758:, to the genus. Prieto-Márquez and colleagues commented on this referral in a 2006 paper, although noted they had not examined the material themselves. They concluded all the specimens, excepting one fragmentary humerus, were too poorly preserved to allow proper comparison with the holotype material. As such, they disagreed with the referral. Of the humerus, they found it distinguishable from that of the SRA material, and so also not referable to the species. First hand study for a 2013 re-evaluation of European lambeosaurine material supported these conclusions. Laurent had, in a 2003 dissertation, referred more French lambeosaurine material to
5673:
5668:
1179:
4758:
4772:
5655:
136:
4765:
5679:
994:
836:
499:
1125:
2269:
1067:
736:
2223:
5662:
691:. All of this material was excavated from a 4 metres (13 ft) by 2.5 metres (8.2 ft) surface, and it is thought to have belonged to a single individual. The holotype is a nearly complete caudally located cervical vertebra (a cervical vertebra located near the base of the neck), specimen number IPS SRA-1. Another cervical, a humerus, and an ulna were designated the
3712:
929:
118:
3700:
902:, was discovered in outcrops of the Conques Formation during the late 1990s. The taxonomic status of this bonebed has fluctuated over time. It has been debated whether the material represents a singular species or a combination of two distinct ones; but today a single, variable species is considered most likely. It was suggested the material likely belong to
2277:
912:. However, due to the lack of any suitable material for comparison and the lack of tsintaosaurin characteristics in the bonebed material, it has been more recently suggested that it should instead be considered indeterminate lambeosaur material. The 2020 hindlimb description allowed for distinction between
3178:
Casanovas-Cladellas, M. L., Santafé-Llopis, J. V., & Pereda-Suberbiola, X. (1997). Nouveaux restes de
Pararhabdodon (Dinosauria, Hadrosauridae) et synthèse de l’ensemble du materiel découvert dans le Cretacé supérieur de Catalogne. In Second European Workshop of Vertebrate Paleontology, Abstracts
964:
level. In 2006, the stratigraphy of the region was re-evaluated by Albert Prieto-Márquez and colleagues, and the dentary's location was corrected to the LL locality, 750 metres (2,460 ft) away from and 9 metres (30 ft) below the SRA locality. They noted that no dentary had been found at the
1044:
Prieto-Márquez returned again to the dentary in 2013, in a study alongside colleagues providing a review and investigation of hadrosaurs from all over Europe. Further preparation of the specimen in the time since his last study regarding it revealed the uniqueness of the dentary had been exaggerated
1040:
were found. As both taxa from the Talarn
Formation uniquely shared traits with the Asian genus, the authors decided to treat the two as one species, as maintaining them as provisionally separate would in their eyes be misleading to non-specialists, who would likely not distinguish that the two taxa
1260:
in
Lambeosaurinae, merely the presence of this in the jugal itself, unknown in the species, and the angular deltopectoral crest is a trait present even in primitive iguanodontians. Additionally, its tooth count was lower than those of known lambeosaurs. In the 2006 re-evaluation of the genus by
2249:
had been scored as having an ancestral hadrosauroid condition for this trait, which had a strong effect on its position. Upon instead coding it for a modified version of the more advanced hadrosaurid condition, the connection between the two species to the exclusion of other lambeosaurines was
1049:
specimens showing the similar condition were found to have been distorted from a similar process. Correcting for the inaccuracies, the Los Llaus dentary is indistinguishable from that of multiple lambeosaurines, and shares no particular connection to
827:
compared to the maxilla of the holotype, which would conflict with belonging to the same species. As the specimen was still recognized as having tsintaosaurin characteristics, the authors still considered it to likely belong to a close relative of
1092:. The holotype specimen is estimated to have been around 6 metres (20 ft) long. Histological analysis indicates the individual was not fully grown, and so the species likely reached similar sizes to North American and Asian relatives such as
3163:
Casanovas-Cladellas, M. L.; Santafé-Llopis, J. V.; Pereda-Suberbiola, X. (1997). "New remains of
Pararhabdodon isonensis (Dinosauria, Hadrosauridae) and a synthesis of the assemblage of material discovered in the Upper Cretaceous of Catalonia".
443:
have been considered at different times to belong to the species, but all of these assignments have more recently been questioned. It was one of the last non-avian dinosaurs known from the fossil record that went extinct during the
2880:
The geologic unit has been variously divided as the Tremp
Formation, composed of several units, or the Tremp Group, composed of several formations. This article will use the Tremp Group terminology throughout for consistency.
3445:
Blanco, Alejandro; Prieto-Márquez, Albert; De
Esteban-Trivigno, Soledad (2015). "Diversity of hadrosauroid dinosaurs from the Late Cretaceous Ibero-Armorican Island (European Archipelago) assessed from dentary morphology".
2299:, Nicholas Longrich and colleagues found a novel arrangement of lambeosaur phylogeny. As opposed to European lambeosaurs being spread across different lineages, all members of the subfamily from the continent — including
1199:- this made it the first such species known from the continent of Europe. Characters of the skeleton supporting this viewpoint were the truncated and rounded anatomy of the articulation of the maxilla to the
1149:
since its remains were first discovered in the mid-1980s. When the first specimens were originally discovered and described by
Casanovas-Cladellas and colleagues, they were not thought to belong a new
2939:"The osteohistology of new remains of Pararhabdodon isonensis sheds light into the life history and paleoecology of this enigmatic European lambeosaurine dinosaur Author links open overlay panel"
459:. Later discoveries of additional material revealed its true nature as a hadrosaur. Its placement within the group remained controversial - in 1999 it was proposed it belonged to the subfamily
1159:, and of an uncertain species. Not long after, they would be recognized of those of a unique species, and in 1993 they were given their modern name. At this point it was still thought to be a
695:
specimens. In 2020, Jesús F. Serrano and colleagues described new hindlimb material from the site, found in 2018. These are thought to belong to the holotype individual. This includes a
3480:
Prieto-Márquez A, Gaete R, Galobart A, Riera V. New data on
European Hadrosauridae (Dinosauria: Ornithopoda) from the latest Cretaceous of Spain. J Vertebr Paleontol. 2007;27(3): 131A.
908:, based on geographic proximity, but this reassignment was abandoned when that genus was recognized as indeterminate. Since then, it has instead been tentatively assigned to the genus
2235:
Prieto-Márquez would return to the issue in 2009 along with
Jonathan R. Wagner. They once again turned to the articulation between the maxilla and the jugal, finding this to link
1167:
closely related to them. This was based on the characteristics of its vertebrae, as the cranial remains had not yet been found. In 1994 these cranial remains would be found - two
1203:, the truncated nature of the back of the maxilla itself, the ventral deflection of the front of the dentary (thought the only known dentary was later referred to a new species,
568:
sp. New remains from this site, excavated in 1990, brought about a reconsideration of the material, and
Casanovas-Cladellas and colleagues named it as the new genus and species
3108:
Casanovas, M.L, Santafé, J.S., Sanz, J.L., and Buscalioni, A.D. (1987). Arcosaurios (Crocodilia, Dinosauria) del Cretácico superior de la Conca de Tremp (Lleida, España) .
3144:
Laurent, Y., LeLoeuff, J., & Buffetaut, E. (1997). Les Hadrosauridae (Dinosauria, Ornithopoda) du Maastrichtien supérieur des Corbières orientales (Aude, France) .
948:, in the 1990s, palaeontologist Marc Boada discovered a new site in the Talarn Formation, bearing dinosaur fossils. From this site, later named Les Llaus (LL), a right
3212:"Diversity, Relationships, and Biogeography of the Lambeosaurine Dinosaurs from the European Archipelago, with Description of the New Aralosaurin Canardia garonnensis"
952:, specimen designation IPS SRA 27, was excavated. In 1997, Casanovas-Cladellas and colleagues stated this dentary was discovered from SRA, the site where the original
6149:
3090:
Casanovas, M.L, Pereda-Suberbiola, X., Santafé, J.V., and Weishampel, D.B. (1999). First lambeosaurine hadrosaurid from Europe: palaeobiogeographical implications.
762:, held at the Musée des Dinosaures d’Espéraza; the referral was made as the genus was at that point the only named lambeosaurine from the continent. Prieto-Márquez
3281:"The osteohistology of new remains of Pararhabdodon isonensis sheds light into the life history and paleoecology of this enigmatic European lambeosaurine dinosaur"
3317:
Laurent, Y. (2002). Les faunes de vertébrés continentaux du Maastrichtien supérieur d'Europe: systématique et biodiversité (Doctoral dissertation, Toulouse 3).
879:, representatives of a single species. The same re-evaluation could not rule out that this single species or either of the separated species were synonyms of
6208:
1226:
was the first known European lambeosaurine was soon challenged, however, in 2001 by Jason Head, in a study re-evaluating the status of another species,
3388:"Ontogeny and taxonomy of the hadrosaur (Dinosauria, Ornithopoda) remains from Basturs Poble bonebed (late early Maastrichtian, Tremp Syncline, Spain)"
1265:
for the first time. This found it to be a non-hadrosaurid hadrosauroid, a similar position as had been argued by Head. The cladogram of Prieto-Márquez
1195:
revised their position for a final time in 1999, when they published a paper arguing for a position as a primitive member of the hadrosaur subfamily
6136:
5672:
613:. The new material, most importantly cranial and mandibular elements (those from the skull) led to Casanovas-Cladellas' team re-classifying it as a
4757:
6228:
1256:, this was based on a rebuttal of arguments put forward in the 1999 study; the present anatomy maxilla-jugal articulation was not considered a
887:
and the other two taxa. The authors refrained from considering any of them representatives of one species pending data from more material. The
5667:
3744:
3563:
445:
4771:
3054:
Prieto-Marquez, A., Gaete, R., Rivas, G., Galobart, Á., and Boada, M. (2006). Hadrosauroid dinosaurs from the Late Cretaceous of Spain:
983:
section on the dentary, which was medially (inwardly) extended. The authors noted the possibility that future discoveries could lead to
3646:"The first duckbill dinosaur (Hadrosauridae: Lambeosaurinae) from Africa and the role of oceanic dispersal in dinosaur biogeography"
463:, making it the first known from the continent of Europe. Later studies questioned this, instead classifying it as a more primitive
2311:. A unique combination of primitive and derived anatomical features, as well as some unique to the group, supported them forming a
5654:
916:
and the Basturs Poble material to be solidified, with the bonebed material showing inconsistent femoral anatomy with the genus.
6033:
2984:
867:
of the two taxa were shown to be found in other hadrosaurs. This left the former with two unique characteristics - both in the
421:
from the skull. Specimens from other sites, including remains from France, a maxilla previously considered the distinct taxon
6056:
6043:
2849:
2254:
as a member of the subfamily as opposed to outside of Hadrosauridae. This relationship was again supported in Prieto-Márquez
1006:
Evidence for this synonymy would later come in a 2009 study, from Prieto-Márquez alongside Jonathan R. Wagner. Material from
6233:
6048:
2890:
Lower Red Unit equivalent to Talarn Formation and Gray Unit equivalent to La Posa Formation under Tremp Group nomenclature
1799:
1461:
3690:
1922:
1213:
821:
721:
713:
705:
697:
685:
677:
669:
661:
415:
135:
3609:
Head, Jason J. (2001). "A reanalysis of the phylogenetic position of Eolambia caroljonesa (Dinosauria, Iguanodontia)".
