2014:
1262:
127:
2032:
1248:
525:
with a robust shaft, and a dorsoventral and very low iliac blade with a long anterior process, exceeding slightly the pubic peduncle. Yet, in 2021, it was found that many of these features fall within the range of variation present in juvenile specimens of
568:
based on a slightly updated data matrix from previous analyses, resulting in a matrix that includes 19 aetosaurs and two outgroup taxa which are scored based on 37 morphological traits.
2171:
354:
was first named by Lúcio
Roberto-da-Silva, Julia B. Desojo, Sérgio F. Cabreira, Alex S. S. Aires, Rodrigo T. Müller, Cristian P. Pacheco and Sérgio Dias-da-Silva in
2112:
2196:
2191:
1367:
Paes-Neto, V.D.; Desojo, J.B.; Brust, A.C.B.; Schultz, C.L.; Da-Rosa, A.A.S.; Soares, M.B. (2021). "Intraspecific variation in the axial skeleton of
2099:
612:
of early aetosaurs that developed into desmatosuchines and typothoracisines later in the
Triassic. Although this clade is placed in a relatively
2186:
1427:
Paes-Neto, V.D.; Desojo, J.B.; Brust, A.C.B.; Ribeiro, A.M.; Schultz, C.L.; Soares, M.B. (2021). "The first braincase of the basal aetosaur
1484:
1311:
Roberto-Da-Silva, L. C.; Desojo, J. B.; Cabreira, S. R. F.; Aires, A. S. S.; Müller, R. T.; Pacheco, C. P.; Dias-Da-Silva, S. R. (2014).
498:
2013:
457:
is known from a relatively small-sized individual with an estimated length of 76 centimetres (30 in), probably a two-year-old
2176:
793:
126:
2181:
958:
941:
1477:
1117:
1014:
917:
721:
583:
762:
704:
1371:(Archosauria: Aetosauria) and its implications for the aetosaur diversity of the Late Triassic of Brazil".
2052:
1086:
658:
2138:
1572:
745:
613:
431:
308:
1134:
355:
244:
2090:
1470:
1267:
902:
687:
486:
458:
2166:
2035:
1448:
1406:
1038:
609:
482:
387:
121:
2143:
517:
is greatly expanded, while the medial portion of scapular blade is expanded anteroposteriorly.
2125:
2117:
1896:
1770:
1398:
1342:
578:
2130:
1920:
1843:
1658:
1440:
1388:
1380:
1332:
1324:
1062:
840:
672:
596:
506:
502:
466:
462:
399:
395:
391:
220:
1871:
1801:
1541:
999:
628:
510:
1985:
1940:
1794:
1708:
624:
620:
293:
17:
1313:"A new aetosaur from the Upper Triassic of the Santa Maria Formation, southern Brazil"
2160:
1963:
1956:
1912:
1904:
1888:
1834:
1815:
1752:
1737:
1729:
1688:
1646:
1452:
1410:
1312:
1253:
1101:
879:
528:
443:
375:
339:
330:
300:
80:
41:
1860:
1808:
1745:
1722:
1668:
1557:
1532:
616:
605:
601:
573:
561:
415:
411:
379:
359:
252:
194:
1444:
1384:
532:. One example is the lateral fossa, which becomes more apparent in individuals of
490:
2075:
2084:
1971:
1787:
1283:
1276:
55:
1978:
1779:
1759:
1681:
1523:
1328:
1243:
862:
643:
383:
100:
65:
1696:
632:
427:
181:
138:
105:
49:
1402:
1346:
2069:
1632:
1610:
1584:
1517:
1493:
478:
474:
371:
367:
296:
207:
158:
95:
90:
75:
70:
60:
1393:
536:
longer than 1 metre (3 ft 3 in)). Furthermore, a braincase of
2104:
1928:
1337:
522:
514:
435:
419:
407:
403:
312:
286:
168:
110:
85:
489:
that widely extend laterally through most of the anterior edge of the
1511:
439:
316:
304:
148:
2046:
587:
423:
289:
1431:(Archosauria: Pseudosuchia) from the Upper Triassic of Brazil".
