3718:
3741:
3733:
3137:
1782:
463:
3726:
4601:
2522:
160:
882:
4587:
130:
4650:
3122:
3130:
1024:
773:
4594:
1882:
and dinosaurs achieved a bipedal posture as their legs increased in size, their hips strengthened, and their spines adapted for dorsoventral flexion. Other adaptations that may have facilitated bipedal locomotion include the development of a chambered heart and lungs with unidirectional airflow (both
1797:
in 1915, he described it as "a well-muscled creature light in weight, possibly bipedal in gait occasionally, and most assuredly swift in movement." Mehl based this description on its long limb bones and deep hip socket, two features which link it with bipedal dinosaurs. Since the 1970s however,
1877:
in birds and crocodilians (most likely), phylogenetic bracketing implies that they inherited this trait from their common ancestor and that this ancestor also had an erect stance. With this reasoning, the first archosaurs are thought to have had the ability to high walk.
921:
vertebrae connecting the spine to the hip, three more than most early archosaurs. The hind legs are about twice as long as the arms and placed close together. Five digits are present on the foot, but the fifth is reduced to a small splint of bone next to the
2270:
Parker WG and
Nesbitt 2013. Cranial remains of Poposaurus gracilis (Pseudosuchia: Poposauroidea) from the Upper Triassic, the distribution of the taxon, and its implications for poposauroid evolution. Geological Society, London, Special Publications 379: 22
509:
based on more complete material from the Popo Agie
Formation, including vertebrae, hips, and limb bones. He cited the holotype as 602, but in fact the holotype is UR 357. Mehl concluded that the ilium described by Lees, UR 358, also belonged to
1894:
was hypothesized in a 2011 study that examined muscle scars on the bones and made inferences based on phylogenetic bracketing. 26 muscles, three ligaments, and two connective tissue structures were described. While the hypothesized muscles of
552:
and theropod dinosaurs, including its hollow leg bones and deep hip socket, but did not consider it a dinosaur because each sacral vertebra supported only one rib (theropods usually have multiple ribs projecting from each sacral vertebra).
1845:
or hip socket faces downward and is positioned directly over the head of the femur. In contrast, dinosaurs have "buttress erect" hip structures in which the acetabulum faces laterally and the head of the femur is angled to fit into it.
2230:
2483:
2478:
584:. To Nopsca, Poposauroidea was one of three suborders that made up the order Ornithopoda. Over the following years, many paleontologists supported this classification. For example, German paleontologist
1903:
is thought to have had a puboischiotibialis muscle, but this muscle is absent in birds and probably non-avian dinosaurs as well. The extensor digitorum brevis was probably present on the foot of
1814:
was expected to be similar in appearance to its relatives. In 2011, the nearly complete subadult specimen YPM VP 057100 was described. The specimen confirmed Mehl's description, revealing that
1853:
and dinosaurs inherited a propensity for erect hind-limb driven movement from an early archosaur ancestor. The posture of this ancestral archosaur can be inferred from a method called
1829:
and early dinosaurs were both bipedal, the method of locomotion evolved independently in each group. The independent origins are shown through several differences in the skeletons of
4774:
1869:, meaning that the first archosaur was the last common ancestor of all birds and crocodilians. All birds have a fixed erect stance, and crocodilians have the ability to
837:
785:
4715:
2508:
1802:
has been considered a pseudosuchian archosaur more closely related to crocodilians than dinosaurs. Most of its close relatives (such as the large-bodied
4794:
4804:
4702:
1919:. The hip origin for this muscle is considered to be the original condition for archosaurs, since it is also seen in birds and non-avian dinosaurs.
1915:
differ from those of crocodilians. For example, the puboischiofemoralis internus muscle originates on the spine in crocodilians and on the hip in
4784:
2555:
442:
was an obligate biped, meaning that it walked on two legs rather than four. However, as a pseudosuchian, it is more closely related to living
518:
as a phytosaur because the shape of its ilium was different and it had more sacral vertebrae fused to the hip. Mehl made comparisons between
4789:
3740:
3717:
4809:
3136:
863:
of
Virginia, marking the first occurrence of this genus from eastern North America. This material was found at the same locality as
833:
2342:(Pseudosuchia: Poposauroidea) and its implications for accessory intervertebral articulation evolution in pseudosuchian archosaurs"
4600:
3732:
375:, an unusual group of Triassic pseudosuchians that includes sail-backed, beaked, and aquatic forms. Fossils have been found in
801:
670:, but Galton interpreted this to be the fused end of the hip's pubis bones. Galton noted similarities between the hips of
2207:
631:
could not have been a more primitive archosaur because it had hollow leg bones and complex vertebrae. He placed it in the
3725:
897:
was about 4–5 metres (13–16 ft) long and weighed 90 to 100 kilograms (200 to 220 lb) as an adult. The body of
840:, Utah. It includes the forelimbs, hind limbs, hips, ribs, dorsal vertebrae, and much of the tail. Another specimen of
4799:
2497:
1924:
1854:
159:
4586:
2548:
2171:, an early saurischian dinosaur from the Triassic of Brazil, with notes on the Herrerasauridae and Poposauridae".
2661:
3121:
2492:
1983:
Gauthier, J.A.; Nesbitt, S.J.; Schachner, E.R.; Bever, G.S.; Joyce, W.G. (2011). "The bipedal stem crocodilian
869:
and was first mentioned in the 1980 paper describing it, where it was tentatively referred to as indeterminate
1943:
also suggested a distinctive form of respiration that involved abdominal muscles. The study hypothesized that
824:
in that it is larger, it does not have a ridge of bone behind the hip socket, and does not have a pit on the
450:
is thought to have evolved this form of locomotion independently, possibly from early archosaurs' ability to
4653:
4024:
1947:
had an ischiotruncus muscle running from the ischium at the back of the hip, across the pubis, and into the
1837:
has the characteristic crurotarsal ankle of pseudosuchians, usually associated with quadrupedal locomotion.
4670:
3211:
3129:
844:
from
Arizona, PEFO 34865, includes not only postcranial remains but also cranial remains, confirming that
734:
404:
1952:
4779:
4769:
4741:
2541:
2503:
1009:
2396:
Schachner, E.R.; Manning, P.L.; Dodson, P. (2011). "Pelvic and hindlimb myology of the basal archosaur
1810:) were obligate quadrupeds that could not walk on two legs. Although the entire skeleton was unknown,
851:
In 2022, the partial remains (fragmentary thoracic vertebrae and part a right humerus) of an immature
4518:
2101:
2039:
644:
620:
501:
468:
315:
277:
4217:
2493:
A photograph of the ilium and sacrum of the Yale specimen, from the blog "What's in John's freezer"?
