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Pannoniasaurus

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539: 530:), as well as a variety of fish and crocodile teeth, were collected from the waste dump of the subterranean Ajka coal mine. The Ajka Coal Formation interdigitates with the CsehbĂĄnya Formation, the depositional environment of the latter was a floodplain, while the Ajka Coal Formation was formed in the accumulation basin of the same river system. Both of these facies were formed in the same paleogeographic area, which itself might have been part of a larger, but isolated landmass. 366: 706: 147: 462: 664: 1694: 1236: 129: 855: 506:, a fragmentary left quadrate. Though all remains (including the holotype) of Pannoniasaurus are isolated bones, the density of the specimens, the various size classes, the large number of similar elements from individual animals, and their unique characters, make it possible to link them together into a single taxon. 693:
a primitive morphology of the complete limbs in correlation with the primitive axial skeleton and pelvis is more probable, but far from certain. It is possible that the retention of a robust sacrum, pelvis and possibly non-paddle-like limbs were used to help to propel the body forward from the bottom
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is known from various material, including 2 isolated premaxillae, 3 maxillae, 2 postorbitofrontals, 2 quadrates, 3 dentaries, 3 splenials, 3 angulars, a coronoid, 2 surangulars, an articular, 91 isolated teeth, 20 cervical, 40 dorsal, 4 sacral, and 18 caudal vertebrae, 34 vertebral fragments, 3 ribs,
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was a medium-sized mosasauroid, estimated to grow up to a maximum of 6 m (20 ft) in length. It exhibited a combination of primitive characteristics, such as having no predental rostrum, the premaxilla-maxilla suture ends anterior to or level with the midline of the fourth maxillary tooth, a
654:
makes it the largest known predator in the waters of this paleoenvironment. Additionally, the crocodile-like flattened skull (as indicated by the premaxilla and maxilla) was a useful adaptation for water-level ambush hunting of terrestrial and shallow water prey.
689:, for example, has an anteriorly oriented, hydropelvic ilium in combination with primitive-looking proximal limb elements, and therefore a flattened, derived distal limb morphology is not entirely impossible for that taxon. For 357:, dubbed "unexpected" because it was discovered in freshwater sediments, unlike other mosasaurs, which were marine predators. It was a medium-sized mosasaur, reaching up to 6 metres (20 ft) in length. 615:
arises (two major lineages of a grade of paddle-bearing marine mosasaurs), is supported, and provides further support for the hypothesis of convergent aquatic adaptations in paddle-bearing mosasaurs.
626:, mosasauroids quickly radiated into a variety of aquatic environments, with some groups reinvading available niches in freshwater habitats, and becoming highly specialized within those ecosystems. 679:
may have looked. It had a primitive vertebral column, a posteriorly oriented ilium and an elongated humerus with a distal epiphysis, all most similar to aigialosaurs. These suggest that
921:"Geochemical study of vertebrate fossils from the Upper Cretaceous (Santonian) CsehbĂĄnya Formation (Hungary): evidence for a freshwater habitat of mosasaurs and pycnodont fish" 646:
was not a seasonal migrant from marine waters into fresh, but rather that ecologically it was a permanent member of a freshwater fauna. With the discovery and description of
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The preserved part of the maxilla bears 12 tooth sockets but the original maxillary tooth count might have been much higher. A number of isolated teeth were attributed to
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nearly straight frontoparietal suture, and a shallow quadrate alar concavity. It also had elongated stapedial pit that was at least three times longer than it was wide.
