49:
73:
363:
353:
in late
Miocene South America along with its relationship to the New World clade may instead indicate the common ancestor of the New World pelicans reached South America via trans-Atlantic dispersal from Africa after diverging from the Old World clade.
711:"First continental vertebrates from the marine Paraná Formation (Late Miocene, Mesopotamia, Argentina): Chronology, biogeography, and paleoenvironments"
772:
345:
before spreading to other continents. Whilst pelicans are traditionally believed to have colonized the
Americas in an eastward direction from
498:
Noriega, Jorge I.; Cenizo, Marcos; Brandoni, Diego; PĂ©rez, Leandro M.; Tineo, David E.; Diederle, Juan M.; Bona, Paula (2023-05-09).
664:"A Sperm Whale (Cetacea: Physeteroidea) from the Paraná Formation (Late Miocene) of Entre RĂs, Argentina. Environment and Taphonomy"
777:
410:
in the middle and late
Miocene. A diverse fauna of marine life is known from the formation, including rays and sharks such as
500:"A new pelican (Aves: Pelecanidae) from the Upper Miocene of Argentina: new clues about the origin of the New World lineages"
188:
319:
pelicans, suggesting it belongs in the New World pelican clade and is closely related to these two species, whilst the
323:
is a sister taxon to the clade including the aforementioned three species. Because the oldest known fossil pelican,
782:
72:
225:. It is the first fossil pelican described from Argentina and the southernmost to be reported from South America.
545:
El Adli, Joseph J.; Wilson
Mantilla, Jeffrey A.; Antar, Mohammed Sameh M.; Gingerich, Philip D. (2021-01-02).
641:"Megascyliorhinus trelewensis (Neoselachii) in the Middle-Upper Miocene of Parana, Central Eastern Argentina"
412:
767:
320:
277:
249:
218:
639:
Cione, Alberto Luis; Mennucci, Jorge; Pé, Leandro; rez; Marí Barla, a Julia (2008-01-01).
168:
439:
cetaceans. Continental vertebrates have also been found in the Paraná Formation, including the rodents
441:
430:
722:
558:
546:
391:
371:
447:
383:
293:
289:
269:
253:
214:
762:
738:
691:
582:
527:
273:
67:
48:
683:
621:
574:
547:"The earliest recorded fossil pelican, recovered from the late Eocene of Wadi Al-Hitan, Egypt"
519:
424:
285:
730:
675:
613:
566:
511:
418:
338:
316:
281:
726:
562:
466:
452:
124:
662:
PĂ©rez, Leandro M.; Cione, Alberto L.; Cozzuol, Mario; Varela, Augusto N. (June 2011).
756:
742:
710:
695:
663:
640:
586:
531:
499:
456:
367:
312:
297:
210:
30:
734:
570:
515:
601:
679:
436:
325:
134:
362:
617:
248:
specimen), which was found in the Cerro La
Matanza locality in Victoria City,
687:
625:
578:
523:
709:
Candela, Adriana M.; Bonini, Ricardo A.; Noriega, Jorge I. (November 2012).
222:
84:
38:
34:
461:
403:
334:
245:
104:
56:
606:
The Annals and
Magazine of Natural History; Zoology, Botany, and Geology
602:"I.—On some fish-remains from the Parana formation, Argentine Republic"
399:
395:
375:
346:
206:
144:
407:
342:
237:
203:
94:
361:
341:) are also restricted to Africa, pelicans likely first emerged in
330:
114:
187:
Noriega, Cenizo, Brandoni, PĂ©rez, Tineo, Diederie & Bona,
645:
Instituto
Superior de Correlacion Geologica, Serie Miscelanea
394:, a shallow inland sea which covered parts of Argentina,
311:
exhibits derived characteristics shared with the extant
390:
was found represents littoral to marine facies of the
256:, the stratigraphic provenance of the specimen.
252:, Argentina. The specific name refers to the
8:
47:
20:
333:, and the closest relatives of pelicans (
479:
304:Classification and evolutionary history
268:falls within the size range of extant
7:
493:
491:
489:
487:
485:
483:
551:Journal of Vertebrate Paleontology
504:Journal of Vertebrate Paleontology
14:
370:, illustrating the extent of the
284:pelicans. It is smaller than the
236:is known from a nearly complete
71:
600:Woodward, Arthur Smith (1900).
1:
773:Fossil taxa described in 2023
735:10.1016/j.geobios.2012.05.003
571:10.1080/02724634.2021.1903910
516:10.1080/02724634.2023.2202702
799:
618:10.1080/00222930008678334
174:
167:
68:Scientific classification
66:
55:
46:
23:
778:Birds described in 2023
680:10.5710/AMGH.v48i3(425)
413:Cosmopolitodus hastalis
16:Extinct species of bird
379:
321:American white pelican
264:The known specimen of
460:and an indeterminate
382:The sediments of the
365:
288:but larger than the
234:Pelecanus paranensis
229:Discovery and naming
199:Pelecanus paranensis
181:Pelecanus paranensis
61:Pelecanus paranensis
25:Pelecanus paranensis
727:2012Geobi..45..515C
563:2021JVPal..41E3910E
250:Entre RĂos Province
219:Entre RĂos Province
380:
349:, the presence of
242:CICYTTP-PV-A-3-277
160:P. paranensis
783:Neogene Argentina
425:Hemipristis serra
286:Dalmatian pelican
195:
194:
790:
747:
746:
706:
700:
699:
659:
653:
652:
636:
630:
629:
597:
591:
590:
542:
536:
535:
495:
442:Plesiacarechimys
431:Megascyliorhinus
419:Otodus megalodon
384:Paraná Formation
374:during the late
358:Paleoenvironment
339:Balaenicipitidae
254:Paraná Formation
215:Paraná Formation
183:
179:
76:
75:
51:
29:Temporal range:
21:
798:
797:
793:
792:
791:
789:
788:
787:
753:
752:
751:
750:
708:
707:
703:
661:
660:
656:
638:
637:
633:
599:
598:
594:
557:(1): e1903910.
