531:
749:
1388:
760:
771:
728:
1584:
739:
1564:
1145:
1574:
1554:
541:
405:– following influxes in previous years, this species, like the yellow-winged darter, has formed temporary breeding colonies. However, since the mid-1990s, the number of these colonies has increased and many have continued to be present from year-to-year, so this species is now better regarded as a successful colonist.
414:
Records of non-native
Odonata species in Britain have been confined to individuals found within heated greenhouses associated with nurseries for aquatic plants. None of these species have been recorded in wild situations or gone on to establish populations in the wild. Details of species which have
116:
A number of other references were used to provide information on specific topics, including rare vagrants, post-1990 additions, predictions, species claimed but not accepted/species of uncertain provenance, non-natives, taxonomic matters and species found only in the
68:
Some of these rare species have not been seen since the 19th
Century; however, the British Odonata list is also currently undergoing a period of unprecedented change, as new species are being discovered for the first time, some going on to become breeding species.
1982:; Grimstone Warren, Norfolk on 21 July; Little Wootton Inclosure New Forest (SZ 227 987) on 6 August; And at the mouth of the Beaulieu River in Hampshire on 10 August. Since 2012 it has probably become established as a breeding species in the Thames Estuary area.
146:
After a period in which the
British Odonata list has been relatively static, since 1990, many new species have been found and some have gone on to become regular breeding species. In chronological order of their first record, these new species are:
2193:
3. The following is a list of species which have previously been predicted to occur in
Britain but have not yet been recorded (see Merritt, Moore & Eversham 1996 pp. 113–114 and Parr 1998, 1999):
3037:
1914:
Willow emerald damselfly, since 2007 this species is established and spreading in south-east
England, especially Suffolk and Essex. It was previously only known from a single record from
3082:
2262:
78:
311:
Many
British Odonata enthusiasts expect further species to be added to the list in the near future. The list below is up to date as of the end of the 2005 breeding season.
2118:
accepted, but the circumstances surrounding the records and/or specimens cast doubt on their natural occurrence, and they are not included in the official
British list:
3092:
2970:
3194:
3247:
3199:
3115:
3057:
530:
3214:
3087:
499:(with the year of the first British record in brackets, except for Red-veined Darter which has occurred as a migrant and sporadic breeder for many decades)
3252:
3067:
2843:
2818:
2755:
2709:
2684:
2659:
2468:
2401:
2376:
2330:
2283:
99:
2584:
3000:
397:– this species has a pattern of establishing small breeding colonies following influxes, but none of these have become permanently established
216:
on 13 June 1996 (Phillips 1997a, 1997b) and now occurring annually; breeding evidence was found at Hayle Kimbro Pool (Jones 2000) and also at
3242:
3138:
3072:
127:'s Odonata fauna is quite different from that of Britain, with many fewer breeding species, but one additional species not found in Britain,
2550:
3052:
2963:
2139:
in early
September 1999 (Parr 2000b). It was not accepted as the species is not a strong migrant and the rig receives weekly supplies from
3158:
1918:
in 1899 (although this record is based on a specimen regarded by some as perhaps not of
British origin, see Gladwin 1997) and a single
135:
3178:
3023:
2900:
Parr, Adrian (1998). "Potential New
Odonata for the British List 1. The Possible Occurrence of Nearctic Species in Western Europe".
2614:
91:
3237:
3173:
3047:
2956:
1975:
478:
The following codes are used to give status details for those species which are not long-established extant breeding species:
3257:
3163:
3097:
3077:
1387:
3168:
3042:
3032:
1971:
since 1998 (Long 2000, Parr 2005). However in 2006 four specimens were seen: In the Adur valley in West Sussex on 13 July
748:
439:
for each breeding species, and an overall status code for species which are not long-established extant breeding species.
2917:
Parr, Adrian (1999). "Potential New Odonata for the British List 2. The Possibility of Vagrant or Colonist Damselflies".
987:
3209:
2203:
428:
2939:
3133:
1490:
682:
442:
The following abbreviations are used to give country-by-country distribution information for the breeding species:
299:
263:
259:
on 17 July 1999 (Dewick and Gerussi 2000) – now a well-established breeding species across much of southern England
759:
2638:
62:
1866:
952:
770:
727:
648:
628:
251:
168:
on 7 August 1995 (Jones 1996), with a small number of further records at scattered locations throughout Britain
509:(with the year of the first record in brackets, except for species with long histories as visitors to Britain)
2211:
1412:
1271:
700:
506:
287:
3204:
2979:
1832:
1583:
1526:
1048:
496:
360:
176:
2580:
2097:
2000:
Yellow-winged darter is listed as a vagrant, but is occasionally subject to large influxes e.g. in 1995.
1100:
970:
918:
774:
243:
on 9 September 1998 – the first record for Europe (Pellow 1999a, 1999b), with several other records in
17:
738:
1729:
1661:
1577:
584:
544:
393:
1644:
1610:
1567:
1557:
1291:
1004:
798:
752:
742:
365:
2636:
Agassiz, D (1981). "Further introduced china mark moths (Lepidoptera: Pyralidae) new to Britain".
2558:
1849:
935:
883:
849:
832:
763:
323:, some do not fall into these two categories. The following species bred in the past but are now
320:
276:
244:
217:
58:
50:
2995:
2919:
2902:
2885:
2864:
2797:
2780:
2734:
2535:
2514:
2493:
2447:
2426:
2355:
2309:
2091:
1. The following species have been claimed but not accepted by the Odonata Records Committee:
1746:
1695:
1563:
1473:
815:
665:
567:
534:
401:
348:
157:
152:
108:
87:
2990:
2222:
2195:
1883:
1509:
1380:
1117:
900:
332:
319:
While most species on the list below are either extant established breeding species or rare
2943:
2606:
2238:
2230:
2160:
2152:
2087:
Rejected species, species of uncertain provenance, predictions and Channel Islands species
1979:
1957:
Southern migrant hawker was known from just a single record, in Kent in 1952, although an
1712:
1360:
1343:
1169:
1024:
866:
731:
432:
424:
415:
occurred in such circumstances can be found in Agassiz 1981, Brooks 1988, and Parr 2000a.