592:
in its own suborder of ornithischians, which he called
Poposauria. In 1930, American paleontologist
3167:
3154:
3088:
3020:
2634:
2527:
2235:(Archosauria: Suchia) based on two new specimens from the Late Triassic of the southwestern U.S.A."
1781:
1214:
609:
493:
462:
1927:
that were even larger than dinosaurs, as their insertion site runs along the entire length of the
4593:
4547:
3884:
2425:
2318:
2253:
2188:
2141:
2119:
2063:
2055:
2004:
1874:
326:
154:
4746:
4728:
4720:
4398:
4362:
3874:
2417:
2373:
860:
797:
698:, and grouped them all in Poposauridae. Like paleontologists before him, Galton distinguished
4733:
2281:
Lucas, Spencer G.; Blodgett, Robert B.; Lichtig, Asher J.; Hunt, Adrian P. (August 2, 2022).
4443:
4312:
4295:
4072:
3244:
2646:
2409:
2363:
2353:
2310:
2245:
2180:
2109:
2047:
1996:
1027:
781:
714:
593:
881:
4418:
4173:
4041:
3962:
3939:
3914:
3772:
3759:
3684:
3672:
3664:
3316:
3039:
2971:
2783:
1667:
1083:
1056:
836:
VP 057100, and informally named "the Yale specimen", was found in the Chinle
Formation of
537:
524:
485:
129:
2447:
2105:
2043:
1899:
share many aspects with those of birds, they are more similar to those of crocodilians.
639:
in its own family, Poposauridae. In the same paper, Colbert described an ilium from the
4623:
4480:
4425:
4383:
4342:
4195:
4166:
4145:
4013:
3894:
3803:
3479:
3281:
3274:
3267:
2998:
2911:
2821:
2748:
2368:
2337:
1807:
1152:
999:
964:, shuvosaurids were bipedal. When the specimen YPM VP 057100 was described by Gauthier
953:
605:
1818:
was indeed bipedal. The skeleton preserves both the fore and hind limbs, showing that
4763:
4459:
4373:
4352:
4248:
4136:
4112:
4102:
4062:
3999:
3955:
3948:
3854:
3844:
3823:
3813:
3627:
3612:
3541:
3521:
3428:
3408:
3334:
3253:
3225:
3198:
3070:
3048:
2991:
2865:
2807:
2299:
2192:
2067:
1537:
1272:
1226:
1128:
1111:
949:
676:
601:
581:
565:
372:
365:
243:
87:
41:
2257:
2008:
745:, leading to the mistaken classification. Long and Murry separated poposaurids like
666:
pseudosuchian. In 1915, Mehl described a "distal femur" in the holotype specimen of
4633:
4473:
4411:
4271:
4257:
4159:
4122:
4092:
3985:
3976:
3904:
3642:
3634:
3598:
3582:
3574:
3565:
3551:
3511:
3490:
3448:
3398:
3307:
3288:
3260:
3077:
2984:
2950:
2943:
2936:
2897:
2800:
2792:
2727:
2714:
2673:
2564:
2429:
2300:"An unusual newly discovered archosaur from the Upper Triassic of Virginia, U.S.A."
1616:
1561:
1489:
1200:
1069:
1023:
981:
973:
945:
902:
766:
694:
682:
655:
640:
577:
573:
561:
489:
396:
358:
300:
256:
227:
146:
17:
2208:"Late Triassic (Carnian and Norian) tetrapods from the southwestern United States"
2449:
Anatomical reconstructions of respiratory morphology and hindlimb musculature in
2282:
1911:
is also similar to those of living crocodilians. Other aspects of the muscles of
4638:
4507:
4452:
4322:
4279:
4264:
4233:
4226:
4203:
4187:
4180:
4152:
3833:
3605:
3531:
3500:
3468:
3458:
3388:
3357:
3350:
3296:
3233:
3184:
3063:
2920:
2886:
2876:
2854:
2814:
2758:
2512:
1866:
1858:
1789:, with leg bones (top), superficial muscles (middle), and deep muscles (bottom).
1633:
1513:
1394:
1370:
1289:
1241:
1042:
1004:
957:
705:
In 1995, paleontologists Robert Long and
Phillip Murry described new fossils of
688:
663:
632:
443:
428:
62:
635:, but because its ilium was distinct from all other archosaurs, Colbert placed
4643:
4628:
4618:
4553:
4524:
4495:
4332:
3925:
3649:
3620:
3590:
3438:
3418:
3218:
2619:
2594:
2517:
1955:, the ischiotruncus would contract and compress the trunk, pumping the lungs.
1842:
1803:
1592:
1346:
1168:
986:
923:
870:
772:
758:
585:
569:
107:
72:
4693:
4536:
4466:
3864:
2768:
2603:
1948:
1870:
1013:
927:
909:
are elongated. The end of the pubis forms a distinct hook that is unique to
865:
733:
from
Poposauridae, claiming that the material used in this assignment was a
726:
613:
560:
was assigned to many different groups of reptiles. Hungarian paleontologist
497:
451:
424:
361:
214:
171:
112:
56:
2421:
2377:
998:
and shuvosaurids than it is to any other pseudosuchian. The analysis found
548:
material could belong to the same animal. Mehl noted similarities between
4687:
4432:
4082:
3992:
3793:
3372:
3009:
2957:
2588:
2000:
1987:: inferring function in fossils and innovation in archosaur locomotion".