1178:
5678:
1437:
2983:
Conti, Simone; Vila, Bernat; Sellés, Albert G.; Galobart, Àngel; Benton, Michael J.; Prieto-Márquez, Albert (2019).
4764:
6218:
6213:
3737:
2241:
2014:
1670:
1348:
1315:
1298:
1016:
999:
871:- and a unique combination of other characteristics, and the latter with only one unique characteristic - in the
467:. In 2009 evidence was put forward that it was indeed a lambeosaurine, and more specifically a close relative of
3622:
2908:
Equivalent to part of the "Lower Red Unit" or "Lower Red Garumnian" unit under the Tremp Formation nomenclature.
875:. As overlapping fossils were not known for the relevant areas, it could not be ruled out that they were indeed
6223:
3844:
1372:
971:
423:
3211:
2937:
Serranoa, Jesús F.; Sellés, Albert G.; Vila, Bernat; Galobart, Àngel; Prieto-Márquez, Albert (February 2021).
3496:
3338:, a new lambeosaurine dinosaur from the Late Cretaceous Ibero-Armorican Island of the European Archipelago".
3334:
Prieto-Márquez, Albert; Fondevilla, Víctor; Sellés, Albert G.; Wagner, Jonathan R.; Galobart; Àngel (2019). "
6203:
1980:
1905:
1041:
were being kept separate to be conservative and not due to strong evidence for two hadrosaurs in the area.
969:
to material known from its original locality. Additionally, they named a genus and species for IPS SRA 27,
6091:
4203:
4055:
3280:
2938:
1823:
647:
In total, known material from the type locality includes: a left and a right maxilla; five cervical, five
399:. The first remains were discovered from the Sant Romà d’Abella fossil locality and assigned to the genus
3558:
Benson, R.B.J.; Brussatte, M.; Xu, X. (2012). Prehistoric Life. London: Dorling Kindersley. pp. 344–345.
6175:
6013:
4569:
3871:
2562:
1775:
1420:
1262:
1245:
899:
795:
in particular because of the rostrocaudally broad rostrocaudal region of the maxilla, a trait unique to
440:
3386:
Fondevilla, V.; Dalla Vecchia, F. M.; Gaete, R.; Galobart, À.; Moncunill-Solé, B.; Köhler, M. (2018).
6123:
5433:
3730:
3657:
3515:
3455:
3399:
3347:
3226:
3009:
1718:
1228:
855:
772:
746:
In their 1997 paper, Laurent and colleagues referred remains from the Le Bexen site of the uppermost
340:
288:
5661:
3279:
Serrano, Jesús F.; Sellés, Albert G.; Vila, Bernat; Galobart, Àngel; Prieto-Márquez, Albert (2020).
3128:
n. gen. n. sp. (Dinosauria). Estudio mofológico, radio-tomográfico y consideraciones biomecanicas .
1874:
1847:
984:
876:
845:
329:
3644:
Longrich, Nicholas R.; Suberbiola, Xabier Pereda; Pyron, R. Alexander; Jalil, Nour-Eddine (2020).
3578:"Global phylogeny of Hadrosauridae (Dinosauria: Ornithopoda) using parsimony and Bayesian methods"
5785:
5396:
3981:
3675:
3626:
3541:
3363:
3300:
3014:
2958:
1058:
was voided, and the specimen is now considered a completely indeterminate lambeosaurine dentary.
945:
319:
130:
6180:
799:
among the tribe. The specimen was discovered at the Serrat del Rostiar 1 locality, part of the
477:. This position has been repeatedly found since, and the group containing them was later named
6162:
6154:
5792:
5531:
5452:
5412:
4296:
4182:
4128:
4099:
3559:
3427:
3254:
2917:
Equivalent to the "Grey Unit" or "Grey Garumnian" unit under the Tremp Formation nomenclature.
2250:
supported. Multiple synapomorphies were found uniting to the two taxa, as well as identifying
1701:
1646:
1396:
860:
850:
800:
523:
6167:
2265:; a diagnosis for the tribe was provided. Their cladogram is reproduced above, on the right.
1045:
significantly by reconstruction of the specimen when it was first prepared in the 1990s. The
803:; this is part of the Tremp Group just like the type locality, but in a lower section of the
5933:
5926:
5736:
5729:
5258:
5169:
5007:
4908:
4894:
4608:
4576:
4211:
4120:
4034:
3969:
3704:
3665:
3618:
3589:
3531:
3523:
3463:
3417:
3407:
3355:
3292:
3244:
3234:
3004:
2996:
2985:"The oldest lambeosaurine dinosaur from Europe: insights into the arrival of Tsintaosaurini"
2950:
2751:
2259:
2063:
1107:
1028:. No material had, as yet, been discovered from the SRA locality permitting comparison with
784:
648:
592:
551:
531:
389:
993:
6018:
5813:
5764:
5627:
5558:
5461:
5281:
5000:
4986:
4965:
4583:
4456:
4223:
3991:
3829:
1751:
1478:
652:
30:
3661:
3519:
3459:
3403:
3351:
3230:
1214:
1124:
835:
822:
498:
6028:
6023:
5961:
5898:
5771:
5620:
5583:
5565:
5510:
5142:
4958:
4951:
4880:
4802:
4789:
4717:
4705:
4674:
4649:
4513:
4435:
4396:
4382:
4259:
4170:
4092:
4070:
4063:
3804:
3422:
3387:
3249:
2355:
2268:
2262:
2000:
1944:
1543:
1196:
956:
remains were found. The following year, they described the specimen and referred it to
573:
478:
460:
267:
254:
212:
735:
6197:
5968:
5912:
5891:
5862:
5778:
5715:
5613:
5606:
5592:
5502:
5482:
5475:
5372:
5322:
5315:
5299:
5274:
5247:
5233:
5204:
5155:
5117:
5082:
5063:
5035:
4979:
4642:
4635:
4590:
4520:
4389:
4248:
4233:
4195:
4106:
3999:
3962:
3955:
3679:
3594:
3577:
3367:
3304:
3166:
Second Workshop of Vertebrate Paleontology, Abstracts (Unpaginated). Espéraza-Quillan
3018:
2962:
2766:
2611:
2532:
2441:
2417:
2393:
2343:
2330:
2228:
2078:
1860:
1528:
1515:
1249:
1208:
1112:
1094:
961:
904:
812:
631:
618:
469:
385:
375:
241:
68:
3630:
3545:
5876:
5852:
5840:
5806:
5799:
5750:
5743:
5549:
5493:
5468:
5440:
5329:
5131:
5096:
5042:
4972:
4944:
4830:
4823:
4663:
4628:
4555:
4548:
4470:
4411:
4366:
4268:
4142:
4135:
4084:
4048:
4006:
3856:
3795:
3716:
2461:
2308:
2304:
2222:
1957:
1257:
1241:
1160:
1146:
1066:
976:
872:
864:
739:
Map of lambeosaur localities in the Pyrenees; MCD 4919 is labelled as belonging to
636:
617:
as opposed to a basal iguanodont. In 1999 it was more specifically identified as a
482:
464:
456:
452:
427:, an additional maxilla from another locality, the material assigned to the genera
296:
196:
3670:
3645:
3527:
3467:
3359:
3296:
3000:
2954:
1171:- and these would be used by the original authors to establish it as a species of
1020:
were examined and compared, and the edentulous slope previously thought unique to
3412:
3239:
979:(unique trait) was used to diagnose the taxon as a new species: a very elongated
587:
Additional material from the type locality was collected in 1994 - including two
6065:
5954:
5919:
5757:
5599:
5572:
5524:
5517:
5350:
5336:
5308:
5288:
5267:
5211:
5190:
5183:
5176:
5110:
5103:
5056:
5021:
4993:
4929:
4922:
4691:
4541:
4527:
4499:
4477:
4419:
4282:
4113:
4077:
4041:
3937:
3924:
3883:
3753:
3124:
Casanovas-Cladellas, M.L., Santafé-Llopis, J.V., and Isidro-Llorens, A. (1993).
2657:
2482:
2369:
2282:
722:
614:
535:
435:
429:
392:
378:
228:
43:
6114:
2258:(2013), where the group containing the two genera was coined as the taxonomic
6007:
5982:
5975:
5947:
5940:
5831:
5820:
5706:
5405:
5385:
5343:
5218:
5070:
4937:
4887:
4852:
4845:
4816:
4656:
4534:
4506:
4375:
4348:
4323:
4163:
4156:
4149:
4027:
1164:
980:
747:
640:
577:
88:
53:
416:
5989:
5905:
5884:
5722:
5419:
5357:
4866:
4838:
4698:
4562:
4492:
4463:
4289:
4275:
4018:
1172:
1155:
1130:
1089:
1086:
883:, as no taxonomically informative areas of the skeleton are known from both
686:
670:
662:
626:
564:
543:
366:
147:
93:
37:
3431:
3258:
928:
895:
to be ruled out, as the femurs of the two taxa were anatomically distinct.
714:
481:. A different position, related to other European lambeosaurs in the group
117:
3210:
Prieto-Márquez, A.; Dalla Vecchia, F. M.; Gaete, R.; Galobart, À. (2013).
706:
698:
6108:
5869:
5538:
5426:
5226:
5197:
5124:
5049:
5014:
4915:
4901:
4859:
4683:
4484:
4449:
4355:
4341:
4303:
3813:
3786:
3777:
2578:
1071:
692:
678:
588:
515:
503:
410:
381:
180:
167:
83:
78:
63:
58:
48:
2276:
6141:
5162:
4597:
2674:
2295:
1168:
949:
600:
559:
555:
514:
began in Spring 1985, at the Sant Romà d’Abella (SRA) locality (in the
98:
73:
3536:
409:
in 1993. Known material includes assorted postcranial remains, mostly
5028:
4621:
3771:
1191:
804:
755:
656:
609:
596:
157:
6085:
1207:, and later declared that of indeterminate lambeosaurine), its tall
965:
SRA locality to compare to the LL specimen, and so they restricted
4873:
4403:
2312:
2275:
2267:
2221:
1200:
1177:
1150:
1123:
1082:
1065:
992:
927:
868:
834:
767:
734:
527:
519:
497:
474:
396:
371:
6128:
4241:
2286:, proposed by Longrich and colleagues to be a close relative of
2272:
Maxilla specimen MCD 4919 from the Serrat del Rostiar 1 locality
751:
629:
from the continent, and the second valid hadrosaur, preceded by
547:
6089:
5694:
4787:
3922:
3764:
3726:
1054:. With this, their reasoning for assignment of the specimen to
936:", considered at different times a distinct taxon, specimen of
3722:
1145:
has been classified in a number of different positions within
839:
A selection of dentary material from the Basturs Poble bonebed
811:
thus extended the range of the genus further within the upper
350:
Casanovas-Cladellas, Santafé-Llopis & Isidro-Llorens, 1993
313:
Casanovas-Cladellas, Santafé-Llopis & Isidro-Llorens, 1993
1116:
was cited as a similar example, being around the same size.