304:
268:
213:
128:
118:
275:
on 30 July 2002 (Nobes 2003); in 2003 & 2004, the species was again seen here and at
2059:
in 1989, although the two 20th Century records may result from accidental introductions.
3005:
2172:
2136:
2105:
1815:
1678:
1308:
1254:
1203:
1144:
720:
523:
450:
353:
337:
236:
209:
200:
189:
1573:
3231:
2187:
in 1926; however Morton was travelling in continental Europe on the date in question.
1967:
may have been this species (Holmes 1993), and the species has been recorded twice in
1915:
1798:
1781:
1627:
1587:
1456:
1237:
185:
38:
2728:
Brook, John; Gill Brook (2004). "Past breeding evidence of Willow Emerald Damselfly
1972:
49:(suborder Anisoptera). Of these, 42 species (17 damselflies and 25 dragonflies) are
2466:(Drury) in Cornwall and Isles of Scilly – The first British and European Records".
2395:
Jones, Steven P. (2000). "First proof of successful breeding by the Lesser Emperor
2180:
1764:
1553:
1546:
1326:
1220:
1186:
292:
231:
2307:
Parr, Adrian (2000a). "An Annotated List of the Odonata of Britain and Ireland".
2124:
2078:
2010:
1432:
540:
436:
377:
369:
2281:
Parr, A. J. (1996). "Dragonfly movement and migration in Britain and Ireland".
283:(Forrest 2005), but there have been no subsequent sightings at either location.
106:
Parr, Adrian (2000a), An Annotated List of the Odonata of Britain and Ireland,
2052:
486:
324:
205:
161:
54:
2445:(Drury) into Cornwall and the Isles of Scilly – The First European Records".
97:
Parr, A. J. (1996), Dragonfly movement and migration in Britain and Ireland,
2324:
Jones, Steven P. (1996). "The first British record of the Scarlet Dragonfly
2184:
2132:
2023:
1137:
1076:
1068:
515:
221:
220:, Cornwall (Pellow 2000) in 1999, and the species appears to be resident at
46:
42:
2179:), only known from a single specimen labelled as having been collected by
2948:
2678:
Gladwin, T.W. (1997). "The error in treating the Green Emerald Damselfly
604:
463:
376:, where the species was recorded from 1820 to 1963, and the other on the
240:
181:
165:
2159:), only known from old specimens with vague data (but recorded from the
344:, first recorded in 1946, and last recorded in 1952. Rediscovered 2010.
2044:
2040:
1964:
454:
272:
193:
124:
34:
30:
2048:
1968:
1959:
1945:
1919:
373:
364:– only ever known from two areas in southern England, one around the
2936:
2491:(Charpentier) Found Breeding in Essex – The First British Records".
2487:
Dewick, Stephen; Richard Gerussi (2000). "Small Red-eyed Damselfly
61:- in respect of the latter, this list follows the decisions of the
2140:
1582:
1572:
1562:
1552:
472:
381:
341:
256:
2267:
2056:
2027:
1923:
280:
225:
83:
2952:
1944:
Yellow-legged dragonfly is known from just a single record, in
196:, which had dark bands on its wings, may have been this species
2703:
Brook, John; Gill Brook (2003). "The Willow Emerald Damselfly
188:
on 16 August 1995, although a darter seen on 29 July 2003 at
2657:
Brooks, S. J. (1988). "Exotic dragonflies in north London".
1935:
Common clubtail was formerly known as club-tailed dragonfly.
495:
to indicate that the species is a recent (i.e. post-1990)
72:
This list is based on the following principal references:
2795:
Parr, Adrian (2005). "Odonata Records Committee update".
2039:
Wandering glider has been recorded only three times – at
2022:
Large white-faced darter has been recorded only once, at
489:
in Britain (with the year of the last record in brackets)
2131:), which was found dead on the Sedco 706 oil rig in the
2707:(Vander Linden) in Kent: a case of mistaken identity".
2374:(Sélys) in Gloucestershire; the first British record".
2114:
2. The following species have been recorded, and their
388:
The following species are sporadic or casual breeders:
2349:
Phillips, John (1997a). "The Lesser Emperor Dragonfly
471:
to indicate that the species' breeding range includes
462:
to indicate that the species' breeding range includes
2068:
Wandering glider was formerly known as globe skimmer.
2862:
Parr, Adrian (2003). "Migrant Dragonflies in 2002".
2077:
Habitat limited to Eastern England and particularly
76:
Merritt, R., N. W. Moore and B. C. Eversham (1996),
3187:
3151:
3124:
3106:
3021:
3014:
2529:Forrest, P. J. (2005). "Southern Emerald Damselfly
2441:Pellow, Keith (1999a). "An influx of Green Darner
2370:Phillips, John (1997b). "Lesser Emperor Dragonfly
2508:Nobes, Geoff (2003). "Southern Emerald Damselfly
2260:Merritt, R.; N. W. Moore; B. C. Eversham (1996).
1991:Vagrant emperor records taken from Silsby (1993).
2420:Pellow, Keith (2000). "Lesser Emperor Dragonfly
2353:in Gloucestershire; The First British Record".
2263:Atlas of the dragonflies of Britain and Ireland
79:Atlas of the dragonflies of Britain and Ireland
2883:(Burmeister) on an Oil Rig in the North Sea".
2749:Holmes, J. D. (1993). "A possible sighting of
2964:
2774:Long, Roger (2000). "Southern Migrant Hawker
2607:"Beautiful 'lost' insect turns up anew in UK"
8:
2462:Pellow, Keith (1999b). "Common Green Darner
2221:4. In addition to the species listed above,
384:where the species was recorded in 1946 only.
2583:. British Dragonfly Society. 11 July 2018.
