: Museum of Comparative Zoology. p. 251.
862:
687:
1678:
736:
1598:
1342:
1042:
919:
666:
521:
347:
252:
205:
60:
1953:
1688:
1668:
661:
1627:
762:
1935:
1888:
1818:
1034:
899:
872:
773:
1783:
1940:
1617:
1443:
1410:
1333:
1277:
1247:
1222:
1073:
1024:
1014:
963:
925:
A List of the Genera of Birds : with an
Indication of the Typical Species of Each Genus
777:
700:
600:
343:
190:
161:
1400:
1097:
395:
196:
1010:
1844:
1831:
1297:
1212:
1203:
1137:
1029:
994:
993:
Meijer, Hanneke J. M.; Sutikna, Thomas; Wahyu
Saptomo, E.; Tocheri, Matthew W. (2022).
845:
324:
1970:
1539:
1527:
1512:
1475:
1420:
1287:
1177:
1132:
1078:
1061:
1046:
866:
708:
582:
328:
248:
184:
48:
1797:
943:
923:
849:
534:
1560:
1506:
683:
339:
153:
1823:
1914:
1870:
1777:
1317:
1307:
1157:
811:
1572:
1187:
939:
670:
1768:
967:
1609:
1581:
704:
297:
77:
1038:
854:(in French). Vol. 1. Paris: F.G. Levrault. p. 583 (Livraison 8).
270:
is long and thick. Juveniles are a duller, browner version of the adult.
1762:
1648:
1494:
1148:
592:
97:
1019:
1862:
1588:
1238:
795:
641:
277:
263:
30:
1634:
1488:
1375:
749:
293:
289:
285:
87:
1875:
1739:
1500:
1120:
308:
228:
127:
117:
475:
South and
Southeast Asia, from India and Sri Lanka to Indonesia
281:
267:
259:
244:
107:
1743:
1525:
1447:
1093:
1089:
865:; Overstreet, L.K.; Dowsett, R.J.; Bruce, M.D. (2011).
631:(Early to Late Pliocene of south Asia and east Africa)
303:
Most storks fly with neck outstretched, but the three
860:
between 1830 and 1831. For the publication date see:
1684:
1674:
1664:
1654:
1623:
1605:
1594:
1556:
968:"Storks, frigatebirds, boobies, darters, cormorants"
280:, building large stick nests in trees. They feed on
1752:
1571:
1547:
1538:
1389:
1366:
1331:
1236:
1201:
1146:
243:). Two species are resident breeders in southern
323:was introduced in 1831 by the French naturalist
1459:
1105:
8:
895:The Helm Dictionary of Scientific Bird Names
871:. Northampton, UK: Aves Press. p. 119.
851:Traité d'Ornithologie, ou Tableau Méthodique
307:storks retract their necks in flight like a
276:storks are gregarious colonial breeders in
1740:
1544:
1535:
1522:
1466:
1452:
1444:
1112:
1098:
1090:
810:The adjutant bird features in the arms of
39:
20:
1077:
1066:Zoological Journal of the Linnean Society
1028:
1018:
928:. London: R. and J.E. Taylor. p. 67.
898:. London: Christopher Helm. p. 222.
531:northern India to mainland southeast Asia
353:The genus contains three extant species.
603:on the Southeast Asian islands of Java (
355:
837:
814:. In the satirical French puppet show
699:(Early Pliocene of Odesa, Ukraine and
974:. International Ornithologists' Union
942:; Cottrell, G. William, eds. (1979).
761:(Notopuro Middle/Late Pleistocene of
387:IUCN status and estimated population
334:meaning "delicate" or "slender" with
7:
1850:c94fea3a-5961-4170-9923-84e078afc227
1436:Twenty extant species in six genera
342:was subsequently designated as the
227:is a genus of very large tropical
14:
1724:
1723:
1706:
1079:10.1111/j.1096-3642.2010.00616.x
972:IOC World Bird List Version 12.2
945:Check-List of Birds of the World
533:
512:
456:
415:
394:
64:
824:was, for a time, depicted as a
703:, Ethiopia to Late Pliocene of
378:Scientific name and subspecies
327:. The genus name combines the
1:
1268:African woolly-necked stork
53:South Luangwa National Park
2008:
1060:Meijer HJ, Due RA (2010).
999:Royal Society Open Science
892:Jobling, James A. (2010).