3623:
10.1671/0272-4634(2001)021[0392:AROTPP]2.0.CO;2
1261:
Albert Prieto-Márquez and colleagues, it was included in a
510:
Excavation of specimens that would later be used to erect
287:
Casanovas-Cladellas, Santafé-Llopis & Isidro-Llorens,
3505:: a new clade of lambeosaurine hadrosaurids from Eurasia"
1106:
living on an island, something generally associated with
1110:, a phenomenon exhibited in other European hadrosaurs;
3205:
863:, another part of the Tremp Group. Previously claimed
815:. A 2019 paper questioned the referral of MCD 4919 to
572:
in 1993. At the time, it was still considered to be a
3688:
3329:
3327:
3325:
3323:
3203:
3201:
3199:
3197:
3195:
3193:
3191:
3189:
3187:
3185:
2904:
2902:
2900:
2898:
2896:
1102:(around 9 metres (30 ft) long). This is despite
891:
himdlimb described in 2020 allowed for synonymy with
303:
6098:
5999:
5850:
5830:
5705:
5582:
5548:
5492:
5451:
5395:
5371:
5298:
5257:
5141:
5080:
4800:
4716:
4673:
4607:
4433:
4365:
4321:
4258:
4222:
4181:
4017:
3980:
3935:
3882:
3855:
3828:
791:was by that point assigned to. It was referable to
621:hadrosaur, in a paper again by Casanovas-Cladellas
451:Initially, the material was thought to belong to a
1252:" (later renamed to Saurolophinae). In regards to
783:in 2013. It possessed traits of the tsintaosaurin
455:dinosaur, or some other similar type of primitive
3381:
3379:
3377:
502:Stratigraphic map of hadrosaur localities in the
3158:
3156:
3154:
1138:was originally thought to be closely related to
849:, from the Blasi 1 locality of the underlying
779:A maxilla, specimen MCD 4919, was referred to
659:; fragmentary rib bones; one end of the right
3738:
3112:. Volumen extraordinario Galve-Tremp:95-110.
819:. The specimen was found to have a differing
766:(2013) used this material to establish a new
364:(meaning "near fluted tooth" in reference to
8:
3274:
3272:
3270:
3268:
2876:
2874:
2872:
2870:
1248:than the split between Lambeosaurinae and "
898:A hadrosaur mega-bonebed, later termed the
6086:
5702:
5691:
5377:
5147:
5088:
4808:
4797:
4784:
4613:
4441:
4333:
4329:
4187:
3947:
3943:
3932:
3919:
3861:
3834:
3825:
3761:
3745:
3731:
3723:
3490:
3488:
3486:
3050:
3048:
1153:at all, but merely to belong to the genus
403:, and later named as the distinct species
116:
20:
3669:
3593:
3582:Zoological Journal of the Linnean Society
3535:
3495:Prieto-Márquez, A.; Wagner, J.R. (2009).
3421:
3411:
3248:
3238:
3046:
3044:
3042:
3040:
3038:
3036:
3034:
3032:
3030:
3028:
3010:1983/be876efb-979c-4237-94f9-5f8d80121f7e
3008:
1219:of the humerus being distally projected.
940:, or indeterminate lambeosaurine specimen
3140:
3138:
3104:
3102:
3100:
3086:
3084:
3082:
3080:
3078:
3076:
3074:
3072:
2323:, instead taking a more basal position.
1240:were found to instead be more primitive
960:, and again stated it was from the same
542:described remains of an ornithopod from
3695:
3120:
3118:
2929:
2866:
932:Holotype dentary and only specimen of "
731:Referred material from other localities
625:. This made it the first member of the
2293:In a 2020 study describing and naming
1629:
1274:
1014:, and material of the Chinese species
599:, two fragmentary ribs, and a partial
2245:. In the 2006 phylogenetic analysis,
446:Cretaceous-Paleogene extinction event
7:
1024:was found to be nearly identical in
603:. The species name was corrected to
6209:Late Cretaceous dinosaurs of Europe
2744:
2649:
2627:
2603:
2555:
2548:
2524:
2474:
2467:
2457:
2433:
2409:
2385:
2361:
2336:
2326:
2056:
2006:
1996:
1972:
1950:
1897:
1890:
1866:
1839:
1815:
1791:
1767:
1743:
1693:
1686:
1662:
1638:
1631:
1521:
1494:
1453:
1412:
1388:
1364:
1340:
1290:
1283:
1276:
859:, from the Blasi 3 locality of the
843:The same 2013 study also evaluated
807:, making it older. The referral to
3611:Journal of Vertebrate Paleontology
3064:Journal of Vertebrate Paleontology
1269:(2006) is seen below on the left:
1232:, as a primitive lambeosaur. Both
14:
997:Diagram of the holotype skull of
5677:
5671:
5666:
5660:
5653:
4770:
4763:
4756:
3710:
3698:
3595:10.1111/j.1096-3642.2009.00617.x
2635:Serrat del Rostiar lambeosaurine
1163:or some other type of primitive
1128:Modern life reconstruction of a
134:
6034:List of informal dinosaur names
2307:clade, which was therein named
538:. In 1987, Casanovas-Cladellas
6044:Timeline of hadrosaur research
2850:Timeline of hadrosaur research
1:
6229:Fossil taxa described in 1993
3671:10.1016/j.cretres.2020.104678
3528:10.1016/j.cretres.2009.06.005
3514:. online preprint (5): 1238.
3468:10.1016/j.cretres.2015.04.001
3360:10.1016/j.cretres.2018.12.002
3297:10.1016/j.cretres.2020.104677
3001:10.1016/j.cretres.2019.104286
2955:10.1016/j.cretres.2020.104677
2319:was not found to be close to
1800:Telmatosaurus transsylvanicus
1462:Telmatosaurus transsylvanicus
1271:
489:History and assigned material
3413:10.1371/journal.pone.0206287
3240:10.1371/journal.pone.0069835
2239:to the Asian lambeosaurine,
1923:Brachylophosaurus canadensis
3576:Prieto-Márquez, A. (2010).
1438:Gilmoreosaurus mongoliensis
584:was in reference to Isona.
494:Sant Romà d’Abella material
6250:
1273:
1032:. However, traits uniting
16:Extinct genus of dinosaurs
5701:
5690:
5651:
5380:
5150:
5091:
4811:
4796:
4783:
4754:
4616:
4444:
4336:
4332:
4190:
3950:
3946:
3931:
3918:
3864:
3837:
3824:
3760:
3503:Tsintaosaurus spinorhinus
2764:
2749:
2742:
2671:
2654:
2647:
2632:
2625:
2608:
2601:
2575:
2560:
2553:
2546:
2529:
2522:
2496:
2479:
2472:
2465:
2455:
2438:
2431:
2414:
2407:
2390:
2383:
2366:
2359:
2341:
2334:
2303:— were found to form one
2242:Tsintaosaurus spinorhinus
2076:
2061:
2054:
2028:
2015:Tsintaosaurus spinorhinus
2011:
2004:
1994:
1977:
1970:
1955:
1948:
1919:
1902:
1895:
1888:
1871:
1864:
1844:
1837:
1820:
1813:
1796:
1789:
1772:
1765:
1748:
1741:
1715:
1698:
1691:
1684:
1671:Probactrosaurus gobiensis
1667:
1660:
1643:
1636:
1541:
1526:
1519:
1499:
1492:
1475:
1458:
1451:
1434:
1417:
1410:
1393:
1386:
1369:
1362:
1349:Probactrosaurus gobiensis
1345:
1338:
1316:Iguanodon atherfieldensis
1312:
1299:Iguanodon bernissartensis
1295:
1288:
1281:
1017:Tsintaosaurus spinorhinus
1000:Tsintaosaurus spinorhinus
667:; a fragment of the left
325:
318:
302:
295:
131:Scientific classification
129:
124:
115:
23:
3130:Paleontologia i Evolució
3060:Koutalisaurus kohlerorum
1373:Ouranosaurus nigeriensis
1074:and a hypothetical adult
972:Koutalisaurus kohlerorum
934:Koutalisaurus kohlerorum
485:, was proposed in 2020.
424:Koutalisaurus kohlerorum
3499:Pararhabdodon isonensis
3126:Pararhabdodon isonensis
3056:Pararhabdodon isonensis
2032:Pararhabdodon isonensis
1981:Jaxartosaurus aralensis
1906:Edmontosaurus annectens
1503:Pararhabdodon isonensis
881:Pararhabdodon isonensis
550:vertebra, some partial
406:Pararhabdodon isonensis
308:Pararhabdodon isonensis
3146:Revue de Paléobiologie
2290:
2273:
2232:
1824:Tethyshadros insularis
1186:
1139:
1075:
1070:Estimated size of the
1003:
941:
840:
743:
570:Pararhabdodon isonense
507:
347:Pararhabdodon isonense
6176:Paleobiology Database
6014:Basturs Poble bonebed
3336:Adynomosaurus arcanus
2563:Basturs lambeosaurine
2279:
2271:
2225:
1776:Bactrosaurus johnsoni
1421:Bactrosaurus johnsoni
1263:phylogenetic analysis
1181:
1127:
1069:
996:
931:
900:Basturs Poble bonebed
838:
738:
501:
441:Basturs Poble bonebed
6234:Ornithischian genera
6036:(contains hadrosaur
5434:Probrachylophosaurus
1719:Eolambia caroljonesa
1229:Eolambia caroljonesa
1189:Casanovas-Cladellas
1182:Life restoration of
987:of their taxon with
856:Arenysaurus ardevoli
773:Canardia garonnensis
558:, and a fragmentary
439:, and the extensive
3662:2021CrRes.12004678L
3650:Cretaceous Research
3520:2009CrRes..30.1238P
3512:Cretaceous Research
3460:2015CrRes..56..447B
3448:Cretaceous Research
3404:2018PLoSO..1306287F
3352:2019CrRes..96...19P
3340:Cretaceous Research
3285:Cretaceous Research
3231:2013PLoSO...869835P
3110:Estudios Geológicos
3092:Geological Magazine
3062:, gen. et sp. nov.