1772: 412:, sourced from a large number of individuals of differing age classes, are known from the alluvial flood-plain deposits that comprise the CsehbĂĄnya Formation. 1681: 519:, being conical and curved posterolingually, bear crowns with fine anastomosing longitudinal striae, and have a strong mesial but weaker labiodistal carina. 920: 1759: 604:, and the clades that include conventional marine mosasaur-grade mosasauroids such as halisaurs, tylosaurs and plioplatecarpines. The concept of a 1826: 976: 650:, mosasauroid evolution is now understood as also having involved important and unsuspected adaptations to freshwater ecosystems. The size of 446: 1671: 731: 146: 1836: 487:
2 humeral fragments, and 4 ilia. Since all remains are isolated bones, the basis for the referral of this material to
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Arambourg, 1952, a pachyostotic marine squamate from the latest Cretaceous phosphates of Morocco and Syria"
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had an overall aigialosaur-like postcranial morphology (including plesiopelvia and plesiopedia). However,
390: 332: 416: 336: 1798: 1235: 962: 932: 885: 816: 639: 277: 805:"The First Freshwater Mosasauroid (Upper Cretaceous, Hungary) and a New Clade of Basal Mosasauroids" 538: 1750: 711: 365: 493: 1697: 901: 141: 1803: 732:"Vertebrate remains from the Upper Cretaceous (Santonian) Ajka Coal Formation, western Hungary" 1785: 1777: 1379: 1322: 844: 800: 754: 663: 461: 1790: 491:
is based on similar methods used by other authors, such as Houssaye et al. in their study of
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Material was copied from this source, which is available under a Creative Commons License.
374: 329: 41: 936: 889: 820: 586:. Tethysaurinae was reconstructed at the base of a clade that includes the aigialosaurs 1520: 1192: 1138: 839: 804: 581: 1820: 1624: 1596: 1554: 1527: 1486: 1442: 1413: 1172: 1131: 1083: 905: 869: 574: 554: 256: 79: 1575: 1469: 1406: 1397: 1345: 1338: 1331: 1311: 1244: 1199: 1165: 1108: 1055: 1034: 1025: 612: 609: 605: 600: 562: 394: 322: 285: 750: 549:
Analysis by MakĂĄdi et al. found three equally most parsimonious trees that placed
944: 829: 1744: 1640: 1633: 1617: 1589: 1568: 1561: 1513: 1458: 1427: 1372: 1264: 1220: 1206: 1183: 1048: 685: 594: 588: 567: 389:) and referred specimens have been collected from the alluvial sediments of the 325: 54: 1735: 1493: 1479: 1280: 1213: 1117: 1097: 1069: 897: 701: 515: 214: 99: 64: 758: 1603: 1420: 1273: 1253: 1124: 1041: 623: 340: 158: 128: 104: 48: 868:
Houssaye A, Bardet N, Rage JC, Suberbiola XP, Bouya B, et al. (2011).
848: 1729: 1145: 1015: 1009: 1003: 985: 450: 382: 243: 227: 198: 178: 94: 89: 74: 69: 59: 675:
It is difficult to estimate how the unknown girdle and limb elements of
1764: 854: 398: 348: 344: 315: 188: 109: 84: 1076: 997: 730:Ɛsi, Attila; Bodor, Emese Réka; Makådi, Låszló; Rabi, Mårton (2016). 168: 1706: 694:
during prey-capture in shallow water, similar to extant crocodiles.
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was then named and described by Makådi, Caldwell & Ɛsi (2012).
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by Makådi, Caldwell & Ɛsi (2012), and then was classified as
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A single vertebra (MTM V.2000.21) was collected in 1999 from the
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from various exposures at the IharkĂșt open-pit bauxite mine,
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Kocsis L, Ɛsi A, Vennemann T, Trueman CN, Palmer MR (2009).
