639:
628:
139:
483:
118:
2165:
369:
2468:
449:) had a great length, extending beyond the second abdominal plate. It was divided into two joints. The first was approximately hastate (with protruding lobes) and was ornamented with fine scales. It was followed by a tubular (tube-shaped) joint that lacked ornamentation. The second joint was less broad and long. The
453:
end (the farthest from the junction point) widens, with a pair of sharp lateral projections ("protuberances"). This gives it a termination finished in three spines similar to those that occur in the genital appendage (of type A) in
343:("primitive") genus that was distinguished from the more derived adelophthalmids by the specialization of its genital operculum (a plate-like segment which contains the genital aperture) and its long and narrow
727:. However, the clade has almost never been used in subsequent studies and lists of eurypterids, and instead, they classify the nanahughmillerids as part of Adelophthalmidae. A derived clade in which
792:), as inferred by Odd Erik Tetlie and Markus Poschmann in 2008, based on the results of a 2008 analysis specifically pertaining to the Adelophthalmoidea and a preceding 2004 analysis.
1446:
390:
was a small eurypterid. The total size of the largest known specimen is estimated at only 6 cm (2.4 in), making it one of the smallest adelopththalmids and eurypterids overall.
530:), including the size of the carapace (broader and shorter than in the latter), the position of the eyes further from the margin (as opposed to the marginal position of
432:) also present three to four parallel lines along the posterior margin, followed by five lines that end in a series of separate and lunate (crescent-shaped) scales.
2531:
542:
based on the size of the carapace and the position of the eyes more or less being similar, as well as the same linear ornamentation. However, while in its
2582:
1383:
slabs that reach a combined thickness of 305 m (1,000 ft). It is also possible to find green shale and very rarely, red shale. This habitat was probably
435:
Its genital operculum (a plate-like segment which contains the genital aperture) is the main characteristic that distinguishes it from the rest of the
2518:
2592:
2587:
2244:
2234:
1756:
1443:
1435:
445:. It possessed two opercular flaps, two protruding extensions lateral to the genital appendage. The genital appendage (which is of type A,
699:
to contain the adelophthalmoids with no or reduced genital spatulae (a long, flat piece in the operculum) and the second to fifth pair of
1574:
1543:
Kjellesvig-Waering, Erik N. (1964). "Eurypterida: Notes on the
Subgenus Hughmilleria (Nanahughmilleria) from the Silurian of New York".
2196:
2211:
780:
below presents the inferred phylogenetic positions of most of the genera included in the three most derived superfamilies of the
1345:) epoch of the Vernon Formation of the New York state. In this formation, fossils of other eurypterids have been found, such as
2455:
2239:
2181:
1401:
2229:
2219:
2173:
408:), the carapace was 18 mm (0.7 in) wide and 13 mm (0.5 in) long. Five parallel lines along the front margin that make up the
138:
579:
should be classified under this new genus. Three years later, Kjellesvig-Waering decided to assign the same species to the
2577:
2191:
2186:
2224:
2201:
758:
638:
2572:
627:
1857:
1749:
1697:"A reappraisal of Eurypterus dumonti Stainier, 1917 and its position within the Adelophthalmidae Tollerton, 1989"
1431:
1396:
588:. In 1966, Kjellesvig-Waering, together with the American paleontologist Kenneth Edward Caster, recognized that
1424:
Cope's Rule and Romer's theory: patterns of diversity and gigantism in eurypterids and
Palaeozoic vertebrates"
2164:
601:
1721:
2486:
1722:"Eurypterid-Associated Biota of the Vernon Formation, Pittsford, New York (Silurian of the United States)"
1635:
Kjellesvig-Waering, Erik N.; Leutze, Willard P. (1966). "Eurypterids from the
Silurian of West Virginia".
470:
Erik Norman
Kjellesvig-Waering predicted that the genital operculum would end up being a feature of great
429:
2544:
1580:
740:
678:
501:
340:
314:
2471:
1742:
1480:
439:(more "advanced") adelopththalmids, showing characteristics indicative of the more basal eurypterid
1360:
534:) and the morphology of the genital appendage. Instead, Ruedemann suggested a relationship between
413:
409:
276:
761:), relatively long and narrow eyes and a complex termination of the genital appendage. Therefore,
1644:
1614:
1552:
1496:
133:
2549:
333:
state. The genus is named after
Pittsford, where the two only known specimens have been found.