2989:Cretaceous Research
2943:Cretaceous Research
1875:Hadrosaurus foulkii
1848:Lophorhothon atopus
1215:deltopectoral crest
920:Relationship with "
846:Blasisaurus canudoi
823:ectopterygoid shelf
607:in 1997 by Laurent
330:Blasisaurus canudoi
5786:Ornithopodichnites
5397:Brachylophosaurini
3982:Hypsilophodontidae
2291:
2274:
2233:
1222:The proposal that
1187:
1140:
1081:would have been a
1076:
1010:, the holotype of
1004:
946:Abella de la Conca
942:
841:
744:
580:, hence the name;
508:
335:Cruzado-Caballero
6191:
6190:
6163:Open Tree of Life
6092:Taxon identifiers
6083:
6082:
6079:
6078:
6075:
6074:
5793:Shiraminesauropus
5686:
5685:
5649:
5648:
5645:
5644:
5641:
5640:
5637:
5636:
5532:Naashoibitosaurus
5453:Austrokritosauria
5413:Brachylophosaurus
5367:
5366:
4779:
4778:
4752:
4751:
4748:
4747:
4744:
4743:
4740:
4739:
4736:
4735:
4429:
4428:
4317:
4316:
4313:
4312:
4297:Transylvanosaurus
4183:Rhabdodontomorpha
4129:Notohypsilophodon
4100:Macrogryphosaurus
3914:
3913:
3910:
3909:
3906:
3905:
3902:
3901:
3564:978-0-7566-9910-9
2840:
2839:
2831:
2830:
2822:
2821:
2813:
2812:
2804:
2803:
2795:
2794:
2786:
2785:
2777:
2776:
2731:
2730:
2722:
2721:
2713:
2712:
2704:
2703:
2695:
2694:
2686:
2685:
2590:
2589:
2511:
2510:
2220:
2219:
2215:
2214:
2206:
2205:
2197:
2196:
2188:
2187:
2179:
2178:
2170:
2169:
2161:
2160:
2152:
2151:
2143:
2142:
2134:
2133:
2125:
2124:
2116:
2115:
2107:
2106:
2098:
2097:
2089:
2088:
2043:
2042:
1934:
1933:
1730:
1729:
1702:Protohadros byrdi
1647:Equijubus normani
1626:
1625:
1617:
1616:
1608:
1607:
1599:
1598:
1590:
1589:
1581:
1580:
1572:
1571:
1563:
1562:
1554:
1553:
1397:Protohadros byrdi
1327:
1326:
944:Near the village
865:diagnostic traits
861:La Posa Formation
801:Conques Formation
357:
356:
351:
343:
291:
6241:
6219:Fossils of Spain
6214:Cretaceous Spain
6184:
6183:
6171:
6170:
6158:
6157:
6145:
6144:
6132:
6131:
6119:
6118:
6117:
6087:
6060:mummy S.M. R4036
5934:Microhadrosaurus
5927:Mandschurosaurus
5737:Camptosaurichnus
5730:Bonaparteichnium
5703:
5696:Related articles
5692:
5681:
5675:
5670:
5665:
5664:
5657:
5378:
5259:Parasaurolophini
5170:Angulomastacator
5148:
5089:
5008:Shuangmiaosaurus
4909:Huehuecanauhtlus
4895:Gongpoquansaurus
4809:
4798:
4785:
4774:
4768:
4767:
4760:
4614:
4609:Hadrosauriformes
4577:Ratchasimasaurus
4442:
4334:
4330:
4212:Muttaburrasaurus
4188:
4121:Muttaburrasaurus
4035:Atlascopcosaurus
3970:Thescelosauridae
3948:
3944:
3933:
3920:
3862:
3835:
3826:
3819:
3818:
3762:
3747:
3740:
3733:
3724:
3715:
3714:
3713:
3703:
3702:
3701:
3694:
3684:
3683:
3673:
3641:
3635:
3634:
3606:
3600:
3599:
3597:
3573:
3567:
3556:
3550:
3549:
3539:
3509:
3492:
3481:
3478:
3472:
3471:
3442:
3436:
3435:
3425:
3415:
3398:(10): e0206287.
3383:
3372:
3371:
3331:
3318:
3315:
3309:
3308:
3276:
3263:
3262:
3252:
3242:
3216:
3207:
3180:
3176:
3170:
3169:
3160:
3149:
3142:
3133:
3122:
3113:
3106:
3095:
3088:
3067:
3052:
3023:
3022:
3012:
2980:
2974:
2973:
2971:
2969:
2934:
2918:
2915:
2909:
2906:
2891:
2888:
2882:
2878:
2752:Parasaurolophini
2745:
2650:
2628:
2604:
2556:
2549:
2525:
2475:
2468:
2458:
2434:
2410:
2386:
2362:
2337:
2327:
2064:Parasaurolophini
2057:
2007:
1997:
1973:
1951:
1898:
1891:
1867:
1840:
1816:
1792:
1768:
1744:
1694:
1687:
1663:
1639:
1632:
1522:
1495:
1454:
1413:
1389:
1365:
1341:
1291:
1284:
1277:
1272:
1216:
1108:insular dwarfism
824:
724:
716:
708:
700:
688:
680:
672:
664:
593:dorsal vertebrae
532:Talarn Formation
418:
390:Upper Cretaceous
349:
334:
305:
286:
279:
266:
253:
240:
227:
211:
195:
139:
138:
120:
110:
40:
29:Temporal range:
21:
6249:
6248:
6244:
6243:
6242:
6240:
6239:
6238:
6224:Tremp Formation
6194:
6193:
6192:
6187:
6179:
6174:
6166:
6161:
6153:
6148:
6140:
6135:
6127:
6122:
6113:
6112:
6107:
6094:
6084:
6071:
6052:mummy AMNH 5060
6019:Dakota (fossil)
5995:
5846:
5826:
5814:Tetrapodosaurus
5765:Hadrosaurichnus
5697:
5682:
5659:
5633:
5628:Shantungosaurus
5578:
5559:Augustynolophus
5544:
5488:
5462:Bonapartesaurus
5447:
5391:
5363:
5294:
5282:Parasaurolophus
5253:
5137:
5085:
5076:
5001:Qianjiangsaurus
4987:Probactrosaurus
4966:Penelopognathus
4805:
4792:
4775:
4762:
4732:
4712:
4669:
4603:
4584:Riabininohadros
4457:Bayannurosaurus
4438:
4425:
4361:
4326:
4309:
4254:
4224:Tenontosauridae
4218:
4177:
4013:
3992:Gideonmantellia
3976:
3940:
3927:
3898:
3878:
3851:
3845:Avemetatarsalia
3830:Avemetatarsalia
3820:
3767:
3766:
3756:
3751:
3721:
3711:
3709:
3699:
3697:
3689:
3687:
3643:
3642:
3638:
3608:
3607:
3603:
3575:
3574:
3570:
3557:
3553:
3507:
3494:
3493:
3484:
3479:
3475:
3444:
3443:
3439:
3385:
3384:
3375:
3333:
3332:
3321:
3316:
3312:
3278:
3277:
3266:
3214:
3209:
3208:
3183:
3177:
3173:
3162:
3161:
3152:
3143:
3136:
3132:26-27:121-131.
3123:
3116:
3107:
3098:
3094:136(2):205-211.
3089:
3070:
3066:26(4): 929-943.
3053:
3026:
2982:
2981:
2977:
2967:
2965:
2936:
2935:
2931:
2927:
2922:
2921:
2916:
2912:
2907:
2894:
2889:
2885:
2879:
2868:
2863:
2858:
2846:
2841:
2832:
2823:
2814:
2805:
2796:
2787:
2778:
2732:
2723:
2714:
2705:
2696:
2687:
2591:
2512:
2216:
2207:
2198:
2189:
2180:
2171:
2162:
2153:
2144:
2135:
2126:
2117:
2108:
2099:
2090:
2044:
1935:
1752:Tanius sinensis
1731:
1627:
1618:
1609:
1600:
1591:
1582:
1573:
1564:
1555:
1479:Tanius sinensis
1328:
1218:
1122:
1064:
926:
826:
733:
726:
718:
710:
702:
690:
682:
674:
666:
506:, including SRA
496:
491:
473:, a genus from
420:
314:
311:
285:
277:
264:
251:
238:
225:
209:
193:
133:
111:
109:
108:
107:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
35:
34:
31:Late Cretaceous
27:
17:
12:
11:
5:
6247:
6245:
6237:
6236:
6231:
6226:
6221:
6216:
6211:
6206:
6204:Lambeosaurines
6196:
6195:
6189:
6188:
6186:
6185:
6172:
6159:
6146:
6133:
6120:
6104:
6102:
6096:
6095:
6090:
6081:
6080:
6077:
6076:
6073:
6072:
6070:
6069:
6062:
6054:
6046:
6041:
6031:
6029:Hypsilophodont
6026:
6024:Hadrosaur diet
6021:
6016:
6011:
6003:
6001:
6000:Other articles
5997:
5996:
5994:
5993:
5986:
5979:
5972:
5965:
5962:Stephanosaurus
5958:
5951:
5944:
5937:
5930:
5923:
5916:
5909:
5902:
5899:Fulgurotherium
5895:
5888:
5881:
5873:
5866:
5858:
5856:
5848:
5847:
5845:
5844:
5836:
5834:
5828:
5827:
5825:
5824:
5817:
5810:
5803:
5796:
5789:
5782:
5775:
5772:Iguanodontipus
5768:
5761:
5754:
5747:
5740:
5733:
5726:
5719:
5711:
5709:
5699:
5698:
5695:
5688:
5687:
5684:
5683:
5652:
5650:
5647:
5646:
5643:
5642:
5639:
5638:
5635:
5634:
5632:
5631:
5624:
5621:Laiyangosaurus
5617:
5610:
5603:
5596:
5588:
5586:
5584:Edmontosaurini
5580:
5579:
5577:
5576:
5569:
5566:Prosaurolophus
5562:
5554:
5552:
5546:
5545:
5543:
5542:
5535:
5528:
5521:
5514:
5511:Coahuilasaurus
5507:
5498:
5496:
5490:
5489:
5487:
5486:
5479:
5472:
5465:
5457:
5455:
5449:
5448:
5446:
5445:
5437:
5430:
5423:
5416:
5409:
5401:
5399:
5393:
5392:
5390:
5389:
5381:
5375:
5369:
5368:
5365:
5364:
5362:
5361:
5354:
5347:
5340:
5333:
5326:
5319:
5312:
5304:
5302:
5296:
5295:
5293:
5292:
5285:
5278:
5271:
5263:
5261:
5255:
5254:
5252:
5251:
5244:
5237:
5230:
5223:
5215:
5208:
5201:
5194:
5187:
5180:
5173:
5166:
5159:
5151:
5145:
5143:Lambeosaurinae
5139:
5138:
5136:
5135:
5128:
5121:
5114:
5107:
5100:
5092:
5086:
5081:
5078:
5077:
5075:
5074:
5067:
5060:
5053:
5046:
5039:
5032:
5025:
5018:
5011:
5004:
4997:
4990:
4983:
4976:
4969:
4962:
4959:Nanyangosaurus
4955:
4952:Nanningosaurus
4948:
4941:
4934:
4926:
4919:
4912:
4905:
4898:
4891:
4884:
4881:Gilmoreosaurus
4877:
4870:
4863:
4856:
4849:
4842:
4835:
4827:
4820:
4812:
4806:
4803:Hadrosauroidea
4801:
4794:
4793:
4790:Hadrosauroidea
4788:
4781:
4780:
4777:
4776:
4755:
4753:
4750:
4749:
4746:
4745:
4742:
4741:
4738:
4737:
4734:
4733:
4731:
4730:
4729:
4728:
4722:
4720:
4718:Hadrosauroidea
4714:
4713:
4711:
4710:
4706:Mantellisaurus
4702:
4695:
4688:
4679:
4677:
4675:Iguanodontidae
4671:
4670:
4668:
4667:
4660:
4653:
4650:Mantellisaurus
4646:
4639:
4632:
4625:
4617:
4611:
4605:
4604:
4602:
4601:
4594:
4587:
4580:
4573:
4566:
4559:
4552:
4545:
4538:
4531:
4524:
4517:
4514:Iguanacolossus
4510:
4503:
4496:
4489:
4481:
4474:
4467:
4460:
4453:
4445:
4439:
4436:Ankylopollexia
4434:
4431:
4430:
4427:
4426:
4424:
4423:
4416:
4408:
4400:
4397:Eousdryosaurus
4393:
4386:
4383:Dysalotosaurus
4379:
4371:
4369:
4363:
4362:
4360:
4359:
4352:
4345:
4337:
4327:
4322:
4319:
4318:
4315:
4314:
4311:
4310:
4308:
4307:
4300:
4293:
4286:
4279:
4272:
4264:
4262:
4260:Rhabdodontidae
4256:
4255:
4253:
4252:
4245:
4238:
4229:
4227:
4220:
4219:
4217:
4216:
4208:
4200:
4191:
4185:
4179:
4178:
4176:
4175:
4171:Weewarrasaurus
4167:
4160:
4153:
4146:
4139:
4132:
4125:
4117:
4110:
4103:
4096:
4093:Leaellynasaura
4089:
4081:
4074:
4071:Gasparinisaura
4067:
4064:Galleonosaurus
4060:
4052:
4045:
4038:
4031:
4023:
4021:
4015:
4014:
4012:
4011:
4003:
3996:
3987:
3985:
3978:
3977:
3975:
3974:
3966:
3959:
3951:
3941:
3936:
3929:
3928:
3923:
3916:
3915:
3912:
3911:
3908:
3907:
3904:
3903:
3900:
3899:
3897:
3896:
3895:
3894:
3888:
3886:
3880:
3879:
3877:
3876:
3875:
3874:
3865:
3859:
3853:
3852:
3850:
3849:
3848:
3847:
3838:
3832:
3822:
3821:
3817:
3816:
3807:
3805:Neornithischia
3798:
3789:
3780:
3774:
3765:
3758:
3757:
3752:
3750:
3749:
3742:
3735:
3727:
3720:
3719:
3707:
3686:
3685:
3636:
3617:(2): 392–396.
