3188:
3183:
1262:
133:
485:
115:
1709:
3211:
1949:
Agnolín, Federico L.; Álvarez
Herrera, Gerardo; Rolando, Mauro Aranciaga; Motta, Matías; Rozadilla, Sebastián; Verdiquio, Lucía; D'Angelo, Julia S.; Moyano-Paz, Damián; Varela, Augusto N.; Sterli, Juliana; Bogan, Sergio; Miner, Santiago; Moreno Rodríguez, Ana; Muñoz, Gonzalo; Isasi, Marcelo P.;
1297:, Soto Acuña, Vargas & Kaluza further elaborated on this, explaining that the discovery of additional fossil material allowed for a rescoring of its characters in their phylogenetic matrix. Based on these updates, they recovered
2028:
Aranciaga
Rolando, Mauro; Cerroni, Mauricio A.; Garcia Marsà, Jordi A.; Agnolín, Federico l.; Motta, Matías J.; Rozadilla, Sebastián; Brisson Eglí, Federico; Novas, Fernando E. (2020-10-14).
3337:
1795:
Frauenfelder, Timothy G.; Bell, Phil R.; Brougham, Tom; Bevitt, Joseph J.; Bicknell, Russell D. C.; Kear, Benjamin P.; Wroe, Stephen; Campione, Nicolás E. (2022-03-28).
3286:
414:
period. The first remains were described in 1996 and often appeared in the literature as the "Argentinian ankylosaur". The fossil material consists of various
512:, within a clade of entirely North American nodosaurids from the middle of the Cretaceous period, in contrast to previous analyses that recovered it in the
1843:
Riguetti, Facundo; Pereda-Suberbiola, Xabier; Ponce, Denis; Salgado, Leonardo; Apesteguía, Sebastián; Rozadilla, Sebastián; Arbour, Victoria (2022-12-31).
3273:
2030:"A new medium-sized abelisaurid (Theropoda, Dinosauria) from the late cretaceous (Maastrichtian) Allen Formation of Northern Patagonia, Argentina"
3187:
3332:
2142:
2087:"A new hadrosaurid (Dinosauria: Ornithischia) from the Late Cretaceous of northern Patagonia and the radiation of South American hadrosaurids"
516:. In either case, these results would suggest that nodosaurids were part of a migration event of North American fauna into South America. The
2085:
Rozadilla, Sebastián; Brissón-Egli, Federico; Lisandro Agnolín, Federico; Aranciaga-Rolando, Alexis Mauro; Novas, Fernando Emilio (2022).
438:
3204:
265:
3352:
3342:
3182:
1845:"A new small-bodied ankylosaurian dinosaur from the Upper Cretaceous of North Patagonia (Río Negro Province, Argentina)"
1797:"New Ankylosaurian Cranial Remains From the Lower Cretaceous (Upper Albian) Toolebuc Formation of Queensland, Australia"
1738:
Rivera-Sylva, H.E.; Frey, E.; Stinnesbeck, W.; Carbot-Chanona, G.; Sanchez-Uribe, I.E.; Guzmán-Gutiérrez, J.R. (2018).
132:
2135:
1274:
452:
as a new genus and species of nodosaurine dinosaurs by
Facundo Riguetti, Xabier Pereda-Suberbiola, Denis Ponce,
3347:
2233:
1360:
457:
3231:
1952:"Fossil vertebrates from the Cerro Fortaleza Formation (Upper Cretaceous), Santa Cruz Province, Argentina"
464:", is derived from "Patago", a reference to the discovery of the taxon in Argentinian Patagonia, and the
3299:
2260:
2128:
288:
399:
3327:
3214:
2029:
1965:
1856:
1751:
122:
Skeletal reconstruction showing known osteoderms (top) and other skeletal remains (bottom) in white
2620:
2517:
1956:
598:
449:
2356:
2106:
2067:
1951:
1880:
1777:
127:
3304:
1740:"Paleodiversity of Late Cretaceous Ankylosauria from Mexico and their phylogenetic significance"
3291:
3113:
2377:
2059:
1931:
1898:
Madzia, D.; Arbour, V.M.; Boyd, C.A.; Farke, A.A.; Cruzado-Caballero, P.; Evans, D.C. (2021).
