520:
738:
1377:
749:
760:
717:
1573:
728:
1553:
1134:
1563:
1543:
530:
394:– following influxes in previous years, this species, like the yellow-winged darter, has formed temporary breeding colonies. However, since the mid-1990s, the number of these colonies has increased and many have continued to be present from year-to-year, so this species is now better regarded as a successful colonist.
403:
Records of non-native
Odonata species in Britain have been confined to individuals found within heated greenhouses associated with nurseries for aquatic plants. None of these species have been recorded in wild situations or gone on to establish populations in the wild. Details of species which have
105:
A number of other references were used to provide information on specific topics, including rare vagrants, post-1990 additions, predictions, species claimed but not accepted/species of uncertain provenance, non-natives, taxonomic matters and species found only in the
57:
Some of these rare species have not been seen since the 19th
Century; however, the British Odonata list is also currently undergoing a period of unprecedented change, as new species are being discovered for the first time, some going on to become breeding species.
1971:; Grimstone Warren, Norfolk on 21 July; Little Wootton Inclosure New Forest (SZ 227 987) on 6 August; And at the mouth of the Beaulieu River in Hampshire on 10 August. Since 2012 it has probably become established as a breeding species in the Thames Estuary area.
135:
After a period in which the
British Odonata list has been relatively static, since 1990, many new species have been found and some have gone on to become regular breeding species. In chronological order of their first record, these new species are:
2182:
3. The following is a list of species which have previously been predicted to occur in
Britain but have not yet been recorded (see Merritt, Moore & Eversham 1996 pp. 113–114 and Parr 1998, 1999):
3026:
1903:
Willow emerald damselfly, since 2007 this species is established and spreading in south-east
England, especially Suffolk and Essex. It was previously only known from a single record from
3071:
2251:
67:
300:
Many
British Odonata enthusiasts expect further species to be added to the list in the near future. The list below is up to date as of the end of the 2005 breeding season.
2107:
accepted, but the circumstances surrounding the records and/or specimens cast doubt on their natural occurrence, and they are not included in the official
British list:
3081:
2959:
3183:
3236:
3188:
3104:
3046:
519:
3203:
3076:
488:(with the year of the first British record in brackets, except for Red-veined Darter which has occurred as a migrant and sporadic breeder for many decades)
3241:
3056:
2832:
2807:
2744:
2698:
2673:
2648:
2457:
2390:
2365:
2319:
2272:
88:
2573:
2989:
386:– this species has a pattern of establishing small breeding colonies following influxes, but none of these have become permanently established
205:
on 13 June 1996 (Phillips 1997a, 1997b) and now occurring annually; breeding evidence was found at Hayle Kimbro Pool (Jones 2000) and also at
3231:
3127:
3061:
116:'s Odonata fauna is quite different from that of Britain, with many fewer breeding species, but one additional species not found in Britain,
2539:
3041:
2952:
2128:
in early
September 1999 (Parr 2000b). It was not accepted as the species is not a strong migrant and the rig receives weekly supplies from
3147:
1907:
in 1899 (although this record is based on a specimen regarded by some as perhaps not of
British origin, see Gladwin 1997) and a single
124:
3167:
3012:
2889:
Parr, Adrian (1998). "Potential New
Odonata for the British List 1. The Possible Occurrence of Nearctic Species in Western Europe".
2603:
80:
3226:
3162:
3036:
2945:
1964:
467:
The following codes are used to give status details for those species which are not long-established extant breeding species:
3246:
3152:
3086:
3066:
1376:
3157:
3031:
3021:
1960:
since 1998 (Long 2000, Parr 2005). However in 2006 four specimens were seen: In the Adur valley in West Sussex on 13 July
737:
428:
for each breeding species, and an overall status code for species which are not long-established extant breeding species.
2906:
Parr, Adrian (1999). "Potential New Odonata for the British List 2. The Possibility of Vagrant or Colonist Damselflies".
976:
3198:
2192:
417:
2928:
3122:
1479:
671:
431:
The following abbreviations are used to give country-by-country distribution information for the breeding species:
288:
252:
248:
on 17 July 1999 (Dewick and Gerussi 2000) – now a well-established breeding species across much of southern England
748:
2627:
51:
1855:
941:
759:
716:
637:
617:
240:
157:
on 7 August 1995 (Jones 1996), with a small number of further records at scattered locations throughout Britain
498:(with the year of the first record in brackets, except for species with long histories as visitors to Britain)
2200:
1401:
1260:
689:
495:
276:
3193:
2968:
1821:
1572:
1515:
1037:
485:
349:
165:
2569:
2086:
1989:
Yellow-winged darter is listed as a vagrant, but is occasionally subject to large influxes e.g. in 1995.
1089:
959:
907:
763:
232:
on 9 September 1998 – the first record for Europe (Pellow 1999a, 1999b), with several other records in
727:
1718:
1650:
1566:
573:
533:
382:
1633:
1599:
1556:
1546:
1280:
993:
787:
741:
731:
354:
2625:
Agassiz, D (1981). "Further introduced china mark moths (Lepidoptera: Pyralidae) new to Britain".
2547:
1838:
924:
872:
838:
821:
752:
312:, some do not fall into these two categories. The following species bred in the past but are now
309:
265:
233:
206:
47:
39:
2984:
2908:
2891:
2874:
2853:
2786:
2769:
2723:
2524:
2503:
2482:
2436:
2415:
2344:
2298:
2080:
1. The following species have been claimed but not accepted by the Odonata Records Committee:
1735:
1684:
1552:
1462:
804:
654:
556:
523:
390:
337:
146:
141:
97:
76:
2979:
2211:
2184:
1872:
1498:
1369:
1106:
889:
321:
308:
While most species on the list below are either extant established breeding species or rare
2932:
2595:
2227:
2219:
2149:
2141:
2076:
Rejected species, species of uncertain provenance, predictions and Channel Islands species
1968:
1946:
Southern migrant hawker was known from just a single record, in Kent in 1952, although an
1701:
1349:
1332:
1158:
1013:
855:
720:
421:
413:
404:
occurred in such circumstances can be found in Agassiz 1981, Brooks 1988, and Parr 2000a.