3601:
3588:(2): 435–502.
3568:
3551:
3482:
3473:
3437:
3373:
3319:
3310:
3264:
3181:
3179:(unpaginated).
3171:
3150:
3134:
3114:
3096:
3068:
3058:revisited and
3024:
2975:
2928:
2926:
2923:
2920:
2919:
2910:
2892:
2883:
2865:
2864:
2862:
2859:
2857:
2854:
2853:
2852:
2845:
2842:
2838:
2837:
2834:
2833:
2829:
2828:
2825:
2824:
2820:
2819:
2816:
2815:
2811:
2810:
2807:
2806:
2802:
2801:
2798:
2797:
2793:
2792:
2789:
2788:
2784:
2783:
2780:
2779:
2775:
2774:
2771:
2770:
2763:
2760:
2759:
2756:
2755:
2748:
2743:
2741:
2738:
2737:
2734:
2733:
2729:
2728:
2725:
2724:
2720:
2719:
2716:
2715:
2711:
2710:
2707:
2706:
2702:
2701:
2698:
2697:
2693:
2692:
2689:
2688:
2684:
2683:
2680:
2679:
2670:
2667:
2666:
2663:
2662:
2653:
2648:
2646:
2643:
2642:
2639:
2638:
2631:
2626:
2624:
2621:
2620:
2617:
2616:
2607:
2602:
2600:
2597:
2596:
2593:
2592:
2588:
2587:
2584:
2583:
2574:
2571:
2570:
2567:
2566:
2559:
2554:
2552:
2547:
2545:
2542:
2541:
2538:
2537:
2528:
2523:
2521:
2518:
2517:
2514:
2513:
2509:
2508:
2505:
2504:
2495:
2492:
2491:
2488:
2487:
2478:
2473:
2471:
2466:
2464:
2456:
2454:
2451:
2450:
2447:
2446:
2437:
2432:
2430:
2427:
2426:
2423:
2422:
2413:
2408:
2406:
2403:
2402:
2399:
2398:
2389:
2384:
2382:
2379:
2378:
2375:
2374:
2365:
2360:
2358:
2356:Lambeosaurinae
2352:
2351:
2348:
2347:
2340:
2335:
2333:
2325:
2263:Tsintaosaurini
2218:
2217:
2213:
2212:
2209:
2208:
2204:
2203:
2200:
2199:
2195:
2194:
2191:
2190:
2186:
2185:
2182:
2181:
2177:
2176:
2173:
2172:
2168:
2167:
2164:
2163:
2159:
2158:
2155:
2154:
2150:
2149:
2146:
2145:
2141:
2140:
2137:
2136:
2132:
2131:
2128:
2127:
2123:
2122:
2119:
2118:
2114:
2113:
2110:
2109:
2105:
2104:
2101:
2100:
2096:
2095:
2092:
2091:
2087:
2086:
2083:
2082:
2075:
2072:
2071:
2068:
2067:
2060:
2055:
2053:
2050:
2049:
2046:
2045:
2041:
2040:
2037:
2036:
2027:
2024:
2023:
2020:
2019:
2010:
2005:
2003:
2001:Tsintaosaurini
1995:
1993:
1990:
1989:
1986:
1985:
1976:
1971:
1969:
1966:
1965:
1962:
1961:
1954:
1949:
1947:
1945:Lambeosaurinae
1941:
1940:
1937:
1936:
1932:
1931:
1928:
1927:
1918:
1915:
1914:
1911:
1910:
1901:
1896:
1894:
1889:
1887:
1884:
1883:
1880:
1879:
1870:
1865:
1863:
1857:
1856:
1853:
1852:
1843:
1838:
1836:
1833:
1832:
1829:
1828:
1819:
1814:
1812:
1809:
1808:
1805:
1804:
1795:
1790:
1788:
1785:
1784:
1781:
1780:
1771:
1766:
1764:
1761:
1760:
1757:
1756:
1747:
1742:
1740:
1737:
1736:
1733:
1732:
1728:
1727:
1724:
1723:
1714:
1711:
1710:
1707:
1706:
1697:
1692:
1690:
1685:
1683:
1680:
1679:
1676:
1675:
1666:
1661:
1659:
1656:
1655:
1652:
1651:
1642:
1637:
1635:
1630:
1628:
1624:
1623:
1620:
1619:
1615:
1614:
1611:
1610:
1606:
1605:
1602:
1601:
1597:
1596:
1593:
1592:
1588:
1587:
1584:
1583:
1579:
1578:
1575:
1574:
1570:
1569:
1566:
1565:
1561:
1560:
1557:
1556:
1552:
1551:
1548:
1547:
1544:Lambeosaurinae
1540:
1537:
1536:
1533:
1532:
1525:
1520:
1518:
1512:
1511:
1508:
1507:
1498:
1493:
1491:
1488:
1487:
1484:
1483:
1474:
1471:
1470:
1467:
1466:
1457:
1452:
1450:
1447:
1446:
1443:
1442:
1433:
1430:
1429:
1426:
1425:
1416:
1411:
1409:
1406:
1405:
1402:
1401:
1392:
1387:
1385:
1382:
1381:
1378:
1377:
1368:
1363:
1361:
1358:
1357:
1354:
1353:
1344:
1339:
1337:
1334:
1333:
1330:
1329:
1325:
1324:
1321:
1320:
1311:
1308:
1307:
1304:
1303:
1294:
1289:
1287:
1282:
1280:
1275:
1212:
1197:Lambeosaurinae
1121:
1120:Classification
1118:
1063:
1060:
1038:T. spinorhinus
1026:T. spinorhinus
985:synonymization
925:
918:
851:Arén Formation
820:
732:
729:
720:
712:
704:
696:
684:
676:
668:
660:
546:, including a
495:
492:
490:
487:
479:Tsintaosaurini
461:Lambeosaurinae
414:
355:
354:
353:
352:
344:
323:
322:
316:
315:
312:
300:
299:
293:
292:
275:
271:
270:
268:Tsintaosaurini
262:
258:
257:
255:Lambeosaurinae
249:
245:
244:
236:
232:
231:
223:
216:
215:
213:Neornithischia
207:
200:
199:
191:
184:
183:
178:
171:
170:
165:
161:
160:
155:
151:
150:
145:
141:
140:
127:
126:
122:
121:
113:
112:
104:
103:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
41:
28:
15:
13:
10:
9:
6:
4:
3:
2:
6246:
6235:
6232:
6230:
6227:
6225:
6222:
6220:
6217:
6215:
6212:
6210:
6207:
6205:
6202:
6201:
6199:
6182:
6177:
6173:
6169:
6164:
6160:
6156:
6151:
6147:
6143:
6138:
6134:
6130:
6125:
6121:
6116:
6110:
6106:
6105:
6103:
6101:
6100:Pararhabdodon
6097:
6093:
6088:
6068:
6067:
6063:
6061:
6059:
6058:Edmontosaurus
6055:
6053:
6051:
6050:Edmontosaurus
6047:
6045:
6042:
6039:
6035:
6032:
6030:
6027:
6025:
6022:
6020:
6017:
6015:
6012:
6010:
6009:
6005:
6004:
6002:
5998:
5992:
5991:
5987:
5985:
5984:
5980:
5978:
5977:
5973:
5971:
5970:
5969:Syngonosaurus
5966:
5964:
5963:
5959:
5957:
5956:
5952:
5950:
5949:
5945:
5943:
5942:
5938:
5936:
5935:
5931:
5929:
5928:
5924:
5922:
5921:
5917:
5915:
5914:
5913:Koutalisaurus
5910:
5908:
5907:
5903:
5901:
5900:
5896:
5894:
5893:
5892:Eucercosaurus
5889:
5887:
5886:
5882:
5879:
5878:
5874:
5872:
5871:
5867:
5865:
5864:
5863:Arstanosaurus
5860:
5859:
5857:
5855:
5854:
5849:
5843:
5842:
5838:
5837:
5835:
5833:
5829:
5823:
5822:
5818:
5816:
5815:
5811:
5809:
5808:
5804:
5802:
5801:
5797:
5795:
5794:
5790:
5788:
5787:
5783:
5781:
5780:
5779:Orcauichnites
5776:
5774:
5773:
5769:
5767:
5766:
5762:
5760:
5759:
5755:
5753:
5752:
5748:
5746:
5745:
5741:
5739:
5738:
5734:
5732:
5731:
5727:
5725:
5724:
5720:
5718:
5717:
5716:Amblydactylus
5713:
5712:
5710:
5708:
5704:
5700:
5693:
5689:
5680:
5676:
5674:
5669:
5663:
5656:
5630:
5629:
5625:
5623:
5622:
5618:
5616:
5615:
5614:Kundurosaurus
5611:
5609:
5608:
5607:Kerberosaurus
5604:
5602:
5601:
5597:
5595:
5594:
5593:Edmontosaurus
5590:
5589:
5587:
5585:
5581:
5575:
5574:
5570:
5568:
5567:
5563:
5561:
5560:
5556:
5555:
5553:
5551:
5547:
5541:
5540:
5536:
5534:
5533:
5529:
5527:
5526:
5522:
5520:
5519:
5515:
5513:
5512:
5508:
5505:
5504:
5503:Anasazisaurus
5500:
5499:
5497:
5495:
5491:
5485:
5484:
5483:Secernosaurus
5480:
5478:
5477:
5476:Kelumapusaura
5473:
5471:
5470:
5466:
5464:
5463:
5459:
5458:
5456:
5454:
5450:
5443:
5442:
5438:
5436:
5435:
5431:
5429:
5428:
5424:
5422:
5421:
5417:
5415:
5414:
5410:
5408:
5407:
5403:
5402:
5400:
5398:
5394:
5388:
5387:
5383:
5382:
5379:
5376:
5374:
5373:Saurolophinae
5370:
5360:
5359:
5355:
5353:
5352:
5348:
5346:
5345:
5341:
5339:
5338:
5334:
5332:
5331:
5327:
5325:
5324:
5323:Hypacrosaurus
5320:
5318:
5317:
5316:Corythosaurus
5313:
5311:
5310:
5306:
5305:
5303:
5301:
5300:Lambeosaurini
5297:
5291:
5290:
5286:
5284:
5283:
5279:
5277:
5276:
5275:Charonosaurus
5272:
5270:
5269:
5265:
5264:
5262:
5260:
5256:
5250:
5249:
5248:Tsintaosaurus
5245:
5243:
5242:
5241:Pararhabdodon
5238:
5236:
5235:
5234:Nipponosaurus
5231:
5229:
5228:
5224:
5221:
5220:
5216:
5214:
5213:
5209:
5207:
5206:
5205:Jaxartosaurus
5202:
5200:
5199:
5195:
5193:
5192:
5188:
5186:
5185:
5181:
5179:
5178:
5174:
5172:
5171:
5167:
5165:
5164:
5160:
5158:
5157:
5156:Adynomosaurus
5153:
5152:
5149:
5146:
5144:
5140:
5134:
5133:
5129:
5127:
5126:
5122:
5120:
5119:
5118:Lapampasaurus
5115:
5113:
5112:
5108:
5106:
5105:
5101:
5099:
5098:
5094:
5093:
5090:
5087:
5084:
5083:Hadrosauridae
5079:
5073:
5072:
5068:
5066:
5065:
5064:Zhanghenglong
5061:
5059:
5058:
5054:
5052:
5051:
5047:
5045:
5044:
5040:
5038:
5037:
5036:Telmatosaurus
5033:
5031:
5030:
5026:
5024:
5023:
5019:
5017:
5016:
5012:
5010:
5009:
5005:
5003:
5002:
4998:
4996:
4995:
4991:
4989:
4988:
4984:
4982:
4981:
4980:Portellsaurus
4977:
4975:
4974:
4970:
4968:
4967:
4963:
4961:
4960:
4956:
4954:
4953:
4949:
4947:
4946:
4942:
4940:
4939:
4935:
4932:
4931:
4927:
4925:
4924:
4920:
4918:
4917:
4913:
4911:
4910:
4906:
4904:
4903:
4899:
4897:
4896:
4892:
4890:
4889:
4885:
4883:
4882:
4878:
4876:
4875:
4871:
4869:
4868:
4864:
4862:
4861:
4857:
4855:
4854:
4850:
4848:
4847:
4843:
4841:
4840:
4836:
4833:
4832:
4828:
4826:
4825:
4821:
4819:
4818:
4814:
4813:
4810:
4807:
4804:
4799:
4795:
4791:
4786:
4782:
4773:
4769:
4766:
4759:
4726:
4725:
4724:
4723:
4721:
4719:
4715:
4708:
4707:
4703:
4701:
4700:
4696:
4694:
4693:
4689:
4686:
4685:
4681:
4680:
4678:
4676:
4672:
4666:
4665:
4661:
4659:
4658:
4654:
4652:
4651:
4647:
4645:
4644:
4643:Jinzhousaurus
4640:
4638:
4637:
4636:Hypselospinus
4633:
4631:
4630:
4626:
4624:
4623:
4619:
4618:
4615:
4612:
4610:
4606:
4600:
4599:
4595:
4593:
4592:
4591:Theiophytalia
4588:
4586:
4585:
4581:
4579:
4578:
4574:
4572:
4571:
4567:
4565:
4564:
4560:
4558:
4557:
4553:
4551:
4550:
4546:
4544:
4543:
4539:
4537:
4536:
4532:
4530:
4529:
4525:
4523:
4522:
4521:Lanzhousaurus
4518:
4516:
4515:
4511:
4509:
4508:
4504:
4502:
4501:
4497:
4495:
4494:
4490:
4487:
4486:
4482:
4480:
4479:
4475:
4473:
4472:
4468:
4466:
4465:
4461:
4459:
4458:
4454:
4452:
4451:
4447:
4446:
4443:
4440:
4437:
4432:
4422:
4421:
4417:
4414:
4413:
4409:
4406:
4405:
4401:
4399:
4398:
4394:
4392:
4391:
4390:Elrhazosaurus
4387:
4385:
4384:
4380:
4378:
4377:
4373:
4372:
4370:
4368:
4364:
4358:
4357:
4353:
4351:
4350:
4346:
4344:
4343:
4339:
4338:
4335:
4331:
4328:
4325:
4320:
4306:
4305:
4301:
4299:
4298:
4294:
4292:
4291:
4287:
4285:
4284:
4280:
4278:
4277:
4273:
4271:
4270:
4266:
4265:
4263:
4261:
4257:
4251:
4250:
4249:Tenontosaurus
4246:
4244:
4243:
4239:
4236:
4235:
4234:Convolosaurus
4231:
4230:
4228:
4225:
4221:
4214:
4213:
4209:
4206:
4205:
4201:
4198:
4197:
4196:Ampelognathus
4193:
4192:
4189:
4186:
4184:
4180:
4173:
4172:
4168:
4166:
4165:
4161:
4159:
4158:
4154:
4152:
4151:
4147:
4145:
4144:
4140:
4138:
4137:
4133:
4131:
4130:
4126:
4123:
4122:
4118:
4116:
4115:
4111:
4109:
4108:
4107:Mahuidacursor
4104:
4102:
4101:
4097:
4095:
4094:
4090:
4087:
4086:
4082:
4080:
4079:
4075:
4073:
4072:
4068:
4066:
4065:
4061:
4058:
4057:
4053:
4051:
4050:
4046:
4044:
4043:
4039:
4037:
4036:
4032:
4030:
4029:
4025:
4024:
4022:
4020:
4016:
4009:
4008:
4004:
4002:
4001:
4000:Hypsilophodon
3997:
3994:
3993:
3989:
3988:
3986:
3983:
3979:
3972:
3971:
3967:
3965:
3964:
3963:Callovosaurus
3960:
3958:
3957:
3956:Burianosaurus
3953:
3952:
3949:
3945:
3942:
3939:
3934:
3930:
3926:
3921:
3917:
3892:
3891:
3890:
3889:
3887:
3885:
3881:
3873:
3869:
3868:
3867:
3866:
3863:
3860:
3858:
3854:
3846:
3842:
3841:
3840:
3839:
3836:
3833:
3831:
3827:
3823:
3815:
3811:
3808:
3806:
3802:
3799:
3797:
3793:
3790:
3788:
3784:
3781:
3779:
3775:
3773:
3769:
3768:
3763:
3759:
3755:
3748:
3743:
3741:
3736:
3734:
3729:
3728:
3725:
3718:
3708:
3706:
3696:
3692:
3681:
3677:
3672:
3667:
3663:
3659:
3655:
3651:
3647:
3640:
3637:
3632:
3628:
3624:
3620:
3616:
3612:
3605:
3602:
3596:
3591:
3587:
3583:
3579:
3572:
3569:
3565:
3561:
3555:
3552:
3547:
3543:
3538:
3533:
3529:
3525:
3521:
3517:
3513:
3506:
3504:
3500:
3491:
3489:
3487:
3483:
3477:
3474:
3469:
3465:
3461:
3457:
3453:
3449:
3441:
3438:
3433:
3429:
3424:
3419:
3414:
3409:
3405:
3401:
3397:
3393:
3389:
3382:
3380:
3378:
3374:
3369:
3365:
3361:
3357:
3353:
3349:
3345:
3341:
3337:
3330:
3328:
3326:
3324:
3320:
3314:
3311:
3306:
3302:
3298:
3294:
3290:
3286:
3282:
3275:
3273:
3271:
3269:
3265:
3260:
3256:
3251:
3246:
3241:
3236:
3232:
3228:
3225:(7): e69835.
