124:
142:
123:
368:, which at the time made it the largest known species of rodent, living or extinct. However, these estimates were criticized as being too large by later studies, which found that
632:
Hopkins, Samantha S. B. (19 February 2008). "Reassessing the Mass of
Exceptionally Large Rodents Using Toothrow Length and Area as Proxies for Body Mass".
871:
820:
896:
891:
484:
Sánchez-Villagra, Marcelo R.; Aguilera, Orangel; Horovitz, Inés (19 September 2003). "The
Anatomy of the World's Largest Extinct Rodent".
594:
Millien, Virginie; Bovy, Helene (17 February 2010). "When teeth and bones disagree: body mass estimation of a giant extinct rodent".
310:
delta around 8 million years ago. It was the second-largest of the roughly seven species of its genus. Like many other rodents,
861:
418:), but the size of this rodent could not easily be determined because it was only known from jaw fragments and the teeth of
372:
had an exceptionally thick femur relative to its body compared to other mammals. Body mass estimates based on skull length,
886:
534:"Resizing the largest known extinct rodents (Caviomorpha: Dinomyidae, Neoepiblemidae) using occipital condyle width"
141:
881:
866:
438:
385:
396:
was considered the largest known rodent at the time of its description, the discovery of a complete skull of
398:
788:
833:
243:
876:
389:
380:
weighed between 150 and 250 kg (330 and 550 lb), making it about the same size as a large
649:
611:
571:
509:
271:
136:
838:
777:
773:
747:
698:
563:
501:
373:
342:
737:
729:
688:
680:
641:
603:
553:
545:
493:
718:"The largest among the smallest: the body mass of the giant rodent Josephoartigasia monesi"
283:
262:
742:
717:
693:
668:
558:
533:
206:
855:
575:
326:
307:
74:
653:
615:
513:
458:
444:
49:
34:
607:
300:
219:
94:
59:
811:
645:
497:
381:
350:
319:
315:
153:
99:
43:
751:
733:
702:
684:
567:
505:
805:
338:
322:
173:
89:
84:
69:
64:
54:
549:
825:
364:
was estimated as being approximately 741 kg based on dimensions of the
346:
104:
79:
30:
303:
193:
183:
163:
782:
365:
786:
353:, in 2000. The new species was later classified with the name
667:
Rinderknecht, Andrés; Blanco, R. Ernesto (22 April 2008).
722:
Proceedings of the Royal
Society B: Biological Sciences
673:
Proceedings of the Royal
Society B: Biological Sciences
795:
376:width, tooth row length, and femur length suggest
589:
587:
585:
422:are unusually small relative to its body size.
402:in 2008 showed this species to be even larger.
527:
525:
523:
357:in honor of palaeontologist Brian Patterson.
627:
625:
8:
783:
122:
20:
741:
692:
557:
479:
477:
410:s discovery (specifically, the species "
431:
716:Millien, Virginie (7 September 2008).
457:Amos, Jonathan (September 18, 2003),
7:
532:Engelman, Russell K. (June 2022).
14:
459:"Giant rodent astonishes science"
872:Miocene mammals of South America
140:
768:McNeill Alexander, R. (2003):
1:
892:Fossil taxa described in 1980
770:A Rodent as Big as a Buffalo
669:"The largest fossil rodent"
913:
897:Taxa named by Álvaro Mones
776:vol. 301, p. 1678-9.
538:Royal Society Open Science
306:that lived in the ancient
608:10.1644/08-MAMM-A-347R1.1
406:was known at the time of
277:
270:
249:
242:
137:Scientific classification
135:
130:
121:
23:
16:Extinct species of rodent
646:10.1644/06-MAMM-A-306.1
498:10.1126/science.1089332
399:Josephoartigasia monesi
734:10.1098/rspb.2008.0087
685:10.1098/rspb.2007.1645
416:Josephoartigasia magna
341:was discovered in the
862:Prehistoric pacaranas
834:Paleobiology Database
797:Phoberomys pattersoni
440:Phoberomys pattersoni
394:Phoberomys pattersoni
362:Phoberomys pattersoni
355:Phoberomys pattersoni
296:Phoberomys pattersoni
256:Phoberomys pattersoni
25:Phoberomys pattersoni
887:Fossils of Venezuela
778:(HTML abstract link)
634:Journal of Mammalogy
596:Journal of Mammalogy
728:(1646): 1953–1955.
550:10.1098/rsos.220370
492:(5640): 1708–1710.