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at the waste dump of the coal mines next to the town of
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Geochemical and isotopic data are most compatible with
292: 1719: 1664: 1541: 1503: 1468: 1441: 1396: 1355: 1321: 1301: 1263: 1243: 1182: 1155: 1107: 1024: 794: 792: 790: 788: 799:MakĂĄdi, L. S.; Caldwell, M. W.; Ɛsi, A. (2012). 786: 784: 782: 780: 778: 776: 774: 772: 770: 768: 725: 723: 970: 401:since the discovery of the locality in 2000. 8: 1682:List of mosasaur-bearing stratigraphic units 622:indicates that, similar to some lineages of 408:Since then, more than one hundred bones of 1707: 1352: 1318: 1260: 1104: 977: 963: 955: 608:clade of aigialosaurs from within which a 127: 31: 838: 828: 136:Known remains and skeletal restorations 719: 642:, and evidence suggests strongly that 925:Palaeogeogr Palaeoclimatol Palaeoecol 7: 498:in 2011. The paratypes are known by 427:. It was described as belonging to 25: 1693: 1692: 1234: 853: 704: 638:having lived in a predominantly 557:. The three trees reconstructed 447:Hungarian Natural History Museum 302:MakĂĄdi, Caldwell & Ɛsi, 2012 145: 931:(3–4) (280 ed.): 532–542. 1: 1827:Fossil taxa described in 2012 1672:Timeline of mosasaur research 751:10.1016/j.cretres.2015.04.014 945:10.1016/j.palaeo.2009.07.009 872:Pachyvaranus crassispondylus 830:10.1371/journal.pone.0051781 445:are currently housed in the 276:MakĂĄdi, Caldwell & Ɛsi, 669:Pannoniasaurus inexpectatus 565:to the clade that includes 544:Pannoniasaurus inexpectatus 471:Pannoniasaurus inexpectatus 436:Pannoniasaurus inexpectatus 403:Pannoniasaurus inexpectatus 371:Pannoniasaurus inexpectatus 354:Pannoniasaurus inexpectatus 297:Pannoniasaurus inexpectatus 18:Pannoniasaurus inexpectatus 1853: 373:, restored with primitive 1690: 1542:Related groups and genera 1232: 992: 898:10.1017/S0016756810000580 291: 284: 142:Scientific classification 140: 135: 126: 34: 27:Extinct genus of lizards 1677:List of mosasaur genera 502:, a left quadrate, and 347:. It contains a single 672: 546: 473: 438:by Ɛsi et al. (2016). 378: 1799:Paleobiology Database 666: 542:Phylogenetic tree of 541: 522:A single vertebra of 464: 368: 640:freshwater ecosystem 553:in a new clade, the 513:that are similar to 369:Skeletal diagram of 361:Discovery and naming 1837:Mosasaurs of Europe 937:2009PPP...280..532K 890:2011GeoM..148..237H 821:2012PLoSO...751781M 739:Cretaceous Research 712:Paleontology portal 465:Holotype quadrate ( 417:Ajka Coal Formation 391:CsehbĂĄnya Formation 337:Ajka Coal Formation 333:CsehbĂĄnya Formation 673: 547: 474: 423:, 20 km from 379: 1814: 1813: 1786:Open Tree of Life 1713:Taxon