435:
420:
392:
191:
138:
102:
97:
82:
77:
67:
2123:
2059:
737:, or a collection of bones belonging to different animals. The pubis of
4707:
3657:
3343:
3055:
2928:
2413:
2358:
2322:
2249:
2184:
1411:
906:
856:
825:
718:
533:
384:
376:
201:
117:
92:
45:
2837:
2582:
1096:
918:
721:. Among the new material were parts of the lower limb, including the
181:
49:
4664:
2314:
2085:
2023:
2114:
2051:
901:
is laterally compressed, with a long and narrow hip structure. The
796:
was again described in 2007, along with two new specimens from the
1928:
1780:
1022:
880:
771:
722:
461:
395:. Except for the skull, most parts of the skeleton are known. The
388:
369:
354:
828:
that fits into the ilium. In 2011, a nearly complete specimen of
368:
of the southwestern and eastern United States. It belongs to the
2212:
Bulletin of the New Mexico Museum of
Natural History and Science
1862:
380:
4668:
3757:
3152:
2712:
2575:
2537:
2533:
1907:, but not in birds. The puboischiofemoralis externus muscle of
627:
and classified it as a theropod dinosaur. Colbert thought that
488:. In 1907, paleontologist J. H. Lees described this fossil, an
952:. It is closely related to other Triassic pseudosuchians like
2479:
A photograph of the Yale specimen, copyrighted Emma
Shachner
1873:
with their limbs erect. If an erect stance is considered
930:
bone extends far from the ankle to form a distinct heel.
2453:(Archosauria: Poposauroidea) and related dinosauriformes
1849:
Although they evolved bipedal locomotion independently,
623:
gave an extensive description of the known material of
1951:
bones of the abdomen. In a form of respiration called
893:
With the tail comprising about half the body length,
4504:
4395:
4379:
4369:
4358:
4348:
4338:
4328:
4318:
4308:
4245:
4214:
4133:
4118:
4108:
4098:
4088:
4078:
4068:
4058:
4010:
3973:
3936:
3921:
3910:
3900:
3890:
3880:
3870:
3860:
3850:
3840:
3829:
3819:
3809:
3799:
3789:
3562:
3547:
3537:
3527:
3517:
3507:
3496:
3486:
3475:
3464:
3454:
3444:
3434:
3424:
3414:
3404:
3394:
3384:
3195:
3180:
3036:
2968:
2908:
2893:
2882:
2872:
2861:
2850:
2780:
2764:
2754:
2744:
307:
292:
4677:
4535:
4503:
4494:
4442:
4394:
4293:
4244:
4213:
4132:
4039:
4009:
3972:
3935:
3770:
3683:
3561:
3371:
3333:
3306:
3243:
3194:
3165:
3087:
3035:
3008:
2967:
2907:
2835:
2779:
2725:
2672:
2645:
2618:
2287:. New Mexico Museum of Natural History and Science.
576:. In 1928, Nopcsa placed it in a new family called
2303:Transactions of the American Philosophical Society
1989:Bulletin of the Peabody Museum of Natural History
654:In his 1977 study of Late Triassic saurischians,
568:dinosaur in 1921, identifying similarities with
411:, was originally the type species of the genus
1841:also has a "pillar erect" stance in which the
2549:
2090:, a new reptile from the Triassic of Wyoming"
8:
838:Grand Staircase–Escalante National Monument
786:Grand Staircase–Escalante National Monument
505:. In 1915, paleontologist M. G. Mehl named
4665:
4500:
4403:
4301:
4051:
4047:
3782:
3778:
3767:
3754:
3377:
3203:
3173:
3162:
3149:
2976:
2843:
2737:
2733:
2722:
2709:
2651:
2624:
2615:
2572:
2556:
2542:
2534:
2456:(Ph.D. thesis). University of Pennsylvania
128:
31:
4775:Late Triassic archosaurs of North America
2367:
2357:
2229:Weinbaum, J.C.; Hungerbühler, A. (2007).
2113:
1978:
1976:
1974:
1972:
1970:
1968:
1939:The 2011 study of the leg musculature of
948:, part of the larger pseudosuchian group
2441:
2439:
2391:
2389:
2387:
702:based on the unique shape of its ilium.
2135:
2133:
2079:
2077:
1964:
980:was placed within Poposauroidea as the
2336:Stefanic, C.M.; Nesbitt, S.J. (2018).
990:and the shuvosaurids. This means that
913:and a few other early pseudosuchians.
472:by J. H. Lees and later identified as
808:was reclassified as a new species of
528:, also from the Triassic of Wyoming.
7:
1785:The hypothesized leg musculature of
1608:
1584:
1577:
1553:
1529:
1505:
1481:
1386:
1362:
1338:
1314:
1264:
1257:
1233:
1221:
1192:
1144:
1103:
1076:
1049:
1038:
496:, and identified it as that of the
419:has been variously classified as a
1887:through phylogenetic bracketing).
848:was a hyper-carnivorous predator.
532:was only known from some teeth, a
25:
1833:and dinosaurs. Unlike dinosaurs,
1822:had much shorter arms than legs.
619:In 1961, American paleontologist
608:dinosaurs. German paleontologist
4795:Prehistoric pseudosuchian genera
4649:
4648:
4599:
4592:
4585:
3739:
3731:
3724:
3716:
3135:
3128:
3120:
2520:
2206:Long, R.A.; Murry, P.A. (1995).
1883:of which are assumed present in
984:to the large-bodied herbivorous
765:, which they held in the family
466:An ilium (hip bone) assigned to
415:. Since it was first described,
158:
4805:Carnian genus first appearances
2400:(archosauria: Poposauroidea)".
1855:extant phylogenetic bracketing
855:were described from the early
1:
4785:Fossil taxa described in 1915
804:of Arizona. Long and Murry's
540:, so Mehl suggested that the
403:, was described and named by
145:(1915) in the collections of
2238:Paläontologische Zeitschrift
2173:Paläontologische Zeitschrift
1793:When M. G. Mehl first named
137:Illustration of UR 357, the
1012:poposauroids. Below is the
994:is more closely related to
612:considered it a very early
492:(part of the hip) from the
407:in 1915. A second species,
4826:
4790:Taxa named by Maurice Mehl
2488:feet, from the same source
944:is a member of the family
580:and a new suborder called
27:Extinct genus of Archosaur
4613:
4582:
4406:
4304:
4054:
4050:
3785:
3781:
3766:
3753:
3713:
3380:
3206:
3176:
3161:
3148:
3117:
2979:
2846:
2740:
2736:
2721:
2708:
2654:
2627:
2614:
2571:
2167:Galton, P.M. (1977). "On
1665:
1630:
1613:
1606:
1589:
1582:
1575:
1558:
1551:
1534:
1527:
1510:
1503:
1486:
1479:
1408:
1391:
1384:
1367:
1360:
1343:
1336:
1319:
1312:
1286:
1269:
1262:
1255:
1238:
1231:
1219:
1197:
1190:
1166:
1149:
1142:
1125:
1108:
1101:
1081:
1074:
1054:
1047:
780:have been found from the
332:
325:
289:
284:
155:Scientific classification
153:
136:
127:
34:
4810:Norian genus extinctions
2446:Schachner, E.R. (2010).