337:An almost complete
280:Dabbenea pattersoni
235:P. pattersoni
882:Neogene Venezuela
849:
848:
789:Taxon identifiers
679:(1637): 923–928.
374:occipital condyle
343:Urumaco Formation
292:
291:
287:
904:
842:
841:
829:
828:
816:
815:
814:
784:
756:
755:
745:
713:
707:
706:
696:
664:
658:
657:
629:
620:
619:
591:
580:
579:
561:
529:
518:
517:
481:
472:
471:
470:
469:
454:
448:
436:
420:Josephoartigasia
404:Josephoartigasia
282:
258:
254:
218:
205:
145:
144:
131:Fossil specimen
126:
116:
46:
29:Temporal range:
21:
912:
911:
907:
906:
905:
903:
902:
901:
867:Miocene rodents
852:
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850:
845:
837:
832:
824:
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763:Further reading
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483:
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467:
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463:BBC News Online
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455:
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437:
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428:
412:Artigasia magna
335:
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27:
17:
12:
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5:
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708:
659:
640:(1): 232–243.
621:
581:
519:
473:
449:
430:
429:
427:
424:
408:P. pattersoni'
334:
331:
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268:
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231:
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207:Neoepiblemidae
201:
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28:
15:
13:
10:
9:
6:
4:
3:
2:
909:
898:
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868:
865:
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840:
835:
831:
827:
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753:
749:
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735:
731:
727:
723:
719:
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695:
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628:
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597:
590:
588:
586:
582:
577:
573:
569:
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560:
555:
551:
547:
544:(6): 220370.
543:
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511:
507:
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491:
487:
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330:
328:
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308:Orinoco River
305:
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276:
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244:Binomial name
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56:
51:
45:
42:9.0–6.8
36:
32:
26:
22:
19:
796:
769:
725:
721:
711:
676:
672:
662:
637:
633:
602:(1): 11–18.
599:
595:
541:
537:
489:
485:
466:, retrieved
462:
452:
439:
434:
419:
415:
411:
407:
403:
397:
393:
386:greater kudu
377:
369:
361:
360:Originally,
359:
354:
336:
320:high-crowned
311:
295:
294:
293:
279:
278:
255:
250:
234:
233:
220:
31:Late Miocene
24:
18:
877:Huayquerian
445:Fossilworks
392:. Although
333:Description
35:Huayquerian
856:Categories
468:2008-03-18
426:References
378:Phoberomys
370:Phoberomys
312:Phoberomys
221:Phoberomys
576:249648588
390:waterbuck
382:antelopes
351:Venezuela
323:premolars
316:herbivore
228:Species:
160:Kingdom:
154:Eukaryota
806:Wikidata
752:18495621
703:18198140
654:84160362
616:74872106
568:35719882
514:33953898
506:14500978
384:such as
339:skeleton
272:Synonyms
200:Family:
194:Rodentia
184:Mammalia
174:Chordata
170:Phylum:
164:Animalia
150:Domain:
826:8530008
812:Q338956
774:Science
743:2596365
694:2599941
559:9198521
486:Science
414:", now
347:Urumaco
301:extinct
253:†
232:†
213:Genus:
190:Order:
180:Class:
111:↓
839:253915
750:
740:
701:
691:
652:
614:
574:
566:
556:
512:
504:
327:molars
314:was a
304:rodent
299:is an
650:S2CID
612:S2CID
572:S2CID
510:S2CID
366:femur
318:with
284:Mones
263:Mones
821:GBIF
748:PMID
699:PMID
564:PMID
502:PMID
447:.org
388:and
325:and
286:1980
265:1980
50:PreꞒ
738:PMC
730:doi
726:275
689:PMC
681:doi
677:275
642:doi
604:doi
554:PMC
546:doi
494:doi
490:301
443:at
345:at
329:.
858::
836::
823::
808::
772:.
746:.
736:.
724:.
720:.
697:.
687:.
675:.
671:.
648:.
638:89
636:.
624:^
610:.
600:91
598:.
584:^
570:.
562:.
552:.
540:.
536:.
522:^
508:.
500:.
488:.
476:^
461:,
349:,
100:Pg
44:Ma
754:.
732::
705:.
683::
656:.
644::
618:.
606::
578:.
548::
542:9
516:.
496::
217:†
204:†
105:N
95:K
90:J
85:T
80:P
75:C
70:D
65:S
60:O
55:Ꞓ
39:~
37:)
33:(
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