identifiers 1704: 1703: 1537: 1536: 1437: 1436: 1380:Plesioplatecarpus 1323:Plioplatecarpinae 1297: 1296: 1230: 1229: 801:Butler, Richard J 671:compared to human 618:The discovery of 441:All specimens of 307: 306: 280: 16:(Redirected from 1844: 1807: 1806: 1794: 1793: 1781: 1780: 1768: 1767: 1755: 1754: 1753: 1740: 1739: 1738: 1708: 1696: 1695: 1665:Related articles 1655:Vallecillosaurus 1357:Plioplatecarpini 1353: 1319: 1261: 1238: 1157:Prognathodontini 1105: 1091:Plesiotylosaurus 979: 972: 965: 956: 949: 948: 916: 910: 909: 865: 859: 857: 852: 842: 832: 796: 763: 762: 736: 727: 714: 709: 708: 707: 294: 275: 268: 255: 242: 226: 213: 150: 149: 131: 121: 51: 40:Temporal range: 32: 21: 1852: 1851: 1847: 1846: 1845: 1843: 1842: 1841: 1832:Russellosaurins 1817: 1816: 1815: 1810: 1802: 1797: 1789: 1784: 1776: 1771: 1763: 1758: 1749: 1748: 1743: 1734: 1733: 1728: 1715: 1705: 1700: 1686: 1660: 1648:Portunatasaurus 1584:Dolichosauridae 1549:Aigialosauridae 1533: 1505:Yaguarasaurinae 1499: 1464: 1433: 1392: 1387:Plioplatecarpus 1366:Latoplatecarpus 1351: 1317: 1303:Russellosaurina 1293: 1288:Phosphorosaurus 1259: 1239: 1226: 1178: 1151: 1103: 1020: 988: 983: 953: 952: 918: 917: 913: 867: 866: 862: 798: 797: 766: 734: 729: 728: 721: 710: 705: 703: 700: 681:P. inexpectatus 677:Pannonisasaurus 661: 632: 536: 496:crassispondylus 459: 363: 330:Late Cretaceous 328:known from the 303: 300: 274: 266: 253: 244:Russellosaurina 240: 224: 211: 144: 122: 120: 119: 118: 117: 112: 107: 102: 97: 92: 87: 82: 77: 72: 67: 62: 57: 47:85.3–83.5  46: 45: 42:Late Cretaceous 38: 28: 23: 22: 15: 12: 11: 5: 1850: 1848: 1840: 1839: 1834: 1829: 1819: 1818: 1812: 1811: 1809: 1808: 1795: 1782: 1769: 1756: 1751:Pannoniasaurus 1741: 1725: 1723: 1721:Pannoniasaurus 1717: 1716: 1711: 1702: 1701: 1691: 1688: 1687: 1685: 1684: 1679: 1674: 1668: 1666: 1662: 1661: 1659: 1658: 1651: 1644: 1637: 1630: 1629: 1628: 1621: 1614: 1607: 1600: 1593: 1581: 1580: 1579: 1572: 1565: 1558: 1545: 1543: 1539: 1538: 1535: 1534: 1532: 1531: 1524: 1521:Russellosaurus 1517: 1509: 1507: 1501: 1500: 1498: 1497: 1490: 1483: 1474: 1472: 1466: 1465: 1463: 1462: 1455: 1452:Pannoniasaurus 1447: 1445: 1439: 1438: 1435: 1434: 1432: 1431: 1424: 1417: 1410: 1402: 1400: 1394: 1393: 1391: 1390: 1383: 1376: 1369: 1361: 1359: 1350: 1349: 1342: 1335: 1327: 1325: 1316: 1315: 1307: 1305: 1299: 1298: 1295: 1294: 1292: 1291: 1284: 1277: 1269: 1267: 1258: 1257: 1249: 1247: 1241: 1240: 1233: 1231: 1228: 1227: 1225: 1224: 1217: 1210: 1203: 1196: 1193:Bentiabasaurus 1188: 1186: 1180: 1179: 1177: 1176: 1169: 1161: 1159: 1153: 1152: 1150: 1149: 1142: 1139:Igdamanosaurus 1135: 1128: 1121: 1113: 1111: 1102: 1101: 1094: 1087: 1080: 1073: 1066: 1059: 1052: 1045: 1038: 1030: 1028: 1022: 1021: 1019: 1018: 1012: 1006: 1000: 993: 990: 989: 984: 982: 981: 974: 967: 959: 951: 950: 911: 884:(2): 237–249. 