293:
257:
202:
117:
107:
264:
on 30 July 2002 (Nobes 2003); in 2003 & 2004, the species was again seen here and at
2048:
in 1989, although the two 20th Century records may result from accidental introductions.
2994:
2161:
2125:
2094:
1804:
1667:
1297:
1243:
1192:
1133:
709:
512:
439:
342:
326:
225:
198:
189:
178:
1562:
3220:
2176:
in 1926; however Morton was travelling in continental Europe on the date in question.
1956:
may have been this species (Holmes 1993), and the species has been recorded twice in
1904:
1787:
1770:
1616:
1576:
1445:
1226:
174:
27:
2717:
Brook, John; Gill Brook (2004). "Past breeding evidence of Willow Emerald Damselfly
1961:
38:(suborder Anisoptera). Of these, 42 species (17 damselflies and 25 dragonflies) are
2455:(Drury) in Cornwall and Isles of Scilly – The first British and European Records".
2384:
Jones, Steven P. (2000). "First proof of successful breeding by the Lesser Emperor
2169:
1753:
1542:
1535:
1315:
1209:
1175:
281:
220:
2296:
Parr, Adrian (2000a). "An Annotated List of the Odonata of Britain and Ireland".
2113:
2067:
1999:
1421:
529:
425:
366:
358:
2270:
Parr, A. J. (1996). "Dragonfly movement and migration in Britain and Ireland".
272:(Forrest 2005), but there have been no subsequent sightings at either location.
95:
Parr, Adrian (2000a), An Annotated List of the Odonata of Britain and Ireland,
2041:
475:
313:
194:
150:
43:
2434:(Drury) into Cornwall and the Isles of Scilly – The First European Records".
86:
Parr, A. J. (1996), Dragonfly movement and migration in Britain and Ireland,
2313:
Jones, Steven P. (1996). "The first British record of the Scarlet Dragonfly
2173:
2121:
2012:
1126:
1065:
1057:
504:
210:
209:, Cornwall (Pellow 2000) in 1999, and the species appears to be resident at
35:
31:
2168:), only known from a single specimen labelled as having been collected by
2937:
2667:
Gladwin, T.W. (1997). "The error in treating the Green Emerald Damselfly
593:
452:
365:, where the species was recorded from 1820 to 1963, and the other on the
229:
170:
154:
2148:), only known from old specimens with vague data (but recorded from the
333:, first recorded in 1946, and last recorded in 1952. Rediscovered 2010.
2033:
2029:
1953:
443:
261:
182:
113:
23:
19:
2037:
1957:
1948:
1934:
1908:
362:
353:– only ever known from two areas in southern England, one around the
2925:
2480:(Charpentier) Found Breeding in Essex – The First British Records".
2476:
Dewick, Stephen; Richard Gerussi (2000). "Small Red-eyed Damselfly
50:- in respect of the latter, this list follows the decisions of the
2129:
1571:
1561:
1551:
1541:
461:
370:
330:
245:
2256:
2045:
2016:
1912:
269:
214:
72:
2941:
1933:
Yellow-legged dragonfly is known from just a single record, in
185:, which had dark bands on its wings, may have been this species
2692:
Brook, John; Gill Brook (2003). "The Willow Emerald Damselfly
177:
on 16 August 1995, although a darter seen on 29 July 2003 at
2646:
Brooks, S. J. (1988). "Exotic dragonflies in north London".
1924:
Common clubtail was formerly known as club-tailed dragonfly.
484:
to indicate that the species is a recent (i.e. post-1990)
61:
This list is based on the following principal references:
2784:
Parr, Adrian (2005). "Odonata Records Committee update".
2028:
Wandering glider has been recorded only three times – at
2011:
Large white-faced darter has been recorded only once, at
478:
in Britain (with the year of the last record in brackets)
2120:), which was found dead on the Sedco 706 oil rig in the
2696:(Vander Linden) in Kent: a case of mistaken identity".
2363:(Sélys) in Gloucestershire; the first British record".
2103:
2. The following species have been recorded, and their
377:
The following species are sporadic or casual breeders:
2338:
Phillips, John (1997a). "The Lesser Emperor Dragonfly
460:
to indicate that the species' breeding range includes
451:
to indicate that the species' breeding range includes
2057:
Wandering glider was formerly known as globe skimmer.
2851:
Parr, Adrian (2003). "Migrant Dragonflies in 2002".
2066:
Habitat limited to Eastern England and particularly
65:
Merritt, R., N. W. Moore and B. C. Eversham (1996),
3176:
3140:
3113:
3095:
3010:
3003:
2518:Forrest, P. J. (2005). "Southern Emerald Damselfly
2430:Pellow, Keith (1999a). "An influx of Green Darner
2359:Phillips, John (1997b). "Lesser Emperor Dragonfly
2497:Nobes, Geoff (2003). "Southern Emerald Damselfly
2249:Merritt, R.; N. W. Moore; B. C. Eversham (1996).
1980:Vagrant emperor records taken from Silsby (1993).
2409:Pellow, Keith (2000). "Lesser Emperor Dragonfly
2342:in Gloucestershire; The First British Record".
2252:Atlas of the dragonflies of Britain and Ireland
68:Atlas of the dragonflies of Britain and Ireland
2872:(Burmeister) on an Oil Rig in the North Sea".
2738:Holmes, J. D. (1993). "A possible sighting of
2953:
2763:Long, Roger (2000). "Southern Migrant Hawker
2596:"Beautiful 'lost' insect turns up anew in UK"
8:
2451:Pellow, Keith (1999b). "Common Green Darner
2210:4. In addition to the species listed above,
373:where the species was recorded in 1946 only.
2572:. British Dragonfly Society. 11 July 2018.