3224:
3220:
3213:
3206:
3204:
3202:
3200:
3198:
3196:
3194:
3192:
3190:
3188:
3186:
3182:
3175:
3172:
3167:
3159:
3157:
3155:
3151:
3147:
3141:
3139:
3135:
3131:
3127:
3121:
3119:
3115:
3111:
3105:
3103:
3101:
3097:
3093:
3087:
3085:
3083:
3081:
3079:
3077:
3075:
3073:
3069:
3065:
3061:
3057:
3051:
3049:
3047:
3045:
3043:
3041:
3039:
3037:
3035:
3033:
3031:
3029:
3025:
3020:
3016:
3011:
3006:
3002:
2998:
2994:
2990:
2986:
2979:
2976:
2964:
2960:
2956:
2952:
2948:
2944:
2940:
2933:
2930:
2924:
2914:
2911:
2905:
2903:
2901:
2899:
2897:
2893:
2887:
2884:
2877:
2875:
2873:
2871:
2867:
2860:
2855:
2851:
2848:
2847:
2843:
2836:
2835:
2827:
2826:
2818:
2817:
2809:
2808:
2800:
2799:
2791:
2790:
2782:
2781:
2773:
2772:
2769:
2768:
2767:Lambeosaurini
2762:
2761:
2758:
2757:
2754:
2753:
2747:
2746:
2740:
2739:
2736:
2735:
2727:
2726:
2718:
2717:
2709:
2708:
2700:
2699:
2691:
2690:
2682:
2681:
2678:
2677:
2676:
2669:
2668:
2665:
2664:
2661:
2660:
2659:
2652:
2651:
2645:
2644:
2641:
2640:
2637:
2636:
2630:
2629:
2623:
2622:
2619:
2618:
2615:
2614:
2613:
2612:Adynomosaurus
2606:
2605:
2599:
2598:
2595:
2594:
2586:
2585:
2582:
2581:
2580:
2573:
2572:
2569:
2568:
2565:
2564:
2558:
2557:
2551:
2550:
2544:
2543:
2540:
2539:
2536:
2535:
2534:
2533:Koutalisaurus
2527:
2526:
2520:
2519:
2516:
2515:
2507:
2506:
2503:
2502:
2501:
2500:Pararhabdodon
2494:
2493:
2490:
2489:
2486:
2485:
2484:
2477:
2476:
2470:
2469:
2463:
2460:
2459:
2453:
2452:
2449:
2448:
2445:
2444:
2443:
2442:Tsintaosaurus
2436:
2435:
2429:
2428:
2425:
2424:
2421:
2420:
2419:
2418:Nipponosaurus
2412:
2411:
2405:
2404:
2401:
2400:
2397:
2396:
2395:
2394:Jaxartosaurus
2388:
2387:
2381:
2380:
2377:
2376:
2373:
2372:
2371:
2364:
2363:
2357:
2354:
2353:
2350:
2349:
2346:
2345:
2344:Saurolophinae
2339:
2338:
2332:
2331:Hadrosauridae
2329:
2328:
2324:
2322:
2321:Pararhabdodon
2318:
2317:Tsintaosaurus
2314:
2310:
2306:
2302:
2301:Pararhabdodon
2298:
2297:
2289:
2288:Pararhabdodon
2285:
2284:
2278:
2270:
2266:
2264:
2261:
2257:
2253:
2252:Pararhabdodon
2248:
2244:
2243:
2238:
2237:Pararhabdodon
2231:
2230:
2229:Tsintaosaurus
2224:
2211:
2210:
2202:
2201:
2193:
2192:
2184:
2183:
2175:
2174:
2166:
2165:
2157:
2156:
2148:
2147:
2139:
2138:
2130:
2129:
2121:
2120:
2112:
2111:
2103:
2102:
2094:
2093:
2085:
2084:
2081:
2080:
2079:Lambeosaurini
2074:
2073:
2070:
2069:
2066:
2065:
2059:
2058:
2052:
2051:
2048:
2047:
2039:
2038:
2035:
2034:
2033:
2026:
2025:
2022:
2021:
2018:
2017:
2016:
2009:
2008:
2002:
1999:
1998:
1992:
1991:
1988:
1987:
1984:
1983:
1982:
1975:
1974:
1968:
1967:
1964:
1963:
1960:
1959:
1953:
1952:
1946:
1943:
1942:
1939:
1938:
1930:
1929:
1926:
1925:
1924:
1917:
1916:
1913:
1912:
1909:
1908:
1907:
1900:
1899:
1893:
1892:
1886:
1885:
1882:
1881:
1878:
1877:
1876:
1869:
1868:
1862:
1861:Hadrosauridae
1859:
1858:
1855:
1854:
1851:
1850:
1849:
1842:
1841:
1835:
1834:
1831:
1830:
1827:
1826:
1825:
1818:
1817:
1811:
1810:
1807:
1806:
1803:
1802:
1801:
1794:
1793:
1787:
1786:
1783:
1782:
1779:
1778:
1777:
1770:
1769:
1763:
1762:
1759:
1758:
1755:
1754:
1753:
1746:
1745:
1739:
1738:
1735:
1734:
1726:
1725:
1722:
1721:
1720:
1713:
1712:
1709:
1708:
1705:
1704:
1703:
1696:
1695:
1689:
1688:
1682:
1681:
1678:
1677:
1674:
1673:
1672:
1665:
1664:
1658:
1657:
1654:
1653:
1650:
1649:
1648:
1641:
1640:
1634:
1633:
1622:
1621:
1613:
1612:
1604:
1603:
1595:
1594:
1586:
1585:
1577:
1576:
1568:
1567:
1559:
1558:
1550:
1549:
1546:
1545:
1539:
1538:
1535:
1534:
1531:
1530:
1529:Hadrosaurinae
1524:
1523:
1517:
1516:Hadrosauridae
1514:
1513:
1510:
1509:
1506:
1505:
1504:
1497:
1496:
1490:
1489:
1486:
1485:
1482:
1481:
1480:
1473:
1472:
1469:
1468:
1465:
1464:
1463:
1456:
1455:
1449:
1448:
1445:
1444:
1441:
1440:
1439:
1432:
1431:
1428:
1427:
1424:
1423:
1422:
1415:
1414:
1408:
1407:
1404:
1403:
1400:
1399:
1398:
1391:
1390:
1384:
1383:
1380:
1379:
1376:
1375:
1374:
1367:
1366:
1360:
1359:
1356:
1355:
1352:
1351:
1350:
1343:
1342:
1336:
1335:
1332:
1331:
1323:
1322:
1319:
1318:
1317:
1310:
1309:
1306:
1305:
1302:
1301:
1300:
1293:
1292:
1286:
1285:
1279:
1278:
1270:
1268:
1264:
1259:
1255:
1254:Pararhabdodon
1251:
1250:Hadrosaurinae
1247:
1243:
1242:hadrosauroids
1239:
1235:
1234:Pararhabdodon
1231:
1230:
1225:
1220:
1217:
1210:
1209:neural spines
1206:
1205:Koutalisaurus
1202:
1198:
1194:
1193:
1185:
1184:Pararhabdodon
1180:
1176:
1174:
1170:
1166:
1162:
1158:
1157:
1152:
1148:
1144:
1143:Pararhabdodon
1137:
1136:Pararhabdodon
1133:
1132:
1126:
1119:
1117:
1115:
1114:
1113:Adynomosaurus
1109:
1105:
1104:Pararhabdodon
1101:
1100:Tsintaosaurus
1097:
1096:
1095:Corythosaurus
1091:
1088:
1084:
1080:
1079:Pararhabdodon
1073:
1068:
1061:
1059:
1057:
1056:Pararhabdodon
1053:
1052:Tsintaosaurus
1048:
1047:Tsintaosaurus
1042:
1039:
1035:
1031:
1030:Pararhabdodon
1027:
1023:
1022:Koutalisaurus
1019:
1018:
1013:
1012:Koutalisaurus
1009:
1008:Pararhabdodon
1002:
1001:
995:
991:
990:
986:
982:
978:
974:
973:
968:
963:
962:stratigraphic
959:
955:
954:Pararhabdodon
951:
947:
939:
935:
930:
923:
922:Koutalisaurus
919:
917:
915:
914:Pararhabdodon
911:
910:Pararhabdodon
907:
906:
905:Koutalisaurus
901:
896:
894:
890:
889:Pararhabdodon
886:
885:Pararhabdodon
882:
878:
874:
870:
866:
862:
858:
857:
852:
848:
847:
837:
833:
831:
825:
818:
814:
813:Maastrichtian
810:
809:Pararhabdodon
806:
802:
798:
794:
793:Pararhabdodon
790:
789:Pararhabdodon
786:
782:
777:
775:
774:
769:
765:
761:
760:Pararhabdodon
757:
753:
749:
742:
741:Pararhabdodon
737:
730:
728:
725:
717:
711:, a complete
709:
701:
694:
689:
681:
673:
665:
658:
654:
650:
645:
643:
642:
638:
634:
633:
632:Telmatosaurus
628:
624:
620:
619:lambeosaurine
616:
612:
611:
606:
602:
598:
595:, a complete
594:
590:
585:
583:
579:
575:
571:
567:
566:
561:
557:
553:
549:
545:
541:
537:
533:
529:
525:
521:
517:
513:
512:Pararhabdodon
505:
500:
493:
488:
486:
484:
480:
476:
472:
471:
470:Tsintaosaurus
466:
462:
458:
457:iguanodontian
454:
449:
447:
442:
438:
437:
432:
431:
426:
425:
419:
413:, as well as
412:
408:
407:
402:
398:
394:
391:
387:
386:Maastrichtian
383:
380:
377:
376:tsintaosaurin
373:
369:
368:
363:
362:
361:Pararhabdodon
348:
345:
342:
338:
332:
331:
327:
326:
324:
321:
317:
310:
309:
301:
298:
294:
290:
284:
283:
282:Pararhabdodon
276:
273:
272:
269:
263:
260:
259:
256:
250:
247:
246:
243:
242:Hadrosauridae
237:
234:
233:
230:
224:
221:
218:
217:
214:
208:
205:
202:
201:
198:
192:
189:
186:
185:
182:
179:
176:
173:
172:
169:
166:
163:
162:
159:
156:
153:
152:
149:
146:
143:
142:
137:
132:
128:
123:
119:
114:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
39:
32:
26:
25:Pararhabdodon
22:
19:
6099:
6064:
6057:
6049:
6037:
6006:
5988:
5981:
5974:
5967:
5960:
5953:
5946:
5939:
5932:
5925:
5918:
5911:
5904:
5897:
5890:
5883:
5877:Claorhynchus
5875:
5868:
5861:
5853:Nomina dubia
5851:
5841:Guegoolithus
5839:
5819:
5812:
5807:Staurichnium
5805:
5800:Sousaichnium
5798:
5791:
5784:
5777:
5770:
5763:
5756:
5751:Dakotasaurus
5749:
5744:Caririchnium
5742:
5735:
5728:
5721:
5714:
5658:
5626:
5619:
5612:
5605:
5598:
5591:
5571:
5564:
5557:
5550:Saurolophini
5537:
5530:
5523:
5516:
5509:
5501:
5494:Kritosaurini
5481:
5474:
5469:Huallasaurus
5467:
5460:
5441:Wulagasaurus
5439:
5432:
5425:
5418:
5411:
5404:
5384:
5356:
5349:
5342:
5335:
5330:Lambeosaurus
5328:
5321:
5314:
5307:
5287:
5280:
5273:
5266:
5246:
5240:
5239:
5232:
5225:
5217:
5210:
5203:
5196:
5189:
5182:
5175:
5168:
5161:
5154:
5132:Yamatosaurus
5130:
5123:
5116:
5109:
5102:
5097:Aquilarhinus
5095:
5069:
5062:
5055:
5048:
5043:Tethyshadros
5041:
5034:
5027:
5020:
5013:
5006:
4999:
4992:
4985:
4978:
4973:Plesiohadros
4971:
4964:
4957:
4950:
4945:Lophorhothon
4943:
4936:
4928:
4921:
4914:
4907:
4900:
4893:
4886:
4879:
4872:
4865:
4858:
4851:
4844:
4837:
4831:Batyrosaurus
4829:
4824:Bactrosaurus
4822:
4815:
4761:
4704:
4697:
4690:
4682:
4664:Ouranosaurus
4662:
4655:
4648:
4641:
4634:
4629:Brighstoneus
4627:
4620:
4596:
4589:
4582:
4575:
4568:
4561:
4556:Osmakasaurus
4554:
4549:Oblitosaurus
4547:
4540:
4533:
4526:
4519:
4512:
4505:
4498:
4491:
4483:
4476:
4471:Camptosaurus
4469:
4462:
4455:
4448:
4418:
4412:Kangnasaurus
4410:
4402:
4395:
4388:
4381:
4374:
4367:Dryosauridae
4354:
4347:
4340:
4302:
4295:
4288:
4281:
4274:
4269:Matheronodon
4267:
4247:
4240:
4232:
4210:
4202:
4194:
4169:
4162:
4155:
4148:
4143:Sektensaurus
4141:
4136:Qantassaurus
4134:
4127:
4119:
4112:
4105:
4098:
4091:
4085:Kangnasaurus
4083:
4076:
4069:
4062:
4054:
4049:Diluvicursor
4047:
4040:
4033:
4026:
4007:Vectidromeus
4005:
3998:
3990:
3968:
3961:
3954:
3872:Ornithischia
3857:Ornithischia
3809:
3800:
3796:Ornithischia
3791:
3782:
3653:
3649:
3639:
3614:
3610:
3604:
3585:
3581:
3571:
3554:
3511:
3502:
3498:
3476:
3451:
3447:
3440:
3395:
3391:
3343:
3339:
3335:
3313:
3288:
3284:
3222:
3218:
3174:
3165:
3148:16:411-423.
