77:
52:
291:
Individuals have a mass of 170–250 g (6.0–8.8 oz), a head and body length of 155–185 mm (6.1–7.3 in) and a forearm length of 109.0–118.5 mm (4.29–4.67 in). It can be differentiated from the
Bismarck masked flying fox by its indistinct facial markings, darker claws, and
328:. Its population is thought to number between 9,000 and 10,000 mature individuals with a declining trend. It is an uncommon species, occurring in a low density throughout its range. There is extensive and ongoing
3492:
3472:
3405:
3091:
3418:
3202:
3067:
3058:
532:
2984:
1396:
1387:
3487:
2916:
1441:
3482:
3002:
2952:
1486:
994:
630:
2934:
1405:
498:
488:
3193:
3444:
2993:
2898:
2889:
2422:
1414:
1012:
666:
2599:
1796:
1495:
1459:
1432:
729:
684:
3423:
2925:
2590:
1477:
1468:
780:
1996:
1960:
1847:
1450:
1152:
1003:
657:
2907:
2880:
1969:
1086:
958:
711:
702:
3248:
3294:
1556:
1423:
771:
693:
675:
639:
1053:
2943:
2494:
2226:
2091:
1619:
1044:
747:
648:
525:
435:"Each flying fox on its own branch: A phylogenetic tree for Pteropus and related genera (Chiroptera: Pteropodidae)"
252:
3211:
2759:
1678:
76:
3497:
2548:
2298:
1987:
1978:
1888:
1519:
1161:
1143:
895:
871:
2503:
2208:
2172:
1110:
967:
236:
3285:
2395:
2127:
2055:
1936:
1912:
1838:
1728:
1637:
1628:
1586:
1274:
1241:
3360:
2732:
2727:
2404:
2307:
2190:
2082:
1595:
1250:
926:
905:
738:
2476:
2458:
2118:
1719:
1669:
1308:
720:
518:
171:
3327:
1355:
976:
378:
3303:
2804:
2786:
2485:
2449:
2334:
2262:
2253:
2028:
1879:
248:
2813:
2675:
2521:
2431:
2316:
2100:
1778:
224:
41:
312:. It is found in forested habitat from sea level to 1,200 m (3,900 ft) above sea level.
2750:
2657:
2512:
2440:
2371:
2244:
2154:
2064:
2037:
1710:
1134:
321:
71:
56:
3477:
3436:
3392:
3161:
3152:
3026:
2741:
2530:
2386:
2361:
2343:
2289:
2280:
2271:
2235:
2136:
2109:
1805:
1787:
1333:
613:
494:
466:
433:
Almeida, Francisca C.; Giannini, Norberto P.; Simmons, Nancy B.; Helgen, Kristofer M. (2014).
247:
had been collected by
Flannery, Lester Seri, T. Heinsohn, and Patricia Ennis on the island of
3128:
2768:
2714:
2566:
2557:
2539:
2467:
2413:
2377:
2325:
2199:
2163:
2046:
1814:
985:
838:
456:
446:
413:
373:
309:
260:
212:
2777:
2666:
2352:
2181:
1862:
1528:
1510:
1298:
1209:
1185:
1062:
847:
355:
510:
3266:
2217:
2073:
578:
296:. The top of its head lacks the yellowish tint seen in the Bismarck masked flying fox.
251:
in 1988. In a 1995 publication, Flannery considered it instead as a subspecies of the
3466:
3431:
3318:
3143:
3119:
3041:
2841:
2799:
1077:
886:
762:
418:
364:
329:
61:
401:
3397:
3383:
3276:
3109:
3082:
3017:
2853:
2581:
1756:
1693:
1610:
1569:
862:
808:
600:
228:
187:
271:(4.7%), along with several physical differences, was enough to warrant elevating
3449:
3176:
2863:
2627:
1224:
1176:
1027:
451:
434:
17:
2640:
1951:
1829:
1652:
1289:
941:
584:
232:
3229:
2690:
1927:
1346:
1265:
1200:
1125:
821:
88:
470:
3377:
3239:
2011:
1903:
1769:
1370:
1101:
917:
572:
566:
560:
554:
293:
244:
148:
108:
3410:
2967:
542:
461:
402:"Type specimens of non-fossil mammals in the Australian Museum, Sydney"
305:
208:
138:
2709:
276:
118:
98:
3354:
325:
3358:
2838:
2624:
1753:
1553:
1330:
805:
597:
514:
128:
3367:
3316:
3274:
3265:
3237:
3228:
3174:
3141:
3117:
3108:
3080:
3039:
3015:
2965:
2861:
2852:
2688:
2638:
2579:
2009:
1949:
1925:
1901:
1860:
1827:
1767:
1691:
1650:
1608:
1567:
1508:
1368:
1344:
1287:
1263:
1222:
1198:
1174:
1123:
1099:
1075:
1025:
939:
915:
884:
860:
819:
760:
611:
379:10.2305/IUCN.UK.2022-1.RLTS.T84883915A209887353.en
332:on Papua New Guinea that is causing habitat loss.
223:The New Ireland masked flying fox was initially
354:Tsang, S.M.; Waldien, D.L.; Pennay, M. (2022).
400:Parnaby, H.; Ingleby, S.; Divljan, A. (2017).