2546:. British Dragonfly Society. Archived from
474:to indicate a former breeding species, now
3007:
2960:
2946:
2938:
2291:
2289:
2287:
420:of each species is given, followed by its
2006:
1998:Highland darter and common darter may be
494:to indicate a species only recorded as a
325:– only ever known from marshes along the
2833:Journal of the British Dragonfly Society
2808:Journal of the British Dragonfly Society
2745:Journal of the British Dragonfly Society
2699:Journal of the British Dragonfly Society
2674:Journal of the British Dragonfly Society
2649:Journal of the British Dragonfly Society
2458:Journal of the British Dragonfly Society
2391:Journal of the British Dragonfly Society
2366:Journal of the British Dragonfly Society
2320:Journal of the British Dragonfly Society
2273:Journal of the British Dragonfly Society
1993:
1975:
1581:
1427:
1383:
1375:
1140:
1132:
1071:
1019:
769:
758:
747:
736:
726:
715:
712:(blue, blue-tailed, and red damselflies)
599:
538:
528:
518:
89:Journal of the British Dragonfly Society
3184:Endangered species of the British Isles
2671:(Vander Linden) as a British species".
2244:
2242:
2238:
2061:
1915:in 1992 (Brook & Brook 2002, 2004).
1898:
42:breeders, and the remainder are either
1928:
3237:Lists of insects in the British Isles
2606:from the original on 27 November 2017
2501:(Fabr.) – The First British Record".
1941:
7:
3189:Endemic species of the British Isles
2826:Merritt, R.; G. S. Vick (1983). "Is
169:), recorded just once to date, near
3204:Birds of Leicestershire and Rutland
2868:Parr, Adrian (2000b). "Blue Dasher
2570:"New Dragonfly Species for Britain"
304:Extinct species and casual breeders
236:that autumn, but none since to date
1984:
1919:
125:List of Odonata species of Ireland
14:
3242:Lists of insects of Great Britain
2576:from the original on 13 July 2019
2052:
2023:
1952:seen in 1992 on the outskirts of
2801:Silsby, J. (1993). "A review of
2522:(Fabr.) at Sandwich Bay, Kent".
1538:(chasers, skimmers, and darters)
284:in South Wales in December 2008.
2413:(Selys) breeding in Cornwall".
1793:Sympetrum striolatum nigrescens
2002:(see Merritt & Vick 1983).
1962:website of reporter with photo
438:to indicate that the species'
1:
3072:Orthoptera and allied insects
2767:in Jersey, Channel Islands".
292:, first (and only) record at
3232:Lists of Odonata by location
2226:) have been recorded in the
977:Scarce blue-tailed damselfly
92:Vol. 12 No. 2 pp. 33–50
34:(suborder Zygoptera) and 36
3199:Flora and fauna of Cornwall
3052:Damselflies and dragonflies
2193:northern white-faced darter
1372:(golden-ringed dragonflies)
341:– only ever known from the
3263:
2721:(Vander Linden) in Kent".
2097:in 2002 – see Parr (2003).
1485:Somatochlora flavomaculata
1016:(white-legged damselflies)
672:Southern emerald damselfly
253:Southern emerald damselfly
3027:Aquatic heteropteran bugs
2975:
2926:British Dragonfly Society
1068:(club-tailed dragonflies)
52:Odonata Records Committee
2805:, the Vagrant Emperor".
1856:Large white-faced darter
1639:Libellula quadrimaculata
1058:Anisoptera (dragonflies)
942:Small red-eyed damselfly
638:Scarce emerald damselfly
618:Willow emerald damselfly
399:Non-native introductions
296:, Suffolk, in July 2018.
241:Small red-eyed damselfly
3227:Lists of insect species
3082:Soldierflies and allies
2870:Pachydiplax longipennis
2201:small emerald damselfly
2118:Pachydiplax longipennis
1861:Leucorrhinia pectoralis
1402:Golden-ringed dragonfly
1380:Golden-ringed dragonfly
1261:Southern migrant hawker
1107:Yellow-legged dragonfly
690:Common winter damselfly
505:Zygoptera (damselflies)
277:Common winter damselfly
2969:Fauna of Great Britain
2931:28 August 2011 at the
2628:Entomologist's Gazette
2166:Somatochlora alpestris
1827:Sympetrum pedemontanum
1741:Sympetrum fonscolombei
1673:Orthetrum coerulescens
1579:
1569:
1559:
1549:
1516:Orange-spotted emerald
1480:Yellow-spotted emerald
1468:Somatochlora metallica
1407:Cordulegaster boltonii
1381:
1138:
1129:(hawkers and emperors)
1038:White-legged damselfly
767:
756:
745:
734:
724:
536:
526:
350:Orange-spotted emerald
289:Yellow-spotted emerald
166:Sympetrum pedemontanum
131:New species since 1990
127:for more information.
18:There are 57 recorded
3247:Dragonflies of Europe
2388:(Sélys) in Britain".
2315:Crocothemis erythraea
2224:Sympetrum meridionale
2197:Leucorrhina rubicunda
2146:Sympetrum meridionale
2087:Two-spotted dragonfly
1690:Crocothemis erythraea
1656:Orthetrum cancellatum
1575:
1565:
1555:
1545:
1424:(emerald dragonflies)
1379:
1136:
1095:Gomphus vulgatissimus
960:Blue-tailed damselfly
913:Enallagma cyathigerum
908:Common blue damselfly
878:Coenagrion pulchellum
844:Coenagrion mercuriale
827:Coenagrion hastulatum
764:Blue-tailed damselfly
762:
751:
740:
730:
719:
596:(emerald damselflies)
532:
522:
345:between 1902 and 1957
101:No. 11 pp. 10–20
2828:Sympetrum nigrescens
2719:Chalcolestes viridis
2694:Chalcolestes viridis
2478:Erythromma viridulum
2093:) was reported from
1967:2 March 2007 at the
1776:Sympetrum striolatum
1759:Sympetrum sanguineum
1719:Yellow-winged darter
1651:Black-tailed skimmer
1567:Black-tailed skimmer
1504:Somatochlora arctica
1043:Platycnemis pennipes
947:Erythromma viridulum
623:Chalcolestes viridis
574:Beautiful demoiselle
562:Calopteryx splendens
534:Beautiful demoiselle
383:Yellow-winged darter
280:, first recorded at
256:, first recorded at
244:, first recorded in
224:, first recorded at
193:, first recorded at
145:, first recorded at
121:Coenagrion lunulatum
3105:Non-marine molluscs
2803:Hemianax ephippiger
2091:Epitheca bimaculata
1724:Sympetrum flaveolum
1634:Four-spotted chaser
1600:Broad-bodied chaser
1557:Four-spotted chaser
1547:Broad-bodied chaser
1355:Hemianax ephippiger
1338:Brachytron pratense
999:Pyrrhosoma nymphula
994:Large red damselfly
895:Coenagrion scitulum
793:Ceriagrion tenellum
788:Small red damselfly
742:Large red damselfly
732:Small red damselfly
2990:Scottish Highlands
2830:a good species?".