2557:. British Dragonfly Society. Archived from
485:to indicate a former breeding species, now
3018:
2971:
2957:
2949:
2302:
2300:
2298:
431:of each species is given, followed by its
2017:
2009:Highland darter and common darter may be
505:to indicate a species only recorded as a
336:– only ever known from marshes along the
2844:Journal of the British Dragonfly Society
2819:Journal of the British Dragonfly Society
2756:Journal of the British Dragonfly Society
2710:Journal of the British Dragonfly Society
2685:Journal of the British Dragonfly Society
2660:Journal of the British Dragonfly Society
2469:Journal of the British Dragonfly Society
2402:Journal of the British Dragonfly Society
2377:Journal of the British Dragonfly Society
2331:Journal of the British Dragonfly Society
2284:Journal of the British Dragonfly Society
2004:
1986:
1592:
1438:
1394:
1386:
1151:
1143:
1082:
1030:
780:
769:
758:
747:
737:
726:
723:(blue, blue-tailed, and red damselflies)
610:
549:
539:
529:
100:Journal of the British Dragonfly Society
3195:Endangered species of the British Isles
2682:(Vander Linden) as a British species".
2255:
2253:
2249:
2072:
1926:in 1992 (Brook & Brook 2002, 2004).
1909:
53:breeders, and the remainder are either
18:List of dragonflies recorded in Britain
1939:
3248:Lists of insects in the British Isles
2617:from the original on 27 November 2017
2512:(Fabr.) – The First British Record".
1952:
7:
3200:Endemic species of the British Isles
2837:Merritt, R.; G. S. Vick (1983). "Is
180:), recorded just once to date, near
3215:Birds of Leicestershire and Rutland
2879:Parr, Adrian (2000b). "Blue Dasher
2581:"New Dragonfly Species for Britain"
315:Extinct species and casual breeders
247:that autumn, but none since to date
1995:
1930:
136:List of Odonata species of Ireland
25:
3253:Lists of insects of Great Britain
2587:from the original on 13 July 2019
2063:
2034:
1963:seen in 1992 on the outskirts of
2812:Silsby, J. (1993). "A review of
2533:(Fabr.) at Sandwich Bay, Kent".
1549:(chasers, skimmers, and darters)
295:in South Wales in December 2008.
2424:(Selys) breeding in Cornwall".
1804:Sympetrum striolatum nigrescens
2013:(see Merritt & Vick 1983).
1973:website of reporter with photo
449:to indicate that the species'
1:
3083:Orthoptera and allied insects
2778:in Jersey, Channel Islands".
303:, first (and only) record at
3243:Lists of Odonata by location
2237:) have been recorded in the
988:Scarce blue-tailed damselfly
103:Vol. 12 No. 2 pp. 33–50
45:(suborder Zygoptera) and 36
3210:Flora and fauna of Cornwall
3063:Damselflies and dragonflies
2204:northern white-faced darter
1383:(golden-ringed dragonflies)
352:– only ever known from the
3274:
2732:(Vander Linden) in Kent".
2108:in 2002 – see Parr (2003).
1496:Somatochlora flavomaculata
1027:(white-legged damselflies)
683:Southern emerald damselfly
264:Southern emerald damselfly
3038:Aquatic heteropteran bugs
2986:
2937:British Dragonfly Society
1079:(club-tailed dragonflies)
63:Odonata Records Committee
2816:, the Vagrant Emperor".
1867:Large white-faced darter
1650:Libellula quadrimaculata
1069:Anisoptera (dragonflies)
953:Small red-eyed damselfly
649:Scarce emerald damselfly
629:Willow emerald damselfly
410:Non-native introductions
307:, Suffolk, in July 2018.
252:Small red-eyed damselfly
3238:Lists of insect species
3093:Soldierflies and allies
2881:Pachydiplax longipennis
2212:small emerald damselfly
2129:Pachydiplax longipennis
1872:Leucorrhinia pectoralis
1413:Golden-ringed dragonfly
1391:Golden-ringed dragonfly
1272:Southern migrant hawker
1118:Yellow-legged dragonfly
701:Common winter damselfly
516:Zygoptera (damselflies)
288:Common winter damselfly
2980:Fauna of Great Britain
2942:28 August 2011 at the
2639:Entomologist's Gazette
2177:Somatochlora alpestris
1838:Sympetrum pedemontanum
1752:Sympetrum fonscolombei
1684:Orthetrum coerulescens
1590:
1580:
1570:
1560:
1527:Orange-spotted emerald
1491:Yellow-spotted emerald
1479:Somatochlora metallica
1418:Cordulegaster boltonii
1392:
1149:
1140:(hawkers and emperors)
1049:White-legged damselfly
778:
767:
756:
745:
735:
547:
537:
361:Orange-spotted emerald
300:Yellow-spotted emerald
177:Sympetrum pedemontanum
142:New species since 1990
138:for more information.
29:There are 57 recorded
3258:Dragonflies of Europe
2399:(Sélys) in Britain".
2326:Crocothemis erythraea
2235:Sympetrum meridionale
2208:Leucorrhina rubicunda
2157:Sympetrum meridionale
2098:Two-spotted dragonfly
1701:Crocothemis erythraea
1667:Orthetrum cancellatum
1586:
1576:
1566:
1556:
1435:(emerald dragonflies)
1390:
1147:
1106:Gomphus vulgatissimus
971:Blue-tailed damselfly
924:Enallagma cyathigerum
919:Common blue damselfly
889:Coenagrion pulchellum
855:Coenagrion mercuriale
838:Coenagrion hastulatum
775:Blue-tailed damselfly
773:
762:
751:
741:
730:
607:(emerald damselflies)
543:
533:
356:between 1902 and 1957
112:No. 11 pp. 10–20
2839:Sympetrum nigrescens
2730:Chalcolestes viridis
2705:Chalcolestes viridis
2489:Erythromma viridulum
2104:) was reported from
1978:2 March 2007 at the
1787:Sympetrum striolatum
1770:Sympetrum sanguineum
1730:Yellow-winged darter
1662:Black-tailed skimmer
1578:Black-tailed skimmer
1515:Somatochlora arctica
1054:Platycnemis pennipes
958:Erythromma viridulum
634:Chalcolestes viridis
585:Beautiful demoiselle
573:Calopteryx splendens
545:Beautiful demoiselle
394:Yellow-winged darter
291:, first recorded at
267:, first recorded at
255:, first recorded in
235:, first recorded at
204:, first recorded at
156:, first recorded at
132:Coenagrion lunulatum
3116:Non-marine molluscs
2814:Hemianax ephippiger
2102:Epitheca bimaculata
1735:Sympetrum flaveolum
1645:Four-spotted chaser
1611:Broad-bodied chaser
1568:Four-spotted chaser
1558:Broad-bodied chaser
1366:Hemianax ephippiger
1349:Brachytron pratense
1010:Pyrrhosoma nymphula
1005:Large red damselfly
906:Coenagrion scitulum
804:Ceriagrion tenellum
799:Small red damselfly
753:Large red damselfly
743:Small red damselfly
3001:Scottish Highlands
2841:a good species?".