860: 815:(12): e51781. 764: 718: 717: 716: 715: 699: 696: 691:Pannoniasaurus 660: 657: 652:Pannoniasaurus 648:Pannoniasaurus 644:Pannoniasaurus 636:Pannoniasaurus 631: 628: 620:Pannoniasaurus 582:Russellosaurus 559:Pannoniasaurus 551:Pannoniasaurus 535: 534:Classification 532: 524:Pannoniasaurus 511:Pannoniasaurus 489:Pannoniasaurus 484:Pannoniasaurus 477:Pannoniasaurus 458: 455: 443:Pannoniasaurus 432:Pannoniasaurus 410:Pannoniasaurus 362: 359: 311:Pannoniasaurus 305: 304: 301: 289: 288: 282: 281: 271:Pannoniasaurus 264: 260: 259: 251: 247: 246: 238: 231: 230: 222: 218: 217: 209: 202: 201: 196: 192: 191: 186: 182: 181: 176: 172: 171: 166: 162: 161: 156: 152: 151: 138: 137: 133: 132: 124: 123: 115: 114: 113: 108: 103: 98: 93: 88: 83: 78: 73: 68: 63: 58: 53: 52: 39: 36:Pannoniasaurus 26: 24: 14: 13: 10: 9: 6: 4: 3: 2: 1849: 1838: 1835: 1833: 1830: 1828: 1825: 1824: 1822: 1805: 1800: 1796: 1792: 1787: 1783: 1779: 1774: 1770: 1766: 1761: 1757: 1752: 1746: 1742: 1737: 1731: 1727: 1726: 1724: 1722: 1718: 1714: 1709: 1699: 1689: 1683: 1680: 1678: 1675: 1673: 1670: 1669: 1667: 1663: 1657: 1656: 1652: 1650: 1649: 1645: 1643: 1642: 1638: 1636: 1635: 1631: 1627: 1626: 1625:Tetrapodophis 1622: 1620: 1619: 1615: 1613: 1612: 1608: 1606: 1605: 1601: 1599: 1598: 1597:Dolichosaurus 1594: 1592: 1591: 1587: 1586: 1585: 1582: 1578: 1577: 1573: 1571: 1570: 1566: 1564: 1563: 1559: 1557: 1556: 1555:Aigialosaurus 1552: 1551: 1550: 1547: 1546: 1544: 1540: 1530: 1529: 1528:Yaguarasaurus 1525: 1523: 1522: 1518: 1516: 1515: 1511: 1510: 1508: 1506: 1502: 1496: 1495: 1491: 1489: 1488: 1487:Taniwhasaurus 1484: 1482: 1481: 1476: 1475: 1473: 1471: 1467: 1461: 1460: 1456: 1454: 1453: 1449: 1448: 1446: 1444: 1443:Tethysaurinae 1440: 1430: 1429: 1425: 1423: 1422: 1418: 1416: 1415: 1414:Goronyosaurus 1411: 1409: 1408: 1404: 1403: 1401: 1399: 1395: 1389: 1388: 1384: 1382: 1381: 1377: 1375: 1374: 1370: 1368: 1367: 1363: 1362: 1360: 1358: 1354: 1348: 1347: 1343: 1341: 1340: 1336: 1334: 1333: 1329: 1328: 1326: 1324: 1320: 1314: 1313: 1309: 1308: 1306: 1304: 1300: 1290: 1289: 1285: 1283: 1282: 1278: 1276: 1275: 1271: 1270: 1268: 1266: 1262: 1256: 1255: 1251: 1250: 1248: 1246: 1242: 1237: 1223: 1222: 1218: 1216: 1215: 1211: 1209: 1208: 1204: 1202: 1201: 1197: 1195: 1194: 1190: 1189: 1187: 1185: 1181: 1175: 1174: 1173:Thalassotitan 1170: 1168: 1167: 1163: 1162: 1160: 1158: 1154: 1148: 1147: 1143: 1141: 1140: 1136: 1134: 1133: 1132:Harranasaurus 1129: 1127: 1126: 1122: 1120: 