2540:"Winter Damselfly
2216:Orthetrum brunneum
1878:Pantala flavescens
1844:Leucorrhinia dubia
1839:White-faced darter
1810:Sympetrum vulgatum
1605:Libellula depressa
1580:
1570:
1560:
1550:
1521:Oxygastra curtisii
1382:
1139:
925:Red-eyed damselfly
873:Variable damselfly
839:Southern damselfly
822:Northern damselfly
810:Coenagrion armatum
768:
757:
753:Red-eyed damselfly
746:
735:
725:
537:
527:
234:South West England
207:Bake Fishing Lakes
3212:
3211:
3136:
3135:
2706:(1&2): 51–54.
2189:Aeshna subarctica
1888:
1887:
1736:Red-veined darter
1685:Scarlet dragonfly
1531:
1530:
1463:Brilliant emerald
1417:
1416:
1365:
1364:
1122:
1121:
1053:
1052:
1009:
1008:
861:Coenagrion puella
805:Norfolk damselfly
705:
704:
655:Emerald damselfly
589:
588:
557:Banded demoiselle
524:Banded demoiselle
424:, details of the
391:Red-veined darter
338:Norfolk damselfly
147:Hayle Kimbro Pool
142:Scarlet dragonfly
3254:
3008:
2962:
2955:
2948:
2939:
2914:
2913:
2903:
2897:
2896:
2886:
2880:
2879:
2865:
2859:
2858:
2848:
2842:
2841:
2823:
2817:
2816:
2798:
2792:
2791:
2781:
2775:
2774:
2760:
2754:
2753:
2735:
2729:
2728:
2714:
2708:
2707:
2689:
2683:
2682:
2664:
2658:
2657:
2643:
2637:
2636:
2622:
2616:
2615:
2613:
2611:
2602:. 21 July 2010.
2592:
2586:
2585:
2583:
2581:
2566:
2560:
2559:
2557:
2555:
2550:on 16 March 2014
2536:
2530:
2529:
2515:
2509:
2508:
2494:
2488:
2487:
2473:
2467:
2466:
2448:
2442:
2441:
2427:
2421:
2420:
2406:
2400:
2399:
2381:
2375:
2374:
2356:
2350:
2349:
2335:
2329:
2328:
2310:
2304:
2303:
2293:
2282:
2281:
2267:
2261:
2260:
2246:
2212:southern skimmer
2185:subarctic darner
2065:
2056:
2027:
2010:
1997:
1988:
1979:
1945:
1932:
1923:
1902:
1873:Wandering glider
1582:
1499:Northern emerald
1428:
1384:
1370:Cordulegastridae
1198:Aeshna isosceles
1141:
1112:Gomphus flavipes
1072:
1020:
982:Ischnura pumilio
965:Ischnura elegans
930:Erythromma najas
890:Dainty damselfly
770:
600:
579:Calopteryx virgo
539:
322:Dainty damselfly
30:, made up of 21
3262:
3261:
3257:
3256:
3255:
3253:
3252:
3251:
3217:
3216:
3213:
3208:
3172:
3132:
3109:
3091:
2999:
2971:
2966:
2933:Wayback Machine
2922:
2917:
2905:
2904:
2900:
2888:
2887:
2883:
2867:
2866:
2862:
2850:
2849:
2845:
2825:
2824:
2820:
2800:
2799:
2795:
2783:
2782:
2778:
2762:
2761:
2757:
2737:
2736:
2732:
2716:
2715:
2711:
2691:
2690:
2686:
2666:
2665:
2661:
2645:
2644:
2640:
2624:
2623:
2619:
2609:
2607:
2594:
2593:
2589:
2579:
2577:
2568:
2567:
2563:
2553:
2551:
2538:
2537:
2533:
2520:Lestes barbarus
2517:
2516:
2512:
2499:Lestes barbarus
2496:
2495:
2491:
2475:
2474:
2470:
2450:
2449:
2445:
2429:
2428:
2424:
2411:Anax parthenope
2408:
2407:
2403:
2386:Anax parthenope
2383:
2382:
2378:
2361:Anax parthenope
2358:
2357:
2353:
2340:Anax parthenope
2337:
2336:
2332:
2312:
2311:
2307:
2295:
2294:
2285:
2269:
2268:
2264:
2248:
2247:
2240:
2236:
2228:Channel Islands
2220:southern darter
2150:Channel Islands
2142:Southern darter
2078:
2073:
2060:
2051:
2044:in 1951 and in
2022:
2005:
1992:
1983:
1974:
1969:Wayback Machine
1940:
1927:
1918:
1897:
1893:
1848:Eng/Scot/Wales
1788:Highland darter
1780:Eng/Scot/Wales
1711:Eng/Scot/Wales
1707:Sympetrum danae
1677:Eng/Scot/Wales
1643:Eng/Scot/Wales
1622:Libellula fulva
1591:Range countries
1588:Scientific name
1540:
1455:Eng/Scot/Wales
1437:Range countries
1434:Scientific name
1426:
1411:Eng/Scot/Wales
1393:Range countries
1390:Scientific name
1374:
1350:Vagrant emperor
1342:Eng/Scot/Wales
1333:Hairy dragonfly
1303:Anax parthenope
1236:Eng/Scot/Wales
1215:Aeshna caerulea
1168:Eng/Scot/Wales
1159:Southern hawker
1150:Range countries
1147:Scientific name
1131:
1090:Common clubtail
1081:Range countries
1078:Scientific name
1070:
1061:
1029:Range countries
1026:Scientific name
1018:
1014:Platycnemididae
1003:Eng/Scot/Wales
969:Eng/Scot/Wales
917:Eng/Scot/Wales
882:Eng/Scot/Wales
865:Eng/Scot/Wales
856:Azure damselfly
779:Range countries
776:Scientific name
721:Azure damselfly
714:
677:Lestes barbarus
664:Eng/Scot/Wales
609:Range countries
606:Scientific name