2536:
2060:
1993:
1985:
1439:
902:
809:
789:
681:
311:
1469:"Phylogeny and palaeoecology of the Adelophthalmoidea (Arthropoda; Chelicerata; Eurypterida)"
420:(rounded protuberances) appear. In the posterior margin, there is a strip of fine triangular
2400:
2114:
2106:
2096:
2088:
2068:
2039:
1955:
1904:
1584:
1488:
1021:
997:
670:
667:
653:
584:
571:
517:
513:
505:
482:
397:
330:
299:
296:
260:
219:
206:
117:
1682:
2429:
2275:
2132:
2073:
2029:
1932:
1919:
1450:
1380:
1125:
421:
1423:
600:
is translated as "wing from
Pittsford", with the first word of the name referring to the
1605:
Kjellesvig-Waering, Erik N. (1961). "The
Silurian Eurypterida of the Welsh Borderland".
1484:
2345:
2303:
2261:
2124:
2051:
2011:
2003:
1927:
1087:
1038:
847:
799:
785:
781:
754:
605:
467:
462:
380:
1696:
368:
2566:
2352:
2324:
2317:
2289:
2268:
2021:
1960:
1950:
1942:
1896:
1886:
1878:
1834:
1798:
1468:
1238:
916:
704:
436:
401:
344:
307:
68:
1500:
692:, but it was considered different enough to represent a new separate genus in 1966.
2310:
2142:
2078:
2034:
1909:
1338:
1173:
1101:
864:
823:
766:
526:
471:
318:
245:
2407:
2384:
2282:
1976:
1869:
1807:
1789:
1522:
1356:
947:
749:
695:
349:
180:
43:
2509:
2373:
2359:
2338:
2331:
2296:
2137:
1825:
1816:
1765:
1492:
1364:
1351:
1342:
1221:
1197:
604:(the location where it was initially found) and the last word composed of the
446:
441:
383:
284:
193:
88:
53:
30:
2450:
2434:
2424:
1783:
1368:
777:
592:
was sufficiently different from the other eurypterids and erected the genus
288:
170:
150:
93:
37:
2503:
2393:
2366:
1334:
1149:
580:
497:
456:
417:
405:
326:
83:
78:
63:
58:
48:
610:
2523:
1648:
1618:
1556:
700:
543:
520:
state. Ruedemann noted several differences between his new species and
425:
98:
73:
1777:
1384:
1376:
696:
450:
160:
2480:
1372:
674:
481:
367:
303:
280:
1704:
Bulletin de l'Institut Royal des
Sciences Naturelles de Belgique
2484:
1738:
404:
placed away from the head margin. In the largest specimen (the
1734:
356:
closest relative. With an estimated length of 6 cm (2.4 in),
1576:
Eurypterid phylogeny with remarks on the origin of arachnids
596:
based on the morphology of its genital appendage. The name
1685:. In World Spider Catalog. Natural History Museum Bern.
757:(shared characteristics different from that of their
500:
and paratype (NYSM 10102 and NYSM 10103, both at the
1683:
A summary list of fossil spiders and their relatives
1662:
1422:
Lamsdell, James C.; Braddy, Simon J. (2009-10-14). "
2493:
2443:
2417:
2383:
2253:
2210:
2172:
2123:
2105:
2087:
2059:
2048:
2020:
2002:
1984:
1973:
1941:
1918:
1895:
1877:
1866:
1850:
1843:
688:was originally described as a species of the genus
496:is only known by two well preserved specimens, the
416:portion, a series of small irregularly distributed
412:can be seen on the surface of the carapace. In the
1681:Dunlop, J. A., Penney, D. & Jekel, D. 2018.
1467:Erik Tetlie, O; Poschmann, Markus (2008-06-01).
784:infraorder of eurypterids (Adelophthalmoidea,
562:could probably represent a late descendant of
400:shield of the head) with elongated and narrow
1750:
8:
1371:of the place consists of dark gray to black
711:-type (hypothetical since the appendages of
486:The holotype (right) and paratype (left) of
575:in 1961, Kjellesvig-Waering suggested that
2481:
2056:
1981:
1874:
1847:
1757:
1743:
1735:
474:importance at least at the generic level.
317:of eurypterids. Fossils of the single and
116:
20:
554:had imbricate scales similar to those of
360:was one of the smallest adelophthalmids.
