118:
572:. The 10.0–13.9 mm (0.39–0.55 in) wing has a convex lower margin and a convex to straight upper margin which meet at the basal end of the samara at a point where it would have connected to the parent plant. Three veins run the length of the wing, arising from the basal wingtip and merging onto the nutlet. Thick veins follow along the upper margin and cross the midline of the wing, while a third, thin vein curves parallel to the lower margin between the midvein and margin.
140:
893:
564:. The nutlets are 3.5 mm–4.5 mm × 2.6 mm–3.0 mm (0.14 in–0.18 in × 0.10 in–0.12 in) with an estimated thickness between 2–2.5 mm (0.079–0.098 in). The tip of the nutlet sports four to five short
864:
428:, having distinct seasonal temperature swings which dipped below freezing in the winters. However, further study has shown the lake system was surrounded by a warm temperate ecosystem that likely had a
549:, in recognition of contribution of the Klondike Mountain Formation specimens. The genus has been occasionally given the vernacular name "winged stranger" as a result of the genus name.
378:
sandstones, siltstones, and conglomerates which preserve either a lake delta environment, or alternatively periodic floods and volcanic mudflows preserved with hot spring activity.
935:
955:
791:"Fossil biotas from the Okanagan Highlands, southern British Columbia and northeastern Washington State: climates and ecosystems across an Eocene landscape"
397:
940:
568:
which are 1.5 mm × 1.5 mm (0.059 in × 0.059 in) with round tips. In the middle of the nutlet tip are two central
950:
925:
735:
945:
489:
960:
328:
for the species plus the White Cliffs. The nut beds are approximately 3 kilometres (1.9 mi) east of the unincorporated community of
627:"Regional and local vegetation community dynamics of the Eocene Okanagan Highlands (British Columbia – Washington State) from palynology"
332:, while the White Cliffs locality is south of Clarno, and both considered to be middle Eocene in age. The dating is based on averaging
970:
920:
139:
472:
389:
442:
It was likely wind-dispersed seed, and possibly part of the canopy flora of the forests, however the affinities of
965:
382:
439:
climate, in which winter temperatures rarely dropped low enough for snow, and which were seasonably equitable.
897:
676:(Betulaceae) from the Middle Eocene Klondike Mountain and Allenby Formations of Northwestern North America".
325:
601:
Manchester, S.R. (1994). "Fruits and Seeds of the Middle Eocene Nut Beds Flora, Clarno
Formation, Oregon".
347:
930:
543:
249:
501:
230:
117:
915:
837:
802:
638:
393:
333:
253:
715:
1. Early Eocene
Lagerstätten of the Okanagan Highlands (British Columbia and Washington State)".
693:
350:
336:
134:
560:
samaras are elongated and flattened sideways, with an elliptical wing that joins to an ovoid
845:
810:
789:
Greenwood, D. R.; Archibald, S. B.; Mathewes, R. W; Moss, P. T. (2005).
769:
711:
Archibald, S.; Greenwood, D.; Smith, R.; Mathewes, R.; Basinger, J. (2011). "Great
Canadian
685:
646:
493:
321:
306:
569:
276:
177:
396:. The formation preserves an upland lake system surrounded by a mixed conifer–broadleaf
841:
806:
744:
Journal of the
Faculty of Science, Hokkaido University. Series 4, Geology and Mineralogy
642:
532:
520:
477:
463:
329:
187:
164:
909:
539:
459:
429:
294:
75:
790:
768:(Report). Bulletin. Vol. 1597. United States Geological Survey. pp. 1–25.
697:
626:
509:
432:
425:
405:
17:
546:
485:
436:
50:
272:
95:
60:
762:
421:
280:
100:
44:
892:
763:
Middle Eocene dicotyledonous plants from
Republic, northeastern Washington
462:
Steven R. Manchester of the
University of Florida. He published his 1994
561:
497:
481:
417:
386:
302:
90:
85:
70:
65:
55:
625:
Moss, P. T.; Greenwood, D. R.; Archibald, S. B. (2005).