540:
had a remarkably similar structure of the basioccipital bone to
2050:
1504:
1466:
426:, a partial right pes, and anterior and mid-dorsal paramedian
1462:
337:
likely represents a juvenile individual of the contemporary
505:. A ventral keel is present in the cervical vertebrae. The
323:
one of the oldest aetosaurs. It contains a single species,
461:
based on the incomplete fusion of neural arches to their
509:
of the anterior and mid-dorsal vertebrae lack a lateral
477:
by a unique combination of characters not controlled by
600:, both of which are also from South America. However,
631:, two derived clades of stagonolepidids. Below is a
259:
2059:
1939:
1870:
1842:
1833:
1769:
1707:
1667:
1656:
1630:
1583:
1556:
1362:
1360:
1358:
1356:
564:analysis was conducted in the first description of
1422:
1420:
1306:
1304:
1302:
1300:
481:, most of which are found in its vertebrae. Its
1478:
8:
1373:Annals of the Brazilian Academy of Sciences
333:to date. Anatomical evidence suggests that
2047:
1880:
1839:
1714:
1673:
1664:
1638:
1627:
1562:
1553:
1501:
1485:
1471:
1463:
370:specimen, a partially articulated partial
31:
1392:
1336:
623:aetosaurs, it is closely related to both
329:, the fifth aetosaur species known from
2172:Late Triassic reptiles of South America
1296:
548:was suggested as a junior synonym of
7:
501:are absent in both cervical and mid-
473:can be distinguished from all other
1109:
1078:
1054:
1030:
1006:
933:
909:
854:
847:
835:
785:
778:
737:
679:
650:
638:
307:. Fossils have been found from the
1433:Journal of Vertebrate Paleontology
25:
2192:Prehistoric pseudosuchian genera
2031:
2030:
2012:
1260:
1246:
125:
2197:Taxa named by Rodrigo T. Müller
794:Aetobarbakinoides brasiliensis
366:. It is known solely from its
1:
2187:Fossil taxa described in 2014
1445:10.1080/02724634.2021.1928681
1385:10.1590/0001-3765202120201239
635:modified from this analysis.
434:which dates back to the late
430:. It was collected from the
2213:
959:Paratypothorax andressorum
942:Rioarribasuchus chamaensis
586:. Close relatives of this
513:. The proximal end of the
2026:
2009:
1883:
1717:
1676:
1641:
1626:
1607:
1565:
1552:
1500:
1329:10.11646/zootaxa.3764.3.1
1131:
1114:
1107:
1083:
1076:
1059:
1052:
1035:
1028:
1011:
1004:
955:
938:
931:
914:
907:
876:
859:
852:
845:
833:
807:
790:
783:
776:
759:
742:
735:
718:
701:
684:
677:
655:
648:
309:Santa Maria Supersequence
258:
251:
122:Scientific classification
120:
34:
27:Extinct genus of reptiles
1118:Desmatosuchus haplocerus
1015:Sierritasuchus macalpini
918:Tecovasuchus chatterjeei
722:Neoaetosauroides engaeus
499:hyposphene articulations
374:. It is composed of the
811:Polesinesuchus aurelioi
763:Stagonolepis robertsoni
705:Coahomasuchus kahleorum
364:Polesinesuchus aurelioi
326:Polesinesuchus aurelioi
264:Polesinesuchus aurelioi
18:Polesinesuchus aurelioi
1429:Aetosauroides scagliai
1379:(supp. 2): e20201239.