544:
543:
539:
510:(5): e2202702.
497:
496:
481:
476:
360:
306:
262:
231:
191:
185:
177:
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163:
70:
42:
27:
17:
12:
11:
5:
796:
794:
786:
785:
780:
775:
770:
765:
755:
754:
749:
748:
721:(6): 515–526.
701:
674:(4): 648–654.
654:
631:
592:
537:
478:
477:
475:
472:
467:Phoenicopterus
392:Paranaense Sea
372:Paranaense Sea
359:
356:
329:, is found in
305:
302:
278:American white
261:
258:
230:
227:
193:
192:
186:
172:
171:
165:
164:
156:
154:
150:
149:
142:
138:
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132:
128:
127:
125:Pelecaniformes
122:
118:
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112:
108:
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102:
98:
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92:
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87:
82:
78:
77:
64:
63:
53:
52:
44:
43:
28:
15:
13:
10:
9:
6:
4:
3:
2:
795:
784:
781:
779:
776:
774:
771:
769:
768:Miocene birds
766:
764:
761:
760:
758:
744:
740:
736:
732:
728:
724:
720:
716:
712:
705:
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697:
693:
689:
685:
681:
677:
673:
669:
665:
658:
655:
650:
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642:
635:
632:
627:
623:
619:
615:
611:
607:
603:
596:
593:
588:
584:
580:
576:
572:
568:
564:
560:
556:
552:
548:
541:
538:
533:
529:
525:
521:
517:
513:
509:
505:
501:
494:
492:
490:
488:
486:
484:
480:
473:
471:
469:
468:
464:of the genus
463:
459:
458:
457:Scirrotherium
454:
450:
449:
448:Cardiatherium
444:
443:
438:
434:
432:
427:
426:
421:
420:
415:
414:
409:
406:and southern
405:
401:
397:
393:
389:
388:P. paranensis
385:
377:
373:
369:
368:South America
364:
357:
355:
352:
351:P. paranensis
348:
344:
340:
336:
332:
328:
327:
322:
318:
314:
310:
309:P. paranensis
303:
301:
299:
295:
291:
287:
283:
279:
275:
271:
267:
266:P. paranensis
259:
257:
255:
251:
247:
243:
239:
235:
228:
226:
224:
220:
216:
212:
211:Upper Miocene
208:
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201:
200:
190:
184:
182:
173:
170:
169:Binomial name
166:
162:
161:
155:
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100:
99:
96:
93:
90:
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83:
80:
79:
74:
69:
65:
62:
58:
54:
50:
45:
40:
36:
32:
26:
22:
19:
718:
714:
704:
671:
667:
657:
648:
644:
634:
609:
605:
595:
554:
550:
540:
507:
503:
465:
455:
453:pampatheriid
446:
440:
433:trelewensis,
429:
423:
417:
411:
387:
381:
350:
324:
308:
307:
265:
263:
241:
233:
232:
198:
197:
196:
180:
175:
159:
158:
145:
60:
31:Late Miocene
24:
18:
668:Ameghiniana
651:(1): 41–49.
612:(31): 1–7.
437:physeteroid
435:as well as
326:Eopelecanus
294:spot-billed
290:pink-backed
270:great white
135:Pelecanidae
757:Categories
474:References
300:pelicans.
274:Australian
763:Pelecanus
743:128431992
696:129118367
688:0002-7014
626:0374-5481
587:236269386
579:0272-4634
532:258605013
524:0272-4634
223:Argentina
209:from the
153:Species:
146:Pelecanus
91:Kingdom:
85:Eukaryota
39:Messinian
35:Tortonian
462:flamingo
404:Paraguay
335:Scopidae
317:Peruvian
282:Peruvian
246:holotype
131:Family:
105:Chordata
101:Phylum:
95:Animalia
81:Domain:
57:Paleoart
723:Bibcode
715:Geobios
559:Bibcode
400:Uruguay
396:Bolivia
376:Miocene
366:Map of
347:Eurasia
213:of the
207:pelican
178:†
157:†
141:Genus:
121:Order:
111:Class:
741:
694:
686:
624:
585:
577:
530:
522:
451:, the
408:Brazil
386:where
378:in red
343:Africa
238:pelvis
204:fossil
739:S2CID
692:S2CID
583:S2CID
528:S2CID
331:Egypt
313:brown
298:brown
244:(the
217:, in
202:is a
684:ISSN
622:ISSN
575:ISSN
520:ISSN
445:and
428:and
337:and
315:and
296:and
280:and
260:Size
189:2023
115:Aves
731:doi
676:doi
614:doi
567:doi
512:doi
470:.
59:of
759::
737:.
729:.
719:45
717:.
713:.
690:.
682:.
672:48
670:.
666:.
649:17
647:.
643:.
620:.
608:.
604:.
581:.
573:.
565:.
555:41
553:.
549:.
526:.
518:.
508:42
506:.
502:.
482:^
422:,
416:,
402:,
398:,
292:,
276:,
272:,
240:,
221:,
745:.
733::
725::
698:.
678::
628:.
616::
610:6
589:.
569::
561::
534:.
514::
41:)
37:-
33:(
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