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swam quadrupedally using its webbed fore and hind feet for propulsion. Phylogenetic studies including molecular evidence suggest a sister relationship between pinnipeds (seals) and ursoids (bears) as well as musteloids (weasels and otters). It had been popularly assumed that land-dwelling mammals had
456:. The palaeobotanical record suggests that the palaeoenvironment around the lake comprised a forest community transitional between a boreal and a conifer–hardwood forest, in a cool temperate, coastal climate with moderate winters.
393:
at some point transitioned to a more marine existence, in essence "returning to the sea" in order to gain some sort of survival advantage. However, fossil evidence of this transition had been weak or contentious. The discovery of
471:
which had run out of fuel several kilometers away from base camp. The brain case was discovered a year later on the first day of the 2008 field expedition by Martin Lipman, the team's photographer.
503:
316:
epoch about 21 to 24 million years ago. Approximately a metre (three feet) in length, the animal possessed only minimal physical adaptations for swimming. Unlike modern pinnipeds, it did not have
705:
495:
397:
is important as it represents a morphological link in early pinniped evolution, and one that appears to morphologically precede the more familiarly structured genus
2416:
2421:
578:
Rybczynski, Natalia; Dawson, Mary R.; Tedford, Richard H. (2009). "A semi-aquatic Arctic mammalian carnivore from the
Miocene epoch and origin of Pinnipedia".
460:
is the first mammalian carnivore found in the
Haughton lake deposits. This also gives an indication that the entire pinniped family may have originated in the
388:
evolution. Its fossil remains demonstrate the presence of enlarged, probably webbed feet, robust forelimbs and an unspecialized tail. This suggests that
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The initial find is credited to field assistant
Elizabeth Ross, and was partly a matter of luck. Ross had been unexpectedly stranded with the team's
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324:-like, albeit more specialized; its skull and teeth are the features that most clearly indicate that it is a seal.
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and her team using surface collection and screening at an early
Miocene lake deposit of the Haughton Formation of
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that provides information about how the seal family returned to the seas, similar to the way that
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and only known specimen is a nearly complete fossilised skeleton. It is being housed at the
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132:
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It is considered to be the most primitive pinnipedimorph yet found. The genus name is an
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639:"Puijila, the walking seal – a beautiful transitional fossil"
496:"Puijila, the walking seal – a beautiful transitional fossil"
403:, despite apparently being a younger genus. In other words,
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709:, the walking seal — a beautiful transitional fossil
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word for a young seal; the species name honours the
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Natalia
Rybczynski at the Canadian Museum of Nature
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531:. Canadian Museum of Nature. Archived from
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417:illuminates the origin of modern birds.
556:(Press release). BBC News. 2009-04-22.
480:
2417:Prehistoric pinnipeds of North America
715:"Fossil seal had the feet of an otter"
525:"Puijila: A prehistoric walking seal"
7:
2422:Prehistoric mammals of North America
506:from the original on 3 November 2012
27:Extinct genus of primitive pinnipeds
554:"'Missing link' fossil seal walked"
529:Puijila: A prehistoric walking seal
560:from the original on 2 August 2014
25:
2447:Chattian genus first appearances
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1711:
1705:
1698:
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149:
699:, a very basal 'pinnipedimorph'
2287:List of fossil pinnipedimorphs
1:
2452:Prehistoric carnivoran genera
2442:Fossil taxa described in 2009
2427:Aquitanian genus extinctions
721:, 25 April 2009, p. 10.
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637:Yong, Ed (22 April 2009).
500:Not Exactly Rocket Science
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347:Canadian Museum of Nature
320:and its overall form was
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146:Scientific classification
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436:This novel species was
312:which lived during the
433:
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2389:Paleobiology Database
759:and their stem-allies
694:Robert Boessenecker,
641:. National Geographic
502:. Discover Magazine.
428:
368:
40:Temporal range: Late
2437:Transitional fossils
689:: A Significant Seal
429:Life restoration of
672:Paleontology portal
600:10.1038/nature07985
592:2009Natur.458.1021R
586:(7241): 1021–1024.
409:transitional fossil
380:which represents a
376:was a semi-aquatic
442:Natalia Rybczynski
438:discovered in 2007
434:
371:
140:Restored skeleton
2432:Miocene pinnipeds
2404:
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2376:Open Tree of Life
2308:Taxon identifiers
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2282:List of pinnipeds
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834:Amphicynodontidae
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16:(Redirected from
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2316:Puijila darwini
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2101:
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1957:
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981:Pinnipediformes
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952:Pinnipedimorpha
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717:by Bob Holmes,
687:Puijila darwini
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458:Puijila darwini
431:Puijila darwini
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374:Puijila darwini
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298:Puijila darwini
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1803:Archaeodobenus
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815:Pan-Pinnipedia
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684:Brian Switek,
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658:External links
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494:(2009-04-22).