2551:"Winter Damselfly
2227:Orthetrum brunneum
1889:Pantala flavescens
1855:Leucorrhinia dubia
1850:White-faced darter
1821:Sympetrum vulgatum
1616:Libellula depressa
1591:
1581:
1571:
1561:
1532:Oxygastra curtisii
1393:
1150:
936:Red-eyed damselfly
884:Variable damselfly
850:Southern damselfly
833:Northern damselfly
821:Coenagrion armatum
779:
768:
764:Red-eyed damselfly
757:
746:
736:
548:
538:
245:South West England
218:Bake Fishing Lakes
3223:
3222:
3147:
3146:
2717:(1&2): 51–54.
2200:Aeshna subarctica
1899:
1898:
1747:Red-veined darter
1696:Scarlet dragonfly
1542:
1541:
1474:Brilliant emerald
1428:
1427:
1376:
1375:
1133:
1132:
1064:
1063:
1020:
1019:
872:Coenagrion puella
816:Norfolk damselfly
716:
715:
666:Emerald damselfly
600:
599:
568:Banded demoiselle
535:Banded demoiselle
435:, details of the
402:Red-veined darter
349:Norfolk damselfly
158:Hayle Kimbro Pool
153:Scarlet dragonfly
16:(Redirected from
3265:
3019:
2973:
2966:
2959:
2950:
2925:
2924:
2914:
2908:
2907:
2897:
2891:
2890:
2876:
2870:
2869:
2859:
2853:
2852:
2834:
2828:
2827:
2809:
2803:
2802:
2792:
2786:
2785:
2771:
2765:
2764:
2746:
2740:
2739:
2725:
2719:
2718:
2700:
2694:
2693:
2675:
2669:
2668:
2654:
2648:
2647:
2633:
2627:
2626:
2624:
2622:
2613:. 21 July 2010.
2603:
2597:
2596:
2594:
2592:
2577:
2571:
2570:
2568:
2566:
2561:on 16 March 2014
2547:
2541:
2540:
2526:
2520:
2519:
2505:
2499:
2498:
2484:
2478:
2477:
2459:
2453:
2452:
2438:
2432:
2431:
2417:
2411:
2410:
2392:
2386:
2385:
2367:
2361:
2360:
2346:
2340:
2339:
2321:
2315:
2314:
2304:
2293:
2292:
2278:
2272:
2271:
2257:
2223:southern skimmer
2196:subarctic darner
2076:
2067:
2038:
2021:
2008:
1999:
1990:
1956:
1943:
1934:
1913:
1884:Wandering glider
1593:
1510:Northern emerald
1439:
1395:
1381:Cordulegastridae
1209:Aeshna isosceles
1152:
1123:Gomphus flavipes
1083:
1031:
993:Ischnura pumilio
976:Ischnura elegans
941:Erythromma najas
901:Dainty damselfly
781:
611:
590:Calopteryx virgo
550:
333:Dainty damselfly
41:, made up of 21
21:
3273:
3272:
3268:
3267:
3266:
3264:
3263:
3262:
3228:
3227:
3224:
3219:
3183:
3143:
3120:
3102:
3010:
2982:
2977:
2944:Wayback Machine
2933:
2928:
2916:
2915:
2911:
2899:
2898:
2894:
2878:
2877:
2873:
2861:
2860:
2856:
2836:
2835:
2831:
2811:
2810:
2806:
2794:
2793:
2789:
2773:
2772:
2768:
2748:
2747:
2743:
2727:
2726:
2722:
2702:
2701:
2697:
2677:
2676:
2672:
2656:
2655:
2651:
2635:
2634:
2630:
2620:
2618:
2605:
2604:
2600:
2590:
2588:
2579:
2578:
2574:
2564:
2562:
2549:
2548:
2544:
2531:Lestes barbarus
2528:
2527:
2523:
2510:Lestes barbarus
2507:
2506:
2502:
2486:
2485:
2481:
2461:
2460:
2456:
2440:
2439:
2435:
2422:Anax parthenope
2419:
2418:
2414:
2397:Anax parthenope
2394:
2393:
2389:
2372:Anax parthenope
2369:
2368:
2364:
2351:Anax parthenope
2348:
2347:
2343:
2323:
2322:
2318:
2306:
2305:
2296:
2280:
2279:
2275:
2259:
2258:
2251:
2247:
2239:Channel Islands
2231:southern darter
2161:Channel Islands
2153:Southern darter
2089:
2084:
2071:
2062:
2055:in 1951 and in
2033:
2016:
2003:
1994:
1985:
1980:Wayback Machine
1951:
1938:
1929:
1908:
1904:
1859:Eng/Scot/Wales
1799:Highland darter
1791:Eng/Scot/Wales
1722:Eng/Scot/Wales
1718:Sympetrum danae
1688:Eng/Scot/Wales
1654:Eng/Scot/Wales
1633:Libellula fulva
1602:Range countries
1599:Scientific name
1551:
1466:Eng/Scot/Wales
1448:Range countries
1445:Scientific name
1437:
1422:Eng/Scot/Wales
1404:Range countries
1401:Scientific name
1385:
1361:Vagrant emperor
1353:Eng/Scot/Wales
1344:Hairy dragonfly
1314:Anax parthenope
1247:Eng/Scot/Wales
1226:Aeshna caerulea
1179:Eng/Scot/Wales
1170:Southern hawker
1161:Range countries
1158:Scientific name
1142:
1101:Common clubtail
1092:Range countries
1089:Scientific name
1081:
1072:
1040:Range countries
1037:Scientific name
1029:
1025:Platycnemididae
1014:Eng/Scot/Wales
980:Eng/Scot/Wales
928:Eng/Scot/Wales
893:Eng/Scot/Wales
876:Eng/Scot/Wales
867:Azure damselfly
790:Range countries
787:Scientific name
732:Azure damselfly
725:
688:Lestes barbarus
675:Eng/Scot/Wales
620:Range countries