369:
355:
105:
80:
424:. Wolfe and Tanai (1987) interpreted the forest climate to have been
362:
341:
446:
with other flowering plants were unknown at the time of description.
413:
298:
290:
849:
814:
664:
Pigg, K. B.; Manchester, S. R.; Wehr, W. C. (2003). "
650:
689:
774:
565:
401:
374:. The beds are composed of silica and calcium carbonate cemented
151:
400:
with nearby volcanism. The pollen flora has notable elements of
505:
375:
531:
roughly translating to "stranger", while
Manchester coined the
865:"Jack Nisbet presents Highland Wonders program about Wes Wehr"
409:
381:
Five additional specimens were studied from a location in the
828:
Archibald, S. B.; et al. (2005). "Wes Wehr dedication".
367:
date. The average of the dates resulted in an age range of
476:. The genus and species were described from a series of
740:(maples) in the Cenozoic of Western North America"
500:specimens. Of the paratypes, one is housed in the
596:
594:
592:
590:
588:
586:
584:
484:specimen OMSI PM018, which was preserved in the
320:has been identified from two locations in the
736:"Systematics, Phylogeny, and Distribution of
365: ± 3.36 million years ago
358: ± 1.31 million years ago
8:
761:Wolfe, J. A.; Wehr, W. C. (1987).
508:collections, with the remaining five in the
344: ± 0.1 million years ago
279:of uncertain affinities, which contains the
620:
618:
616:
116:
31:
773:
324:, the leaf layer in the Clarno nut beds,
936:Flora of the Northwestern United States
678:International Journal of Plant Sciences
580:
490:Oregon Museum of Science and Industry
7:
956:Prehistoric plants of North America
297:sediments exposed in north central
830:Canadian Journal of Earth Sciences
795:Canadian Journal of Earth Sciences
631:Canadian Journal of Earth Sciences
25:
891:
504:collections, a second is in the
408:, and distinct trace amounts of
138:
734:Wolfe, J.A.; Tanai, T. (1987).
941:Extinct flora of North America
863:Julie Vanderwal (2019-02-27).
1:
951:Prehistoric angiosperm genera
926:Fossil taxa described in 1994
313:Distribution and paleoecology
946:Eocene life of North America
289:. It is known from isolated
961:Klondike Mountain Formation
603:Palaeontographica Americana
473:Palaeontographica Americana
390:Klondike Mountain Formation
987:
512:paleontology collections.
458:specimens were studied by
450:History and classification
971:Enigmatic angiosperm taxa
383:Eocene Okanagan Highlands
267:, sometimes known as the
236:
229:
135:Scientific classification
133:
124:
115:
34:
921:Plants described in 1994
519:is a combination of the
339:which yielded an age of
871:. Wenatchee, Washington
372: million years ago
305:-age fossils found in
27:Fossil genus of plants
502:University of Florida
902:at Wikimedia Commons
385:, an outcrop of the
334:zircon fission track
869:The Wenatchee World
842:2005CaJES..42..115A
807:2005CaJES..42..167G
643:2005CaJES..42..187M
558:Pteronepelys wehrii
527:meaning "wing" and
488:collections of the
456:Pteronepelys wehrii
318:Pteronepelys wehrii
286:Pteronepelys wehrii
243:Pteronepelys wehrii
127:Pteronepelys wehrii
18:Pteronepelys wehrii
433:upper microthermal
351:radiometric dating
348:Argon–argon dating
337:radiometric dating
896:Media related to
717:Geoscience Canada
437:lower mesothermal
260:
259:
210:
16:(Redirected from
978:
966:Clarno Formation
895:
880:
879:
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819:
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731:
725:
724:
708:
702:
701:
661:
655:
654:
622:
611:
610:
598:
533:specific epithet
494:Portland, Oregon
464:type description
373:
366:
359:
353:which yielded a
345:
322:Clarno Formation
245:
241:
209:Manchester, 1994
208:
201:
143:
142:
120:
110:
47:
40:Temporal range:
32:
21:
986:
985:
981:
980:
979:
977:
976:
975:
906:
905:
888:
883:
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862:
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857:
850:10.1139/E05-013
827:
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822:
815:10.1139/e04-100
788:
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783:
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733:
732:
728:
710:
709:
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663:
662:
658:
651:10.1139/E04-095
624:
623:
614:
600:
599:
582:
578:
555:
515:The genus name
470:in the Journal
452:
368:
361:
354:
340:
315:
293:seeds found in
277:flowering plant
269:winged stranger
256:
247:
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11:
5:
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886:External links
884:
882:
881:
855:
836:(2): 115–117.