413:Africa south of the Sahara
1992:Taxa named by René Lesson
1719:
1704:
1534:
1521:
1483:
1434:
1258:Asian woolly-necked stork
1127:
599:storks survived into the
509:
453:
391:
386:
383:
380:
377:
374:
211:
204:
181:
176:
159:
152:
61:Scientific classification
59:
47:
38:
23:
1478:and their extinct allies
822:Valéry Giscard d'Estaing
791:Leptoptilos siwalicensis
731:(Beglia Late Miocene of
711:, Tanzania) – includes
696:Leptoptilos pliocenicus
679:Leptoptilos patagonicus
595:of Afro-Eurasia. Giant
338:meaning "feather". The
235:. The name means thin (
1987:Taxa described in 1831
1659:Palaeoephippiorhynchus
966:, eds. (August 2022).
404:Leptoptilos crumenifer
1949:Paleobiology Database
719:, may be the same as
648:, may be the same as
628:Leptoptilos falconeri
588:Leptoptilos falconeri
466:Leptoptilos javanicus
29:Temporal range: Late
1845:Fauna Europaea (new)
745:Leptoptilos robustus
646:Cryptociconia indica
644:, India) – formerly
315:Taxonomy and species
296:. They are frequent
231:, commonly known as
1353:Saddle-billed stork
1168:Yellow-billed stork
1020:10.1098/rsos.220435
1011:2022RSOS....920435M
920:Gray, George Robert
637:Leptoptilos indicus
527:(Gmelin, JF,, 1789)
371:
1343:Black-necked stork
962:; Donsker, David;
780:, Kenya: Louchart
765:, Java, Indonesia)
728:Leptoptilos richae
673:, Liaoning, China)
667:Middle Pleistocene
640:(Late Pliocene of
581:There is an ample
522:Leptoptilos dubius
409:(Lesson, RP, 1831)
356:
348:George Robert Gray
253:Sub-Saharan Africa
1964:
1963:
1936:Open Tree of Life
1746:Taxon identifiers
1737:
1736:
1715:
1714:
1702:
1701:
1698:
1697:
1441:
1440:
964:Rasmussen, Pamela
905:978-1-4081-2501-4
878:978-0-9568611-1-5
774:Ngorora Formation
758:Leptoptilos titan
571:
570:
528:
510:Greater adjutant
472:
471:(Horsfield, 1821)
410:
384:Size and ecology
369:
220:
219:
148:
1999:
1957:
1956:
1944:
1943:
1931:
1930:
1918:
1917:
1915:NHMSYS0020930222
1905:
1904:
1892:
1891:
1879:
1878:
1866:
1865:
1853:
1852:
1840:
1839:
1827:
1826:
1814:
1813:
1801:
1800:
1788:
1787:
1786:
1773:
1772:
1771:
1741:
1727:
1726:
1710:
1686:
1676:
1666:
1656:
1625:
1618:Ephippiorhynchus
1607:
1596:
1558:
1545:
1536:
1523:
1468:
1461:
1454:
1445:
1411:Greater adjutant
1334:Ephippiorhynchus
1223:African openbill
1114:
1107:
1100:
1091:
1084:
1083:
1081:
1057:
1051:
1050:
1032:
1022:
990:
984:
983:
981:
979:
956:
950:
949:
936:
930:
929:
916:
910:
909:
889:
883:
882:
855:
842:
778:Baringo District
776:Late Miocene of
688:Valdés Peninsula
601:Late Pleistocene
562:
537:
526:
516:
500:
470:
460:
454:Lesser adjutant
444:
419:
408:
398:
372:
370:– three species
364:
344:greater adjutant
143:
69:
68:
43:
21:
2007:
2006:
2002:
2001:
2000:
1998:
1997:
1996:
1967:
1966:
1965:
1960:
1952:
1947:
1939:
1934:
1926:
1921:
1913:
1908:
1900:
1895:
1887:
1882:
1874:
1869:
1861:
1856:
1848:
1843:
1835:
1830:
1822:
1817:
1809:
1804:
1796:
1791:
1782:
1781:
1776:
1767:
1766:
1761:
1748:
1738:
1733:
1711:
1694:
1567:
1530:
1517:
1479:
1472:
1442:
1437:
1430:
1401:Lesser adjutant
1385:
1362:
1357:E. senegalensis
1327:
1232:
1227:A. lamelligerus
1197:
1182:M. leucocephala
1142:
1138:List of species
1123:
1118:
1088:
1087:
1059:
1058:
1054:
992:
991:
987:
977:
975:
958:
957:
953:
938:
937:
933:
918:
917:
913:
906:
891:
890:
886:
879:
863:Dickinson, E.C.