1872:
1818:
1769:
1593:
1261:
2578:
2525:
2363:
2098:
2049:
2041:
2008:
1973:
1921:
1911:
1864:
1808:
1759:
1599:
484:
453:
423:
419:
3043:
3007:
2947:
2913:
2906:
2799:
2740:
2612:
2509:
2425:
2385:
2218:
1713:
1615:
1566:
1452:
1322:
1282:
1020:
622:
411:
391:
365:
351:
343:
331:
241:
30:
1273:
A study published in 2024 by Agnolín and colleagues reviewed vertebrate fossils from the
1969:
1860:
1755:
3106:
3092:
3078:
2986:
2959:
2855:
2836:
2751:
2697:
2562:
2532:
2502:
2465:
2441:
1926:
1899:
1687:
1663:
1658:
1648:
1637:
1501:
1336:
948:
911:
513:
465:
374:
1708:
3321:
3069:
2979:
2972:
2924:
2868:
2815:
2781:
2682:
2628:
2603:
2585:
2488:
2434:
2407:
2399:
2110:
2086:
2071:
1884:
1781:
1702:
1627:
1621:
1587:
1412:
1293:
1287:
1138:
852:
680:
663:
501:
407:
339:
72:
40:
114:
3254:
3099:
3085:
3014:
2898:
2822:
2767:
2718:
2690:
2675:
2662:
2635:
2554:
2539:
2495:
2475:
2342:
2324:
2302:
2272:
2245:
2193:
2151:
1669:
1653:
1643:
1632:
1610:
1516:
1312:
1121:
934:
704:
646:
565:
527:
509:
395:
324:
225:
193:
2102:
1977:
1868:
2045:
1277:
and similar
Patagonian formations. They mentioned new findings that suggest that
3134:
3051:
3000:
2940:
2891:
2733:
2643:
2592:
2392:
2370:
2202:
1844:
1698:
1575:
1570:
1469:
361:
209:
47:
2013:
1992:
3127:
3120:
3058:
2933:
2884:
2876:
2829:
2774:
2760:
2726:
2704:
2570:
2414:
2349:
1813:
1796:
1764:
1739:
1693:
1605:
1003:
972:
821:
804:
752:
548:
415:
92:
57:
2063:
1876:
1822:
1773:
3141:
3021:
2808:
2788:
2650:
2547:
2449:
1681:
1675:
1581:
1395:
1077:
869:
728:
517:
403:
347:
335:
144:
97:
36:
3265:
1993:"A new look at the first dinosaur discovered in Antarctica: reappraisal of
1935:
3248:
3148:
2184:
2175:
1302:
489:
434:
327:
177:
164:
87:
82:
67:
62:
52:
3278:
2054:
1916:
3036:
3028:
372:
is a very small ankylosaur, comparable in size to the dwarf nodosaurid
320:
102:
77:
476:, referring to the large crests on its cervical osteoderms and femur.
2169:
427:
154:
3225:
2993:
1991:
Soto Acuña, Sergio; Vargas, Alexander O.; Kaluza, Jonatan (2024).
1707:
1569:
of Río Negro
Province, Argentina. Many other dinosaurs, including
1260:
483:
473:
1305:
ankylosaurs. Their results are displayed in the cladogram below:
360:, known from a partial skeleton. While originally described as a
3155:
3229:
2162:
2124:
2120:
460:, Sebastián Rozadilla, and Victoria Arbour. The generic name, "
1352:
520:
below displays the results of their phylogenetic analyses.
1900:"The phylogenetic nomenclature of ornithischian dinosaurs"
1301:
1291:
rather than nodosaurids. In their 2024 redescription of
441:-SM-78, is represented by a cervical half-ring element.
1705:
mammals have also been described from the formation.
468:
word "pelta", meaning "shield". The specific name, "
3238:
3068:
2958:
2923:
2854:
2798:
2750:
2661:
2602:
2474:
2463:
2424:
2322:
2271:
2244:
2217:
402:, Argentina. This locality is dated to the upper
1667:), have been named from the formation. Birds (
390:fossil material was found in sediments of the
2136:
1838:
1836:
1834:
1832:
8:
3338:Late Cretaceous dinosaurs of South America
3226:
2964:
2860:
2667:
2480:
2471:
2334:
2330:
2319:
2250:
2223:
2214:
2159:
2143:
2129:
2121:
113:
20:
2053:
2012:
1925:
1915:
1812:
1763:
364:, later discoveries provided support for
2034:Journal of South American Earth Sciences
1269:(as a parankylosaur) compared to a human
1730:
444:New remains described in 2022 allowed
378:, about 2 m (6.6 ft) long.
7:
319:(meaning "Patagonian shield") is an
2091:Journal of Systematic Palaeontology
1849:Journal of Systematic Palaeontology
1494:
1444:
1437:
1328:
1318:
1308:
1113:
1093:
1069:
1045:
995:
988:
964:
940:
903:
844:
837:
796:
744:
720:
696:
638:
614:
590:
540:
533:
523:
504:, Riguetti et al. (2022) recovered
1565:is known from the Late Cretaceous
14:
3210:
3209:
3186:
3181:
1997:(Ankylosauria: Parankylosauria)"
394:(Salitral Moreno locality) near
131:
1744:Swiss Journal of Palaeontology
1285:, similar to ankylosaurs like
1:
3333:Fossil taxa described in 2022
2103:10.1080/14772019.2021.2020917
1978:10.1016/j.cretres.2023.105735
1869:10.1080/14772019.2022.2137441
16:Genus of nodosaurid dinosaurs
2046:10.1016/j.jsames.2020.102915
1950:Novas, Fernando E. (2024).
3369:
2014:10.12429/j.advps.2023.0036
1801:Frontiers in Earth Science
368:affinities for the taxon.