1008:to be successively more
556:In the following years,
484:were found in 1904 near
2484:An articulated pair of
2338:"The axial skeleton of
2142:"The Triassic reptile,
1923:is thought to have had
1890:The leg musculature of
741:was in fact a pubis of
647:, which he assigned to
3212:Benggwigwishingasuchus
2140:Colbert, E.H. (1961).
2094:The Journal of Geology
2032:The Journal of Geology
2028:, a Wyoming phytosaur"
1790:
1035:
890:
806:Lythrosuchus langstoni
792:The known material of
789:
753:, and the newly named
522:and the earlier named
514:. He did not classify
477:
405:Maurice Goldsmith Mehl
4742:Paleobiology Database
2504:Paleobiology Database
2402:Journal of Morphology
1861:represented today by
1784:
1026:
884:
775:
480:The first remains of
465:
2340:Poposaurus langstoni
2298:R. E. Weems (1980).
2169:Staurikosaums pricei
2001:10.3374/014.052.0102
889:relative to a human.
645:Howard County, Texas
621:Edwin Harris Colbert
564:classified it as an
502:Paleorhinus bransoni
469:Paleorhinus bransoni
339:Long and Murry, 1995
3168:Paracrocodylomorpha
3155:Paracrocodylomorpha
3089:Paracrocodylomorpha
2528:Paleontology portal
2451:Poposaurus gracilis
2398:Poposaurus gracilis
2233:Poposaurus gracilis
2106:1915JG.....23..516M
2088:Poposaurus gracilis
2084:Mehl, M.G. (1915).
2044:1907JG.....15..121L
2022:Lees, J.H. (1907).
1985:Poposaurus gracilis
1953:cuirassal breathing
1857:. Archosauria is a
1215:Paracrocodylomorpha
778:Poposaurus gracilis
610:Friedrich von Huene
494:Popo Agie Formation
474:Poposaurus gracilis
446:than to dinosaurs.
143:Poposaurus gracilis
18:Poposaurus gracilis
4800:Late Triassic life
3885:Pseudhesperosuchus
2414:10.1002/jmor.10997
2359:10.7717/peerj.4235
2250:10.1007/BF02988388
2185:10.1007/BF02986571
1791:
1036:
972:was included in a
891:
790:
536:, and part of the
478:
4757:
4756:
4729:Open Tree of Life
4671:Taxon identifiers
4662:
4661:
4609:
4608:
4580:
4579:
4576:
4575:
4572:
4571:
4568:
4567:
4564:
4563:
4490:
4489:
4399:Shartegosuchoidea
4363:Pachycheilosuchus
4289:
4288:
4035:
4034:
3875:Phyllodontosuchus
3749:
3748:
3711:
3710:
3707:
3706:
3703:
3702:
3367:
3366:
3144:
3143:
3115:
3114:
3111:
3110:
3107:
3106:
3031:
3030:
2831:
2830:
2704:
2703:
2700:
2699:
2696:
2695:
2692:
2691:
2408:(12): 1464–1491.
2150:Fieldiana Geology
1768:
1767:
1759:
1758:
1750:
1749:
1741:
1740:
1732:
1731:
1723:
1722:
1714:
1713:
1705:
1704:
1696:
1695:
1687:
1686:
1678:
1677:
1654:
1653:
1645:
1644:
1468:
1467:
1459:
1458:
1450:
1449:
1441:
1440:
1432:
1431:
1423:
1422:
1301:
1300:
1179:
1178:
861:Doswell Formation
800:of Texas and the
798:Tecovas Formation
346:
345:
340:
319:
314:(Long and Murry,
304:
280:
16:(Redirected from
4817:
4750:
4749:
4737:
4736:
4724:
4723:
4711:
4710:
4698:
4697:
4696:
4666:
4652:
4651:
4604:
4603:
4596:
4590:
4589:
4506:
4501:
4444:Shartegosuchidae
4404:
4397:
4381:
4371:
4360:
4350:
4340:
4330:
4320:
4313:Dianchungosaurus
4310:
4302:
4296:Mesoeucrocodylia
4247:
4216:
4135:
4120:
4110:
4100:
4090:
4080:
4073:Eopneumatosuchus
4070:
4060:
4052:
4048:
4012:
3975:
3938:
3923:
3912:
3902:
3892:
3882:
3872:
3862:
3852:
3842:
3831:
3821:
3811:
3801:
3791:
3783:
3779:
3768:
3755:
3744:
3743:
3736:
3735:
3729:
3728:
3721:
3720:
3564:
3549:
3539:
3529:
3519:
3509:
3498:
3488:
3477:
3466:
3456:
3446:
3436:
3426:
3416:
3406:
3396:
3386:
3378:
3245:Ctenosauriscidae
3204:
3197:
3182:
3174:
3163:
3150:
3139:
3133:
3132:
3125:
3124:
3038:
2977:
2970:
2910:
2895:
2884:
2874:
2863:
2852:
2844:
2782:
2766:
2756:
2746:
2738:
2734:
2723:
2710:
2662:Archosauromorpha
2652:
2647:Archosauromorpha
2625:
2616:
2609:
2608:
2573:
2558:
2551:
2544:
2535:
2530:
2525:
2524:
2523:
2466:
2465:
2463:
2461:
2443:
2434:
2433:
2393:
2382:
2381:
2371:
2361:
2333:
2327:
2326:
2295:
2289:
2288:
2278:
2272:
2268:
2262:
2261:
2226:
2220:
2219:
2203:
2197:
2196:
2179:(3–4): 234–245.