1369:Aetosauroides scagliai
1087:Acaenasuchus geoffreyi
659:Aetosauroides scagliai
582:, known from the same
2177:Santa Maria Formation
2139:Paleobiology Database
746:Calyptosuchus wellesi
572:was recovered as the
544:. For these reasons,
432:Santa Maria Formation
1135:Desmatosuchus smalli
584:geological formation
1268:Paleontology portal
903:Paratypothoracisini
688:Aetosaurus ferratus
386:elements including
1039:Longosuchus meadei
483:cervical vertebrae
2182:Fossils of Brazil
2154:
2153:
2126:Open Tree of Life
2053:Taxon identifiers
2044:
2043:
2022:
2021:
2007:
2006:
2003:
2002:
1999:
1998:
1995:
1994:
1944:(Desmatosuchinae
1897:Aetobarbakinoides
1829:
1828:
1825:
1824:
1771:Paratypothoracini
1622:
1621:
1618:
1617:
1603:
1602:
1236:
1235:
1227:
1226:
1218:
1217:
1209:
1208:
1200:
1199:
1191:
1190:
1182:
1181:
1173:
1172:
1164:
1163:
1155:
1154:
1146:
1145:
988:
987:
979:
978:
970:
969:
891:
890:
822:
821:
579:Aetobarbakinoides
278:
277:
269:Roberto-da-Silva
247:
240:Roberto-da-Silva
48:231.4–225.9
16:(Redirected from
2204:
2147:
2146:
2134:
2133:
2121:
2120:
2108:
2107:
2095:
2094:
2093:
2080:
2079:
2078:
2048:
2034:
2033:
2017:
2016:
1921:Neoaetosauroides
1881:
1844:Stagonolepidinae
1840:
1715:
1674:
1665:
1659:Stagonolepididae
1639:
1628:
1563:
1554:
1547:
1546:
1502:
1487:
1480:
1473:
1464:
1457:
1456:
1424:
1415:
1414:
1396:
1364:
1351:
1350:
1340:
1308:
1270:
1265:
1264:
1263:
1256:
1251:
1250:
1249:
1110:
1079:
1063:Lucasuchus hunti
1055:
1031:
1007:
934:
910:
855:
848:
841:Typothoracisinae
836:
786:
779:
738:
680:
673:Stagonolepididae
651:
639:
629:typothoracisines
597:Neoaetosauroides
521:possess a short
503:dorsal vertebrae
467:vertebral column
400:caudal vertebrae
261:
239:
232:
221:Stagonolepididae
219:
206:
130:
129:
115:
52:
40:Temporal range:
32:
21:
2212:
2211:
2207:
2206:
2205:
2203:
2202:
2201:
2157:
2156:
2155:
2150:
2142:
2137:
2129:
2124:
2116:
2111:
2103:
2098:
2089:
2088:
2083:
2074:
2073:
2068:
2055:
2045:
2040:
2018:
2011:
1991:
1945:
1943:
1935:
1874:
1872:Desmatosuchinae
1866:
1821:
1802:Rioarribasuchus
1765:
1703:
1661:
1652:
1635:
1614:
1599:
1579:
1548:
1542:Aetosauriformes
1507:
1506:
1496:
1491:
1461:
1460:
1439:(2): e1928681.
1426:
1425:
1418:
1366:
1365:
1354:
1310:
1309:
1298:
1293:
1266:
1261:
1259:
1252:
1247:
1245:
1242:
1237:
1228:
1219:
1210:
1201:
1192:
1183:
1174:
1165:
1156:
1147:
1000:Desmatosuchinae
989:
980:
971:
892:
823:
625:desmatosuchines
619:position among
608:representing a
604:is found to be
558:
487:prezygapophyses
452:
349:
319:stages, making
299:known from the
274:
267:
238:
230:
217:
204:
124:
116:
114:
113:
108:
103:
98:
93:
88:
83:
78:
73:
68:
63:
58:
47:
46:
44:
38:
28:
23:
22:
15:
12:
11:
5:
2210:
2208:
2200:
2199:
2194:
2189:
2184:
2179:
2174:
2169:
2159:
2158:
2152:
2151:
2149:
2148:
2135:
2122:
2109:
2096:
2091:Polesinesuchus
2081:
2065:
2063:
2061:Polesinesuchus
2057:
2056:
2051:
2042:
2041:
2039:
2038:
2027:
2024:
2023:
2020:
2019:
2010:
2008:
2005:
2004:
2001:
2000:
1997:
1996:
1993:
1992:
1990:
1989:
1986:Sierritasuchus
1982:
1975:
1968:
1960:
1952:
1950:
1941:Desmatosuchini
1937:
1936:
1934:
1933:
1925:
1917:
1909:
1901:
1893:
1884:
1878:
1868:
1867:
1865:
1864:
1857:
1854:Polesinesuchus
1849:
1847:
1837:
1831:
1830:
1827:
1826:
1823:
1822:
1820:
1819:
1812:
1805:
1798:
1795:Paratypothorax
1791:
1784:
1775:
1773:
1767:
1766:
1764:
1763:
1756:
1749:
1742:
1734:
1726:
1718:
1712:
1709:Typothoracinae
1705:
1704:
1702:
1701:
1693:
1685:
1677:
1671:
1662:
1657:
1654:
1653:
1651:
1650:
1642:
1636:
1631:
1624:
1623:
1620:
1619:
1616:
1615:
1608:
1605:
1604:
1601:
1600:
1598:
1597:
1596:
1595:
1589:
1587:
1581:
1580:
1578:
1577:
1576:
1575:
1566:
1560:
1550:
1549:
1545:
1544:
1535:
1526:
1520:
1514:
1505:
1498:
1497:
1492:
1490:
1489:
1482:
1475:
1467:
1459:
1458:
1416:
1352:
1323:(3): 240–278.