3145:
3129:
3125:
3109:
3091:
3063:
3059:
3055:
2992:
2988:
2978:
2966:. Retrieved
2946:
2942:
2932:
2913:
2886:
2765:
2750:
2673:
2672:
2656:
2655:
2634:
2633:
2610:
2609:
2577:
2576:
2561:
2531:
2530:
2499:
2498:
2497:
2481:
2480:
2462:Arenysaurini
2440:
2439:
2416:
2415:
2392:
2391:
2368:
2367:
2342:
2320:
2316:
2309:Arenysaurini
2305:monophyletic
2300:
2294:
2292:
2287:
2281:
2255:
2251:
2246:
2240:
2236:
2234:
2227:
2077:
2062:
2031:
2030:
2029:
2013:
2012:
1979:
1978:
1958:Aralosaurini
1956:
1921:
1920:
1904:
1903:
1873:
1872:
1846:
1845:
1822:
1821:
1798:
1797:
1774:
1773:
1750:
1749:
1717:
1716:
1700:
1699:
1669:
1668:
1645:
1644:
1542:
1527:
1502:
1501:
1500:
1477:
1476:
1460:
1459:
1436:
1435:
1419:
1418:
1395:
1394:
1371:
1370:
1347:
1346:
1314:
1313:
1297:
1296:
1266:
1258:synapomorphy
1253:
1237:
1233:
1227:
1224:P. isonensis
1223:
1221:
1204:
1190:
1188:
1183:
1161:rhabdodontid
1154:
1147:Iguanodontia
1142:
1141:
1135:
1129:
1111:
1103:
1099:
1093:
1078:
1077:
1055:
1051:
1046:
1043:
1037:
1034:P. isonensis
1033:
1029:
1025:
1021:
1015:
1011:
1007:
1005:
998:
989:P. isonensis
988:
977:autapomorphy
970:
967:P. isonensis
966:
958:P. isonensis
957:
953:
943:
938:P. isonensis
937:
933:
921:
913:
909:
903:
897:
892:
888:
884:
880:
854:
844:
842:
830:P. isonensis
829:
817:P. isonensis
816:
808:
797:P. isonensis
796:
792:
788:
781:P. isonensis
780:
778:
771:
763:
759:
745:
740:
703:, a partial
655:vertebra; a
646:
639:
630:
622:
608:
604:
586:
581:
576:-like basal
569:
563:
539:
511:
509:
483:Arenysaurini
468:
465:hadrosauroid
453:rhabdodontid
450:
434:
428:
422:
405:
404:
400:
365:
360:
359:
358:
346:
336:
328:
307:
306:
297:Type species
281:
280:
219:
203:
197:Ornithischia
187:
174:
24:
18:
6066:Willinakaqe
6038:nomina nuda
5955:Siluosaurus
5920:Loncosaurus
5758:Dinehichnus
5600:Kamuysaurus
5573:Saurolophus
5525:Kritosaurus
5518:Gryposaurus
5351:Sahaliyania
5337:Magnapaulia
5309:Amurosaurus
5289:Tlatolophus
5268:Adelolophus
5212:Kazaklambia
5191:Blasisaurus
5184:Arenysaurus
5177:Aralosaurus
5111:Hadrosaurus
5104:Eotrachodon
5057:Yunganglong
5022:Sirindhorna
4994:Protohadros
4930:Koshisaurus
4923:Jintasaurus
4692:Comptonatus
4542:Napaisaurus
4528:Lurdusaurus
4500:Fukuisaurus
4478:Cedrorestes
4420:Valdosaurus
4283:Pareisactus
4114:Morrosaurus
4078:Isasicursor
4042:Chakisaurus
3938:Ornithopoda
3925:Ornithopoda
3884:Ornithopoda
3754:Ornithopoda
3454:: 447–457.
2658:Blasisaurus
2483:Arenysaurus
2370:Aralosaurus
2283:Arenysaurus
2280:Fossils of
2247:P. isonenis
1087:quadrupedal
1062:Description
893:Arenysaurus
754:, southern
723:haemal arch
615:hadrosaurid
536:Tremp Group
436:Arenysaurus
430:Blasisaurus
393:Tremp Group
384:, from the
379:hadrosaurid
248:Subfamily:
229:Ornithopoda
6198:Categories
6008:Arkharavia
5983:Tichosteus
5976:Thespesius
5948:Pteropelyx
5941:Orthomerus
5821:Wintonopus
5406:Acristavus
5386:Barsboldia
5344:Olorotitan
5219:Latirhinus
5071:Zuoyunlong
4938:Levnesovia
4888:Gobihadros
4853:Datonglong
4846:Claosaurus
4817:Altirhinus
4727:see below↓
4657:Morelladon
4535:Magnamanus
4507:Hippodraco
4376:Dryosaurus
4349:Hesperonyx
4324:Dryomorpha
4164:Trinisaura
4157:Tietasaura
4150:Talenkauen
4028:Anabisetia
3893:see below↓
3787:Dinosauria
3656:: 104678.
3537:2152/41080
3291:: 104677.
2995:: 104286.
2949:: 104677.
2856:References
1165:iguanodont
981:edentulous
877:synonymous
748:Cretaceous
651:, and one
641:Orthomerus
578:iguanodont
574:rhabdodont
181:Dinosauria
5990:Trachodon
5906:Glishades
5885:Diclonius
5723:Anomoepus
5707:Ichnotaxa
5420:Maiasaura
5358:Velafrons
4867:Equijubus
4839:Choyrodon
4699:Iguanodon
4563:Planicoxa
4493:Dakotadon
4464:Calvarius
4290:Rhabdodon
4276:Mochlodon
4019:Elasmaria
3770:Kingdom:
3705:Dinosaurs
3680:228807024
3368:134582286
3346:: 19–37.
3305:225110719
3019:208195457
2963:225110719
2925:Citations
2226:Skull of
1173:hadrosaur
1156:Rhabdodon
1131:Rhabdodon
1090:herbivore
627:subfamily
605:isonensis
565:Rhabdodon
544:Catalonia
530:) of the
411:vertebrae
401:Rhabdodon
367:Rhabdodon
154:Kingdom:
148:Eukaryota
125:Maxillae
6115:Q3493376
6109:Wikidata
5870:Cionodon
5539:Rhinorex
5427:Ornatops
5227:Minqaria
5198:Canardia
5125:Malefica
5050:Xuwulong
5015:Siamodon
4916:Jeyawati
4902:Gonkoken
4860:Eolambia
4684:Barilium
4485:Cumnoria
4450:Barilium
4356:Owenodon
4342:Draconyx
4304:Zalmoxes
4204:Fostoria
4056:Fostoria
3814:Cerapoda
3778:Chordata
3776:Phylum:
3772:Animalia
3631:85705882
3546:85081036
3432:30379888
3392:PLOS ONE
3259:23922815
3219:PLOS ONE
2844:See also
2579:Canardia
1238:Eolambia
1134:, which
1072:holotype
719:, and a
693:paratype
635:and the
589:maxillae
582:isonense
548:cervical
516:Pyrenees
504:Pyrenees
417:maxillae
382:dinosaur
320:Synonyms
235:Family:
168:Chordata
164:Phylum:
158:Animalia
144:Domain:
36:66
6168:4947941
6155:1407345
6142:4823291
6129:4532047
5163:Ajnabia
4598:Uteodon
3691:Portals
3658:Bibcode
3516:Bibcode
3456:Bibcode
3423:6209292
3400:Bibcode
3348:Bibcode
3250:3724916
3227:Bibcode
2675:Ajnabia
2296:Ajnabia
1169:maxilla
1083:bipedal
950:dentary
873:frontal
687:humerus
671:scapula
663:ischium
637:dubious
601:ischium
560:scapula
556:humerus
552:dorsals
534:in the
370:) is a
274:Genus:
261:Tribe:
105:↓
5832:Ootaxa
5029:Tanius
4622:Bolong
3678:
3629:
3562:
3544:
3430:
3420:
3366:
3303:
3257:
3247:
3017:
2961:
2256:et al.
1267:et al.
1211:, and
1192:et al.
975:. One
853:, and
805:strata
787:which
764:et al.
756:France
715:fibula
683:and a
657:sacrum
653:caudal
649:dorsal
623:et al.
610:et al.
597:sacrum
591:, two
540:et al.
524:Lleida
337:et al.
6181:57426
6150:IRMNG
4874:Fylax
4404:Iyuku
3810:Clade
3801:Clade
3792:Clade
3783:Clade
3717:Spain
3676:S2CID
3627:S2CID
3542:S2CID
3508:(PDF)
3364:S2CID
3301:S2CID
3215:(PDF)
3015:S2CID
2968:3 May
2959:S2CID
2861:Notes
2313:clade
2260:tribe
1246:basal
1244:more
1201:jugal
1151:taxon
1036:with
869:jugal
785:tribe
768:taxon
707:tibia
699:femur
675:, an
562:, as
528:Spain
520:Isona
518:near
475:China
397:Spain
388:-age
372:genus
220:Clade
204:Clade
188:Clade
175:Clade
6137:GBIF
4570:Proa
4242:Iani
3870:see
3843:see
3560:ISBN
3501:and
3428:PMID
3255:PMID
2970:2021
1236:and
1098:and
752:Aude
679:ulna
554:, a
433:and
341:2010
289:1993
44:PreꞒ
6124:EoL
3666:doi
3654:120
3619:doi
3590:doi
3586:159
3532:hdl
3524:doi
3464:doi
3418:PMC
3408:doi
3356:doi
3293:doi
3289:118
3245:PMC
3235:doi
3005:hdl
2997:doi
2993:107
2951:doi
2947:118
1175:.
750:of
395:of
374:of
6200::
6178::
6165::
6152::
6139::
6126::
6111::
3812::
3803::
3794::
3785::
3674:.
3664:.
3652:.
3648:.
3625:.
3615:21
3613:.
3584:.
3580:.
3540:.
3530:.
3522:.
3510:.
3485:^
3462:.
3452:53
3450:.
3426:.
3416:.
3406:.
3396:13
3394:.
3390:.
3376:^
3362:.
3354:.
3344:96
3342:.
3322:^
3299:.
3287:.
3283:.
3267:^
3253:.
3243:.
3233:.
3221:.
3217:.
3184:^
3153:^
3137:^
3117:^
3099:^
3071:^
3027:^
3013:.
3003:.
2991:.
2987:.
2957:.
2945:.
2941:.
2895:^
2869:^
2315:.
832:.
776:.
770:,
727:.
644:.
526:,
522:,
448:.
339:,
333:?
222::
206::
190::
177::
94:Pg
38:Ma
33:,
6040:)
5880:?
5506:?
5444:?
5222:?
4933:?
4834:?
4709:?
4687:?
4488:?
4415:?
4407:?
4237:?
4226:?
4215:?
4207:?
4199:?
4174:?
4124:?
4088:?
4059:?
4010:?
3995:?
3984:?
3973:?
3746:e
3739:t
3732:v
3693::
3682:.
3668::
3660::
3633:.
3621::
3598:.
3592::
3566:.
3548:.
3534::
3526::
3518::
3497:"
3470:.
3466::
3458::
3434:.
3410::
3402::
3370:.
3358::
3350::
3307:.
3295::
3261:.
3237::
3229::
3223:8
3168:.
3021:.
3007::
2999::
2972:.
2953::
1085:-
924:"
304:†
278:†
265:†
252:†
239:†
226:†
210:†
194:†
99:N
89:K
84:J
79:T
74:P
69:C
64:D
59:S
54:O
49:Ꞓ
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