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384:link in early
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2190:Arctocephalus
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1936:Dusignathinae
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1883:Proneotherium
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1565:
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1546:
1540:
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1532:
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1522:
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1506:
1500:
1499:
1492:
1490:
1489:
1485:
1484:
1482:
1480:
1476:
1470:
1469:
1468:Sarmatonectes
1462:
1460:
1459:
1452:
1450:
1449:
1442:
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1439:
1432:
1430:
1429:
1422:
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1399:
1392:
1390:
1389:
1382:
1381:
1378:
1375:
1372:
1367:
1357:
1356:
1355:Virginiaphoca
1349:
1347:
1346:
1339:
1337:
1336:
1329:
1327:
1326:
1322:
1320:
1319:
1315:
1313:
1312:
1311:Leptonychotes
1308:
1306:
1305:
1301:
1299:
1298:
1291:
1289:
1288:
1281:
1279:
1278:
1277:Australophoca
1271:
1269:
1268:
1261:
1260:
1258:
1256:
1252:
1246:
1245:
1241:
1239:
1238:
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1158:
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1134:
1132:
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1114:
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1111:
1104:
1103:
1101:
1099:
1095:
1089:
1088:
1081:
1080:
1077:
1074:
1071:
1066:
1060:
1059:
1052:
1051:
1048:
1045:
1042:
1037:
1034:
1031:
1026:
1020:
1019:
1012:
1010:
1009:
1008:Pinnarctidion
1002:
1000:
999:
998:Pacificotaria
992:
991:
988:
985:
982:
977:
971:
970:
963:
962:
959:
956:
953:
948:
938:
937:
933:
931:
930:
926:
924:
923:
922:Potamotherium
919:
917:
916:
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911:
909:
907:
900:
894:
893:
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847:
845:
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840:
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837:
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828:
824:
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811:
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797:
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766:
763:
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743:
738:
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730:
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719:New Scientist
716:
713:
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546:
535:on 2015-06-21
534:
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420:
418:
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415:
414:Archaeopteryx
410:
406:
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401:
396:
391:
387:
383:
382:morphological
379:
375:
367:
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344:
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325:
323:
319:
315:
311:
307:
304:
300:
299:
289:
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279:
277:
268:
265:
264:Binomial name
261:
257:
256:
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247:
246:
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238:
232:
231:
224:
221:
220:
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211:
208:
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168:
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161:
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147:
143:
138:
134:
129:
115:
110:
105:
100:
95:
90:
85:
80:
75:
70:
65:
60:
54:
47:
43:
37:
33:
30:
19:
2315:
2226:
2219:
2212:
2205:
2195:
2188:
2168:
2163:Proterozetes
2161:
2153:Oriensarctos
2151:
2141:
2123:Thalassoleon
2121:
2096:Pithanotaria
2094:
2084:
2055:
2041:Panotariidae
2020:
2012:Protodobenus
2010:
2000:
1990:
1980:
1973:
1950:
1943:
1911:
1901:
1891:
1881:
1871:
1863:Pelagiarctos
1861:
1851:
1841:
1831:
1821:
1811:
1801:
1774:
1769:Desmatophoca
1767:
1760:
1753:
1704:
1634:
1627:
1617:
1607:
1600:
1593:
1585:Histriophoca
1583:
1576:
1569:
1559:
1536:
1526:
1516:
1509:Cystophorini
1496:
1486:
1466:
1456:
1446:
1436:
1426:
1416:
1406:
1396:
1386:
1353:
1343:
1333:
1323:
1316:
1309:
1302:
1295:
1285:
1275:
1265:
1242:
1235:
1212:
1202:
1192:
1182:
1174:Palmidophoca