617:Scientific name
609:
594:Eng/Scot/Wales
577:Eng/Scot/Wales
559:Range countries
556:Scientific name
528:
519:
437:range countries
433:scientific name
425:taxonomic order
423:The list is in
421:
412:
317:
305:Carlton Marshes
269:Winterton Dunes
214:Gloucestershire
144:
129:Irish Damselfly
119:Channel Islands
23:
22:
15:
12:
11:
5:
3271:
3269:
3261:
3260:
3255:
3250:
3245:
3240:
3230:
3229:
3221:
3220:
3218:
3217:
3212:
3207:
3205:Birds of Wales
3202:
3197:
3191:
3189:
3185:
3184:
3182:
3181:
3176:
3171:
3166:
3161:
3155:
3153:
3149:
3148:
3145:
3144:
3142:
3141:
3136:
3130:
3128:
3122:
3121:
3119:
3118:
3112:
3110:
3104:
3103:
3101:
3100:
3095:
3090:
3085:
3080:
3075:
3070:
3065:
3060:
3055:
3050:
3045:
3040:
3035:
3029:
3027:
3016:
3012:
3011:
3009:
3008:
3003:
2998:
2993:
2987:
2984:
2983:
2978:
2976:
2975:
2968:
2961:
2953:
2947:
2946:
2932:
2931:External links
2929:
2927:
2926:
2909:
2892:
2871:
2854:
2829:
2804:
2787:
2776:Aeshna affinis
2766:
2751:Aeshna affinis
2741:
2720:
2695:
2680:Lestes viridis
2670:
2649:
2628:
2598:
2572:
2553:Sympecma fusca
2542:
2521:
2500:
2479:
2454:
2433:
2412:
2387:
2362:
2341:
2316:
2294:
2273:
2248:
2246:
2243:
2241:(Parr 2000a).
2191:
2190:
2189:
2188:
2173:Alpine emerald
2167:
2166:
2165:
2164:
2147:
2146:
2145:
2144:
2137:Shetland Isles
2116:identification
2112:
2111:
2110:
2109:
2106:Leicestershire
2088:
2085:
2083:
2082:
2069:
2060:
2031:
2014:
2001:
1992:
1983:
1949:
1936:
1927:
1905:
1903:
1900:
1897:
1896:
1893:
1891:
1886:
1880:
1879:
1876:
1874:
1869:
1863:
1862:
1860:
1857:
1852:
1846:
1845:
1842:
1840:
1835:
1829:
1828:
1825:
1823:
1818:
1816:Vagrant darter
1812:
1811:
1809:
1806:
1801:
1795:
1794:
1792:
1789:
1784:
1778:
1777:
1775:
1772:
1767:
1761:
1760:
1757:
1754:
1749:
1743:
1742:
1739:
1737:
1732:
1726:
1725:
1723:
1720:
1715:
1709:
1708:
1705:
1703:
1698:
1692:
1691:
1689:
1686:
1681:
1679:Keeled skimmer
1675:
1674:
1672:
1669:
1664:
1658:
1657:
1655:
1652:
1647:
1641:
1640:
1638:
1635:
1630:
1624:
1623:
1621:
1618:
1613:
1607:
1606:
1603:
1600:
1597:
1550:
1543:
1540:
1539:
1536:
1534:
1529:
1523:
1522:
1520:
1517:
1512:
1506:
1505:
1503:
1500:
1498:
1493:
1487:
1486:
1484:
1481:
1476:
1470:
1469:
1467:
1464:
1462:Cordulia aenea
1459:
1453:
1452:
1449:
1446:
1443:
1436:
1429:
1426:
1425:
1423:
1420:
1415:
1409:
1408:
1405:
1402:
1399:
1384:
1377:
1374:
1373:
1370:
1368:
1363:
1357:
1356:
1354:
1351:
1346:
1340:
1339:
1336:
1334:
1329:
1323:
1322:
1319:
1316:
1311:
1309:Lesser emperor
1305:
1304:
1302:
1299:
1297:Anax imperator
1294:
1288:
1287:
1281:
1279:
1277:Aeshna affinis
1274:
1268:
1267:
1265:
1262:
1257:
1255:Migrant hawker
1251:
1250:
1248:
1245:
1240:
1234:
1233:
1231:
1228:
1223:
1217:
1216:
1214:
1211:
1206:
1204:Norfolk hawker
1200:
1199:
1197:
1194:
1192:Aeshna grandis
1189:
1183:
1182:
1180:
1177:
1172:
1166:
1165:
1162:
1159:
1156:
1141:
1134:
1131:
1130:
1127:
1125:
1120:
1114:
1113:
1111:
1108:
1103:
1097:
1096:
1093:
1090:
1087:
1080:
1073:
1071:
1065:
1062:
1061:
1059:
1056:
1051:
1045:
1044:
1041:
1038:
1035:
1028:
1021:
1018:
1017:
1015:
1012:
1007:
1001:
1000:
998:
995:
990:
984:
983:
981:
978:
973:
967:
966:
963:
960:
955:
949:
948:
946:
943:
938:
932:
931:
929:
926:
921:
915:
914:
911:
908:
903:
897:
896:
894:
891:
886:
880:
879:
877:
874:
869:
863:
862:
860:
857:
852:
846:
845:
843:
840:
835:
829:
828:
825:
823:
818:
812:
811:
809:
806:
801:
795:
794:
791:
788:
785:
724:
721:Coenagrionidae
717:
714:
713:
710:
708:
706:Sympecma fusca
703:
697:
696:
693:
690:
685:
679:
678:
676:
673:
668:
662:
661:
659:
656:
651:
645:
644:
638:
636:
631:
625:
624:
621:
618:
615:
608:
601:
598:
597:
595:
592:
587:
581:
580:
578:
575:
570:
564:
563:
560:
557:
554:
527:
524:Calopterygidae
520:
518:
512:
511:
510:
500:
490:
476:
475:
466:
457:
451:breeding range
420:
417:
411:
408:
407:
406:
398:
386:
385:
357:
354:Norfolk Broads
345:
338:Thames Estuary
316:
313:
309:
308:
296:
284:
260:
248:
237:Penlee Battery
228:
210:Forest of Dean
201:Lesser emperor
197:
190:Hickling Broad
169:
143:
140:
114:
113:
104:
95:
24:
14:
13:
10:
9:
6:
4:
3:
2:
3270:
3259:
3256:
3254:
3251:
3249:
3246:
3244:
3241:
3239:
3236:
3235:
3233:
3226:
3216:
3213:
3211:
3208:
3206:
3203:
3201:
3198:
3196:
3193:
3192:
3190:
3186:
3180:
3177:
3175:
3172:
3170:
3167:
3165:
3162:
3160:
3157:
3156:
3154:
3150:
3140:
3137:
3135:
3132:
3131:
3129:
3127:
3123:
3117:
3114:
3113:
3111:
3109:
3105:
3099:
3096:
3094:
3091:
3089:
3086:
3084:
3081:
3079:
3076:
3074:
3071:
3069:
3066:
3064:
3061:
3059:
3058:Conopid flies
3056:
3054:
3051:
3049:
3046:
3044:
3041:
3039:
3036:
3034:
3031:
3030:
3028:
3026:
3025:
3020:
3017:
3015:Invertebrates
3013:
3007:
3004:
3002:
2999:
2997:
2994:
2992:
2989:
2988:
2985:
2981:
2974:
2969:
2967:
2962:
2960:
2955:
2954:
2951:
2945:
2941:
2938:
2935:
2934:
2930:
2922:
2921:
2913:
2910:
2905:
2904:
2896:
2893:
2888:
2887:
2882:
2875:
2872:
2867:
2866:
2858:
2855:
2850:
2846:
2845:
2840:
2833:
2830:
2825:
2821:
2820:
2815:
2808:
2805:
2800:
2799:
2791:
2788:
2783:
2782:
2777:
2770:
2767:
2762:
2758:
2757:
2752:
2745:
2742:
2737:
2736:
2731:
2724:
2721:
2716:
2712:
2711:
2706:
2699:
2696:
2691:
2687:
2686:
2681:
2674:
2671:
2666:
2662:
2661:
2653:
2650:
2645:
2641:
2640:
2632:
2629:
2616:
2612:
2608:
2602:
2599:
2586:
2582:
2576:
2573:
2560:
2556:
2554:
2546:
2543:
2538:
2537:
2532:
2525:
2522:
2517:
2516:
2511:
2504:
2501:
2496:
2495:
2490:
2483:
2480:
2475:
2471:
2470:
2465:
2458:
2455:
2450:
2449:
2444:
2437:
2434:
2429:
2428:
2423:
2416:
2413:
2408:
2404:
2403:
2398:
2391:
2388:
2383:
2379:
2378:
2373:
2366:
2363:
2358:
2357:
2352:
2345:
2342:
2337:
2333:
2332:
2327:
2320:
2317:
2312:
2311:
2303:
2301:
2299:
2295:
2290:
2286:
2285:
2277:
2274:
2269:
2265:
2264:
2256:
2254:
2250:
2244:
2242:
2240:
2236:
2232:
2228:
2224:
2219:
2217:
2216:Lestes virens
2213:
2209:
2205:
2201:
2197:
2186:
2182:
2178:
2174:
2171:
2170:
2169:
2168:
2162:
2158:
2154:
2151:
2150:
2149:
2148:
2142:
2138:
2134:
2130:
2126:
2123:
2122:
2121:
2120:
2119:
2117:
2107:
2103:
2099:
2096:
2095:
2094:
2093:
2092:
2086:
2080:
2075:
2074:
2070:
2066:
2065:
2061:
2058:
2054:
2050:
2046:
2042:
2037:
2036:
2032:
2029:
2025:
2020:
2019:
2015:
2012:
2007:
2006:
2002:
1998:
1997:
1993:
1989:
1988:
1984:
1981:
1977:
1974:
1970:
1966:
1962:
1961:
1955:
1954:
1950:
1947:
1942:
1941:
1937:
1933:
1932:
1928:
1925:
1922:collected in
1921:
1917:
1916:Hertfordshire
1912:
1911:
1907:
1906:
1901:
1894:
1892:
1890:
1887:
1885:
1882:
1881:
1877:
1875:
1873:
1870:
1868:
1865:
1864:
1861:
1858:
1856:
1853:
1851:
1848:
1847:
1843:
1841:
1839:
1836:
1834:
1833:Banded darter
1831:
1830:
1826:
1824:
1822:
1819:
1817:
1814:
1813:
1810:
1807:
1805:
1802:
1800:
1797:
1796:
1793:
1790:
1788:
1785:
1783:
1782:Common darter
1780:
1779:
1776:
1773:
1771:
1768:
1766:
1763:
1762:
1758:
1755:
1753:
1750:
1748:
1745:
1744:
1740:
1738:
1736:
1733:
1731:
1728:
1727:
1724:
1721:
1719:
1716:
1714:
1711:
1710:
1706:
1704:
1702:
1699:
1697:
1694:
1693:
1690:
1687:
1685:
1682:
1680:
1677:
1676:
1673:
1670:
1668:
1665:
1663:
1660:
1659:
1656:
1653:
1651:
1648:
1646:
1643:
1642:
1639:
1636:
1634:
1631:
1629:
1628:Scarce chaser
1626:
1625:
1622:
1619:
1617:
1614:
1612:
1609:
1608:
1604:
1601:
1598:
1595:
1594:
1589:
1588:Common darter
1585:
1579:
1575:
1569:
1565:
1559:
1555:
1548:
1544:
1537:
1535:
1533:
1530:
1528:
1525:
1524:
1521:
1518:
1516:
1513:
1511:
1508:
1507:
1504:
1501:
1499:
1497:
1494:
1492:
1489:
1488:
1485:
1482:
1480:
1477:
1475:
1472:
1471:
1468:
1465:
1463:
1460:
1458:
1457:Downy emerald
1455:
1454:
1450:
1447:
1444:
1441:
1440:
1434:
1430:
1424:
1421:
1419:
1416:
1414:
1411:
1410:
1406:
1403:
1400:
1397:
1396:
1389:
1382:
1378:
1371:
1369:
1367:
1364:
1362:
1359:
1358:
1355:
1352:
1350:
1347:
1345:
1342:
1341:
1337:
1335:
1333:
1330:
1328:
1325:
1324:
1320:
1317:
1315:
1312:
1310:
1307:
1306:
1303:
1300:
1298:
1295:
1293:
1290:
1289:
1286:
1282:
1280:
1278:
1275:
1273:
1270:
1269:
1266:
1263:
1261:
1258:
1256:
1253:
1252:
1249:
1246:
1244:
1243:Aeshna juncea
1241:
1239:
1238:Common hawker
1236:
1235:
1232:
1229:
1227:
1224:
1222:
1219:
1218:
1215:
1212:
1210:
1207:
1205:
1202:
1201:
1198:
1195:
1193:
1190:
1188:
1185:
1184:
1181:
1178:
1176:
1175:Aeshna cyanea
1173:
1171:
1168:
1167:
1163:
1160:
1157:
1154:
1153:
1148:Common hawker
1146:
1139:
1135:
1128:
1126:
1124:
1121:
1119:
1116:
1115:
1112:
1109:
1107:
1104:
1102:
1099:
1098:
1094:
1091:
1088:
1085:
1084:
1078:
1074:
1070:
1066:
1060:
1057:
1055:
1052:
1050:
1047:
1046:
1042:
1039:
1036:
1033:
1032:
1026:
1022:
1016:
1013:
1011:
1008:
1006:
1003:
1002:
999:
996:
994:
991:
989:
986:
985:
982:
979:
977:
974:
972:
969:
968:
964:
961:
959:
956:
954:
951:
950:
947:
944:
942:
939:
937:
934:
933:
930:
927:
925:
922:
920:
917:
916:
912:
909:
907:
904:
902:
899:
898:
895:
892:
890:
887:
885:
882:
881:
878:
875:
873:
870:
868:
865:
864:
861:
858:
856:
853:
851:
848:
847:
844:
841:
839:
836:
834:
831:
830:
826:
824:
822:
819:
817:
814:
813:
810:
807:
805:
802:
800:
797:
796:
792:
789:
786:
783:
782:
776:
772:
765:
761:
754:
750:
744:
740:
733:
729:
722:
718:
711:
709:
707:
704:
702:
699:
698:
694:
691:
689:
686:
684:
681:
680:
677:
674:
672:
671:Lestes sponsa
669:
667:
664:
663:
660:
657:
655:
652:
650:
647:
646:
643:
639:
637:
635:
632:
630:
627:
626:
622:
619:
616:
613:
612:
606:
602:
596:
593:
591:
588:
586:
583:
582:
579:
576:
574:
571:
569:
566:
565:
561:
558:
555:
552:
551:
546:
542:
536:
532:
526:(demoiselles)
525:
521:
517:
513:
508:
504:
501:
498:
494:
491:
488:
484:
481:
480:
479:
474:
470:
467:
465:
461:
458:
456:
452:
448:
445:
444:
443:
440:
438:
434:
430:
426:
418:
416:
409:
404:
403:
399:
396:
395:
391:
390:
389:
383:
379:
375:
371:
367:
363:
362:
358:
355:
351:
350:
346:
343:
339:
335:
334:
330:
329:
328:
326:
322:
314:
312:
306:
302:
301:
297:
294:
290:
289:
285:
282:
278:
274:
270:
266:
265:
261:
258:
254:
253:
249:
246:
242:
238:
234:
233:
229:
227:
223:
219:
215:
211:
207:
203:
202:
198:
195:
191:
187:
186:Monmouthshire
183:
179:
178:
173:
172:Banded darter
170:
167:
163:
159:
155:
154:
150:
149:
148:
141:
139:
137:
133:
130:
126:
122:
120:
111:
110:
105:
102:
101:
96:
93:
92:0-11-701561-X
89:
85:
81:
80:
75:
74:
73:
70:
66:
64:
60:
56:
52:
48:
44:
40:
36:
32:
27:
19:
3225:
3125:
3107:
3062:
3022:
2918:
2912:
2901:
2895:
2884:
2880:
2874:
2868:(18): 18–24.
2863:
2857:
2848:
2842:
2838:
2832:
2823:
2817:
2813:
2807:
2796:
2790:
2779:
2775:
2769:
2760:
2754:
2750:
2744:
2733:
2729:
2723:
2714:
2708:
2704:
2698:
2689:
2683:
2679:
2673:
2664:
2658:
2652:
2643:
2637:
2631:
2619:. Retrieved
2610:
2601:
2589:. Retrieved
2575:
2563:. Retrieved
2559:the original
2552:
2545:
2539:(24): 24–25.
2534:
2530:
2524:
2513:
2509:
2503:
2492:
2488:
2482:
2473:
2467:
2463:
2457:
2446:
2442:
2436:
2425:
2421:
2415:
2406:
2400:
2396:
2390:
2381:
2375:
2371:
2365:
2354:
2350:
2344:
2335:
2329:
2325:
2319:
2313:(11): 10–20.
2308:
2288:
2282:
2276:
2261:
2234:
2226:
2220:
2215:
2207:
2199:
2192:
2181:K. J. Morton
2176:
2156:
2128:
2115:
2113:
2101:
2090:
2073:
2064:
2035:
2018:
2005:
1996:
1987:
1958:
1953:
1940:
1931:
1910:
1888:
1871:
1854:
1837:
1820:
1803:
1786:
1769:
1765:Ruddy darter
1751:
1734:
1717:
1713:Black darter
1700:
1683:
1666:
1649:
1632:
1615:
1605:Status code
1547:Libellulidae
1531:
1514:
1495:
1478:
1461:
1451:Status code
1417:
1407:Status code
1365:
1348:
1331:
1327:Green darner
1313:
1296:
1284:
1276:
1260:Aeshna mixta
1259:
1242:
1225:
1221:Azure hawker
1208:
1191:
1187:Brown hawker
1174:
1164:Status code
1122:
1105:
1095:Status Code
1053:
1043:Status code
1009:
992:
975:
957:
940:
923:
905:
888:
871:
854:
837:
820:
803:
793:Status code
705:
687:
670:
654:Lestes dryas
653:
641:
633:
623:Status code
589:
572:
562:Status code
502:
492:
482:
477:
468:
459:
446:
441:
429:English name
422:
413:
400:
392:
387:
359:
347:
331:
318:
310:
298:
293:Tonna, Neath
286:
277:Sandwich Bay
262:
250:
232:Green darner
230:
199:
175:
171:
151:
145:
131:
123:
115:
107:
98:
77:
71:
67:
57:species, or
28:
26:
3152:Vertebrates
3126:Crustaceans
3088:Shield bugs
3053:Butterflies
2923:(8): 21–25.