526:
8:
3355:
3271:
3234:
3114:
2858:
2849:
2835:
2635:
2621:
1764:
1750:
1564:
1550:
1341:
1327:
816:
802:
608:
594:
533:
519:
511:
50:
31:
460:
450:
417:
377:
3203:East African little collared fruit bat (
482:
480:
259:). However, a 2014 study found that the
3092:Veldkamp's dwarf epauletted fruit bat (
341:
349:
347:
345:
231:and J. P. White, who regarded it as a
3059:Hayman's dwarf epauletted fruit bat (
2917:Lesser Angolan epauletted fruit bat (
439:Molecular Phylogenetics and Evolution
304:The New Ireland masked flying fox is
7:
3068:Peters' dwarf epauletted fruit bat (
2985:Buettikofer's epauletted fruit bat (
1487:New Britain naked-backed fruit bat (
995:Nusatenggara short-nosed fruit bat (
631:Broad-striped tube-nosed fruit bat (
2935:East African epauletted fruit bat (
1406:Philippine naked-backed fruit bat (
1397:Beaufort's naked-backed fruit bat (
1388:Andersen's naked-backed fruit bat (
365:IUCN Red List of Threatened Species
2423:Little golden-mantled flying fox (
1442:Solomon's naked-backed fruit bat (
1415:Halmahera naked-backed fruit bat (
1013:Indonesian short-nosed fruit bat (
667:Round-eared tube-nosed fruit bat (
25:
3493:Endemic fauna of Papua New Guinea
3003:Franquet's epauletted fruit bat (
2953:Wahlberg's epauletted fruit bat (
2600:Sulawesi stripe-faced fruit bat (
1797:Giant golden-crowned flying fox (
1496:Greenish naked-backed fruit bat (
1460:Moluccan naked-backed fruit bat (
1433:Sulawesi naked-backed fruit bat (
730:Philippine tube-nosed fruit bat (
493:. CSIRO Publishing. p. 358.
490:Mammals of the South-west Pacific
487:Lavery, T.; Flannery, T. (2023).
3473:IUCN Red List vulnerable species
2926:Ethiopian epauletted fruit bat (
2591:Mindoro stripe-faced fruit bat (
1478:Western naked-backed fruit bat (
1469:Panniet naked-backed fruit bat (
781:Steadfast tube-nosed fruit bat (
419:10.3853/j.2201-4349.69.2017.1653
406:Records of the Australian Museum
308:to the island of New Ireland in
75:
2899:Peters's epauletted fruit bat (
2890:Ansell's epauletted fruit bat (
2146:New Ireland masked flying fox (
1997:New Georgian monkey-faced Bat (
1961:Bougainville monkey-faced Bat (
1848:Small white-winged flying fox (
1451:Lesser naked-backed fruit bat (
1004:Greater short-nosed fruit bat (
658:Mountain tube-nosed fruit bat (
320:As of 2022, it is considered a
2908:Gambian epauletted fruit bat (
2881:Angolan epauletted fruit bat (
1970:Guadalcanal monkey-faced Bat (
959:Lesser short-nosed fruit bat (
712:Demonic tube-nosed fruit bat (
703:Malaita tube-nosed fruit bat (
685:Keast's tube-nosed fruit bat (
35:New Ireland masked flying fox
1:
3194:São Tomé collared fruit bat (
1424:Biak naked-backed fruit bat (
772:Lesser tube-nosed fruit bat (
694:Island tube-nosed fruit bat (
676:Dragon tube-nosed fruit bat (
640:Common tube-nosed fruit bat (
199:New Ireland masked flying fox
3184:(Little collared fruit bats)
2944:Minor epauletted fruit bat (
2495:Philippine gray flying fox (
2092:Bismarck masked flying fox (
1153:Ratanaworabhan's fruit bat (
748:Umboi tube-nosed fruit bat (
257:Pteropus capistratus ennisae
3212:Little collared fruit bat (
1870:(Straw-coloured fruit bats)
1679:New Caledonia blossom bat (
1087:Fischer's pygmy fruit bat (
452:10.1016/j.ympev.2014.03.009
241:Pteropus temminckii ennisae
3514:
3488:Taxa named by Tim Flannery
2549:New Caledonia flying fox (
2299:Black-bearded flying fox (
1988:Montane monkey-faced Bat (
1979:Greater monkey-faced Bat (
1889:Straw-coloured fruit bat (
1520:Sulawesi harpy fruit bat (
1162:White-collared fruit bat (
1144:Javan tailless fruit bat (
1054:Rickart's dyak fruit bat (
896:Spotted-winged fruit bat (
872:Mindanao pygmy fruit bat (
283:is Patricia "Tish" Ennis.
253:Bismarck masked flying fox
3483:Mammals described in 1991
3286:Short-palated fruit bat (
2848:
2834:
2634:
2620:
2396:Geelvink Bay flying fox (
2227:Livingstone's fruit bat (
2128:Large Samoan flying fox (
2056:Small Samoan flying fox (
1937:Small-toothed fruit bat (
1913:Fijian monkey-faced Bat (
1839:White-winged flying fox (
1763:
1749:
1729:Moss-forest blossom bat (
1629:Black-bellied fruit bat (
1620:Fardoulis's blossom bat (
1587:Long-tongued nectar bat (
1577:(Long-tongued fruit bats)
1563:
1549:
1340:
1326:
1242:Greater musky fruit bat (
1045:Brooks's dyak fruit bat (
815:
801:
649:Pallas's tube-nosed bat (
607:
593:
549:
541:Extant species of family
372:: e.T84883915A209887353.