2164:
2158:
2157:
2137:
2128:
2127:
2117:
2081:
2072:
2071:
2019:
2013:
2012:
1980:
1925:adductor muscles
1609:
1585:
1578:
1554:
1530:
1506:
1482:
1387:
1363:
1339:
1315:
1265:
1258:
1234:
1222:
1193:
1145:
1104:
1077:
1050:
1039:
1028:Life restoration
1002:and the aquatic
802:Petrified Forest
782:Chinle Formation
715:Chinle Formation
594:Oliver Perry Hay
338:
313:
309:
298:
294:
275:
268:
255:
242:
163:
162:
132:
122:
59:
40:Temporal range:
32:
21:
4825:
4824:
4820:
4819:
4818:
4816:
4815:
4814:
4760:
4759:
4758:
4753:
4745:
4740:
4732:
4727:
4719:
4714:
4706:
4701:
4692:
4691:
4686:
4673:
4663:
4658:
4605:
4598:
4584:
4560:
4531:
4486:
4438:
4419:Shantungosuchus
4390:
4298:
4285:
4240:
4209:
4174:Hemiprotosuchus
4128:
4044:
4042:Crocodyliformes
4031:
4005:
3968:
3963:Terrestrisuchus
3940:Saltoposuchidae
3931:
3915:Syntomiprosopus
3775:
3773:Crocodylomorpha
3762:
3760:Crocodylomorpha
3745:
3738:
3730:
3723:
3715:
3699:
3685:Crocodylomorpha
3679:
3673:Vytshegdosuchus
3665:Vjushkovisaurus
3557:
3363:
3329:
3317:Dolichobrachium
3302:
3239:
3190:
3170:
3157:
3140:
3127:
3119:
3103:
3083:
3040:Gracilisuchidae
3027:
3004:
2972:Aetosauriformes
2963:
2903:
2840:
2827:
2784:Ornithosuchidae
2775:
2730:
2717:
2688:
2668:
2641:
2610:
2578:
2577:
2567:
2562:
2526:
2521:
2519:
2475:
2470:
2469:
2459:
2457:
2445:
2444:
2437:
2395:
2394:
2385:
2335:
2334:
2330:
2315:10.2307/1006472
2297:
2296:
2292:
2284:FOSSIL RECORD 8
2280:
2279:
2275:
2269:
2265:
2231:"A revision of
2228:
2227:
2223:
2205:
2204:
2200:
2166:
2165:
2161:
2139:
2138:
2131:
2083:
2082:
2075:
2021:
2020:
2016:
1982:
1981:
1966:
1961:
1937:
1808:ctenosauriscids
1779:
1774:
1769:
1760:
1751:
1742:
1733:
1724:
1715:
1706:
1697:
1688:
1679:
1668:Crocodylomorpha
1655:
1646:
1469:
1460:
1451:
1442:
1433:
1424:
1302:
1180:
1084:Ornithosuchidae
1057:Avemetatarsalia
1000:ctenosauriscids
954:ctenosauriscids
939:
933:
879:
729:. They removed
546:Dolichobrachium
538:pectoral girdle
530:Dolichobrachium
525:Dolichobrachium
486:Lander, Wyoming
460:
274:
266:
253:
240:
157:
123:
121:
120:
115:
110:
105:
100:
95:
90:
85:
80:
75:
70:
65:
54:
53:
38:
28:
23:
22:
15:
12:
11:
5:
4823:
4821:
4813:
4812:
4807:
4802:
4797:
4792:
4787:
4782:
4777:
4772:
4762:
4761:
4755:
4754:
4752:
4751:
4738:
4725:
4712:
4699:
4683:
4681:
4675:
4674:
4669:
4660:
4659:
4657:
4656:
4646:
4641:
4636:
4631:
4626:
4624:Prestosuchidae
4621:
4614:
4611:
4610:
4607:
4606:
4583:
4581:
4578:
4577:
4574:
4573:
4570:
4569:
4566:
4565:
4562:
4561:
4559:
4558:
4557:
4556:
4550:
4541:
4539:
4533:
4532:
4530:
4529:
4528:
4527:
4521:
4512:
4510:
4498:
4492:
4491:
4488:
4487:
4485:
4484:
4481:Shartegosuchus
4477:
4470:
4463:
4456:
4448:
4446:
4440:
4439:
4437:
4436:
4429:
4426:Sichuanosuchus
4422:
4415:
4407:
4401:
4392:
4391:
4389:
4388:
4384:Thalattosuchia
4377:
4367:
4356:
4346:
4343:Lusitanisuchus
4336:
4326:
4316:
4305:
4299:
4294:
4291:
4290:
4287:
4286:
4284:
4283:
4276:
4268:
4261:
4253:
4251:
4242:
4241:
4239:
4238:
4230:
4222:
4220:
4218:Notochampsidae
4211:
4210:
4208:
4207:
4200:
4196:Pedeticosaurus
4192:
4184:
4177:
4170:
4167:Erythrochampsa
4163:
4156:
4149:
4146:Coloradisuchus
4141:
4139:
4130:
4129:
4127:
4126:
4116:
4106:
4096:
4086:
4076:
4066:
4055:
4045:
4040:
4037:
4036:
4033:
4032:
4030:
4029:
4028:
4027:
4025:Thalattosuchia
4018:
4016:
4014:Thalattosuchia
4007:
4006:
4004:
4003:
3996:
3989:
3981:
3979:
3970:
3969:
3967:
3966:
3959:
3952:
3944:
3942:
3933:
3932:
3930:
3929:
3919:
3908:
3898:
3895:Redondavenator
3888:
3878:
3868:
3858:
3848:
3838:
3827:
3817:
3807:
3804:Dibothrosuchus
3797:
3786:
3776:
3771:
3764:
3763:
3758:
3751:
3750:
3747:
3746:
3714:
3712:
3709:
3708:
3705:
3704:
3701:
3700:
3698:
3697:
3696:
3695:
3689:
3687:
3681:
3680:
3678:
3677:
3669:
3661:
3654:
3646:
3639:
3631:
3624:
3617:
3609:
3602:
3595:
3587:
3579:
3570:
3568:
3559:
3558:
3556:
3555:
3545:
3535:
3525:
3515:
3505:
3494:
3484:
3480:Incertovenator
3473:
3462:
3452:
3442:
3432:
3422:
3412:
3402:
3392:
3381:
3375:
3369:
3368:
3365:
3364:
3362:
3361:
3354:
3347:
3339:
3337:
3331:
3330:
3328:
3327:
3320:
3312:
3310:
3304:
3303:
3301:
3300:
3293:
3285:
3282:Hypselorhachis
3278:
3275:Ctenosauriscus
3271:
3268:Bystrowisuchus
3264:
3257:
3249:
3247:
3241:
3240:
3238:
3237:
3230:
3222:
3215:
3207:
3201:
3192:
3191:
3189:
3188:
3177:
3171:
3166:
3159:
3158:
3153:
3146:
3145:
3142:
3141:
3118:
3116:
3113:
3112:
3109:
3108:
3105:
3104:
3102:
3101:
3100:
3099:
3093:
3091:
3085:
3084:
3082:
3081:
3074:
3067:
3060:
3052:
3044:
3042:
3033:
3032:
3029:
3028:
3026:
3025:
3024:
3023:
3014:
3012:
3006:
3005:
3003:
3002:
2999:Revueltosaurus
2995:
2988:
2980:
2974:
2965:
2964:
2962:
2961:
2954:
2947:
2940:
2933:
2925:
2916:
2914:
2912:Erpetosuchidae
2905:
2904:
2902:
2901:
2891:
2880:
2870:
2859:
2847:
2841:
2836:
2833:
2832:
2829:
2828:
2826:
2825:
2822:Venaticosuchus
2818:
2811:
2804:
2797:
2788:
2786:
2777:
2776:
2774:
2773:
2762:
2752:
2749:Dasygnathoides
2741:
2731:
2726:
2719:
2718:
2713:
2706:
2705:
2702:
2701:
2698:
2697:
2694:
2693:
2690:
2689:
2687:
2686:
2685:
2684:
2678:
2676:
2670:
2669:
2667:
2666:
2665:
2664:
2655:
2649:
2643:
2642:
2640:
2639:
2638:
2637:
2628:
2622:
2612:
2611:
2607:
2606:
2597:
2591:
2585:
2576:
2569:
2568:
2563:
2561:
2560:
2553:
2546:
2538:
2532:
2531:
2516:
2515:
2506:
2495:
2490:
2481:
2474:
2473:External links
2471:
2468:
2467:
2435:
2383:
2328:
2305:. New Series.