504:). In 1921, the American paleontologist
1538:
1536:
1412:
743:(more "primitive") group consisting of
1523:"A recurrent Pittsford (Salina) fauna"
1355:, as well as indeterminate species of
325:, have been discovered in deposits of
1568:
1566:
239:Kjellesvig-Waering & Leutze, 1966
7:
1677:
1675:
1630:
1628:
1516:
1514:
1512:
1510:
1462:
1460:
1458:
1418:
1416:
753:. This clade is backed by a pair of
715:are unknown). This family contained
1473:Journal of Systematic Palaeontology
1213:
1189:
1165:
1141:
1117:
1093:
1013:
989:
939:
932:
908:
898:
839:
815:
805:
795:
550:had the same linear ornamentation,
1695:Tetlie, O.E.; van Roy, P. (2006).
739:is better supported, as well as a
14:
2583:Silurian animals of North America
1333:fossils have been recovered from
2467:
2466:
2163:
1583:. pp. 1β344. Archived from
637:
626:
569:In the description of the genus
137:
2456:Timeline of eurypterid research
1402:Timeline of eurypterid research
125:Restoration of the carapace of
1527:New York State Museum Bulletin
651:(top) and the closely related
287:, an extinct group of aquatic
1:
2593:Fossil taxa described in 1966
666:is classified as part of the
647:Type A genital appendages of
558:. Even so, he suggested that
428:(the dorsal part of the body
295:is classified as part of the
2588:Eurypterids of North America
447:assumed to represent females
16:Genus of arthropods (fossil)
2609:
1521:Ruedemann, Rudolf (1921).
654:Nanahughmilleria norvegica
2464:
2161:
1858:List of eurypterid genera
1772:
1726:The Paleobiology Database
1573:Tetlie, Odd Erik (2004).
1493:10.1017/S1477201907002416
1444:Supplementary information
1434::10.1098/rsbl.2009.0700.
1397:List of eurypterid genera
1347:Eurypterus pittsfordensis
1235:
1218:
1211:
1194:
1187:
1170:
1163:
1146:
1139:
1122:
1115:
1098:
1091:
1035:
1018:
1011:
994:
987:
961:
944:
937:
930:
913:
906:
896:
861:
844:
837:
820:
813:
803:
703:(of the prosoma, "head")
649:Pittsfordipterus phelpsae
255:Pittsfordipterus phelpsae
251:
244:
134:Scientific classification
132:
124:
115:
23:
1352:Mixopterus multispinosus
1637:Journal of Paleontology
1607:Journal of Paleontology
1545:Journal of Paleontology
769:(closest relative) of
759:latest common ancestor
508:described the species
490:
396:had a broad carapace (
376:
2545:Paleobiology Database
1581:University of Bristol
1337:deposits of the Late
677:("group") within the
510:Hughmilleria phelpsae
502:New York State Museum
485:
371:
2578:Silurian eurypterids
1485:2008JSPal...6..237T
478:History of research
372:Size comparison of
1986:Onychopterelloidea
1667:www.wiktionary.org
1449:2018-02-28 at the
491:
377:
329:age in Pittsford,
2573:Adelophthalmoidea
2560:
2559:
2487:Taxon identifiers
2478:
2477:
2159:
2158:
2155:
2154:
2151:
2150:
2107:Adelophthalmoidea
2089:Waeringopteroidea
2061:Carcinosomatoidea
1994:Onychopterellidae
1969:
1968:
1322:
1321:
1313:
1312:
1304:
1303:
1295:
1294:
1286:
1285:
1277:
1276:
1268:
1267:
1259:
1258:
1250:
1249:
1077:
1076:
1068:
1067:
1059:
1058:
1050:
1049:
976:
975:
903:Adelophthalmoidea
885:
884:
876:
875:
810:Waeringopteroidea
682:Adelophthalmoidea
524:(type species of
312:Adelophthalmoidea
268:
267:
240:
207:Adelophthalmoidea
36:421.3β418.7
2600:
2553:
2552:
2540:
2539:
2527:
2526:
2514:
2513:
2512:
2495:Pittsfordipterus
2482:
2470:
2469:
2444:Related articles
2401:Merostomichnites
2167:
2115:Adelophthalmidae
2097:Waeringopteridae
2069:Carcinosomatidae
2057:
2040:Strobilopteridae
1982:
1956:Hibbertopteridae
1905:Parastylonuridae
1875:
1848:
1759:
1752:
1745:
1736:
1730:
1729:
1718:
1712:
1711:
1701:
1692:
1686:
1679:
1670:
1659:
1653:
1652:
1643:(5): 1109β1122.