1295:
1294:
1292:
1289:
1288:
1287:
1280:
1272:
1271:
1257:
1241:
1238:
1234:
1233:
1230:
1229:
1225:
1224:
1221:
1220:
1216:
1215:
1212:
1211:
1207:
1206:
1203:
1202:
1198:
1197:
1194:
1193:
1189:
1188:
1185:
1184:
1180:
1179:
1176:
1175:
1171:
1170:
1167:
1166:
1162:
1161:
1158:
1157:
1153:
1152:
1149:
1148:
1144:
1143:
1140:
1139:
1130:
1127:
1126:
1123:
1122:
1113:
1108:
1106:
1096:
1095:
1092:
1091:
1082:
1077:
1075:
1072:
1071:
1068:
1067:
1058:
1053:
1051:
1048:
1047:
1044:
1043:
1034:
1029:
1027:
1024:
1023:
1020:
1019:
1010:
1005:
1003:
995:
994:
991:
990:
986:
985:
982:
981:
977:
976:
973:
972:
968:
967:
964:
963:
954:
951:
950:
947:
946:
937:
932:
930:
927:
926:
923:
922:
913:
908:
906:
898:
897:
894:
893:
889:
888:
885:
884:
875:
872:
871:
868:
867:
858:
853:
851:
846:
844:
834:
832:
829:
828:
825:
824:
820:
819:
816:
815:
806:
803:
802:
799:
798:
789:
784:
782:
777:
775:
772:
771:
768:
767:
758:
755:
754:
751:
750:
741:
736:
734:
731:
730:
727:
726:
717:
714:
713:
710:
709:
700:
697:
696:
693:
692:
683:
678:
676:
668:
667:
664:
663:
654:
649:
647:
637:
621:stagonolepidid
570:Polesinesuchus
566:Polesinesuchus
557:
556:Classification
554:
546:Polesinesuchus
542:Polesinesuchus
519:Polesinesuchus
495:Polesinesuchus
471:Polesinesuchus
455:Polesinesuchus
451:
448:
442:stages of the
352:Polesinesuchus
348:
345:
335:Polesinesuchus
321:Polesinesuchus
294:stagonolepidid
282:Polesinesuchus
276:
275:
268:
256:
255:
249:
248:
235:Polesinesuchus
228:
224:
223:
215:
211:
210:
202:
198:
197:
192:
185:
184:
179:
172:
171:
166:
162:
161:
156:
152:
151:
146:
142:
141:
136:
132:
131:
118:
117:
109:
104:
99:
94:
89:
84:
79:
74:
69:
64:
59:
54:
53:
39:
36:Polesinesuchus
26:
24:
14:
13:
10:
9:
6:
4:
3:
2:
2209:
2198:
2195:
2193:
2190:
2188:
2185:
2183:
2180:
2178:
2175:
2173:
2170:
2168:
2165:
2164:
2162:
2145:
2140:
2136:
2132:
2127:
2123:
2119:
2114:
2110:
2106:
2101:
2097:
2092:
2086:
2082:
2077:
2071:
2067:
2066:
2064:
2062:
2058:
2054:
2049:
2037:
2029:
2028:
2025:
2015:
1988:
1987:
1983:
1981:
1980:
1976:
1974:
1973:
1969:
1966:
1965:
1964:Gorgetosuchus
1961:
1959:
1958:
1957:Desmatosuchus
1954:
1953:
1951:
1948:
1947:sensu stricto
1942:
1938:
1931:
1930:
1926:
1923:
1922:
1918:
1915:
1914:
1913:Chilenosuchus
1910:
1907:
1906:
1905:Calyptosuchus
1902:
1899:
1898:
1894:
1891:
1890:
1889:Adamanasuchus
1886:
1885:
1882:
1879:
1877:
1873:
1869:
1863:
1862:
1858:
1856:
1855:
1851:
1850:
1848:
1845:
1841:
1838:
1836:
1835:Desmatosuchia
1832:
1818:
1817:
1816:Venkatasuchus
1813:
1811:
1810:
1806:
1804:
1803:
1799:
1797:
1796:
1792:
1790:
1789:
1785:
1782:
1781:
1777:
1776:
1774:
1772:
1768:
1762:
1761:
1757:
1755:
1754:
1753:Redondasuchus
1750:
1748:
1747:
1743:
1740:
1739:
1738:Gorgetosuchus
1735:
1732:
1731:
1730:Chilenosuchus