861:
856:Published in 8
844:
843:
839:
834:
808:
735:, Tunisia, and
733:Bled ed Douarah
682:(Puerto Madryn
662:Leptoptilos lĂĽi
579:
573:
566:
560:
551:
550:
545:
544:
532:
525:
504:
498:
489:
488:
483:
482:
469:
448:
442:
433:
432:
427:
426:
414:
407:
317:
288:, young birds,
258:These are huge
194:
188:
172:
169:
142:
63:
34:
27:
17:
12:
11:
5:
2005:
2003:
1995:
1994:
1989:
1984:
1979:
1969:
1968:
1962:
1961:
1959:
1958:
1945:
1932:
1919:
1906:
1893:
1880:
1867:
1854:
1841:
1832:Fauna Europaea
1828:
1815:
1802:
1789:
1774:
1758:
1756:
1750:
1749:
1744:
1735:
1734:
1732:
1731:
1720:
1717:
1716:
1713:
1712:
1705:
1703:
1700:
1699:
1696:
1695:
1693:
1692:
1682:
1672:
1662:
1652:
1645:
1638:
1631:
1621:
1614:
1603:
1592:
1585:
1577:
1575:
1569:
1568:
1566:
1565:
1553:
1551:
1549:incertae sedis
1542:
1532:
1531:
1526:
1519:
1518:
1516:
1515:
1509:
1503:
1497:
1491:
1484:
1481:
1480:
1473:
1471:
1470:
1463:
1456:
1448:
1439:
1438:
1435:
1432:
1431:
1429:
1428:
1418:
1408:
1397:
1395:
1387:
1386:
1384:
1383:
1372:
1370:
1364:
1363:
1361:
1360:
1350:
1339:
1337:
1329:
1328:
1326:
1325:
1315:
1305:
1298:Oriental stork
1295:
1285:
1275:
1272:C. microscelis
1265:
1255:
1244:
1242:
1234:
1233:
1231:
1230:
1220:
1213:Asian openbill
1209:
1207:
1199:
1198:
1196:
1195:
1185:
1175:
1165:
1154:
1152:
1144:
1143:
1141:
1140:
1135:
1128:
1125:
1124:
1119:
1117:
1116:
1109:
1102:
1094:
1086:
1085:
1072:(4): 707–724.
1052:
985:
951:
931:
911:
904:
884:
877:
836:
835:
833:
830:
817:Le BĂ©bĂŞte Show
807:
804:
786:
785:
766:
753:
748:(Pleistocene,
740:
723:
691:
674:
657:
632:
611:) and Flores (
578:
575:
569:
568:
561: NT
556:
538:
529:
518:
507:
506:
499: NT
494:
476:
473:
462:
451:
450:
443: LC
438:
420:
411:
400:
392:Marabou stork
389:
388:
385:
382:
379:
376:
316:
313:
218:
217:
209:
208:
202:
201:
179:
178:
174:
173:
170:
157:
156:
150:
149:
135:
131:
130:
125:
121:
120:
115:
111:
110:
105:
101:
100:
95:
91:
90:
85:
81:
80:
75:
71:
70:
57:
56:
45:
44:
36:
35:
28:
16:Genus of birds
15:
13:
10:
9:
6:
4:
3:
2:
2004:
1993:
1990:
1988:
1985:
1983:
1980:
1978:
1975:
1974:
1972:
1955:
1950:
1946:
1942:
1937:
1933:
1929:
1924:
1920:
1916:
1911:
1907:
1903:
1898:
1894:
1890:
1885:
1881:
1877:
1872:
1868:
1864:
1859:
1855:
1851:
1846:
1842:
1838:
1833:
1829:
1825:
1820:
1816:
1812:
1807:
1803:
1799:
1794:
1790:
1785:
1779:
1775:
1770:
1764:
1760:
1759:
1757:
1755:
1751:
1747:
1742:
1730:
1722:
1721:
1718:
1709:
1691:
1690:
1683:
1681:
1680:
1673:
1671:
1670:
1663:
1661:
1660:
1653:
1651:
1650:
1646:
1644:
1643:
1639:
1637:
1636:
1632:
1630:
1629:
1622:
1620:
1619:
1615:
1612:
1611:
1604:
1601:
1600:
1593:
1591:
1590:
1586:
1584:
1583:
1579:
1578:
1576:
1574:
1570:
1563:
1562:
1555:
1554:
1552:
1550:
1546:
1543:
1541:
1540:Ciconiiformes
1537:
1533:
1529:
1528:Ciconiiformes
1524:
1520:
1514:
1513:Ciconiiformes
1510:
1508:
1504:
1502:
1498:
1496:
1492:
1490:
1486:
1485:
1482:
1477:
1469:
1464:
1462:
1457:
1455:
1450:
1449:
1446:
1433:
1426:
1425:L. crumenifer
1422:
1421:Marabou stork
1419:
1416:
1412:
1409:
1406:
1402:
1399:
1398:
1396:
1394:
1393:
1388:
1381:
1377:
1374:
1373:
1371:
1369:
1365:
1358:
1354:
1351:
1348:
1344:
1341:
1340:
1338:
1336:
1335:
1330:
1323:
1319:
1316:
1313:
1309:
1306:
1303:
1299:
1296:
1293:
1289:
1288:Maguari stork
1286:
1283:
1279:
1278:Storm's stork
1276:
1273:
1269:
1266:
1263:
1259:
1256:
1253:
1249:
1248:Abdim's stork
1246:
1245:
1243:
1241:
1240:
1235:
1228:
1224:
1221:
1218:
1214:
1211:
1210:
1208:
1206:
1205:
1200:
1193:
1189:
1186:
1183:
1179:
1178:Painted stork
1176:
1173:
1169:
1166:
1163:
1159:
1156:
1155:
1153:
1151:
1150:
1145:
1139:
1136:
1134:
1133:Ciconiiformes
1130:
1129:
1126:
1122:
1115:
1110:
1108:
1103:
1101:
1096:
1095:
1092:
1080:
1075:
1071:
1067:
1063:
1056:
1053:
1048:
1044:
1040:
1036:
1031:
1026:
1021:
1016:
1012:
1008:
1005:(7): 220435.
1004:
1000:
996:
989:
986:
973:
969:
965:
961:
955:
952:
947:
946:
941:
935:
932:
927:
926:
921:
915:
912:
907:
901:
897:
896:
888:
885:
880:
874:
870:
869:
864:
859:
853:
852:
847:
841:
838:
831:
829:
827:
823:
819:
818:
813:
805:
803:
801:
797:
793:
792:
783:
779:
775:
771:
767:
764:
760:
759:
754:
751:
747:
746:
741:
738:
734:
730:
729:
724:
722:
718:
714:
710:
706:
702:
698:
697:
692:
689:
685:
681:
680:
675:
672:
668:
664:
663:
658:
655:
651:
647:
643:
639:
638:
633:
630:
629:
624:
623:
622:
620:
619:
615:
610:
606:
602:
598:
594:
590:
589:
584:
583:fossil record
576:
574:
567:
564:
563:
557:
554:
548:
542:
539:
536:
530:
524:
523:
519:
517:
515:
508:
505:
502:
501:
495:
492:
486:
480:
477:
474:
468:
467:
463:
461:
459:
452:
449:
446:
445:
439:
436:
430:
424:
421:
418:
412:
406:
405:
401:
399:
397:
390:
373:
367:
362:
361:
354:
351:
349:
345:
341:
337:
333:
330:
329:Ancient Greek
326:
322:
314:
312:
310:
306:
301:
299:
295:
291:
287:
283:
279:
275:
271:
269:
265:
261:
256:
254:
250:
249:marabou stork
246:
242:
238:
234:
230:
226:
225:
216:
215:
214:Cryptociconia
210:
207:
203:
200:
199:
198:
193:
192:
187:
186:
185:L. crumenifer
180:
175:
168:
164:
163:
158:
155:
151:
146:
141:
140:
136:
133:
132:
129:
126:
123:
122:
119:
118:Ciconiiformes
116:
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92:
89:
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83:
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72:
67:
62:
58:
54:
50:
49:Marabou stork
46:
42:
37:
32:
26:
22:
19:
1753:
1687:
1679:Pelargosteon
1677:
1667:
1657:
1647:
1641:
1640:
1633:
1626:
1616:
1608:
1597:
1587:
1580:
1561:Sanshuiornis
1559:
1548:
1507:Aequornithes
1424:
1414:
1405:L. javanicus
1404:
1391:
1390:
1379:
1367:
1356:
1347:E. asiaticus
1346:
1332:
1321:
1311:
1301:
1291:
1281:
1271:
1262:C. episcopus
1261:
1251:
1237:
1226:
1216:
1202:
1192:M. americana
1191:
1181:
1171:
1161:
1147:
1069:
1065:
1055:
1002:
998:
988:
976:. Retrieved
971:
954:
944:
934:
924:
914:
894:
887:
867:
857:
850:
846:Lesson, René
840:
825:
815:
809:
799:
789:
787:
781:
769:
756:
752:, Indonesia)
743:
737:Wadi Moghara
726:
721:L. falconeri
720:
716:
712:
694:
690:, Argentina)
684:Late Miocene
677:
660:
653:
650:L. falconeri
649:
645:
635:
626:
617:
613:
608:
604:
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586:
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511:
503:
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490:
484:
478:
464:
455:
447:
441:
434:
428:
422:
402:
393:
375:Common name
359:
358:
352:
340:type species
335:
331:
320:
318:
304:
302:
273:
272:
266:. The huge
257:
251:is found in
240:
236:
232:
223:
222:
221:
213:
212:
197:L. javanicus
195:
189:
183:
182:
171:Latham, 1790
166:
162:Ardea argala
160:
154:Type species
138:
137:
24:
18:
1982:Bird genera
1977:Leptoptilos
1871:iNaturalist
1798:Leptoptilos
1784:Leptoptilos
1778:Wikispecies
1754:Leptoptilos
1642:Leptoptilos
1599:Ciconiopsis
1392:Leptoptilos
1380:J. mycteria
1318:Black stork
1308:White stork
1302:C. boyciana
1217:A. oscitans
1158:Milky stork
978:16 November
960:Gill, Frank
940:Mayr, Ernst
826:Leptoptilos
812:Baron Sinha
770:Leptoptilos
707:, Chad and
597:Leptoptilos
360:Leptoptilos
325:René Lesson
321:Leptoptilos
305:Leptoptilos
274:Leptoptilos
239:) feather (
224:Leptoptilos
167:Ardea dubia
139:Leptoptilos
25:Leptoptilos
1971:Categories
1689:Prociconia
1669:Pelargodes
1573:Ciconiidae
1474:Genera of
1312:C. ciconia
1292:C. maguari
1252:C. abdimii
1188:Wood stork
1162:M. cinerea
858:livraisons
832:References
806:In culture
671:Jinniushan
652:(Louchart
366:Lesson, RP
319:The genus
298:scavengers
247:, and the
145:Lesson, RP
128:Ciconiidae
1628:Grallavis
1610:Eociconia
1582:Anastomus
1487:Kingdom:
1415:L. dubius
1282:C. stormi
1204:Anastomus
1047:250459008
794:from the
763:Watualang
739:, Egypt?)
717:falconeri
705:Koro Toro
591:from the
233:adjutants
191:L. dubius
84:Kingdom:
78:Eukaryota
55:, Zambia
33:to Recent
1763:Wikidata
1729:Category
1649:Mycteria
1495:Chordata
1493:Phylum:
1489:Animalia
1322:C. nigra
1149:Mycteria
1039:35845853
922:(1840).
848:(1831).
618:robustus
593:Pliocene
278:wetlands
206:Synonyms
177:Species
124:Family:
98:Chordata
94:Phylum:
88:Animalia
74:Domain:
1889:1309209
1863:2481944
1769:Q577346
1589:Ciconia
1511:Order:
1505:Clade:
1499:Class:
1239:Ciconia
1172:M. ibis
1131:Order:
1030:9277297
1007:Bibcode
802:2005).
796:Siwalik
709:Olduvai
642:Siwalik
577:Fossils
547:Habitat
485:Habitat
429:Habitat
294:rodents
290:lizards
286:insects
264:vulture
134:Genus:
114:Order:
104:Class:
31:Miocene
1954:105607
1941:464690
1902:174918
1635:Jabiru
1476:storks
1376:Jabiru
1368:Jabiru
1121:Storks
1045:
1037:
1027:
902:
875:
800:et al.
782:et al.
750:Flores
701:Urugus
654:et al.
381:Range
368:, 1831
357:Genus
336:ptilon
332:leptos
241:ptilos
229:storks
147:, 1831
1928:33588
1884:IRMNG
1837:96675
1811:90158
1043:S2CID
784:2005)
772:sp. (
656:2005)
609:titan
309:heron
282:frogs
260:birds
237:lepto
51:near
1923:NCBI
1897:ITIS
1876:4748
1858:GBIF
1824:5DB9
1806:BOLD
1501:Aves
1035:PMID
980:2022
900:ISBN
873:ISBN
715:cf.
553:Diet
541:Size
491:Diet
479:Size
435:Diet
423:Size
292:and
268:bill
245:Asia
108:Aves
1910:NBN
1819:CoL
1793:ADW
1074:doi
1070:160
1025:PMC
1015:doi
686:of
669:of
621:).
346:by
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970:.
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820:,
713:L.
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363:–
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