1632:
1623:
1622:
1602:
1596:
1595:
1593:
1592:
1570:
1561:
1560:
1540:
1531:
1530:
1518:
1505:
1504:
1464:
1453:
1430:: rsbl20090700.
1420:
1331:Pittsfordipterus
1214:
1190:
1166:
1142:
1118:
1094:
1022:Parahughmilleria
1014:
998:Nanahughmilleria
990:
965:Pittsfordipterus
940:
933:
909:
899:
840:
816:
806:
796:
790:waeringopteroids
763:Pittsfordipterus
745:Pittsfordipterus
733:Parahughmilleria
729:Nanahughmilleria
725:Parahughmilleria
721:Pittsfordipterus
713:Pittsfordipterus
671:Adelophthalmidae
664:Pittsfordipterus
641:
630:
598:Pittsfordipterus
594:Pittsfordipterus
590:H. (N.) phelpsae
585:Nanahughmilleria
572:Parahughmilleria
538:and the species
514:Vernon Formation
506:Rudolf Ruedemann
494:Pittsfordipterus
394:Pittsfordipterus
388:Pittsfordipterus
358:Pittsfordipterus
355:
337:Pittsfordipterus
300:Adelophthalmidae
293:Pittsfordipterus
272:Pittsfordipterus
238:
234:Pittsfordipterus
231:
220:Adelophthalmidae
218:
205:
192:
142:
141:
120:
110:
40:
29:Temporal range:
25:Pittsfordipterus
21:
2608:
2607:
2603:
2602:
2601:
2599:
2598:
2597:
2563:
2562:
2561:
2556:
2548:
2543:
2535:
2530:
2522:
2517:
2508:
2507:
2502:
2489:
2479:
2474:
2460:
2439:
2430:Chasmataspidida
2413:
2379:
2276:Campylocephalus
2249:
2206:
2168:
2147:
2133:Hughmilleriidae
2119:
2101:
2083:
2074:Megalograptidae
2050:
2044:
2030:Dolichopteridae
2016:
2004:Moselopteroidea
1998:
1975:
1965:
1951:Drepanopteridae
1937:
1933:Hardieopteridae
1920:Kokomopteroidea
1914:
1891:
1868:
1862:
1839:
1768:
1763:
1733:
1720:
1719:
1715:
1699:
1694:
1693:
1689:
1680:
1673:
1660:
1656:
1634:
1633:
1626:
1604:
1603:
1599:
1590:
1588:
1572:
1571:
1564:
1542:
1541:
1534:
1520:
1519:
1508:
1466:
1465:
1456:
1451:Wayback Machine
1428:Biology Letters
1421:
1414:
1410:
1393:
1328:
1323:
1314:
1305:
1296:
1287:
1278:
1269:
1260:
1251:
1126:Herefordopterus
1078:
1069:
1060:
1051:
977:
886:
877:
737:Adelopththalmus
717:Nanahughmileria
661:
660:
659:
658:
644:
643:
642:
633:
632:
631:
620:
480:
466:. The American
379:Like the other
366:
353:
264:
258:
237:
229:
216:
203:
190:
136:
111:
109:
108:
107:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
35:
34:
27:
17:
12:
11:
5:
2606:
2604:
2596:
2595:
2590:
2585:
2580:
2575:
2565:
2564:
2558:
2557:
2555:
2554:
2541:
2528:
2515:
2499:
2497:
2491:
2490:
2485:
2476:
2475:
2465:
2462:
2461:
2459:
2458:
2453:
2447:
2445:
2441:
2440:
2438:
2437:
2432:
2427:
2421:
2419:
2418:Related groups
2415:
2414:
2412:
2411:
2404:
2397:
2389:
2387:
2381:
2380:
2378:
2377:
2370:
2363:
2356:
2349:
2346:Onychopterella
2342:
2335:
2328:
2321:
2314:
2307:
2304:Hibbertopterus
2300:
2293:
2286:
2279:
2272:
2265:
2262:Adelophthalmus
2257:
2255:
2254:Notable genera
2251:
2250:
2248:
2247:
2242:
2237:
2232:
2227:
2222:
2216:
2214:
2208:
2207:
2205:
2204:
2199:
2194:
2189:
2184:
2178:
2176:
2170:
2169:
2162:
2160:
2157:
2156:
2153:
2152:
2149:
2148:
2146:
2145:
2140:
2135:
2129:
2127:
2125:Pterygotioidea
2121:
2120:
2118:
2117:
2111:
2109:
2103:
2102:
2100:
2099:
2093:
2091:
2085:
2084:
2082:
2081:
2076:
2071:
2065:
2063:
2054:
2052:Diploperculata
2046:
2045:
2043:
2042:
2037:
2032:
2026:
2024:
2018:
2017:
2015:
2014:
2012:Moselopteridae
2008:
2006:
2000:
1999:
1997:
1996:
1990:
1988:
1979:
1971:
1970:
1967:
1966:
1964:
1963:
1958:
1953:
1947:
1945:
1939:
1938:
1936:
1935:
1930:
1928:Kokomopteridae
1924:
1922:
1916:
1915:
1913:
1912:
1907:
1901:
1899:
1893:
1892:
1890:
1889:
1883:
1881:
1879:Rhenopteroidea
1872:
1864:
1863:
1861:
1860:
1854:
1852:
1845:
1841:
1840:
1838:
1837:
1828:
1819:
1810:
1801:
1792:
1786:
1780:
1773:
1770:
1769:
1764:
1762:
1761:
1754:
1747:
1739:
1732:
1731:
1713:
1687:
1671:
1654:
1624:
1613:(4): 789β835.
1597:
1562:
1551:(2): 410β412.
1532:
1506:
1479:(2): 237β249.
1454:
1411:
1409:
1406:
1405:
1404:
1399:
1392:
1389:
1375:with abundant
1327:
1324:
1320:
1319:
1316:
1315:
1311:
1310:
1307:
1306:
1302:
1301:
1298:
1297:
1293:
1292:
1289:
1288:
1284:
1283:
1280:
1279:
1275:
1274:
1271:
1270:
1266:
1265:
1262:
1261:
1257:
1256:
1253:
1252:
1248:
1247:
1244:
1243:
1234:
1231:
1230:
1227:
1226:
1217:
1212:
1210:
1207:
1206:
1203:
1202:
1193:
1188:
1186:
1183:
1182:
1179:
1178:
1169:
1164:
1162:
1159:
1158:
1155:
1154:
1145:
1140:
1138:
1135:
1134:
1131:
1130:
1121:
1116:
1114:
1111:
1110:
1107:
1106:
1097:
1092:
1090:
1088:Pterygotioidea
1084:
1083:
1080:
1079:
1075:
1074:
1071:
1070:
1066:
1065:
1062:
1061:
1057:
1056:
1053:
1052:
1048:
1047:
1044:
1043:
1039:Adelophthalmus
1034:
1031:
1030:
1027:
1026:
1017:
1012:
1010:
1007:
1006:
1003:
1002:
993:
988:
986:
983:
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979:
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974:
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943:
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887:
883:
882:
879:
878:
874:
873:
870:
869:
860:
857:
856:
853:
852:
848:Waeringopterus
843:
838:
836:
833:
832:
829:
828:
819:
814:
812:
804:
802:
800:Diploperculata
794:
786:Pterygotioidea
782:Diploperculata
755:synapomorphies
731:is closest to
646:
645:
636:
635:
634:
625:
624:
623:
622:
621:
619:
618:Classification
616:
479:
476:
468:paleontologist
463:Adelophthalmus
381:adelophthalmid
365:
362:
266:
265:
259:
249:
248:
242:
241:
227:
223:
222:
214:
210:
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201:
197:
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188:
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178:
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130:
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122:
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113:
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102:
97:
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67:
62:
57:
52:
47:
42:
41:
28:
15:
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10:
9:
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2605:
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2498:
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2457:
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2423:
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2416:
2410:
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2405:
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2388:
2386:
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2376:
2375:
2371:
2369:
2368:
2364:
2362:
2361:
2357:
2355:
2354:
2353:Pentecopterus
2350:
2348:
2347:
2343:
2341:
2340:
2336:
2334:
2333:
2329:
2327:
2326:
2325:Megalograptus
2322:
2320:
2319:
2318:Jaekelopterus
2315:
2313:
2312:
2308:
2306:
2305:
2301:
2299:
2298:
2294:
2292:
2291:
2290:Drepanopterus
2287:
2285:
2284:
2280:
2278:
2277:
2273:
2271:
2270:
2269:Brachyopterus
2266:
2264:
2263:
2259:
2258:
2256:
2252:
2246:
2245:South America
2243:
2241:
2238:
2236:
2235:North America
2233:
2231:
2228:
2226:
2223:
2221:
2218:
2217:
2215:
2213:
2209:
2203:
2200:
2198:
2197:Carboniferous
2195:
2193:
2190:
2188:
2185:
2183:
2180:
2179:
2177:
2175:
2174:Geochronology
2171:
2166:
2144:
2141:
2139:
2136:
2134:
2131:
2130:
2128:
2126:
2122:
2116:
2113:
2112:
2110:
2108:
2104:
2098:
2095:
2094:
2092:
2090:
2086:
2080:
2077:
2075:
2072:
2070:
2067:
2066:
2064:
2062:
2058:
2055:
2053:
2047:
2041:
2038:
2036:
2033:
2031:
2028:
2027:
2025:
2023:
2022:Eurypteroidea
2019:
2013:
2010:
2009:
2007:
2005:
2001:
1995:
1992:
1991:
1989:
1987:
1983:
1980:
1978:
1972:
1962:
1961:Mycteroptidae
1959:
1957:
1954:
1952:
1949:
1948:
1946:
1944:
1943:Mycteropoidea
1940:
1934:
1931:
1929:
1926:
1925:
1923:
1921:
1917:
1911:
1908:
1906:
1903:
1902:
1900:
1898:
1897:Stylonuroidea