3199:
3179:
2967:
2863:
2670:
2483:
2337:
2333:
2318:
2299:
2253:
2226:
2213:
2158:
2001:Advances in Polar Science
1814:10.3389/feart.2022.803505
1765:10.1007/s13358-018-0153-1
1514:
1499:
1492:
1466:
1449:
1442:
1435:
1409:
1392:
1375:
1350:
1333:
1326:
1316:
1281:may actually represent a
1275:Cerro Fortaleza Formation
1135:
1118:
1111:
1098:
1091:
1074:
1067:
1050:
1043:
1017:
1000:
993:
986:
969:
962:
945:
938:
908:
901:
866:
849:
842:
835:
818:
801:
794:
749:
742:
725:
718:
701:
694:
677:
660:
643:
636:
619:
612:
595:
588:
562:
545:
538:
531:
294:
287:
128:Scientific classification
126:
121:
112:
23:
350:. The genus contains a
1995:Antarctopelta oliveroi
1721:
1712:Dinosaur fauna of the
1685:), rhynchocephalians (
1270:
497:
472:", means "crested" in
3300:Paleobiology Database
1720:in red, center right)
1711:
1264:
502:phylogenetic analyses
487:
307:Riguetti et al., 2022
3353:Cretaceous Argentina
3343:Fossils of Argentina
1100:"Paw Paw scuteling"
458:Sebastián Apesteguía
446:Patagopelta cristata
382:Discovery and naming
301:Patagopelta cristata
1970:2024CrRes.15405735A
1957:Cretaceous Research
1917:10.7717/peerj.12362
1861:2022JSPal..2037441R
1756:2018SwJP..137...83R
508:as a member of the
1722:
1271:
498:
400:Río Negro Province
3315:
3314:
3232:Taxon identifiers
3223:
3222:
3195:
3194:
3177:
3176:
3173:
3172:
3169:
3168:
3165:
3164:
3114:Nodocephalosaurus
2850:
2849:
2846:
2845:
2459:
2458:
2378:Serendipaceratops
2314:
2313:
2310:
2309:
2295:
2294:
2291:
2290:
2097:(17): 1207–1235.
1594:Panamericansaurus
1554:
1553:
1545:
1544:
1536:
1535:
1527:
1526:
1481:
1480:
1424:
1423:
1258:
1257:
1249:
1248:
1240:
1239:
1231:
1230:
1222:
1221:
1213:
1212:
1204:
1203:
1195:
1194:
1186:
1185:
1177:
1176:
1168:
1167:
1159:
1158:
1150:
1149:
1032:
1031:
924:
923:
890:
889:
881:
880:
783:
782:
774:
773:
765:
764:
577:
576:
312:
311:
268:
264:Riguetti et al.,
3360:
3308:
3307:
3295:
3294:
3282:
3281:
3269:
3268:
3259:
3258:
3257:
3227:
3213:
3212:
3190:
3185:
2965:
2861:
2668:
2579:Propanoplosaurus
2526:Glyptodontopelta
2481:
2472:
2364:Priodontognathus
2335:
2331:
2320:
2251:
2224:
2215:
2208:
2207:
2160:
2145:
2138:
2131:
2122:
2115:
2114:
2082:
2076:
2075:
2057:
2025:
2019:
2018:
2016:
1988:
1982:
1981:
1946:
1940:
1939:
1929:
1919:
1895:
1889:
1888:
1840:
1827:
1826:
1816:
1792:
1786:
1785:
1767:
1735:
1691:), plesiosaurs (
1600:Pellegrinisaurus
1495:
1445:
1438:
1364:
1329:
1319:
1309:
1283:parankylosaurian
1114:
1094:
1070:
1046:
996:
989:
965:
941:
904:
845:
838:
797:
745:
721:
697:
639:
615:
591:
541:
534:
524:
454:Leonardo Salgado
366:parankylosaurian
303:
299:
280:P. cristata
263:
256:
245:
240:
224:
208:
192:
136:
135:
117:
107:
44:
29:Temporal range:
21:
3368:
3367:
3363:
3362:
3361:
3359:
3358:
3357:
3348:Allen Formation
3318:
3317:
3316:
3311:
3303:
3298:
3290:
3285:
3277:
3272:
3264:
3262:
3253:
3252:
3247:
3234:
3224:
3219:
3191:
3161:
3064:
3044:Tianzhenosaurus
3008:Minotaurasaurus
2954:
2948:Zhongyuansaurus
2919:
2914:Zhejiangosaurus
2907:Liaoningosaurus
2842:
2800:Struthiosaurini
2794:
2746:
2741:Tatankacephalus
2657:
2613:Gargoyleosaurus
2598:
2510:Gargoyleosaurus
2468:
2455:
2426:Parankylosauria
2420:
2386:Sinankylosaurus
2327:
2306:
2287:
2267:
2240:
2234:Avemetatarsalia
2219:Avemetatarsalia
2209:
2165:
2164:
2154:
2149:
2119:
2118:
2084:
2083:
2079:
2027:
2026:
2022:
1990:
1989:
1985:
1948:
1947:
1943:
1897:
1896:
1892:
1842:
1841:
1830:
1794:
1793:
1789:
1737:
1736:
1732:
1727:
1714:Allen Formation
1679:), pterosaurs (
1616:Bonapartesaurus
1567:Allen Formation
1560:
1555:
1546:
1537:
1528:
1482:
1453:Liaoningosaurus
1425:
1358:
1323:Parankylosauria
1259:
1250:
1241:
1232:
1223:
1214:
1205:
1196:
1187:
1178:
1169:
1160:
1151:
1033:
1021:Tatankacephalus
925:
891:
882:
784:
775:
766:
623:Gargoyleosaurus
578:
496:as a nodosaurid
482:
426:vertebrae, and
412:Late Cretaceous
392:Allen Formation
384:
344:Allen Formation
332:Late Cretaceous
308:
305:
297:
296:
283:
262:
254:
243:
242:Parankylosauria
238:
222:
206:
190:
130:
108:
106:
105:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
34:
33:
31:Late Cretaceous
27:
17:
12:
11:
5:
3366:
3364:
3356:
3355:
3350:
3345:
3340:
3335:
3330:
3320:
3319:
3313:
3312:
3310:
3309:
3296:
3283:
3270:
3260:
3244:
3242:
3236:
3235:
3230:
3221:
3220:
3218:
3217:
3207:
3200:
3197:
3196:
3193:
3192:
3180:
3178:
3175:
3174:
3171:
3170:
3167:
3166:
3163:
3162:
3160:
3159:
3152:
3145:
3138:
3131:
3124:
3117:
3110:
3107:Euoplocephalus
3103:
3096:
3093:Anodontosaurus
3089:
3082:
3079:Akainacephalus
3074:
3072:
3066:
3065:
3063:
3062:
3055:
3048:
3040:
3033:
3025:
3018:
3011:
3004:
2997:
2990:
2987:Crichtonsaurus
2983:
2976:
2968:
2962:
2960:Ankylosaurinae
2956:
2955:
2953:
2952:
2944:
2937:
2929:
2927:
2921:
2920:
2918:
2917:
2910:
2903:
2895:
2888:
2881:
2873:
2864:
2858:
2856:Ankylosauridae
2852:
2851:
2848:
2847:
2844:
2843:
2841:
2840:
2837:Struthiosaurus
2833:
2826:
2819:
2812:
2804:
2802:
2796:
2795:
2793:
2792:
2785:
2778:
2771:
2764:
2756:
2754:
2752:Panoplosaurini
2748:
2747:
2745:
2744:
2737:
2730:
2723:
2715:
2708:
2701:
2698:Niobrarasaurus
2694:
2687:
2679:
2671:
2665:
2659:
2658:
2656:
2655:
2647:
2640:
2632:
2625:
2617:
2608:
2606:
2600:
2599:
2597:
2596:
2589:
2582:
2575:
2567:
2563:Polacanthoides
2559:
2551:
2544:
2536:
2533:Horshamosaurus
2529:
2522:
2514:
2506:
2503:Dongyangopelta
2499:
2492:
2484:
2478:
2469:
2466:Euankylosauria
2464:
2461:
2460:
2457:
2456:
2454:
2453:
2446:
2442:Kunbarrasaurus
2438:
2430:
2428:
2422:
2421:
2419:
2418:
2411:
2404:
2396:
2389:
2382:
2374:
2367:
2360:
2353:
2346:
2338:
2328:
2323:
2316:
2315:
2312:
2311:
2308:
2307:
2300:
2297:
2296:
2293:
2292:
2289:
2288:
2286:
2285:
2284:
2283:
2277:
2275:
2269:
2268:
2266:
2265:
2264:
2263:
2254:
2248:
2242:
2241:
2239:
2238:
2237:
2236:
2227:
2221:
2211:
2210:
2206:
2205:
2196:
2187:
2178:
2172:
2163:
2156:
2155:
2150:
2148:
2147:
2140:
2133:
2125:
2117:
2116:
2077:
2020:
1983:
1941:
1890:
1855:(1): 2137441.
1828:
1787:
1729:
1728:
1726:
1723:
1688:Lamarquesaurus
1664:Bonapartenykus
1659:alvarezsaurids
1649:dromaeosaurids
1559:
1556:
1552:
1551:
1548:
1547:
1543:
1542:
1539:
1538:
1534:
1533:
1530:
1529:
1525:
1524:
1521:
1520:
1513:
1510:
1509:
1506:
1505:
1502:Ankylosauridae
1498:
1493:
1491:
1488:
1487:
1484:
1483:
1479:
1478:
1475:
1474:
1465:
1462:
1461:
1458:
1457:
1448:
1443:
1441:
1436:
1434:
1431:
1430:
1427:
1426:
1422:
1421:
1418:
1417:
1408:
1405:
1404:
1401:
1400:
1391:
1388:
1387:
1384:
1383:
1374:
1371:
1370:
1367:
1366:
1356:Kunbarrasaurus
1349:
1346:
1345:
1342:
1341:
1337:Kunbarrasaurus
1332:
1327:
1325:
1317:
1315:
1307:
1256:
1255:
1252:
1251:
1247:
1246:
1243:
1242:
1238:
1237:
1234:
1233:
1229:
1228:
1225:
1224:
1220:
1219:
1216:
1215:
1211:
1210:
1207:
1206:
1202:
1201:
1198:
1197:
1193:
1192:
1189:
1188:
1184:
1183:
1180:
1179:
1175:
1174:
1171:
1170:
1166:
1165:
1162:
1161:
1157:
1156:
1153:
1152:
1148:
1147:
1144:
1143:
1134:
1131:
1130:
1127:
1126:
1117:
1112:
1110:
1107:
1106:
1103:
1102:
1097:
1092:
1090:
1087:
1086:
1083:
1082:
1073:
1068:
1066:
1063:
1062:
1059:
1058:
1049:
1044:
1042:
1039:
1038:
1035:
1034:
1030:
1029:
1026:
1025:
1016:
1013:
1012:
1009:
1008:
999:
994:
992:
987:
985:
982:
981:
978:
977:
968:
963:
961:
958:
957:
954:
953:
949:Niobrarasaurus
944:
939:
937:
931:
930:
927:
926:
922:
921:
918:
917:
912:Struthiosaurus
907:
902:
900:
897:
896:
893:
892:
888:
887:
884:
883:
879:
878:
875:
874:
865:
862:
861:
858:
857:
848:
843:
841:
836:
834:
831:
830:
827:
826:
817:
814:
813:
810:
809:
800:
795:
793:
790:
789:
786:
785:
781:
780:
777:
776:
772:
771:
768:
767:
763:
762:
759:
758:
748:
743:
741:
738:
737:
734:
733:
724:
719:
717:
714:
713:
710:
709:
700:
695:
693:
690:
689:
686:
685:
676:
673:
672:
669:
668:
659:
656:
655:
652:
651:
642:
637:
635:
632:
631:
628:
627:
618:
613:
611:
608:
607:
604:
603:
594:
589:
587:
584:
583:
580:
579:
575:
574:
571:
570:
561:
558:
557:
554:
553:
544:
539:
537:
532:
530:
522:
514:Panoplosaurini
481:
480:Classification
478:
383:
380:
375:Struthiosaurus
352:single species
310:
309:
306:
292:
291:
285:
284:
276:
274:
270:
269:
252:
248:
247:
236:
229:
228:
220:
213:
212:
204:
197:
196:
188:
181:
180:
175:
168:
167:
162:
158:
157:
152:
148:
147:
142:
138:
137:
124:
123:
119:
118:
110:
109:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
45:
28:
15:
13:
10:
9:
6:
4:
3:
2:
3365:
3354:
3351:
3349:
3346:
3344:
3341:
3339:
3336:
3334:
3331:
3329:
3326:
3325:
3323:
3306:
3301:
3297:
3293:
3288:
3284:
3280:
3275:
3271:
3267:
3261:
3256:
3250:
3246:
3245:
3243:
3241:
3237:
3233:
3228:
3216:
3208:
3206:
3202:
3201:
3198:
3189:
3184:
3158:
3157:
3153:
3151:
3150:
3146:
3144:
3143:
3139:
3137:
3136:
3132:
3130:
3129:
3125:
3123:
3122:
3118:
3116:
3115:
3111:
3109:
3108:
3104:
3102:
3101:
3097:
3095:
3094:
3090:
3088:
3087:
3083:
3081:
3080:
3076:
3075:
3073:
3071:
3070:Ankylosaurini
3067:
3061:
3060:
3056:
3054:
3053:
3049:
3046:
3045:
3041:
3039:
3038:
3034:
3031:
3030:
3026:
3024:
3023:
3019:
3017:
3016:
3012:
3010:
3009:
3005:
3003:
3002:
2998:
2996:
2995:
2991:
2989:
2988:
2984:
2982:
2981:
2980:Crichtonpelta
2977:
2975:
2974:
2973:Bissektipelta
2970:
2969:
2966:
2963:
2961:
2957:
2950:
2949:
2945:
2943:
2942:
2938:
2936:
2935:
2931:
2930:
2928:
2926:
2925:Shamosaurinae
2922:
2916:
2915:
2911:
2909:
2908:
2904:
2901:
2900:
2896:
2894:
2893:
2889:
2887:
2886:
2882:
2879:
2878:
2874:
2871:
2870:
2869:Ahshislepelta
2866:
2865:
2862:
2859:
2857:
2853:
2839:
2838:
2834:
2832:
2831:
2827:
2825:
2824:
2820:
2818:
2817:
2816:Hungarosaurus
2813:
2811:
2810:
2806:
2805:
2803:
2801:
2797:
2791:
2790:
2786:
2784:
2783:
2782:Panoplosaurus
2779:
2777:
2776:
2772:
2770:
2769:
2765:
2763:
2762:
2758:
2757:
2755:
2753:
2749:
2743:
2742:
2738:
2736:
2735:
2731:
2729:
2728:
2724:
2721:
2720:
2716:
2714:
2713:
2709:
2707:
2706:
2702:
2700:
2699:
2695:
2693:
2692:
2688:
2685:
2684:
2683:Ahshislepelta
2680:
2678:
2677:
2673:
2672:
2669:
2666:
2664:
2660:
2653:
2652:
2648:
2646:
2645:
2641:
2638:
2637:
2633:
2631:
2630:
2629:Hoplitosaurus
2626:
2623:
2622:
2618:
2615:
2614:
2610:
2609:
2607:
2605:
2604:Polacanthinae
2601:
2595:
2594:
2590:
2588:
2587:
2586:Rhadinosaurus
2583:
2581:
2580:
2576:
2573:
2572:
2568:
2565:
2564:
2560:
2557:
2556:
2552:
2550:
2549:
2545:
2542:
2541:
2537:
2535:
2534:
2530:
2528:
2527:
2523:
2520:
2519:
2515:
2512:
2511:
2507:
2505:
2504:
2500:
2498:
2497:
2493:
2491:
2490:
2489:Acanthopholis
2486:
2485:
2482:
2479:
2477:
2473:
2470:
2467:
2462:
2452:
2451:
2447:
2444:
2443:
2439:
2437:
2436:
2435:Antarctopelta
2432:
2431:
2429:
2427:
2423:
2417:
2416:
2412:
2410:
2409:
2408:Tianchisaurus
2405:
2402:
2401:
2400:Stegosaurides
2397:
2395:
2394:
2390:
2388:
2387:
2383:
2380:
2379:
2375:
2373:
2372:
2368:
2366:
2365:
2361:
2359:
2358:
2354:
2352:
2351:
2347:
2345:
2344:
2340:
2339:
2336:
2332:
2329:
2326:
2321:
2317:
2305:
2304:
2298:
2281:
2280:
2279:
2278:
2276:
2274:
2270:
2262:
2258:
2257:
2256:
2255:
2252:
2249:
2247:
2243:
2235:
2231:
2230:
2229:
2228:
2225:
2222:
2220:
2216:
2212:
2204:
2200:
2197:
2195:
2191:
2188:
2186:
2182:
2179:
2177:
2173:
2171:
2167:
2166:
2161:
2157:
2153:
2146:
2141:
2139:
2134:
2132:
2127:
2126:
2123:
2112:
2108:
2104:
2100:
2096:
2092:
2088:
2081:
2078:
2073:
2069:
2065:
2061:
2056:
2051:
2047:
2043:
2039:
2035:
2031:
2024:
2021:
2015:
2010:
2007:(1): 78–107.