598:
583:Eng/Scot/Wales
566:Eng/Scot/Wales
548:Range countries
545:Scientific name
517:
508:
426:range countries
422:scientific name
414:taxonomic order
412:The list is in
410:
401:
306:
294:Carlton Marshes
258:Winterton Dunes
203:Gloucestershire
133:
118:Irish Damselfly
108:Channel Islands
12:
11:
5:
3260:
3258:
3250:
3249:
3244:
3239:
3234:
3229:
3219:
3218:
3210:
3209:
3207:
3206:
3201:
3196:
3194:Birds of Wales
3191:
3186:
3180:
3178:
3174:
3173:
3171:
3170:
3165:
3160:
3155:
3150:
3144:
3142:
3138:
3137:
3134:
3133:
3131:
3130:
3125:
3119:
3117:
3111:
3110:
3108:
3107:
3101:
3099:
3093:
3092:
3090:
3089:
3084:
3079:
3074:
3069:
3064:
3059:
3054:
3049:
3044:
3039:
3034:
3029:
3024:
3018:
3016:
3005:
3001:
3000:
2998:
2997:
2992:
2987:
2982:
2976:
2973:
2972:
2967:
2965:
2964:
2957:
2950:
2942:
2936:
2935:
2921:
2920:External links
2918:
2916:
2915:
2898:
2881:
2860:
2843:
2818:
2793:
2776:
2765:Aeshna affinis
2755:
2740:Aeshna affinis
2730:
2709:
2684:
2669:Lestes viridis
2659:
2638:
2617:
2587:
2561:
2542:Sympecma fusca
2531:
2510:
2489:
2468:
2443:
2422:
2401:
2376:
2351:
2330:
2305:
2283:
2262:
2237:
2235:
2232:
2230:(Parr 2000a).
2180:
2179:
2178:
2177:
2162:Alpine emerald
2156:
2155:
2154:
2153:
2136:
2135:
2134:
2133:
2126:Shetland Isles
2105:identification
2101:
2100:
2099:
2098:
2095:Leicestershire
2077:
2074:
2072:
2071:
2058:
2049:
2020:
2003:
1990:
1981:
1972:
1938:
1925:
1916:
1894:
1892:
1889:
1886:
1885:
1882:
1880:
1875:
1869:
1868:
1865:
1863:
1858:
1852:
1851:
1849:
1846:
1841:
1835:
1834:
1831:
1829:
1824:
1818:
1817:
1814:
1812:
1807:
1805:Vagrant darter
1801:
1800:
1798:
1795:
1790:
1784:
1783:
1781:
1778:
1773:
1767:
1766:
1764:
1761:
1756:
1750:
1749:
1746:
1743:
1738:
1732:
1731:
1728:
1726:
1721:
1715:
1714:
1712:
1709:
1704:
1698:
1697:
1694:
1692:
1687:
1681:
1680:
1678:
1675:
1670:
1668:Keeled skimmer
1664:
1663:
1661:
1658:
1653:
1647:
1646:
1644:
1641:
1636:
1630:
1629:
1627:
1624:
1619:
1613:
1612:
1610:
1607:
1602:
1596:
1595:
1592:
1589:
1586:
1539:
1532:
1529:
1528:
1525:
1523:
1518:
1512:
1511:
1509:
1506:
1501:
1495:
1494:
1492:
1489:
1487:
1482:
1476:
1475:
1473:
1470:
1465:
1459:
1458:
1456:
1453:
1451:Cordulia aenea
1448:
1442:
1441:
1438:
1435:
1432:
1425:
1418:
1415:
1414:
1412:
1409:
1404:
1398:
1397:
1394:
1391:
1388:
1373:
1366:
1363:
1362:
1359:
1357:
1352:
1346:
1345:
1343:
1340:
1335:
1329:
1328:
1325:
1323:
1318:
1312:
1311:
1308:
1305:
1300:
1298:Lesser emperor
1294:
1293:
1291:
1288:
1286:Anax imperator
1283:
1277:
1276:
1270:
1268:
1266:Aeshna affinis
1263:
1257:
1256:
1254:
1251:
1246:
1244:Migrant hawker
1240:
1239:
1237:
1234:
1229:
1223:
1222:
1220:
1217:
1212:
1206:
1205:
1203:
1200:
1195:
1193:Norfolk hawker
1189:
1188:
1186:
1183:
1181:Aeshna grandis
1178:
1172:
1171:
1169:
1166:
1161:
1155:
1154:
1151:
1148:
1145:
1130:
1123:
1120:
1119:
1116:
1114:
1109:
1103:
1102:
1100:
1097:
1092:
1086:
1085:
1082:
1079:
1076:
1069:
1062:
1060:
1054:
1051:
1050:
1048:
1045:
1040:
1034:
1033:
1030:
1027:
1024:
1017:
1010:
1007:
1006:
1004:
1001:
996:
990:
989:
987:
984:
979:
973:
972:
970:
967:
962:
956:
955:
952:
949:
944:
938:
937:
935:
932:
927:
921:
920:
918:
915:
910:
904:
903:
900:
897:
892:
886:
885:
883:
880:
875:
869:
868:
866:
863:
858:
852:
851:
849:
846:
841:
835:
834:
832:
829:
824:
818:
817:
814:
812:
807:
801:
800:
798:
795:
790:
784:
783:
780:
777:
774:
713:
710:Coenagrionidae
706:
703:
702:
699:
697:
695:Sympecma fusca
692:
686:
685:
682:
679:
674:
668:
667:
665:
662:
657:
651:
650:
648:
645:
640:
634:
633:
627:
625:
620:
614:
613:
610:
607:
604:
597:
590:
587:
586:
584:
581:
576:
570:
569:
567:
564:
559:
553:
552:
549:
546:
543:
516:
513:Calopterygidae
509:
507:
501:
500:
499:
489:
479:
465:
464:
455:
446:
440:breeding range
409:
406:
400:
397:
396:
395:
387:
375:
374:
346:
343:Norfolk Broads
334:
327:Thames Estuary
305:
302:
298:
297:
285:
273:
249:
237:
226:Penlee Battery
217:
199:Forest of Dean
190:Lesser emperor
186:
179:Hickling Broad
158:
132:
129:
103:
102:
93:
84:
13:
10:
9:
6:
4:
3:
2:
3259:
3248:
3245:
3243:
3240:
3238:
3235:
3233:
3230:
3228:
3225:
3224:
3222:
3215:
3205:
3202:
3200:
3197:
3195:
3192:
3190:
3187:
3185:
3182:
3181:
3179:
3175:
3169:
3166:
3164:
3161:
3159:
3156:
3154:
3151:
3149:
3146:
3145:
3143:
3139:
3129:
3126:
3124:
3121:
3120:
3118:
3116:
3112:
3106:
3103:
3102:
3100:
3098:
3094:
3088:
3085:
3083:
3080:
3078:
3075:
3073:
3070:
3068:
3065:
3063:
3060:
3058:
3055:
3053:
3050:
3048:
3047:Conopid flies
3045:
3043:
3040:
3038:
3035:
3033:
3030:
3028:
3025:
3023:
3020:
3019:
3017:
3015:
3014:
3009:
3006:
3004:Invertebrates
3002:
2996:
2993:
2991:
2988:
2986:
2983:
2981:
2978:
2977:
2974:
2970:
2963:
2958:
2956:
2951:
2949:
2944:
2943:
2940:
2934:
2930:
2927:
2924:
2923:
2919:
2911:
2910:
2902:
2899:
2894:
2893:
2885:
2882:
2877:
2876:
2871:
2864:
2861:
2856:
2855:
2847:
2844:
2839:
2835:
2834:
2829:
2822:
2819:
2814:
2810:
2809:
2804:
2797:
2794:
2789:
2788:
2780:
2777:
2772:
2771:
2766:
2759:
2756:
2751:
2747:
2746:
2741:
2734:
2731:
2726:
2725:
2720:
2713:
2710:
2705:
2701:
2700:
2695:
2688:
2685:
2680:
2676:
2675:
2670:
2663:
2660:
2655:
2651:
2650:
2642:
2639:
2634:
2630:
2629:
2621:
2618:
2605:
2601:
2597:
2591:
2588:
2575:
2571:
2565:
2562:
2549:
2545:
2543:
2535:
2532:
2527:
2526:
2521:
2514:
2511:
2506:
2505:
2500:
2493:
2490:
2485:
2484:
2479:
2472:
2469:
2464:
2460:
2459:
2454:
2447:
2444:
2439:
2438:
2433:
2426:
2423:
2418:
2417:
2412:
2405:
2402:
2397:
2393:
2392:
2387:
2380:
2377:
2372:
2368:
2367:
2362:
2355:
2352:
2347:
2346:
2341:
2334:
2331:
2326:
2322:
2321:
2316:
2309:
2306:
2301:
2300:
2292:
2290:
2288:
2284:
2279:
2275:
2274:
2266:
2263:
2258:
2254:
2253:
2245:
2243:
2239:
2233:
2231:
2229:
2225:
2221:
2217:
2213:
2208:
2206:
2205:Lestes virens
2202:
2198:
2194:
2190:
2186:
2175:
2171:
2167:
2163:
2160:
2159:
2158:
2157:
2151:
2147:
2143:
2140:
2139:
2138:
2137:
2131:
2127:
2123:
2119:
2115:
2112:
2111:
2110:
2109:
2108:
2106:
2096:
2092:
2088:
2085:
2084:
2083:
2082:
2081:
2075:
2069:
2064:
2063:
2059:
2055:
2054:
2050:
2047:
2043:
2039:
2035:
2031:
2026:
2025:
2021:
2018:
2014:
2009:
2008:
2004:
2001:
1996:
1995:
1991:
1987:
1986:
1982:
1978:
1977:
1973:
1970:
1966:
1963:
1959:
1955:
1951:
1950:
1944:
1943:
1939:
1936:
1931:
1930:
1926:
1922:
1921:
1917:
1914:
1911:collected in
1910:
1906:
1905:Hertfordshire
1901:
1900:
1896:
1895:
1890:
1883:
1881:
1879:
1876:
1874:
1871:
1870:
1866:
1864:
1862:
1859:
1857:
1854:
1853:
1850:
1847:
1845:
1842:
1840:
1837:
1836:
1832:
1830:
1828:
1825:
1823:
1822:Banded darter
1820:
1819:
1815:
1813:
1811:
1808:
1806:
1803:
1802:
1799:
1796:
1794:
1791:
1789:
1786:
1785:
1782:
1779:
1777:
1774:
1772:
1771:Common darter
1769:
1768:
1765:
1762:
1760:
1757:
1755:
1752:
1751:
1747:
1744:
1742:
1739:
1737:
1734:
1733:
1729:
1727:
1725:
1722:
1720:
1717:
1716:
1713:
1710:
1708:
1705:
1703:
1700:
1699:
1695:
1693:
1691:
1688:
1686:
1683:
1682:
1679:
1676:
1674:
1671:
1669:
1666:
1665:
1662:
1659:
1657:
1654:
1652:
1649:
1648:
1645:
1642:
1640:
1637:
1635:
1632:
1631:
1628:
1625:
1623:
1620:
1618:
1617:Scarce chaser
1615:
1614:
1611:
1608:
1606:
1603:
1601:
1598:
1597:
1593:
1590:
1587:
1584:
1583:
1578:
1577:Common darter
1574:
1568:
1564:
1558:
1554:
1548:
1544:
1537:
1533:
1526:
1524:
1522:
1519:
1517:
1514:
1513:
1510:
1507:
1505:
1502:
1500:
1497:
1496:
1493:
1490:
1488:
1486:
1483:
1481:
1478:
1477:
1474:
1471:
1469:
1466:
1464:
1461:
1460:
1457:
1454:
1452:
1449:
1447:
1446:Downy emerald
1444:
1443:
1439:
1436:
1433:
1430:
1429:
1423:
1419:
1413:
1410:
1408:
1405:
1403:
1400:
1399:
1395:
1392:
1389:
1386:
1385:
1378:
1371:
1367:
1360:
1358:
1356:
1353:
1351:
1348:
1347:
1344:
1341:
1339:
1336:
1334:
1331:
1330:
1326:
1324:
1322:
1319:
1317:
1314:
1313:
1309:
1306:
1304:
1301:
1299:
1296:
1295:
1292:
1289:
1287:
1284:
1282:
1279:
1278:
1275:
1271:
1269:
1267:
1264:
1262:
1259:
1258:
1255:
1252:
1250:
1247:
1245:
1242:
1241:
1238:
1235:
1233:
1232:Aeshna juncea
1230:
1228:
1227:Common hawker
1225:
1224:
1221:
1218:
1216:
1213:
1211:
1208:
1207:
1204:
1201:
1199:
1196:
1194:
1191:
1190:
1187:
1184:
1182:
1179:
1177:
1174:
1173:
1170:
1167:
1165:
1164:Aeshna cyanea
1162:
1160:
1157:
1156:
1152:
1149:
1146:
1143:
1142:
1137:Common hawker
1135:
1128:
1124:
1117:
1115:
1113:
1110:
1108:
1105:
1104:
1101:
1098:
1096:
1093:
1091:
1088:
1087:
1083:
1080:
1077:
1074:
1073:
1067:
1063:
1059:
1055:
1049:
1046:
1044:
1041:
1039:
1036:
1035:
1031:
1028:
1025:
1022:
1021:
1015:
1011:
1005:
1002:
1000:
997:
995:
992:
991:
988:
985:
983:
980:
978:
975:
974:
971:
968:
966:
963:
961:
958:
957:
953:
950:
948:
945:
943:
940:
939:
936:
933:
931:
928:
926:
923:
922:
919:
916:
914:
911:
909:
906:
905:
901:
898:
896:
893:
891:
888:
887:
884:
881:
879:
876:
874:
871:
870:
867:
864:
862:
859:
857:
854:
853:
850:
847:
845:
842:
840:
837:
836:
833:
830:
828:
825:
823:
820:
819:
815:
813:
811:
808:
806:
803:
802:
799:
796:
794:
791:
789:
786:
785:
781:
778:
775:
772:
771:
765:
761:
754:
750:
743:
739:
733:
729:
722:
718:
711:
707:
700:
698:
696:
693:
691:
688:
687:
683:
680:
678:
675:
673:
670:
669:
666:
663:
661:
660:Lestes sponsa
658:
656:
653:
652:
649:
646:
644:
641:
639:
636:
635:
632:
628:
626:
624:
621:
619:
616:
615:
611:
608:
605:
602:
601:
595:
591:
585:
582:
580:
577:
575:
572:
571:
568:
565:
563:
560:
558:
555:
554:
550:
547:
544:
541:
540:
535:
531:
525:
521:
515:(demoiselles)
514:
510:
506:
502:
497:
493:
490:
487:
483:
480:
477:
473:
470:
469:
468:
463:
459:
456:
454:
450:
447:
445:
441:
437:
434:
433:
432:
429:
427:
423:
419:
415:
407:
405:
398:
393:
392:
388:
385:
384:
380:
379:
378:
372:
368:
364:
360:
356:
352:
351:
347:
344:
340:
339:
335:
332:
328:
324:
323:
319:
318:
317:
315:
311:
303:
301:
295:
291:
290:
286:
283:
279:
278:
274:
271:
267:
263:
259:
255:
254:
250:
247:
243:
242:
238:
235:
231:
227:
223:
222:
218:
216:
212:
208:
204:
200:
196:
192:
191:
187:
184:
180:
176:
175:Monmouthshire
172:
168:
167:
162:
161:Banded darter
159:
156:
152:
148:
144:
143:
139:
138:
137:
130:
128:
126:
122:
119:
115:
111:
109:
100:
99:
94:
91:
90:
85:
82:
81:0-11-701561-X
78:
74:
70:
69:
64:
63:
62:
59:
55:
53:
49:
45:
41:
37:
33:
29:
25:
21:
16:
3214:
3114:
3096:
3051:
3011:
2907:
2901:
2890:
2884:
2873:
2869:
2863:
2857:(18): 18–24.
2852:
2846:
2837:
2831:
2827:
2821:
2812:
2806:
2802:
2796:
2785:
2779:
2768:
2764:
2758:
2749:
2743:
2739:
2733:
2722:
2718:
2712:
2703:
2697:
2693:
2687:
2678:
2672:
2668:
2662:
2653:
2647:
2641:
2632:
2626:
2620:
2608:. Retrieved
2599:
2590:
2578:. Retrieved
2564:
2552:. Retrieved
2548:the original
2541:
2534:
2528:(24): 24–25.
2523:
2519:
2513:
2502:
2498:
2492:
2481:
2477:
2471:
2462:
2456:
2452:
2446:
2435:
2431:
2425:
2414:
2410:
2404:
2395:
2389:
2385:
2379:
2370:
2364:
2360:
2354:
2343:
2339:
2333:
2324:
2318:
2314:
2308:
2302:(11): 10–20.
2297:
2277:
2271:
2265:
2250:
2223:
2215:
2209:
2204:
2196:
2188:
2181:
2170:K. J. Morton
2165:
2145:
2117:
2104:
2102:
2090:
2079:
2062:
2053:
2024:
2007:
1994:
1985:
1976:
1947:
1942:
1929:
1920:
1899:
1877:
1860:
1843:
1826:
1809:
1792:
1775:
1758:
1754:Ruddy darter
1740:
1723:
1706:
1702:Black darter
1689:
1672:
1655:
1638:
1621:
1604:
1594:Status code
1536:Libellulidae
1520:
1503:
1484:
1467:
1450:
1440:Status code
1406:
1396:Status code
1354:
1337:
1320:
1316:Green darner
1302:
1285:
1273:
1265:
1249:Aeshna mixta
1248:
1231:
1214:
1210:Azure hawker
1197:
1180:
1176:Brown hawker
1163:
1153:Status code
1111:
1094:
1084:Status Code
1042:
1032:Status code
998:
981:
964:
946:
929:
912:
894:
877:
860:
843:
826:
809:
792:
782:Status code
694:
676:
659:
643:Lestes dryas
642:
630:
622:
612:Status code
578:
561:
551:Status code
491:
481:
471:
466:
457:
448:
435:
430:
418:English name
411:
402:
389:
381:
376:
348:
336:
320:
307:
299:
287:
282:Tonna, Neath
275:
266:Sandwich Bay
251:
239:
221:Green darner
219:
188:
164:
160:
140:
134:
120:
112:
104:
96:
87:
66:
60:
56:
46:species, or
17:
15:
3141:Vertebrates
3115:Crustaceans
3077:Shield bugs
3042:Butterflies
2912:(8): 21–25.