1119: 1115: 1114: 1112: 1110: 1106: 1100: 1099: 1095: 1093: 1092: 1088: 1086: 1085: 1084:Megapterygius 1081: 1079: 1078: 1074: 1072: 1071: 1067: 1065: 1064: 1060: 1058: 1057: 1053: 1051: 1050: 1046: 1044: 1043: 1039: 1037: 1036: 1032: 1031: 1029: 1027: 1023: 1017: 1013: 1011: 1007: 1005: 1001: 999: 995: 994: 991: 987: 980: 975: 973: 968: 966: 961: 960: 957: 946: 942: 938: 934: 930: 926: 922: 915: 912: 907: 903: 899: 895: 891: 887: 883: 879: 875: 873: 870:"A review of 864: 861: 856: 850: 846: 841: 836: 831: 826: 822: 818: 814: 810: 806: 802: 795: 793: 791: 789: 787: 785: 783: 781: 779: 777: 775: 773: 771: 769: 765: 760: 756: 752: 748: 744: 740: 733: 726: 724: 720: 713: 702: 697: 695: 692: 688: 687: 682: 678: 670: 665: 658: 656: 653: 649: 645: 641: 637: 629: 627: 625: 621: 616: 614: 611: 607: 603: 602: 597: 596: 591: 590: 585: 583: 578: 576: 575:Yaguarasaurus 571: 569: 564: 560: 556: 555:Tethysaurinae 552: 545: 540: 533: 531: 529: 528:MTM V.2000.21 525: 520: 518: 517: 512: 507: 505: 504:MTM 2007.31.1 501: 497: 495: 490: 485: 481: 478: 472: 468: 467:MTM 2011.43.1 463: 456: 454: 452: 448: 444: 439: 437: 433: 430: 426: 422: 418: 413: 411: 406: 404: 400: 396: 392: 388: 387:MTM 2011.43.1 384: 376: 372: 367: 360: 358: 356: 355: 350: 346: 342: 338: 334: 331: 327: 324: 320: 317: 313: 312: 299: 298: 290: 287: 283: 279: 273: 272: 265: 262: 261: 258: 257:Tethysaurinae 252: 249: 248: 245: 239: 236: 233: 232: 229: 223: 220: 219: 216: 210: 207: 204: 203: 200: 197: 194: 193: 190: 187: 184: 183: 180: 177: 174: 173: 170: 167: 164: 163: 160: 157: 154: 153: 148: 143: 139: 134: 130: 125: 111: 106: 101: 96: 91: 86: 81: 76: 71: 66: 61: 56: 50: 43: 37: 33: 30: 19: 1720: 1653: 1646: 1639: 1632: 1623: 1616: 1609: 1602: 1595: 1588: 1576:Opetiosaurus 1574: 1567: 1560: 1553: 1526: 1519: 1512: 1492: 1485: 1478: 1470:Tylosaurinae 1457: 1451: 1450: 1426: 1419: 1412: 1407:Gavialimimus 1405: 1398:Selmasaurini 1385: 1378: 1371: 1364: 1346:Sarabosaurus 1344: 1339:Ectenosaurus 1337: 1332:Angolasaurus 1330: 1312:Haasiasaurus 1310: 1286: 1279: 1272: 1252: 1245:Halisaurinae 1219: 1212: 1205: 1200:Eremiasaurus 1198: 1191: 1171: 1166:Prognathodon 1164: 1144: 1137: 1130: 1123: 1116: 1109:Globidensini 1096: 1089: 1082: 1075: 1068: 1061: 1056:Gnathomortis 1054: 1047: 1040: 1035:Amphekepubis 1033: 1026:Mosasaurinae 928: 924: 914: 881: 877: 871: 863: 812: 808: 742: 738: 690: 684: 680: 676: 674: 668: 651: 647: 643: 635: 633: 630:Paleobiology 619: 617: 613:Mosasauridae 610:polyphyletic 606:monophyletic 601:Haasiasaurus 599: 593: 587: 580: 573: 566: 563:sister taxon 558: 550: 548: 543: 527: 523: 521: 514: 510: 508: 503: 500:MTM V.01.115 499: 494:Pachyvaranus 492: 488: 483: 