1172:
1162:
1152:
1142:
1135:
1128:
1118:
1108:
1087:Magnotherium
1085:
1056:
1018:Pteronarctos
1016:
1006:
996:
967:
934:
928:
927:
920:
913:
906:Semantoridae
890:
883:
878:Pachycynodon
876:
869:
862:
857:Amphicynodon
855:
850:Amphicticeps
848:
841:
798:Infraorder:
706:
696:
686:
643:. Retrieved
632:
583:
579:
573:
562:. Retrieved
548:
537:. Retrieved
533:the original
528:
519:
508:. Retrieved
499:
466:
457:
446:Devon Island
435:
430:
412:
404:
398:
394:
389:
373:
372:
326:
297:
296:
295:
283:
275:
270:
254:
253:
236:
229:
228:
216:Semantoridae
35:
29:
2086:Callorhinus
2077:Callorhinae
1952:Gomphotaria
1945:Dusignathus
1913:Titanotaria
1903:Pseudotaria
1833:Nanodobenus
1578:Halichoerus
1479:Erignathini
1438:Monotherium
1428:Monachopsis
1388:Cryptophoca
1325:Ommatophoca
1255:Lobodontini
1214:Terranectes
1204:Pristiphoca
1144:Neomonachus
1120:Auroraphoca
1058:Devinophoca
969:Enaliarctos
400:Enaliarctos
369:Restoration
282:Rybczynski
235:Rybczynski
51:24–21
2411:Categories
2228:Phocarctos
2207:Hydrarctos
2197:Arctophoca
2143:Eumetopias
2022:Valenictus
1963:Odobeninae
1924:Neodobenia
1893:Prototaria
1853:Osodobenus
1843:Neotherium
1813:Imagotaria
1787:Odobenidae
1762:Atopotarus
1755:Allodesmus
1731:Otarioidea
1668:see below↓
1659:Otarioidea
1602:Pagophilus
1561:Batavipusa
1538:Pachyphoca
1518:Cystophora
1498:Platyphoca
1488:Erignathus
1448:Phocanella
1418:Leptophoca
1398:Frisiphoca
1335:Piscophoca
1287:Hadrokirus
1237:Callophoca
1225:Miroungini
1194:Pontophoca
1154:Eomonachus
1130:Messiphoca
1070:Monachinae
1030:Pinnipedia
915:Necromites
871:Kolponomos
794:Caniformia
792:Suborder:
755:Genera of
645:10 January
564:2014-10-07
539:2014-10-07
510:2014-10-07
475:References
361:Background
336:naturalist
2134:Zalophini
2107:Otariinae
2068:Otariidae
2002:Pliopedia
1992:Ontocetus
1619:Planopusa
1595:Nanophoca
1297:Homiphoca
1267:Acrophoca
1184:Pliophoca
1164:Noriphoca
1110:Afrophoca
1098:Monachini
892:Wangictis
864:Drassonax
800:Arctoidea
788:Carnivora
768:Kingdom:
757:pinnipeds
704:Ed Yong,
608:0028-0836
421:Discovery
378:carnivore
329:Inuktitut
248:Species:
203:Carnivora
169:Kingdom:
163:Eukaryota
42:Oligocene
2355:42399745
2339:BioLib:
2325:Wikidata
2214:Neophoca
2181:Otariini
2170:Zalophus
1982:Odobenus
1873:Pontolis
1776:Eodesmus
1629:Praepusa
1571:Gryphoca
1528:Miophoca
1458:Prophoca
1371:Phocinae
1304:Hydrurga
1244:Mirounga
1137:Monachus
1041:Phocidae
936:Semantor
885:Parictis
843:Allocyon
782:Mammalia
776:Chordata
774:Phylum:
770:Animalia
616:19396145
558:Archived
504:Archived
386:pinniped
343:holotype
318:flippers
310:pinniped
308:of stem-
209:Family:
193:Mammalia
183:Chordata
179:Phylum:
173:Animalia
159:Domain:
44:- Early
2381:6145502
2368:4941721
2331:Q895480
2057:Eotaria
1975:Aivukus
1549:Phocini
1318:Lobodon
929:Puijila
786:Order:
780:Class:
707:Puijila
697:Puijila
624:4371413
588:Bibcode
492:Ed Yong
450:Nunavut
405:Puijila
395:Puijila
390:Puijila
355:Ontario
333:English
314:Miocene
306:species
303:extinct
273:†
252:†
230:Puijila
222:Genus:
199:Order:
189:Class:
120:↓
46:Miocene
36:Puijila
2394:146165
2342:760994
2221:Otaria
622:
614:
606:
580:Nature
462:Arctic
454:Canada
351:Ottawa
341:. The
301:is an
2272:Lists
1609:Phoca
1408:Kawas
620:S2CID
407:is a
322:otter
284:et al
237:et al
2363:GBIF
1636:Pusa
647:2024
612:PMID
604:ISSN
288:2009
241:2009
59:PreꞒ
2350:EoL
596:doi
584:458
469:ATV
448:in
440:by
349:in
286:.,
239:.,
2413::
2391::
2378::
2365::
2352::
2327::
618:.
610:.
602:.
594:.
582:.
527:.
498:.
483:^
464:.
452:,
357:.
353:,
109:Pg
53:Ma
48:,
2203:†
2159:†
2149:†
2119:†
2092:†
2053:†
2018:†
2008:†
1998:†
1988:†
1971:†
1933:†
1909:†
1899:†
1889:†
1879:†
1869:†
1859:†
1849:†
1839:†
1829:†
1819:†
1809:†
1799:†
1743:†
1625:†
1615:†
1591:†
1567:†
1557:†
1534:†
1524:†
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1464:†
1454:†
1444:†
1434:†
1424:†
1414:†
1404:†
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1384:†
1351:†
1341:†
1331:†
1293:†
1283:†
1273:†
1263:†
1233:†
1210:†
1200:†
1190:†
1180:†
1170:†
1160:†
1150:†
1126:†
1116:†
1106:†
1083:†
1054:†
1014:†
1004:†
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903:†
831:†
748:e
741:t
734:v
711:.
701:.
691:.
649:.
626:.
598::
590::
567:.
542:.
513:.
226:†
213:†
114:N
104:K
99:J
94:T
89:P
84:C
79:D
74:S
69:O
64:Ꞓ
20:)
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