1894:
1888:
1887:Rhenopteridae
1885:
1884:
1882:
1880:
1876:
1873:
1871:
1865:
1859:
1856:
1855:
1853:
1849:
1846:
1842:
1836:
1835:Sclerophorata
1832:
1829:
1827:
1823:
1820:
1818:
1814:
1811:
1809:
1805:
1802:
1800:
1799:Euchelicerata
1796:
1793:
1791:
1787:
1785:
1781:
1779:
1775:
1774:
1771:
1767:
1760:
1755:
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1587:on 2021-07-30
1586:
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1239:Jaekelopterus
1233:
1232:
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1225:
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650:
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629:
617:
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612:
608:word ΟΟΞ΅ΟΟΞ½ (
607:
603:
602:type locality
599:
595:
591:
587:
586:
582:
578:
574:
573:
567:
565:
564:H. shawangunk
561:
557:
553:
549:
548:H. shawangunk
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540:H. shawangunk
537:
533:
529:
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459:
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452:
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427:
423:
419:
415:
411:
410:ornamentation
407:
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395:
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389:
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382:
375:
370:
363:
361:
359:
352:
351:
346:
342:
338:
334:
332:
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324:
320:
316:
313:
310:("advanced")
309:
305:
301:
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278:
274:
273:
262:
257:
256:
250:
247:
243:
236:
235:
228:
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211:
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202:
200:Superfamily:
199:
198:
195:
189:
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185:
182:
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176:
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172:
169:
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159:
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95:
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2323:
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2311:Hughmilleria
2309:
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2281:
2274:
2267:
2260:
2143:Pterygotidae
2079:Mixopteridae
2035:Eurypteridae
1910:Stylonuridae
1830:
1821:
1812:
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1794:
1725:
1716:
1707:
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1600:
1589:. Retrieved
1585:the original
1575:
1548:
1544:
1526:
1476:
1472:
1427:
1361:leperditiids
1357:phyllocarids
1350:
1346:
1330:
1329:
1326:Paleoecology
1237:
1236:
1220:
1219:
1196:
1195:
1174:Erettopterus
1172:
1171:
1148:
1147:
1124:
1123:
1102:Hughmilleria
1100:
1099:
1037:
1036:
1020:
1019:
996:
995:
964:
963:
962:
946:
945:
915:
914:
865:Grossopterus
863:
862:
846:
845:
824:Orcanopterus
822:
821:
775:
770:
767:sister group
762:
748:
744:
736:
732:
728:
724:
723:and perhaps
720:
716:
712:
709:Hughmilleria
708:
694:
690:Hughmilleria
689:
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663:
662:
652:
648:
609:
597:
593:
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559:
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547:
539:
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531:
527:Hughmilleria
525:
521:
509:
493:
492:
487:
472:phylogenetic
461:
455:
440:
434:
393:
392:
387:
378:
373:
357:
348:
336:
335:
322:
319:type species
292:
275:("wing from
271:
270:
269:
254:
253:
246:Type species
233:
232:
126:
24:
18:
2408:Palmichnium
2385:Ichnogenera
2283:Carcinosoma
1977:Eurypterina
1870:Stylonurina
1808:Prosomapoda
1790:Chelicerata
1788:Subphylum:
1766:Eurypterids
1661:Meaning of
1438:1744-9561.