2906:(4): 18–21.
2826:(2): 47–50.
2692:(2): 50–51.
2476:(1): 21–22.
2464:Anax junius
2443:Anax junius
2430:(9): 28–29.
2409:(1): 20–23.
2384:(1): 22–24.
2359:(2): 40–41.
2338:(1): 11–12.
2328:(Brullé)".
2291:(2): 33–50.
2125:Blue dasher
2079:East Anglia
2011:conspecific
1433:Corduliidae
1332:Anax junius
1285:?RC (2012)
378:River Tamar
370:Moors River
366:River Stour
47:dragonflies
43:damselflies
3232:Categories
3159:Amphibians
3068:Hoverflies
2889:(10): 3–5.
2753:in Avon".
2738:(21): 3–5.
2667:(1): 9–12.
2518:(18): 3–6.
2245:References
2053:Lancashire
1774:Eng/Wales
1671:Eng/Wales
1620:Eng/Wales
1538:Ex (1963)
1321:RC (1996)
1301:Eng/Wales
1264:Eng/Wales
1196:Eng/Wales
1110:Eng/Wales
1058:Eng/Wales
997:Eng/Wales
965:RC (1999)
945:Eng/Wales
913:RC (2010)
859:Eng/Wales
827:Ex (1957)
808:Eng/Wales
642:RC (2007)
206:Cinderford
162:The Lizard
2851:(1): 7–8.
2801:(26): 28.
2497:(9): 3–4.
2451:(6): 3–7.
2185:Inverness
2133:North Sea
2047:in 1823,
2024:Sheerness
1895:V (1823)
1878:V (1859)
1844:V (1995)
1707:V (1995)
1502:V (2018)
1483:Eng/Scot
1372:V (1903)
1338:V (1998)
1283:V (1952)
1138:Aeshnidae
1129:V (1818)
1077:Gomphidae
1067:Suborder
712:V (2008)
695:V (2002)
640:V (1899)
514:Suborder
453:includes
222:Dungeness
208:, in the
3188:See also
3179:Reptiles
3139:Woodlice
3108:Molluscs
3073:Mayflies
2996:Scotland
2940:Archived
2784:(9): 81.
2763:: 17–18.
2646:: 21–26.
2615:Archived
2611:BBC News
2585:Archived
2565:15 March
2135:off the
2030:in 1859.
1976:Archived
1948:in 1818.
605:Lestidae
497:colonist
464:Scotland
419:The list
372:in east
321:vagrants
241:Cornwall
182:Tredegar
166:Cornwall
59:vagrants
51:resident
3174:Mammals
3048:Beetles
3024:Insects
2991:England
2920:Atropos
2903:Atropos
2886:Atropos
2865:Atropos
2798:Atropos
2781:Atropos
2735:Atropos
2621:20 June
2591:15 June
2536:Atropos
2515:Atropos
2494:Atropos
2448:Atropos
2427:Atropos
2356:Atropos
2310:Atropos
2045:Norfolk
2041:Horning
1965:Bristol
1596:Species
1545:Family
1442:Species
1431:Family
1398:Species
1379:Family
1292:Emperor
1155:Species
1136:Family
1086:Species
1075:Family
1034:Species
1023:Family
784:Species
719:Family
614:Species
603:Family
553:Species
522:Family
507:vagrant
487:extinct
455:England
325:extinct
273:Norfolk
194:Norfolk
125:Ireland
109:Atropos
55:extinct
39:Britain
35:Odonata
31:species
2229:) and
2210:) and
2049:Bolton
1969:Jersey
1960:Aeshna
1946:Sussex
1920:exuvia
777:female
427:. The
374:Dorset
134:– see
90:
3164:Birds
3134:Crabs
3098:Wasps
3078:Moths
3006:Wales
2141:Texas
1902:Notes
1808:Scot
1519:Scot
1230:Scot
842:Scot
473:Wales
469:Wales
382:Devon
342:Essex
257:Essex
3169:Fish
3043:Bees
3033:Ants
2623:2018
2593:2019
2567:2014
2268:HMSO
2057:Kent
2028:Kent
1924:Kent
1756:Eng
1637:Eng
1318:Eng
1213:Eng
962:Eng
910:Eng
766:male
755:male
734:male
692:Eng
658:Eng
460:Scot
368:and
281:Kent
226:Kent
88:ISBN
84:HMSO
2218:).
2202:),
2183:in
1759:RC
447:Eng
380:in
340:in
65:.
37:in
33:of
3234::
2847:.
2822:.
2759:.
2715:19
2713:.
2690:13
2688:.
2663:.
2644:32
2642:.
2609:.
2474:15
2472:.
2407:16
2405:.
2382:13
2380:.
2336:12
2334:.
2297:^
2289:12
2287:.
2266:.
2252:^
2163:).
2051:,
2043:,
2026:,
1827:V
1741:V
493:RC
483:Ex
327::
279:,
271:,
239:,
224:,
212:,
192:,
184:,
164:,
160:,
121:.
82:,
2972:e
2965:t
2958:v
2849:1
2824:9
2761:9
2665:4
2625:.
2595:.
2569:.
2555:"
2270:.
2233:(
2225:(
2214:(
2206:(
2198:(
2175:(
2155:(
2143:.
2127:(
2100:(
2081:.
503:V
174:(
94:)
86:(
20:)
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