2006:
2002:
1998:
1996:
1987:
1984:
1979:
1975:
1971:
1967:
1963:
1959:
1958:
1953:
1945:
1942:
1937:
1933:
1928:
1923:
1918:
1913:
1909:
1905:
1901:
1894:
1891:
1886:
1882:
1878:
1874:
1870:
1866:
1862:
1858:
1854:
1850:
1846:
1839:
1837:
1835:
1833:
1829:
1824:
1820:
1815:
1810:
1806:
1802:
1798:
1791:
1788:
1783:
1779:
1775:
1771:
1766:
1761:
1757:
1753:
1749:
1745:
1741:
1734:
1731:
1724:
1719:
1715:
1710:
1706:
1704:
1703:gondwanathere
1700:
1696:
1695:
1690:
1689:
1684:
1683:
1678:
1677:
1672:
1671:
1666:
1665:
1660:
1656:
1655:
1650:
1646:
1645:
1640:
1639:
1634:
1630:
1629:
1628:Lapampasaurus
1624:
1623:
1622:Kelumapusaura
1618:
1617:
1612:
1608:
1607:
1602:
1601:
1596:
1595:
1590:
1589:
1588:Menucocelsior
1584:
1583:
1578:
1577:
1572:
1568:
1564:
1557:
1550:
1549:
1541:
1540:
1532:
1531:
1523:
1522:
1519:
1518:
1512:
1511:
1508:
1507:
1504:
1503:
1497:
1496:
1490:
1489:
1486:
1485:
1477:
1476:
1473:
1472:
1471:
1464:
1463:
1460:
1459:
1456:
1455:
1454:
1447:
1446:
1440:
1439:
1433:
1432:
1429:
1428:
1420:
1419:
1416:
1415:
1414:
1413:Antarctopelta
1407:
1406:
1403:
1402:
1399:
1398:
1397:
1390:
1389:
1386:
1385:
1382:
1381:
1380:
1373:
1372:
1369:
1368:
1365:
1362:
1357:
1354:
1348:
1347:
1344:
1343:
1340:
1339:
1338:
1331:
1330:
1324:
1321:
1320:
1314:
1311:
1310:
1306:
1304:
1300:
1296:
1295:
1294:Antarctopelta
1290:
1289:
1288:Antarctopelta
1284:
1280:
1276:
1268:
1263:
1254:
1253:
1245:
1244:
1236:
1235:
1227:
1226:
1218:
1217:
1209:
1208:
1200:
1199:
1191:
1190:
1182:
1181:
1173:
1172:
1164:
1163:
1155:
1154:
1146:
1145:
1142:
1141:
1140:
1139:Ahshislepelta
1133:
1132:
1129:
1128:
1125:
1124:
1123:
1116:
1115:
1109:
1108:
1105:
1104:
1101:
1096:
1095:
1089:
1088:
1085:
1084:
1081:
1080:
1079:
1072:
1071:
1065:
1064:
1061:
1060:
1057:
1056:
1055:
1048:
1047:
1041:
1040:
1037:
1036:
1028:
1027:
1024:
1023:
1022:
1015:
1014:
1011:
1010:
1007:
1006:
1005:
998:
997:
991:
990:
984:
983:
980:
979:
976:
975:
974:
967:
966:
960:
959:
956:
955:
952:
951:
950:
943:
942:
936:
933:
932:
929:
928:
920:
919:
916:
914:
913:
906:
905:
899:
898:
895:
894:
886:
885:
877:
876:
873:
872:
871:
864:
863:
860:
859:
856:
855:
854:
853:Hungarosaurus
847:
846:
840:
839:
833:
832:
829:
828:
825:
824:
823:
816:
815:
812:
811:
808:
807:
806:
799:
798:
792:
791:
788:
787:
779:
778:
770:
769:
761:
760:
757:
755:
754:
747:
746:
740:
739:
736:
735:
732:
731:
730:
723:
722:
716:
715:
712:
711:
708:
707:
706:
699:
698:
692:
691:
688:
687:
684:
683:
682:
681:Panoplosaurus
675:
674:
671:
670:
667:
666:
665:
664:Hoplitosaurus
658:
657:
654:
653:
650:
649:
648:
641:
640:
634:
633:
630:
629:
626:
625:
624:
617:
616:
610:
609:
606:
605:
602:
601:
600:
593:
592:
586:
585:
582:
581:
573:
572:
569:
568:
567:
560:
559:
556:
555:
552:
551:
550:
543:
542:
536:
535:
529:
526:
525:
521:
519:
515:
511:
507:
503:
495:
491:
486:
479:
477:
475:
471:
467:
463:
459:
455:
451:
447:
442:
440:
436:
433:
429:
425:
421:
417:
413:
409:
408:Maastrichtian
405:
401:
397:
393:
389:
381:
379:
377:
376:
371:
367:
363:
359:
358:
353:
349:
345:
341:
340:Maastrichtian
337:
333:
329:
326:
325:ankylosaurian
322:
318:
317:
304:
302:
293:
290:
289:Binomial name
286:
282:
281:
275:
272:
271:
267:
261:
260:
253:
250:
249:
246:
237:
234:
231:
230:
227:
221:
218:
215:
214:
211:
205:
202:
199:
198:
195:
189:
186:
183:
182:
179:
176:
173:
170:
169:
166:
163:
160:
159:
156:
153:
150:
149:
146:
143:
140:
139:
134:
129:
125:
120:
116:
111:
104:
99:
94:
89:
84:
79:
74:
69:
64:
59:
54:
49:
42:
41:Maastrichtian
38:
32:
26:
22:
19:
3239:
3154:
3147:
3140:
3133:
3126:
3119:
3112:
3105:
3100:Dyoplosaurus
3098:
3091:
3086:Ankylosaurus
3084:
3077:
3057:
3050:
3042:
3035:
3027:
3020:
3015:Pinacosaurus
3013:
3006:
2999:
2992:
2985:
2978:
2971:
2946:
2939:
2932:
2912:
2905:
2899:Hylaeosaurus
2897:
2890:
2883:
2875:
2867:
2835:
2828:
2823:Pawpawsaurus
2821:
2814:
2807:
2787:
2780:
2773:
2768:Denversaurus
2766:
2759:
2739:
2732:
2725:
2719:Peloroplites
2717:
2711:
2710:
2703:
2696:
2691:Borealopelta
2689:
2681:
2676:Acantholipan
2674:
2663:Nodosaurinae
2649:
2642:
2636:Peloroplites
2634:
2627:
2619:
2611:
2591:
2584:
2577:
2569:
2561:
2555:Mymoorapelta
2553:
2546:
2540:Hylaeosaurus
2538:
2531:
2524:
2516:
2508:
2501:
2496:Anoplosaurus
2494:
2487:
2476:Nodosauridae
2448:
2440:
2433:
2413:
2406:
2398:
2391:
2384:
2376:
2369:
2362:
2355:
2348:
2343:Cryptosaurus
2341:
2325:Ankylosauria
2303:Ankylosauria
2301:
2273:Ankylosauria
2261:Ornithischia
2246:Ornithischia
2198:
2194:Ornithischia
2189:
2180:
2152:Ankylosauria
2094:
2090:
2080:
2055:11336/150468
2037:
2033:
2023:
2004:
2000:
1994:
1986:
1961:
1955:
1944:
1907:
1903:
1893:
1852:
1848:
1804:
1800:
1790:
1750:(1): 83–93.
1747:
1743:
1733:
1717:
1692:
1686:
1680:
1674:
1670:Lamarqueavis
1668:
1662:
1654:Austroraptor
1652:
1644:Quilmesaurus
1642:
1636:
1633:abelisaurids
1626:
1620:
1614:
1611:hadrosaurids
1604:
1598:
1592:
1586:
1580:
1574:
1562:
1561:
1558:Paleoecology
1517:Nodosauridae
1515:
1500:
1468:
1467:
1451:
1450:
1411:
1410:
1394:
1393:
1378:
1377:
1376:
1355:
1351:
1335:
1334:
1313:Ankylosauria
1298:
1292:
1286:
1278:
1272:
1266:
1137:
1136:
1122:Borealopelta
1120:
1119:
1099:
1076:
1075:
1053:
1052:
1051:
1019:
1018:
1002:
1001:
971:
970:
947:
946:
935:Nodosaurinae
910:
909:
868:
867:
851:
850:
820:
819:
803:
802:
751:
750:
727:
726:
705:Mymoorapelta
703:
702:
679:
678:
662:
661:
647:Peloroplites
645:
644:
621:
620:
597:
596:
566:Hylaeosaurus
564:
563:
547:
546:
528:Nodosauridae
510:Nodosaurinae
505:
499:
493:
488:Speculative
469:
461:
445:
443:
431:
410:ages of the
396:General Roca
387:
385:
373:
369:
356:
355:
315:
314:
313:
300:
295:
279:
278:
258:
257:
232:
226:Ankylosauria
216:
200:
194:Ornithischia
184:
171:
24:
18:
3328:Nodosaurids
3240:Patagopelta
3135:Scolosaurus
3052:Tsagantegia
3001:Jinyunpelta
2941:Shamosaurus
2892:Chuanqilong
2734:Silvisaurus
2712:Patagopelta
2644:Polacanthus
2593:Sauroplites
2393:Spicomellus
2371:Sarcolestes
2203:Thyreophora
1718:Patagopelta
1699:dryolestoid
1576:Aeolosaurus
1571:titanosaurs
1563:Patagopelta
1470:Chuanqilong
1379:Patagopelta
1299:Patagopelta
1279:Patagopelta
1267:Patagopelta
1054:Patagopelta
506:Patagopelta
494:Patagopelta
490:restoration
462:Patagopelta
432:Patagopelta
418:, a tooth,
388:Patagopelta
370:Patagopelta
362:nodosaurine
357:P. cristata
316:Patagopelta
259:Patagopelta
210:Thyreophora
25:Patagopelta
3322:Categories
3255:Q115547839
3203:See also:
3128:Platypelta
3121:Oohkotokia
3059:Zaraapelta
2934:Gobisaurus
2885:Cedarpelta
2877:Aletopelta
2830:Stegopelta
2775:Edmontonia
2761:Animantarx
2727:Sauropelta
2705:Nodosaurus
2571:Priconodon
2415:Vectipelta
2350:Dracopelta
2282:see below↓
2185:Dinosauria
2040:: 102915.