177:
170:
72:Scientific classification
70:
48:
39:
34:
3249:D'Anchieta's fruit bat (
2760:Leschenault's rousette (
2405:Grey-headed flying fox (
2308:Black-eared flying fox (
2191:Ontong Java flying fox (
2083:Ashy-headed flying fox (
1660:(Long-tailed fruit bats)
1596:Long-tongued fruit bat (
1378:(Bare-backed fruit bats)
1251:Lesser musky fruit bat (
927:Black-capped fruit bat (
739:Eastern tube-nosed Bat (
3295:Campo-Ma’an fruit bat (
2504:Temminck's flying fox (
2477:Little red flying fox (
2459:Madagascan flying fox (
2209:Andersen's flying fox (
2173:Gilliard's flying fox (
2119:Spectacled flying fox (
1720:Halmahera blossom bat (
1670:Long-tailed fruit bat (
1111:Salim Ali's fruit bat (
968:Horsfield's fruit bat (
721:Lesser tube-nosed bat (
621:(tube-nosed fruit bats)
3049:(Dwarf epauleted bats)
2871:(Epauleted fruit bats)
2805:Long-haired rousette (
2787:Bare-backed rousette (
2486:Seychelles fruit bat (
2450:Rodrigues flying fox (
2335:Mauritian flying fox (
2263:Big-eared flying fox (
2029:Admiralty flying fox (
1880:Madagascan fruit bat (
1638:Woodford's fruit bat (
1275:Blanford's fruit bat (
949:(Dog-faced fruit bats)
906:Balionycteris seimundi
275:to species level. The
219:Taxonomy and etymology
2814:Madagascan rousette (
2733:Geoffroy's rousette (
2698:(rousette fruit bats)
2676:Philippine dawn bat (
2522:Vanikoro flying fox (
2432:Solomons flying fox (
2317:Caroline flying fox (
2101:Moluccan flying fox (
1779:Sulawesi flying fox (
279:for the species name
237:Temminck's flying fox
3328:Zenker's fruit bat (
2994:Dobson's fruit bat (
2658:Greater nectar bat (
2513:Insular flying fox (
2441:Rennell flying fox (
2245:Okinawa flying fox (
2155:Nicobar flying fox (
2065:Vanuatu flying fox (
2038:Aldabra flying fox (
1711:Common blossom bat (
1356:Bulmer's fruit bat (
1135:Tailless fruit bat (
977:Peters's fruit bat (
3304:Pohle's fruit bat (
3153:Azagnyi fruit bat (
3027:Hammer-headed Bat (
2816:R. madagascariensis
2751:Egyptian rousette (
2742:Sulawesi rousette (
2531:Kosrae flying fox (
2387:Masked flying fox (
2362:Ornate flying fox (
2344:Temotu flying fox (
2290:Indian flying fox (
2281:Mariana fruit bat (
2272:Lesser flying fox (
2254:Lyle's flying fox (
2236:Lombok flying fox (
2137:Ryukyu flying fox (
2110:Makira flying fox (
1850:D. microleucopterus
1806:Palawan fruit bat (
1788:Talaud flying fox (
211:that is endemic to
42:Conservation status
3129:Angolan rousette (
2769:Linduan Rousette (
2735:R. amplexicaudatus
2715:Manado fruit bat (
2567:Dwarf flying fox (
2558:Pemba flying fox (
2540:Large flying fox (
2468:Samoa flying fox (
2414:Bonin flying fox (
2378:Pelew flying fox (
2326:Great flying fox (
2200:Small flying fox (
2164:Banks flying fox (
2047:Black flying fox (
1815:Sunda flying fox (
1232:(Musky fruit bats)
1035:(Dayak fruit bats)
986:Minute fruit bat (
839:Borneo fruit bat (
829:(Pygmy fruit bats)
322:vulnerable species
207:) is a species of
3460:
3459:
3361:Taxon identifiers
3352:
3351:
3348:
3347:
3344:
3343:
3340:
3339:
3297:C. campomaanensis
3261:
3260:
3224:
3223:
3187:
3104:
3103:
3052:
2978:
2874:
2830:
2829:
2826:
2825:
2778:Comoro rousette (
2701:
2667:Cave nectar bat (
2651:
2648:(Dawn fruit bats)
2616:
2615:
2612:
2611:
2353:Ceram fruit bat (
2182:Gray flying fox (
2121:P. conspicillatus
2022:
1873:
1745:
1744:
1741:
1740:
1704:
1663:
1580:
1545:
1544:
1541:
1540:
1529:Harpy fruit bat (
1381:
1334:Harpyionycterinae
1322:
1321:
1318:
1317:
1299:Swift fruit bat (
1235:
1210:Dusky fruit bat (
1188:O. cartilagonodus
1186:Luzon fruit bat (
1063:Dayak fruit bat (
1038:
1015:C. titthaecheilus
952:
929:C. melanocephalus
848:Pygmy fruit bat (
832:
797:
796:
793:
792:
624:
300:Range and habitat
195:
194:
190:& White, 1991
65:
16:(Redirected from
3505:
3453:
3452:
3440:
3439:
3427:
3426:
3414:
3413:
3401:
3400:
3388:
3387:
3386:
3369:Pteropus ennisae
3356:
3272:
3235:
3196:M. brachycephala
3186:
3185:
3181:
3162:Woermann's bat (
3115:
3051:
3050:
3046:
2977:
2976:
2975:(Epauleted bats)
2972:
2873:
2872:
2868:
2859:
2850:
2836:
2700:
2699:
2695:
2650:
2649:
2645:
2636:
2622:
2488:P. seychellensis
2407:P. poliocephalus
2328:P. neohibernicus
2218:Kei flying fox (
2103:P. chrysoproctus
2074:Aru flying fox (
2031:P. admiralitatum
2021:
2020:
2016:
1872:
1871:
1867:
1765:
1751:
1703:
1702:
1698:
1681:N. neocaledonica
1662:
1661:
1657:
1579:
1578:
1574:
1565:
1551:
1380:
1379:
1375:
1342:
1328:
1234:
1233:
1229:
1037:
1036:
1032:
951:
950:
946:
831:
830:
826:
817:
803:
623:
622:
618:
609:
595:
535:
528:
521:
512:
505:
504:
484:
475:
474:
464:
454:
430:
424:
423:
421:
397:
391:
390:
388:
386:
381:
358:Pteropus ennisae
351:
310:Papua New Guinea
294:flight membranes
261:genetic distance
213:Papua New Guinea
204:Pteropus ennisae
183:
181:Pteropus ennisae
80:
79:
59:
54:
53:
32:
21:
18:Pteropus ennisae
3513:
3512:
3508:
3507:
3506:
3504:
3503:
3502:
3498:Bats of Oceania
3463:
3462:
3461:
3456:
3448:
3443:
3435:
3430:
3422:
3417:
3409:
3404:
3396:
3391:
3382:
3381:
3376:
3363:
3353:
3336:
3312:
3257:
3220:
3183:
3182:
3180:
3170:
3137:
3100:
3076:
3048:
3047:
3045:
3035:
3011:
2987:E. buettikoferi
2974:
2973:
2971:
2961:
2870:
2869:
2867:
2844:
2822:
2762:R. leschenaulti
2697:
2696:
2694:
2684:
2647:
2646:
2644:
2630:
2608:
2575:
2524:P. tuberculatus
2346:P. nitendiensis
2229:P. livingstonii
2175:P. gilliardorum
2018:
2017:
2015:
2005:
1945:
1921:
1897:
1869:
1868:
1866:
1856:
1823:
1759:
1737:
1700:
1699:
1697:
1687:
1659:
1658:
1656:
1646:
1604:
1576:
1575:
1573:
1559:
1537:
1511:Harpyionycteris
1504:
1471:D. pannietensis
1377:
1376:
1374:
1364:
1336:
1314:
1283:
1259:
1231:
1230:
1228:
1218:
1194:
1170:
1119:
1095:
1071:
1034:
1033:
1031:
1021:
997:C. nusatenggara
948:
947:
945:
935:
911:
880:
874:A. paucidentata
856:
828:
827:
825:
811:
789:
756:
620:
619:
617:
603:
589:
545:
539:
509:
508:
501:
486:
485:
478:
432:
431:
427:
399:
398:
394:
384:
382:
353:
352:
343:
338:
318:
302:
289:
221:
191:
185:
179:
166:
163:P. ennisae
74:
66:
55:
51:
44:
28:
23:
22:
15:
12:
11:
5:
3511:
3509:
3501:
3500:
3495:
3490:
3485:
3480:
3475:
3465:
3464:
3458:
3457:
3455:
3454:
3441:
3428:
3415:
3402:
3389:
3373:
3371:
3365:
3364:
3359:
3350:
3349:
3346:
3345:
3342:
3341:
3338:
3337:
3335:
3334:
3324:
3322:
3314:
3313:
3311:
3310:
3301:
3292:
3282:
3280:
3269:
3267:Scotonycterini
3263:
3262:
3259:
3258:
3256:
3255:
3245:
3243:
3232:
3226:
3225:
3222:
3221:
3219:
3218:
3209:
3200:
3190:
3188:
3172:
3171:
3169:
3168:
3159:
3149:
3147:
3139:
3138:
3136:
3135:
3125:
3123:
3112:
3106:
3105:
3102:
3101:
3099:
3098:
3088:
3086:
3078:
3077:
3075:
3074:
3065:
3061:M. intermedius
3055:
3053:
3037:
3036:
3034:
3033:
3023:
3021:
3013:
3012:
3010:
3009:
3000:
2991:
2981:
2979:
2963:
2962:
2960:
2959:
2950:
2941:
2932:
2923:
2914:
2905:
2896:
2887:
2877:
2875:
2856:
2846:
2845:
2839:
2832:
2831:
2828:
2827:
2824:
2823:
2821:
2820:
2811:
2794:
2793:
2784:
2775:
2766:
2757:
2753:R. aegyptiacus
2748:
2739:
2722:
2721:
2704:
2702:
2686:
2685:
2683:
2682:
2673:
2664:
2654:
2652:
2632:
2631:
2625:
2618:
2617:
2614:
2613:
2610:
2609:
2607:
2606:
2597:
2593:S. mindorensis
2587:
2585:
2577:
2576:
2574:
2573:
2564:
2555:
2546:
2537:
2528:
2519:
2510:
2501:
2492:
2483:
2474:
2465:
2456:
2452:P. rodricensis
2447:
2438:
2429:
2420:
2411:
2402:
2393:
2384:
2375:
2368:
2359:
2350:
2341:
2332:
2323:
2314:
2305:
2301:P. melanopogon
2296:
2287:
2278:
2269:
2260:
2251:
2247:P. loochoensis
2242:
2238:P. lombocensis
2233:
2224:
2215:
2211:P. intermedius
2206:
2202:P. hypomelanus
2197:
2188:
2179:
2170:
2161:
2152:
2143:
2134:
2125:
2116:
2107:
2098:
2094:P. capistratus
2089:
2080:
2071:
2062:
2053:
2044:
2040:P. aldabrensis
2035:
2025:
2023:
2019:(flying foxes)
2007:
2006:
2004:
2003:
1994:
1985:
1976:
1967:
1957:
1955:
1947:
1946:
1944:
1943:
1933:
1931:
1923:
1922:
1920:
1919:
1909:
1907:
1899:
1898:
1896:
1895:
1886:
1876:
1874:
1858:
1857:
1855:
1854:
1845:
1841:D. leucopterus
1835:
1833:
1825:
1824:
1822:
1821:
1812:
1803:
1794:
1785:
1775:
1773:
1761:
1760:
1754:
1747:
1746:
1743:
1742:
1739:
1738:
1736:
1735:
1726:
1717:
1707:
1705:
1701:(blossom bats)
1689:
1688:
1686:
1685:
1676:
1666:
1664:
1648:
1647:
1645:
1644:
1635:
1626:
1616:
1614:
1606:
1605:
1603:
1602:
1593:
1583:
1581:
1561:
1560:
1557:Macroglossinae
1554:
1547:
1546:
1543:
1542:
1539:
1538:
1536:
1535:
1526:
1516:
1514:
1506:
1505:
1503:
1502:
1493:
1484:
1475:
1466:
1462:D. moluccensis
1457:
1448:
1439:
1430:
1421:
1412:
1403:
1394:
1384:
1382:
1366:
1365:
1363:
1362:
1352:
1350:
1338:
1337:
1331:
1324:
1323:
1320:
1319:
1316:
1315:
1313:
1312:
1305:
1295:
1293:
1285:
1284:
1282:
1281:
1271:
1269:
1261:
1260:
1258:
1257:
1248:
1238:
1236:
1220:
1219:
1217:
1216:
1206:
1204:
1196:
1195:
1193:
1192:
1182:
1180:
1172:
1171:
1169:
1168:
1159:
1150:
1141:
1131:
1129:
1121:
1120:
1118:
1117:
1107:
1105:
1097:
1096:
1094:
1093:
1083:
1081:
1073:
1072:
1070:
1069:
1060:
1051:
1041:
1039:
1023:
1022:
1020:
1019:
1010:
1001:
992:
983:
979:C. luzoniensis
974:
970:C. horsfieldii
965:
955:
953:
937:
936:
934:
933:
923:
921:
913:
912:
910:
909:
902:
892:
890:
882:
881:
879:
878:
868:
866:
858:
857:
855:
854:
845:
835:
833:
813:
812:
806:
799:
798:
795:
794:
791:
790:
788:
787:
778:
768:
766:
758:
757:
755:
754:
745:
736:
727:
718:
709:
705:N. malaitensis
700:
691:
682:
673:
664:
655:
646:
637:
627:
625:
605:
604:
598:
591:
590:
588:
587:
581:
579:Laurasiatheria
575:
569:
563:
557:
550:
547:
546:
540:
538:
537:
530:
523:
515:
507:
506:
499:
476:
425:
392:
340:
339:
337:
334:
317:
314:
301:
298:
288:
285:
265:P. capistratus
220:
217:
193:
192:
186:
175:
174:
168:
167:
160:
158:
154:
153:
146:
142:
141:
136:
132:
131:
126:
122:
121:
116:
112:
111:
106:
102:
101:
96:
92:
91:
86:
82:
81:
68:
67:
49:
46:
45:
40:
37:
36:
27:Species of bat
26:
24:
14:
13:
10:
9:
6:
4:
3:
2:
3510:
3499:
3496:
3494:
3491:
3489:
3486:
3484:
3481:
3479:
3476:
3474:
3471:
3470:
3468:
3451:
3446:
3442:
3438:
3433:
3429:
3425:
3420:
3416:
3412:
3407:
3403:
3399:
3394:
3390:
3385:
3379:
3375:
3374:
3372:
3370:
3366:
3362:
3357:
3333:
3331:
3326:
3325:
3323:
3321:
3320:
3319:Scotonycteris
3315:
3309:
3307:
3302:
3300:
3298:
3293:
3291:
3289:
3284:
3283:
3281:
3279:
3278:
3273:
3270:
3268:
3264:
3254:
3252:
3247:
3246:
3244:
3242:
3241:
3236:
3233:
3231:
3227:
3217:
3215:
3210:
3208:
3206:
3201:
3199:
3197:
3192:
3191:
3189:
3179:
3178:
3173:
3167:
3165:
3160:
3158:
3156:
3151:
3150:
3148:
3146:
3145:
3144:Megaloglossus
3140:
3134:
3132:
3131:L. angolensis
3127:
3126:
3124:
3122:
3121:
3120:Lissonycteris
3116:
3113:
3111:
3107:
3097:
3095:
3094:N. veldkampii
3090:
3089:
3087:
3085:
3084:
3079:
3073:
3071:
3066:
3064:
3062:
3057:
3056:
3054:
3044:
3043:
3042:Micropteropus
3038:
3032:
3030:
3029:H. monstrosus
3025:
3024:
3022:
3020:
3019:
3014:
3008:
3006:
3001:
2999:
2997:
2992:
2990:
2988:
2983:
2982:
2980:
2970:
2969:
2964:
2958:
2956:
2951:
2949:
2947:
2942:
2940:
2938:
2933:
2931:
2929:
2924:
2922:
2920:
2915:
2913:
2911:
2906:
2904:
2902:
2897:
2895:
2893:
2888:
2886:
2884:
2883:E. angolensis
2879:
2878:
2876:
2866:
2865:
2860:
2857:
2855:
2851:
2847:
2843:
2842:Epomophorinae
2837:
2833:
2819:
2817:
2812:
2810:
2808:
2802:
2801:
2800:Stenonycteris
2796:
2795:
2792:
2790:
2789:R. spinalatus
2785:
2783:
2781:
2780:R. obliviosus
2776:
2774:
2772:
2771:R. linduensis
2767:
2765:
2763:
2758:
2756:
2754:
2749:
2747:
2745:
2744:R. celebensis
2740:
2738:
2736:
2730:
2729:
2724:
2723:
2720:
2718:
2712:
2711:
2706:
2705:
2703:
2693:
2692:
2687:
2681:
2679:
2674:
2672:
2670:
2665:
2663:
2661:
2656:
2655:
2653:
2643:
2642:
2637:
2633:
2629:
2623:
2619:
2605:
2603:
2598:
2596:
2594:
2589:
2588:
2586:
2584:
2583:
2578:
2572:
2570:
2565:
2563:
2561:
2560:P. voeltzkowi
2556:
2554:
2552:
2547:
2545:
2543:
2538:
2536:
2534:
2529:
2527:
2525:
2520:
2518:
2516:
2511:
2509:
2507:
2506:P. temminckii
2502:
2500:
2498:
2493:
2491:
2489:
2484:
2482:
2480:
2479:P. scapulatus
2475:
2473:
2471:
2466:
2464:
2462:
2457:
2455:
2453:
2448:
2446:
2444:
2439:
2437:
2435:
2430:
2428:
2426:
2421:
2419:
2417:
2412:
2410:
2408:
2403:
2401:
2399:
2394:
2392:
2390:
2389:P. personatus
2385:
2383:
2381:
2380:P. pelewensis
2376:
2374:
2373:
2369:
2367:
2365:
2360:
2358:
2356:
2351:
2349:
2347:
2342:
2340:
2338:
2333:
2331:
2329:
2324:
2322:
2320:
2319:P. molossinus
2315:
2313:
2311:
2306:
2304:
2302:
2297:
2295:
2293:
2288:
2286:
2284:
2279:
2277:
2275:
2270:
2268:
2266:
2261:
2259:
2257:
2252:
2250:
2248:
2243:
2241:
2239:
2234:
2232:
2230:
2225:
2223:
2221:
2216:
2214:
2212:
2207:
2205:
2203:
2198:
2196:
2194:
2189:
2187:
2185:
2180:
2178:
2176:
2171:
2169:
2167:
2162:
2160:
2158:
2153:
2151:
2149:
2144:
2142:
2140:
2139:P. dasymallus
2135:
2133:
2131:
2126:
2124:
2122:
2117:
2115:
2113:
2108:
2106:
2104:
2099:
2097:
2095:
2090:
2088:
2086:
2081:
2079:
2077:
2072:
2070:
2068:
2063:
2061:
2059:
2054:
2052:
2050:
2045:
2043:
2041:
2036:
2034:
2032:
2027:
2026:
2024:
2014:
2013:
2008:
2002:
2000:
1995:
1993:
1991:
1986:
1984:
1982:
1977:
1975:
1973:
1968:
1966:
1964:
1959:
1958:
1956:
1954:
1953:
1948:
1942:
1940:
1935:
1934:
1932:
1930:
1929:
1924:
1918:
1916:
1911:
1910:
1908:
1906:
1905:
1900:
1894:
1892:
1887:
1885:
1883:
1878:
1877:
1875:
1865:
1864:
1859:
1853:
1851:
1846:
1844:
1842:
1837:
1836:
1834:
1832:
1831:
1826:
1820:
1818:
1813:
1811:
1809:
1804:
1802:
1800:
1795:
1793:
1791:
1786:
1784:
1782:
1781:A. celebensis
1777:
1776:
1774:
1772:
1771:
1766:
1762:
1758:
1752:
1748:
1734:
1732:
1727:
1725:
1723:
1718:
1716:
1714:
1709:
1708:
1706:
1696:
1695:
1690:
1684:
1682:
1677:
1675:
1673:
1672:N. macdonaldi
1668:
1667:
1665:
1655:
1654:
1649:
1643:
1641:
1636:
1634:
1632:
1627:
1625:
1623:
1622:M. fardoulisi
1618:
1617:
1615:
1613:
1612:
1607:
1601:
1599:
1594:
1592:
1590:
1585:
1584:
1582:
1572:
1571:
1566:
1562:
1558:
1552:
1548:
1534:
1532:
1531:H. whiteheadi
1527:
1525:
1523:
1522:H. celebensis
1518:
1517:
1515:
1513:
1512:
1507:
1501:
1499:
1494:
1492:
1490:
1489:D. praedatrix
1485:
1483:
1481:
1476:
1474:
1472:
1467:
1465:
1463:
1458:
1456:
1454:
1449:
1447:
1445:
1440:
1438:
1436:
1431:
1429:
1427:
1422:
1420:
1418:
1413:
1411:
1409:
1404:
1402:
1400:
1395:
1393:
1391:
1386:
1385:
1383:
1373:
1372:
1367:
1361:
1359:
1354:
1353:
1351:
1349:
1348:
1343:
1339:
1335:
1329:
1325:
1311:
1310:
1309:T. suhaniahae
1306:
1304:
1302:
1301:T. nigrescens
1297:
1296:
1294:
1292:
1291:
1286:
1280:
1278:
1273:
1272:
1270:
1268:
1267:
1262:
1256:
1254:
1249:
1247:
1245:
1240:
1239:
1237:
1227:
1226:
1221:
1215:
1213:
1208:
1207:
1205:
1203:
1202:
1197:
1191:
1189:
1184:
1183:
1181:
1179:
1178:
1173:
1167:
1165:
1160:
1158:
1156:
1151:
1149:
1147:
1142:
1140:
1138:
1133:
1132:
1130:
1128:
1127:
1122:
1116:
1114:
1109:
1108:
1106:
1104:
1103:
1098:
1092:
1090:
1085:
1084:
1082:
1080:
1079:
1078:Haplonycteris
1074:
1068:
1066:
1061:
1059:
1057:
1052:
1050:
1048:
1043:
1042:
1040:
1030:
1029:
1024:
1018:
1016:
1011:
1009:
1007:
1002:
1000:
998:
993:
991:
989:
984:
982:
980:
975:
973:
971:
966:
964:
962:
961:C. brachyotis
957:
956:
954:
944:
943:
938:
932:
930:
925:
924:
922:
920:
919:
914:
908:
907:
903:
901:
899:
894:
893:
891:
889:
888:
887:Balionycteris
883:
877:
875:
870:
869:
867:
865:
864:
859:
853:
851:
846:
844:
842:
837:
836:
834:
824:
823:
818:
814:
810:
804:
800:
786:
784:
779:
777:
775:
770:
769:
767:
765:
764:
763:Paranyctimene
759:
753:
751:
746:
744:
742:
737:
735:
733:
728:
726:
724:
719:
717:
715:
710:
708:
706:
701:
699:
697:
692:
690:
688:
683:
681:
679:
678:N. draconilla
674:
672:
670:
665:
663:
661:
656:
654:
652:
651:N. cephalotes
647:
645:
643:
642:N. albiventer
638:
636:
634:
629:
628:
626:
616:
615:
610:
606:
602:
596:
592:
586:
582:
580:
576:
574:
570:
568:
564:
562:
558:
556:
552:
551:
548:
544:
536:
531:
529:
524:
522:
517:
516:
513:
502:
500:9781486312641
496:
492:
491:
483:
481:
477:
472:
468:
463:
458:
453:
448:
444:
440:
436:
429:
426:
420:
415:
411:
407:
403:
396:
393:
380:
375:
371:
367:
366:
361:
359:
350:
348:
346:
342:
335:
333:
331:
330:deforestation
327:
323:
315:
313:
311:
307:
299:
297:
295:
286:
284:
282:
278:
274:
270:
269:P. c. ennisae
266:
262:
258:
254:
250:
246:
242:
238:
234:
230:
226:
218:
216:
214:
210:
206:
205:
200:
189:
184:
182:
176:
173:
172:Binomial name
169:
165:
164:
159:
156:
155:
152:
151:
147:
144:
143:
140:
137:
134:
133:
130:
127:
124:
123:
120:
117:
114:
113:
110:
107:
104:
103:
100:
97:
94:
93:
90:
87:
84:
83:
78:
73:
69:
63:
58:
47:
43:
38:
33:
30:
19:
3368:
3329:
3317:
3305:
3296:
3287:
3277:Casinycteris
3275:
3251:P. anchietae
3250:
3238:
3213:
3204:
3195:
3175:
3164:M. woermanni
3163:
3154:
3142:
3130:
3118:
3110:Myonycterini
3093:
3083:Nanonycteris
3081:
3069:
3060:
3040:
3028:
3018:Hypsignathus
3016:
3005:E. franqueti
3004:
2995:
2986:
2966:
2955:E. wahlbergi
2954:
2945:
2936:
2927:
2918:
2910:E. gambianus
2909:
2901:E. crypturus
2900:
2891:
2882:
2862:
2854:Epomophorini
2815:
2806:
2797:
2788:
2779:
2770:
2761:
2752:
2743:
2734:
2725:
2716:
2707:
2689:
2677:
2668:
2659:
2639:
2601:
2592:
2582:Styloctenium
2580:
2569:P. woodfordi
2568:
2559:
2550:
2541:
2532:
2523:
2514:
2505:
2497:P. speciosus
2496:
2487:
2478:
2470:P. samoensis
2469:
2460:
2451:
2442:
2433:
2424:
2416:P. pselaphon
2415:
2406:
2397:
2388:
2379:
2372:P. pelagicus
2370:
2363:
2354:
2345:
2336:
2327:
2318:
2310:P. melanotus
2309:
2300:
2291:
2283:P. mariannus
2282:
2274:P. mahaganus
2273:
2264:
2255:
2246:
2237:
2228:
2219:
2210:
2201:
2192:
2183:
2174:
2165:
2156:
2147:
2145:
2138:
2129:
2120:
2111:
2102:
2093:
2084:
2075:
2067:P. anetianus
2066:
2058:P. allenorum
2057:
2048:
2039:
2030:
2010:
1998:
1989:
1981:P. flanneryi
1980:
1971:
1962:
1950:
1938:
1926:
1915:M. acrodonta
1914:
1902:
1890:
1882:E. dupreanum
1881:
1861:
1849:
1840:
1828:
1816:
1807:
1798:
1789:
1780:
1768:
1757:Pteropodinae
1730:
1722:S. carolinae
1721:
1713:S. australis
1712:
1694:Syconycteris
1692:
1680:
1671:
1651:
1640:M. woodfordi
1639:
1630:
1621:
1611:Melonycteris
1609:
1597:
1588:
1570:Macroglossus
1568:
1530:
1521:
1509:
1497:
1488:
1479:
1470:
1461:
1452:
1443:
1434:
1425:
1417:D. crenulata
1416:
1407:
1399:D. beauforti
1398:
1390:D. anderseni
1389:
1369:
1357:
1345:
1307:
1300:
1288:
1277:S. blanfordi
1276:
1264:
1252:
1243:
1223:
1211:
1199:
1187:
1175:
1163:
1154:
1145:
1137:M. ecaudatus
1136:
1124:
1113:L. salimalii
1112:
1100:
1088:
1076:
1065:D. spadiceus
1064:
1055:
1046:
1026:
1014:
1005:
996:
987:
978:
969:
960:
940:
928:
916:
904:
897:
885:
873:
863:Alionycteris
861:
849:
840:
820:
809:Cynopterinae
782:
773:
761:
750:N. vizcaccia
749:
741:N. robinsoni
740:
731:
722:
713:
704:
695:
686:
677:
668:
659:
650:
641:
632:
612:
601:Nyctimeninae
577:Superorder:
571:Infraclass:
543:Pteropodidae
489:
442:
438:
428:
409:
405:
395:
383:. Retrieved
369:
363:
357:
319:
316:Conservation
303:
290:
280:
272:
268:
264:
256:
240:
229:Tim Flannery
222:
203:
202:
198:
196:
180:
178:
162:
161:
149:
139:Pteropodidae
29:
3306:C. ophiodon
3288:C. argynnis
3214:M. torquata
3177:Myonycteris
3070:M. pusillus
2928:E. labiatus
2864:Epomophorus
2628:Rousettinae
2602:S. wallacei
2542:P. vampyrus
2515:P. tonganus
2443:P. rennelli
2355:P. ocularis
2265:P. macrotis
2220:P. keyensis
2193:P. howensis
2166:P. fundatus
2157:P. faunulus
2112:P. cognatus
2085:P. caniceps
2076:P. aruensis
1830:Desmalopex
1817:A. mackloti
1808:A. leucotis
1631:M. melanops
1598:M. sobrinus
1408:D. chapmani
1358:A. bulmerae
1225:Ptenochirus
1177:Otopteropus
1164:M. wetmorei
1155:M. niphanae
1146:M. kusnotoi
1089:H. fischeri
1056:D. rickarti
1028:Dyacopterus
898:B. maculata
841:A. aequalis
669:N. cyclotis
462:11336/12485
385:28 December
287:Description
249:New Ireland
227:in 1991 by
3467:Categories
3384:Q117349977
3330:S. zenkeri
3205:M. relicta
3155:M. azagnyi
2996:E. dobsoni
2937:E. minimus
2919:E. grandis
2892:E. anselli
2840:Subfamily
2807:R. lanosus
2678:E. robusta
2669:E. spelaea
2641:Eonycteris
2626:Subfamily
2551:P. vetulus
2533:P. ualanus
2434:P. rayneri
2425:P. pumilus
2364:P. ornatus
2184:P. griseus
2148:P. ennisae
1990:P. pulchra
1952:Pteralopex
1799:A. jubatus
1790:A. humilis
1755:Subfamily
1653:Notopteris
1589:M. minimus
1555:Subfamily
1498:D. viridis
1480:D. peronii
1444:D. inermis
1435:D. exoleta
1332:Subfamily
1290:Thoopterus
1047:D. brooksi
988:C. minutus
942:Cynopterus
807:Subfamily
723:N. minutus
714:N. masalai
660:N. certans
599:Subfamily
585:Chiroptera
412:(5): 388.
336:References
273:P. ennisae
233:subspecies
129:Chiroptera
57:Vulnerable
3230:Plerotini
2798:Subgenus
2728:Rousettus
2726:Subgenus
2717:R. bidens
2708:Subgenus
2691:Rousettus
2398:P. pohlei
2292:P. medius
2049:P. alecto
1972:P. atrata
1963:P. anceps
1939:N. frosti
1928:Neopteryx
1891:E. helvum
1731:S. hobbit
1426:D. emersa
1347:Aproteles
1266:Sphaerias
1244:P. jagori
1212:P. lucasi
1201:Penthetor
1126:Megaerops
1006:C. sphinx
850:A. alecto
822:Aethalops
774:P. raptor
732:N. rabori
687:N. keasti
614:Nyctimene
553:Kingdom:
445:: 83–95.
225:described
157:Species:
95:Kingdom:
89:Eukaryota
3478:Pteropus
3437:84883915
3411:11116721
3378:Wikidata
3240:Plerotes
2946:E. minor
2660:E. major
2461:P. rufus
2337:P. niger
2256:P. lylei
2012:Pteropus
1904:Mirimiri
1770:Acerodon
1453:D. minor
1371:Dobsonia
1253:P. minor
1102:Latidens
918:Chironax
783:P. tenax
696:N. major
633:N. aello
573:Eutheria
567:Mammalia
561:Chordata
559:Phylum:
555:Animalia
471:24662680
263:between
245:holotype
188:Flannery
150:Pteropus
135:Family:
119:Mammalia
109:Chordata
105:Phylum:
99:Animalia
85:Domain:
62:IUCN 3.1
3450:1004462
3424:1145690
2968:Epomops
2130:P. coxi
1999:P. taki
1863:Eidolon
583:Order:
565:Class:
324:by the
306:endemic
292:darker
281:ennisae
243:). The
209:megabat
145:Genus:
125:Order:
115:Class:
60: (
2710:Boneia
497:
469:
277:eponym
3398:7WP9B
3432:IUCN
3419:ITIS
3406:GBIF
495:ISBN
467:PMID
387:2023
370:2022
326:IUCN
267:and
197:The
3445:MDD
3393:CoL
457:hdl
447:doi
414:doi
374:doi
235:of
3469::
3447::
3434::
3421::
3408::
3395::
3380::
2803::
2731::
2713::
479:^
465:.
455:.
443:77
441:.
437:.
410:69
408:.
404:.
368:.
362:.
344:^
215:.
3332:)
3308:)
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534:e
527:t
520:v
503:.
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416::
389:.
376::
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356:"
255:(
239:(
201:(
64:)
20:)
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