1727:
1725:
1724:
1720:
1719:
1716:
1713:
1710:
1706:
1699:
1698:
1694:
1691:
1690:
1689:Coahomasuchus
1686:
1684:
1683:
1679:
1678:
1675:
1672:
1670:
1666:
1663:
1660:
1655:
1649:
1648:
1647:Aetosauroides
1644:
1643:
1640:
1637:
1634:
1629:
1625:
1613:
1612:
1606:
1593:
1592:
1591:
1590:
1588:
1586:
1582:
1574:
1570:
1569:
1568:
1567:
1564:
1561:
1559:
1555:
1551:
1543:
1539:
1536:
1534:
1530:
1527:
1525:
1521:
1519:
1515:
1513:
1509:
1508:
1503:
1499:
1495:
1488:
1483:
1481:
1476:
1474:
1469:
1468:
1465:
1454:
1450:
1446:
1442:
1438:
1434:
1430:
1423:
1421:
1417:
1412:
1408:
1404:
1400:
1395:
1390:
1386:
1382:
1378:
1374:
1370:
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1129:
1128:
1125:
1124:
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1120:
1119:
1112:
1111:
1104:
1103:
1102:Desmatosuchus
1098:
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1088:
1081:
1080:
1074:
1073:
1070:
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919:
912:
911:
904:
900:
899:
896:
895:
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883:
882:
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880:Redondasuchus
874:
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869:
866:
865:
864:
857:
856:
850:
849:
842:
838:
837:
831:
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592:Aetosauroides
589:
585:
581:
580:
575:
571:
567:
563:
555:
553:
551:
550:Aetosauroides
547:
543:
539:
538:Aetosauroides
535:
534:Aetosauroides
531:
530:
529:Aetosauroides
524:
520:
516:
512:
508:
504:
500:
496:
492:
488:
484:
480:
476:
472:
468:
465:in the whole
464:
460:
456:
449:
447:
445:
444:Late Triassic
441:
438:and earliest
437:
433:
429:
425:
421:
417:
413:
409:
405:
401:
397:
393:
389:
385:
381:
377:
376:parietal bone
373:
369:
365:
361:
357:
353:
346:
344:
342:
341:
340:Aetosauroides
336:
332:
331:South America
328:
327:
322:
318:
314:
310:
306:
302:
301:Late Triassic
298:
295:
291:
288:
284:
283:
272:
266:
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257:
254:
250:
246:
243:
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128:
123:
119:
112:
107:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
51:
43:
42:Late Triassic
37:
33:
30:
19:
2060:
1984:
1977:
1970:
1962:
1955:
1946:
1927:
1919:
1911:
1903:
1895:
1887:
1875:
1861:Stagonolepis
1859:
1853:
1852:
1814:
1809:Tecovasuchus
1807:
1800:
1793:
1786:
1778:
1758:
1751:
1746:Kryphioparma
1744:
1736:
1728:
1723:Apachesuchus
1721:
1695:
1687:
1680:
1669:Aetosaurinae
1645:
1609:
1573:Pseudosuchia
1558:Pseudosuchia
1537:
1533:Pseudosuchia
1528:
1436:
1432:
1428:
1394:11336/150258
1376:
1372:
1368:
1320:
1316:
1282:
1275:
1133:
1132:
1116:
1115:
1100:
1085:
1084:
1061:
1060:
1037:
1036:
1013:
1012:
957:
956:
940:
939:
916:
915:
878:
877:
861:
860:
810:
809:
808:
792:
791:
761:
760:
744:
743:
720:
719:
703:
702:
686:
685:
657:
656:
617:phylogenetic
606:paraphyletic
602:Aetosaurinae
595:
591:
577:
574:sister taxon
569:
565:
562:phylogenetic
559:
549:
545:
541:
537:
533:
527:
518:
494:
470:
454:
453:
363:
360:type species
351:
350:
338:
334:
325:
324:
320:
311:of the late
303:of southern
281:
280:
279:
270:
263:
262:
253:Type species
241:
234:
233:
195:Pseudosuchia
188:
175:
35:
29:
2085:Wikispecies
1972:Longosuchus
1788:Kocurypelta
1338:11336/20729
1284:Ixalerpeton
1277:Buriolestes
491:diapophyses
450:Description
384:postcranial
382:as well as
182:Archosauria
2161:Categories
1979:Lucasuchus
1876:sensu lato
1780:Garzapelta
1760:Typothorax
1682:Aetosaurus
1633:Aetosauria
1611:Aetosauria
1594:see below↓
1585:Aetosauria
1524:Sauropsida
1494:Aetosauria
1291:References
863:Typothorax
644:Aetosauria
428:osteoderms
315:and early
208:Aetosauria
2167:Aetosaurs
2076:Q15726984
1697:Stenomyti
1510:Kingdom:
1453:237518035
1411:237372648
633:cladogram
590:included
475:aetosaurs
380:braincase
347:Discovery
145:Kingdom:
139:Eukaryota
2070:Wikidata
2036:Category
1518:Chordata
1516:Phylum:
1512:Animalia
1403:34468486
1347:24870635
1240:See also
479:ontogeny
459:juvenile
404:scapulae
388:cervical
378:and the
372:skeleton
368:holotype
358:and the
297:aetosaur
214:Family:
169:Reptilia
159:Chordata
155:Phylum:
149:Animalia
135:Domain:
2131:6149724
2118:1483593
2105:8438088
1929:Scutarx
1522:Class:
1317:Zootaxa
1105:
1002:
905:
843:
675:
646:
523:humerus
515:scapula
436:Carnian
420:ischium
408:humerus
402:, both
313:Carnian
287:extinct
227:Genus:
201:Order:
165:Class:
2144:333722
1451:
1409:
1401:
1345:
1099:
998:
901:
839:
671:
642:
507:centra
463:centra
440:Norian
410:, the
396:sacral
392:dorsal
317:Norian
305:Brazil
285:is an
271:et al.
242:et al.
2113:IRMNG
1538:Clade
1529:Clade
1449:S2CID
1407:S2CID
614:basal
610:grade
588:clade
511:fossa
493:. In
485:show
424:tibia
416:pubis
412:ilium
290:genus
189:Clade
176:Clade
2100:GBIF
1571:see
1399:PMID
1343:PMID
1321:3764
627:and
594:and
422:and
406:, a
398:and
356:2014
273:2014
245:2014
56:PreꞒ
1441:doi
1389:hdl
1381:doi
1333:hdl
1325:doi
576:of
362:is
292:of
2163::
2141::
2128::
2115::
2102::
2087::
2072::
1540::
1531::
1447:.
1437:41
1435:.
1419:^
1405:.
1397:.
1387:.
1377:93
1375:.
1355:^
1341:.
1331:.
1319:.
1315:.
1299:^
560:A
552:.
497:,
469:.
446:.
418:,
414:,
394:,
390:,
343:.
191::
178::
106:Pg
50:Ma
1967:?
1949:)
1932:?
1924:?
1916:?
1908:?
1900:?
1892:?
1846:?
1783:?
1741:?
1733:?
1711:?
1700:?
1692:?
1486:e
1479:t
1472:v
1455:.
1443::
1413:.
1391::
1383::
1349:.
1335::
1327::
260:†
231:†
218:†
205:†
111:N
101:K
96:J
91:T
86:P
81:C
76:D
71:S
66:O
61:Ꞓ
45:~
20:)
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