1964:. 105735.
1910:: e12362.
1725:References
1694:Kawanectes
1606:Rocasaurus
1004:Sauropelta
973:Nodosaurus
822:Animantarx
805:Stegopelta
753:Edmontonia
549:Cedarpelta
437:specimen,
416:osteoderms
178:Dinosauria
3142:Talarurus
3022:Saichania
2809:Europelta
2789:Texasetes
2651:Taohelong
2548:Invictarx
2450:Stegouros
2168:Kingdom:
2111:247122005
2072:225123133
2064:0895-9811
1885:254212751
1877:1477-2019
1823:2296-6463
1782:134924657
1774:1664-2376
1682:Aerotitan
1676:Limenavis
1582:Bonatitan
1396:Stegouros
1303:Gondwanan
1078:Texasetes
870:Europelta
729:Taohelong
518:cladogram
500:In their
450:described
406:to lower
404:Campanian
348:Argentina
336:Campanian
330:from the
323:genus of
273:Species:
244: (?)
151:Kingdom:
145:Eukaryota
37:Campanian
3292:11925108
3279:12382882
3263:BioLib:
3249:Wikidata
3215:Category
3205:Timeline
3149:Ziapelta
2621:Gastonia
2518:Gastonia
2176:Chordata
2174:Phylum:
2170:Animalia
1936:34966571
1265:Size of
599:Gastonia
470:cristata
435:holotype
328:dinosaur
165:Chordata
161:Phylum:
155:Animalia
141:Domain:
3266:1963186
3037:Tarchia
3029:Shanxia
1966:Bibcode
1927:8667728
1857:Bibcode
1752:Bibcode
1697:), and
1657:), and
1363:P40536)
338:–lower
334:(upper
321:extinct
298:†
277:†
251:Genus:
3305:466647
2109:
2070:
2062:
1934:
1924:
1883:
1875:
1821:
1780:
1772:
1638:Niebla
1625:, and
1603:, and
448:to be
430:. The
428:femora
424:caudal
420:dorsal
3287:IRMNG
2994:Datai
2357:Minmi
2199:Clade
2190:Clade
2181:Clade
2107:S2CID
2068:S2CID
1904:PeerJ
1881:S2CID
1778:S2CID
915:spp.
756:spp.
474:Latin
466:Greek
233:Clade
217:Clade
201:Clade
185:Clade
172:Clade
3274:GBIF
3156:Zuul
2259:see
2232:see
2060:ISSN
1932:PMID
1873:ISSN
1819:ISSN
1770:ISSN
1701:and
1673:and
1641:and
1361:SAMA
439:MPCA
422:and
386:The
266:2022
48:PreꞒ
2099:doi
2050:hdl
2042:doi
2038:105
2009:doi
1974:doi
1962:154
1922:PMC
1912:doi
1865:doi
1809:doi
1760:doi
1748:137
1647:),
1631:),
1609:),
1353:cf.
492:of
346:of
3324::
3302::
3289::
3276::
3251::
2201::
2192::
2183::
2105:.
2095:19
2093:.
2089:.
2066:.
2058:.
2048:.
2036:.
2032:.
2005:35
2003:.
1999:.
1972:.
1960:.
1954:.
1930:.
1920:.
1906:.
1902:.
1879:.
1871:.
1863:.
1853:20
1851:.
1847:.
1831:^
1817:.
1807:.
1805:10
1803:.
1799:.
1776:.
1768:.
1758:.
1746:.
1742:.
1619:,
1597:,
1591:,
1585:,
1579:,
456:,
398:,
354:,
342:)
235::
219::
203::
187::
174::
98:Pg
3047:?
3032:?
2951:?
2902:?
2880:?
2872:?
2722:?
2686:?
2654:?
2639:?
2624:?
2616:?
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2558:?
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2521:?
2513:?
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2403:?
2381:?
2144:e
2137:t
2130:v
2113:.
2101::
2074:.
2052::
2044::
2017:.
2011::
1980:.
1976::
1968::
1938:.
1914::
1908:9
1887:.
1867::
1859::
1825:.
1811::
1784:.
1762::
1754::
1716:(
1661:(
1651:(
1635:(
1613:(
1573:(
1359:(
255:†
239:†
223:†
207:†
191:†
103:N
93:K
88:J
83:T
78:P
73:C
68:D
63:S
58:O
53:Ꞓ
43:)
39:–
35:(
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