1365:cephalopods
948:Bassipterus
771:Bassipterus
750:Bassipterus
707:(limbs) of
686:P. phelpsae
679:superfamily
673:, the only
577:H. phelpsae
560:H. phelpsae
556:H. socialis
552:H. phelpsae
536:H. phelpsae
532:H. socialis
522:H. socialis
488:P. phelpsae
384:eurypterids
374:P. phelpsae
364:Description
350:Bassipterus
323:P. phelpsae
315:superfamily
302:, the only
194:Eurypterida
181:Chelicerata
177:Subphylum:
127:P. phelpsae
2567:Categories
2374:Stylonurus
2360:Pterygotus
2339:Mixopterus
2332:Megarachne
2297:Eurypterus
2182:Ordovician
2138:Slimonidae
2049:Infraorder
1826:Dekatriata
1817:Planaterga
1784:Arthropoda
1591:2018-09-09
1529:: 205β222.
1442:19828493.
1408:References
1343:Ludfordian
1222:Acutiramus
1198:Pterygotus
705:appendages
442:Eurypterus
289:arthropods
285:eurypterid
171:Arthropoda
31:Ludfordian
2451:Metastoma
2435:Xiphosura
2425:Arachnida
2212:Geography
1776:Kingdom:
1369:lithology
778:cladogram
614:, wing).
512:from the
418:tubercles
414:posterior
277:Pittsford
261:Ruedemann
157:Kingdom:
151:Eukaryota
2510:Q7199502
2504:Wikidata
2472:Category
2394:Arcuites
2367:Slimonia
2192:Devonian
2187:Silurian
1974:Suborder
1867:Suborder
1844:Taxonomy
1782:Phylum:
1778:Animalia
1710:: 79β90.
1501:59488956
1447:Archived
1391:See also
1385:lagoonal
1381:dolomite
1335:Silurian
1150:Slimonia
788:and the
701:prosomal
657:(bottom)
581:subgenus
518:New York
498:holotype
457:Slimonia
430:segments
426:tergites
406:paratype
347:, being
331:New York
327:Silurian
279:") is a
213:Family:
167:Phylum:
161:Animalia
147:Domain:
2537:1116887
2524:4650075
2240:Oceania
2202:Permian
1649:1301985
1619:1301214
1579:(PhD).
1557:1301566
1481:Bibcode
765:is the
544:ventral
516:of the
437:derived
308:derived
306:in the
226:Genus:
187:Order:
105:↓
2230:Europe
2220:Africa
1851:Genera
1663:pterus
1647:
1617:
1555:
1499:
1377:gypsum
1367:. The
1339:Ludlow
697:thesis
668:family
611:pteron
546:part,
451:distal
424:. The
422:scales
398:dorsal
339:was a
297:family
263:, 1921
2550:18991
2532:IRMNG
1831:Clade
1822:Clade
1813:Clade
1804:Clade
1795:Clade
1700:(PDF)
1645:JSTOR
1615:JSTOR
1553:JSTOR
1497:S2CID
1373:shale
741:basal
675:clade
606:Greek
354:'
341:basal
304:clade
281:genus
2519:GBIF
2225:Asia
1440:PMID
1436:ISSN
1379:and
1363:and
776:The
747:and
735:and
460:and
402:eyes
345:eyes
44:Preκ
1489:doi
1432:doi
1349:or
283:of
2569::
2547::
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2521::
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1833::
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1815::
1806::
1797::
1724:.
1708:76
1706:.
1702:.
1674:^
1665:.
1641:40
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1627:^
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1565:^
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94:Pg
38:Ma
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1559:.
1503:.
1491::
1483::
1477:6
1341:(
252:β
230:β
217:β
204:β
191:β
99:N
89:K
84:J
79:T
74:P
69:C
64:D
59:S
54:O
49:κ
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