2895:(4): 18–21.
2815:(2): 47–50.
2681:(2): 50–51.
2465:(1): 21–22.
2453:Anax junius
2432:Anax junius
2419:(9): 28–29.
2398:(1): 20–23.
2373:(1): 22–24.
2348:(2): 40–41.
2327:(1): 11–12.
2317:(Brullé)".
2280:(2): 33–50.
2114:Blue dasher
2068:East Anglia
2000:conspecific
1422:Corduliidae
1321:Anax junius
1274:?RC (2012)
367:River Tamar
359:Moors River
355:River Stour
36:dragonflies
32:damselflies
3221:Categories
3148:Amphibians
3057:Hoverflies
2878:(10): 3–5.
2742:in Avon".
2727:(21): 3–5.
2656:(1): 9–12.
2507:(18): 3–6.
2234:References
2042:Lancashire
1763:Eng/Wales
1660:Eng/Wales
1609:Eng/Wales
1527:Ex (1963)
1310:RC (1996)
1290:Eng/Wales
1253:Eng/Wales
1185:Eng/Wales
1099:Eng/Wales
1047:Eng/Wales
986:Eng/Wales
954:RC (1999)
934:Eng/Wales
902:RC (2010)
848:Eng/Wales
816:Ex (1957)
797:Eng/Wales
631:RC (2007)
195:Cinderford
151:The Lizard
2840:(1): 7–8.
2790:(26): 28.
2486:(9): 3–4.
2440:(6): 3–7.
2174:Inverness
2122:North Sea
2036:in 1823,
2013:Sheerness
1884:V (1823)
1867:V (1859)
1833:V (1995)
1696:V (1995)
1491:V (2018)
1472:Eng/Scot
1361:V (1903)
1327:V (1998)
1272:V (1952)
1127:Aeshnidae
1118:V (1818)
1066:Gomphidae
1056:Suborder
701:V (2008)
684:V (2002)
629:V (1899)
503:Suborder
442:includes
211:Dungeness
197:, in the
3177:See also
3168:Reptiles
3128:Woodlice
3097:Molluscs
3062:Mayflies
2985:Scotland
2929:Archived
2773:(9): 81.
2752:: 17–18.
2635:: 21–26.
2604:Archived
2600:BBC News
2574:Archived
2554:15 March
2124:off the
2019:in 1859.
1965:Archived
1937:in 1818.
594:Lestidae
486:colonist
453:Scotland
408:The list
361:in east
310:vagrants
230:Cornwall
171:Tredegar
155:Cornwall
48:vagrants
40:resident
3163:Mammals
3037:Beetles
3013:Insects
2980:England
2909:Atropos
2892:Atropos
2875:Atropos
2854:Atropos
2787:Atropos
2770:Atropos
2724:Atropos
2610:20 June
2580:15 June
2525:Atropos
2504:Atropos
2483:Atropos
2437:Atropos
2416:Atropos
2345:Atropos
2299:Atropos
2034:Norfolk
2030:Horning
1954:Bristol
1585:Species
1534:Family
1431:Species
1420:Family
1387:Species
1368:Family
1281:Emperor
1144:Species
1125:Family
1075:Species
1064:Family
1023:Species
1012:Family
773:Species
708:Family
603:Species
592:Family
542:Species
511:Family
496:vagrant
476:extinct
444:England
314:extinct
262:Norfolk
183:Norfolk
114:Ireland
98:Atropos
44:extinct
28:Britain
24:Odonata
20:species
2218:) and
2199:) and
2038:Bolton
1958:Jersey
1949:Aeshna
1935:Sussex
1909:exuvia
766:female
416:. The
363:Dorset
123:– see
79:
3153:Birds
3123:Crabs
3087:Wasps
3067:Moths
2995:Wales
2130:Texas
1891:Notes
1797:Scot
1508:Scot
1219:Scot
831:Scot
462:Wales
458:Wales
371:Devon
331:Essex
246:Essex
3158:Fish
3032:Bees
3022:Ants
2612:2018
2582:2019
2556:2014
2257:HMSO
2046:Kent
2017:Kent
1913:Kent
1745:Eng
1626:Eng
1307:Eng
1202:Eng
951:Eng
899:Eng
755:male
744:male
723:male
681:Eng
647:Eng
449:Scot
357:and
270:Kent
215:Kent
77:ISBN
73:HMSO
2207:).
2191:),
2172:in
1748:RC
436:Eng
369:in
329:in
54:.
26:in
22:of
3223::
2836:.
2811:.
2748:.
2704:19
2702:.
2679:13
2677:.
2652:.
2633:32
2631:.
2598:.
2463:15
2461:.
2396:16
2394:.
2371:13
2369:.
2325:12
2323:.
2286:^
2278:12
2276:.
2255:.
2241:^
2152:).
2040:,
2032:,
2015:,
1816:V
1730:V
482:RC
472:Ex
316::
268:,
260:,
228:,
213:,
201:,
181:,
173:,
153:,
149:,
110:.
71:,
2961:e
2954:t
2947:v
2838:1
2813:9
2750:9
2654:4
2614:.
2584:.
2558:.
2544:"
2259:.
2222:(
2214:(
2203:(
2195:(
2187:(
2164:(
2144:(
2132:.
2116:(
2089:(
2070:.
492:V
163:(
83:)
75:(
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