820:
801:(2): 167–185.
781:
753:
726:
703:
690:10.1086/376816
684:(5): 807–822.
674:Palaeocarpinus
656:
637:(2): 187–204.
612:
609:: 21, 106–107.
579:
577:
574:
554:
551:
523:word elements
486:paleobotanical
478:type specimens
451:
448:
330:Clarno, Oregon
314:
311:
258:
257:
248:
234:
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227:
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222:P. wehrii
218:
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189:incertae sedis
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931:Eocene plants
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775:10.3133/b1597
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764:
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749:
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727:
723:(4): 155–164.
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541:
540:paleobotanist
537:
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511:
507:
503:
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487:
483:
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460:paleobotanist
457:
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327:
326:type locality
323:
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296:
295:middle Eocene
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231:Binomial name
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165:Tracheophytes
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62:
57:
52:
46:
43:49.5–44
37:
33:
30:
19:
899:Pteronepelys
898:
873:. Retrieved
868:
858:
833:
829:
823:
798:
794:
784:
756:
747:
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720:
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713:Lagerstätten
712:
706:
681:
677:
673:
669:
665:
659:
634:
630:
606:
602:
557:
556:
538:in honor of
535:
528:
524:
517:Pteronepelys
516:
514:
510:Burke Museum
496:, and seven
471:
467:
455:
453:
444:Pteronepelys
443:
441:
426:microthermal
406:golden larch
380:
317:
316:
285:
284:
268:
264:Pteronepelys
263:
262:
261:
242:
237:
221:
220:
204:Pteronepelys
203:
202:
188:
171:
158:
126:
36:Pteronepelys
35:
29:
916:Angiosperms
750:(1): 1–246.
553:Description
547:Wesley Wehr
178:Angiosperms
910:Categories
875:2021-11-07
576:References
376:tuffaceous
307:Washington
250:Manchester
468:P. wehrii
283:species,
275:genus of
215:Species:
148:Kingdom:
698:19802370
670:Carpinus
562:endocarp
498:paratype
482:holotype
394:Republic
387:Ypresian
303:Ypresian
271:, is an
838:Bibcode
803:Bibcode
666:Corylus
639:Bibcode
418:cypress
273:extinct
240:†
219:†
196:Genus:
184:Order:
152:Plantae
125:Fossil
696:
672:, and
570:styles
566:sepals
544:artist
536:wehrii
529:epelys
525:Pteron
480:, the
420:, and
414:spruce
398:forest
309:, US.
299:Oregon
291:fossil
129:fruit
766:(PDF)
694:S2CID
521:Greek
430:mesic
402:birch
356:36.38
172:Clade
159:Clade
738:Acer
542:and
506:OMSI
466:for
454:The
422:palm
404:and
363:46.8
346:and
342:43.7
301:and
254:1994
51:PreꞒ
846:doi
811:doi
770:doi
686:doi
682:164
647:doi
492:in
435:to
410:fir
392:in
360:to
281:one
912::
867:.
844:.
834:42
832:.
809:.
799:42
797:.
793:.
748:22
746:.
742:.
721:38
719:.
692:.
680:.
668:,
645:.
635:42
633:.
629:.
615:^
607:58
605:.
583:^
416:,
412:,
252:,
174::
161::
101:Pg
45:Ma
878:.
852:.
848::
840::
817:.
813::
805::
778:.
772::
700:.
688::
653:.
649::
641::
370:2
200:†
106:N
96:K
91:J
86:T
81:P
76:C
71:D
66:S
61:O
56:Ꞓ
20:)
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