482: 476: 475: 470: 466: 442: 440: 435: 431: 414: 409: 407: 402: 395:Bakony Hills 386: 380: 377:-style limbs 370: 353: 352: 323:tethysaurine 310: 309: 308: 296: 295: 286:Type species 270: 269: 234: 228:Mosasauridae 205: 35: 29: 1745:Wikispecies 1641:Judeasaurus 1634:Adriosaurus 1618:Pontosaurus 1590:Coniasaurus 1569:Komensaurus 1562:Carsosaurus 1514:Romeosaurus 1459:Tethysaurus 1428:Selmasaurus 1373:Platecarpus 1265:Halisaurini 1221:Plotosaurus 1207:Moanasaurus 1184:Mosasaurini 1063:Jormungandr 1049:Dallasaurus 745:: 228–238. 686:Dallasaurus 595:Komensaurus 589:Carsosaurus 577:columbianus 568:Tethysaurus 457:Description 453:, Hungary. 375:aigialosaur 326:mosasauroid 250:Subfamily: 1821:Categories 1611:Primitivus 1494:Tylosaurus 1480:Kaikaifilu 1281:Halisaurus 1214:Mosasaurus 1118:Carinodens 1098:Stelladens 1070:Kourisodon 698:References 516:Halisaurus 397:, Western 343:stage) of 215:Mosasauria 1604:Kaganaias 1421:Khinjaria 1274:Eonatator 1254:Pluridens 1125:Globidens 1042:Clidastes 996:Kingdom: 986:Mosasaurs 906:128706999 759:0195-6671 624:cetaceans 341:Santonian 165:Kingdom: 159:Eukaryota 1736:Q1397354 1730:Wikidata 1698:Category 1146:Xenodens 1016:Squamata 1010:Reptilia 1004:Chordata 1002:Phylum: 998:Animalia 878:Geol Mag 849:23284766 809:PLOS ONE 667:Size of 451:Budapest 383:holotype 221:Family: 199:Squamata 189:Reptilia 179:Chordata 175:Phylum: 169:Animalia 155:Domain: 1791:6158065 1778:1483230 1765:8666962 1014:Order: 1008:Class: 933:Bibcode 886:Bibcode 840:3526648 817:Bibcode 803:(ed.). 570:nopcsai 561:as the 425:IharkĂșt 399:Hungary 349:species 345:Hungary 316:extinct 263:Genus: 195:Order: 185:Class: 116:↓ 1804:253722 1077:Liodon 904:  847:  837:  757:  598:, and 584:coheni 579:, and 314:is an 1773:IRMNG 902:S2CID 735:(PDF) 659:Limbs 469:) of 319:genus 235:Clade 206:Clade 1760:GBIF 845:PMID 755:ISSN 421:Ajka 381:The 335:and 278:2012 55:PreꞒ 941:doi 929:280 894:doi 882:148 835:PMC 825:doi 747:doi 429:cf. 321:of 1823:: 1801:: 1788:: 1775:: 1762:: 1747:: 1732:: 939:. 927:. 923:. 900:. 892:. 880:. 876:. 843:. 833:. 823:. 811:. 807:. 767:^ 753:. 743:57 741:. 737:. 722:^ 592:, 572:, 449:, 351:, 237:: 208:: 105:Pg 49:Ma 44:, 1477:? 978:e 971:t 964:v 947:. 943:: 935:: 908:. 896:: 888:: 851:. 827:: 819:: 813:7 761:. 749:: 526:( 385:( 339:( 293:† 267:† 254:† 241:† 225:† 212:† 110:N 100:K 95:J 90:T 85:P 80:C 75:D 70:S 65:O 60:Ꞓ 20:)

Index

Pannoniasaurus inexpectatus
Late Cretaceous
Ma
PreꞒ
Ꞓ
O
S
D
C
P
T
J
K
Pg
N

Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Reptilia
Squamata
Mosasauria
Mosasauridae
Russellosaurina
Tethysaurinae
Pannoniasaurus
2012
Type species

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