2290:
2273:
2263:
2244:(2): 131–145.
2221:
2198:
2159:
2129:
2115:10.1086/622268
2100:(6): 516–522.
2073:
2052:10.1086/621382
2038:(2): 121–151.
2024:"The skull of
2014:
1995:(1): 107–126.
1963:
1962:
1960:
1957:
1936:
1933:
1778:
1775:
1773:
1770:
1766:
1765:
1762:
1761:
1757:
1756:
1753:
1752:
1748:
1747:
1744:
1743:
1739:
1738:
1735:
1734:
1730:
1729:
1726:
1725:
1721:
1720:
1717:
1716:
1712:
1711:
1708:
1707:
1703:
1702:
1699:
1698:
1694:
1693:
1690:
1689:
1685:
1684:
1681:
1680:
1676:
1675:
1672:
1671:
1664:
1661:
1660:
1657:
1656:
1652:
1651:
1648:
1647:
1643:
1642:
1639:
1638:
1629:
1626:
1625:
1622:
1621:
1612:
1607:
1605:
1602:
1601:
1598:
1597:
1588:
1583:
1581:
1576:
1574:
1571:
1570:
1567:
1566:
1557:
1552:
1550:
1547:
1546:
1543:
1542:
1533:
1528:
1526:
1523:
1522:
1519:
1518:
1509:
1504:
1502:
1499:
1498:
1495:
1494:
1485:
1480:
1478:
1475:
1474:
1471:
1470:
1466:
1465:
1462:
1461:
1457:
1456:
1453:
1452:
1448:
1447:
1444:
1443:
1439:
1438:
1435:
1434:
1430:
1429:
1426:
1425:
1421:
1420:
1417:
1416:
1407:
1404:
1403:
1400:
1399:
1390:
1385:
1383:
1380:
1379:
1376:
1375:
1366:
1361:
1359:
1356:
1355:
1352:
1351:
1342:
1337:
1335:
1332:
1331:
1328:
1327:
1318:
1313:
1311:
1308:
1307:
1304:
1303:
1299:
1298:
1295:
1294:
1285:
1282:
1281:
1278:
1277:
1268:
1263:
1261:
1256:
1254:
1251:
1250:
1247:
1246:
1237:
1232:
1230:
1220:
1218:
1210:
1209:
1206:
1205:
1196:
1191:
1189:
1186:
1185:
1182:
1181:
1177:
1176:
1173:
1172:
1165:
1162:
1161:
1158:
1157:
1153:Revueltosaurus
1148:
1143:
1141:
1138:
1137:
1134:
1133:
1124:
1121:
1120:
1117:
1116:
1107:
1102:
1100:
1092:
1091:
1088:
1087:
1080:
1075:
1073:
1065:
1064:
1061:
1060:
1053:
1048:
1046:
1037:
1016:from Gauthier
938:
937:Classification
935:
878:
875:
713:quarry in the
606:sauropodomorph
604:, a family of
459:
456:
353:is an extinct
344:
343:
342:
341:
330:
329:
323:
322:
321:
320:
305:
287:
286:
282:
281:
264:
260:
259:
251:
247:
246:
238:
231:
230:
225:
218:
217:
212:
205:
204:
199:
195:
194:
189:
185:
184:
179:
175:
174:
169:
165:
164:
151:
150:
134:
133:
125:
124:
116:
111:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
60:
39:
26:
24:
14:
13:
10:
9:
6:
4:
3:
2:
4822:
4811:
4808:
4806:
4803:
4801:
4798:
4796:
4793:
4791:
4788:
4786:
4783:
4781:
4778:
4776:
4773:
4771:
4768:
4767:
4765:
4748:
4743:
4739:
4735:
4730:
4726:
4722:
4717:
4713:
4709:
4704:
4700:
4695:
4689:
4685:
4684:
4682:
4680:
4676:
4672:
4667:
4655:
4647:
4645:
4642:
4640:
4637:
4635:
4632:
4630:
4627:
4625:
4622:
4620:
4616:
4615:
4612:
4602:
4597:
4595:
4588:
4555:
4551:
4549:
4545:
4544:
4543:
4542:
4540:
4538:
4534:
4526:
4522:
4520:
4516:
4515:
4514:
4513:
4511:
4509:
4502:
4499:
4497:
4493:
4483:
4482:
4478:
4476:
4475:
4471:
4469:
4468:
4464:
4462:
4461:
4460:Fruitachampsa
4457:
4455:
4454:
4450:
4449:
4447:
4445:
4441:
4435:
4434:
4430:
4428:
4427:
4423:
4421:
4420:
4416:
4414:
4413:
4409:
4408:
4405:
4402:
4400:
4393:
4386:
4385:
4378:
4376:
4375:
4374:Titanochampsa
4368:
4365:
4364:
4357:
4355:
4354:
4353:Neuquensuchus
4347:
4345:
4344:
4337:
4335:
4334:
4327:
4325:
4324:
4317:
4315:
4314:
4307:
4306:
4303:
4300:
4297:
4292:
4282:
4281:
4277:
4274:
4273:
4269:
4267:
4266:
4262:
4260:
4259:
4255:
4254:
4252:
4250:
4249:Gobiosuchidae
4243:
4236:
4235:
4231:
4229:
4228:
4224:
4223:
4221:
4219:
4212:
4206:
4205:
4201:
4198:
4197:
4193:
4190:
4189:
4185:
4183:
4182:
4178:
4176:
4175:
4171:
4169:
4168:
4164:
4162:
4161:
4157:
4155:
4154:
4150:
4148:
4147:
4143:
4142:
4140:
4138:
4137:Protosuchidae
4131:
4125:
4124:
4117:
4115:
4114:
4113:Stegomosuchus
4107:
4105:
4104:
4103:Platyognathus
4097:
4095:
4094:
4087:
4085:
4084:
4077:
4075:
4074:
4067:
4065:
4064:
4063:Entradasuchus
4057:
4056:
4053:
4049:
4046:
4043:
4038:
4026:
4022:
4021:
4020:
4019:
4017:
4015:
4008:
4002:
4001:
4000:Macelognathus
3997:
3995:
3994:
3990:
3988:
3987:
3983:
3982:
3980:
3978:
3971:
3965:
3964:
3960:
3958:
3957:
3956:Saltoposuchus
3953:
3951:
3950:
3949:Litargosuchus
3946:
3945:
3943:
3941:
3934:
3928:
3927:
3920:
3917:
3916:
3909:
3907:
3906:
3899:
3897:
3896:
3889:
3887:
3886:
3879:
3877:
3876:
3869:
3867:
3866:
3859:
3857:
3856:
3855:Kayentasuchus
3849:
3847:
3846:
3845:Junggarsuchus
3839:
3836:
3835:
3828:
3826:
3825:
3824:Hesperosuchus
3818:
3816:
3815:
3814:Dromicosuchus
3808:
3806:
3805:
3798:
3796:
3795:
3788:
3787:
3784:
3780:
3777:
3774:
3769:
3765:
3761:
3756:
3752:
3742:
3737:
3734:
3727:
3719:
3693:
3692:
3691:
3690:
3688:
3686:
3682:
3675:
3674:
3670:
3667:
3666:
3662:
3660:
3659:
3655:
3652:
3651:
3647:
3645:
3644:
3640:
3637:
3636:
3632:
3630:
3629:
3628:Scolotosuchus
3625:
3623:
3622:
3618:
3615:
3614:
3613:Procerosuchus
3610:
3608:
3607:
3603:
3601:
3600:
3596:
3593:
3592:
3588:
3585:
3584:
3580:
3577:
3576:
3572:
3571:
3569:
3567:
3560:
3554:
3553:
3546:
3544:
3543:
3542:Stagonosuchus
3536:
3534:
3533:
3526:
3524:
3523:
3522:Procerosuchus
3516:
3514:
3513:
3506:
3503:
3502:
3495:
3493:
3492:
3485:
3482:
3481:
3474:
3471:
3470:
3463:
3461:
3460:
3453:
3451:
3450:
3443:
3441:
3440:
3433:
3431:
3430:
3429:Decuriasuchus
3423:
3421:
3420:
3413:
3411:
3410:
3409:Batrachotomus
3403:
3401:
3400:
3393:
3391:
3390:
3383:
3382:
3379:
3376:
3374:
3370:
3360:
3359:
3355:
3353:
3352:
3348:
3346:
3345:
3341:
3340:
3338:
3336:
3335:Shuvosauridae
3332:
3326:
3325:
3321:
3319:
3318:
3314:
3313:
3311:
3309:
3305:
3299:
3298:
3294:
3291:
3290:
3286:
3284:
3283:
3279:
3277:
3276:
3272:
3270:
3269:
3265:
3263:
3262:
3258:
3256:
3255:
3254:Arizonasaurus
3251:
3250:
3248:
3246:
3242:
3236:
3235:
3231:
3228:
3227:
3226:Schultzsuchus
3223:
3221:
3220:
3216:
3214:
3213:
3209:
3208:
3205:
3202:
3200:
3199:Poposauroidea
3193:
3187:
3186:
3179:
3178:
3175:
3172:
3169:
3164:
3160:
3156:
3151:
3147:
3138:
3134:
3131:
3123:
3097:
3096:
3095:
3094:
3092:
3090:
3086:
3080:
3079:
3075:
3073:
3072:
3071:Turfanosuchus
3068:
3066:
3065:
3061:
3058:
3057:
3053:
3051:
3050:
3049:Gracilisuchus
3046:
3045:
3043:
3041:
3034:
3022:
3018:
3017:
3016:
3015:
3013:
3011:
3007:
3001:
3000:
2996:
2994:
2993:
2992:Euscolosuchus
2989:
2987:
2986:
2982:
2981:
2978:
2975:
2973:
2966:
2960:
2959:
2955:
2953:
2952:
2948:
2946:
2945:
2941:
2939:
2938:
2934:
2931:
2930:
2926:
2923:
2922:
2918:
2917:
2915:
2913:
2906:
2900:
2899:
2892:
2889:
2888:
2881:
2879:
2878:
2871:
2868:
2867:
2866:Koilamasuchus
2860:
2857:
2856:
2849:
2848:
2845:
2842:
2839:
2834:
2824:
2823:
2819:
2817:
2816:
2812:
2810:
2809:
2808:Ornithosuchus
2805:
2803:
2802:
2798:
2795:
2794:
2790:
2789:
2787:
2785:
2778:
2771:
2770:
2763:
2761:
2760:
2753:
2751:
2750:
2743:
2742:
2739:
2735:
2732:
2729:
2724:
2720:
2716:
2711:
2707:
2682:
2681:
2680:
2679:
2677:
2675:
2671:
2663:
2659:
2658:
2657:
2656:
2653:
2650:
2648:
2644:
2636:
2632:
2631:
2630:
2629:
2626:
2623:
2621:
2617:
2613:
2605:
2601:
2598:
2596:
2592:
2590:
2586:
2584:
2580:
2579:
2574:
2570:
2566:
2559:
2554:
2552:
2547:
2545:
2540:
2539:
2536:
2529:
2518:
2514:
2510:
2507:
2505:
2501:
2500:
2496:
2494:
2491:
2489:
2487:
2482:
2480:
2477:
2476:
2472:
2455:
2454:
2450:
2442:
2440:
2436:
2431:
2427:
2423:
2419:
2415:
2411:
2407:
2403:
2399:
2392:
2390:
2388:
2384:
2379:
2375:
2370:
2365:
2360:
2355:
2351:
2347:
2343:
2341:
2332:
2329:
2324:
2320:
2316:
2312:
2308:
2304:
2301:
2294:
2291:
2286:
2285:
2277:
2274:
2267:
2264:
2259:
2255:
2251:
2247:
2243:
2239:
2236:
2234:
2225:
2222:
2217:
2213:
2209:
2202:
2199:
2194:
2190:
2186:
2182:
2178:
2174:
2170:
2163:
2160:
2155:
2151:
2147:
2145:
2136:
2134:
2130:
2125:
2121:
2116:
2111:
2107:
2103:
2099:
2095:
2091:
2089:
2080:
2078:
2074:
2069:
2065:
2061:
2057:
2053:
2049:
2045:
2041:
2037:
2033:
2029:
2027:
2018:
2015:
2010:
2006:
2002:
1998:
1994:
1990:
1986:
1979:
1977:
1975:
1973:
1971:
1969:
1965:
1958:
1956:
1954:
1950:
1946:
1942:
1934:
1932:
1930:
1926:
1922:
1918:
1914:
1910:
1906:
1902:
1898:
1893:
1888:
1886:
1881:
1876:
1872:
1868:
1864:
1860:
1856:
1852:
1847:
1844:
1840:
1836:
1832:
1828:
1823:
1821:
1817:
1813:
1809:
1805:
1801:
1796:
1788:
1783:
1776:
1771:
1764:
1763:
1755:
1754:
1746:
1745:
1737:
1736:
1728:
1727:
1719:
1718:
1710:
1709:
1701:
1700:
1692:
1691:
1683:
1682:
1674:
1673:
1670:
1669:
1663:
1662:
1659:
1658:
1650:
1649:
1641:
1640:
1637:
1636:
1635:
1628:
1627:
1624:
1623:
1620:
1619:
1618:
1611:
1610:
1604:
1603:
1600:
1599:
1596:
1595:
1594:
1587:
1586:
1580:
1579:
1573:
1572:
1569:
1568:
1565:
1564:
1563:
1556:
1555:
1549:
1548:
1545:
1544:
1541:
1540:
1539:
1538:Batrachotomus
1532:
1531:
1525:
1524:
1521:
1520:
1517:
1516:
1515:
1508:
1507:
1501:
1500:
1497:
1496:
1493:
1492:
1491:
1484:
1483:
1477:
1476:
1473:
1472:
1464:
1463:
1455:
1454:
1446:
1445:
1437:
1436:
1428:
1427:
1419:
1418:
1415:
1414:
1413:
1406:
1405:
1402:
1401:
1398:
1397:
1396:
1389:
1388:
1382:
1381:
1378:
1377:
1374:
1373:
1372:
1365:
1364:
1358:
1357:
1354:
1353:
1350:
1349:
1348:
1341:
1340:
1334:
1333:
1330:
1329:
1326:
1325:
1324:
1317:
1316:
1310:
1309:
1306:
1305:
1297:
1296:
1293:
1292:
1291:
1284:
1283:
1280:
1279:
1276:
1275:
1274:
1273:Arizonasaurus
1267:
1266:
1260:
1259:
1253:
1252:
1249:
1248:
1245:
1244:
1243:
1236:
1235:
1228:
1227:Poposauroidea
1224:
1223:
1216:
1212:
1211:
1208:
1207:
1204:
1203:
1202:
1195:
1194:
1188:
1187:
1184:
1183:
1175:
1174:
1171:
1170:
1164:
1163:
1160:
1159:
1156:
1155:
1154:
1147:
1146:
1140:
1139:
1136:
1135:
1132:
1131:
1130:
1129:Turfanosuchus
1123:
1122:
1119:
1118:
1115:
1114:
1113:
1112:Gracilisuchus
1106:
1105:
1098:
1094:
1093:
1090:
1089:
1086:
1085:
1079:
1078:
1071:
1067:
1066:
1063:
1062:
1059:
1058:
1052:
1051:
1044:
1041:
1040:
1033:
1029:
1025:
1021:
1019:
1015:
1011:
1007:
1006:
1001:
997:
993:
989:
988:
983:
979:
975:
971:
967:
963:
959:
955:
951:
950:Poposauroidea
947:
943:
936:
934:
931:
929:
925:
920:
916:
912:
908:
904:
900:
896:
888:
883:
876:
874:
872:
868:
867:
862:
858:
854:
849:
847:
843:
839:
835:
831:
827:
823:
820:differs from
819:
815:
811:
807:
803:
799:
795:
787:
783:
779:
774:
770:
768:
764:
760:
756:
752:
748:
744:
740:
736:
732:
728:
724:
720:
716:
712:
708:
703:
701:
697:
696:
691:
690:
685:
684:
679:
678:
677:Arizonasaurus
673:
669:
665:
661:
658:reclassified
657:
652:
650:
646:
642:
638:
634:
630:
626:
622:
617:
615:
611:
607:
603:
602:Anchisauridae
599:
595:
591:
587:
583:
582:Poposauroidea
579:
575:
571:
567:
566:ornithischian
563:
559:
554:
551:
547:
543:
539:
535:
531:
527:
526:
521:
517:
513:
508:
504:
503:
499:
495:
491:
487:
483:
475:
471:
470:
464:
457:
455:
453:
449:
445:
441:
437:
432:
430:
426:
422:
418:
414:
410:
406:
402:
398:
394:
390:
386:
382:
378:
374:
373:Poposauroidea
371:
367:
366:Late Triassic
363:
360:
359:pseudosuchian
356:
352:
351:
337:
334:
333:
331:
328:
324:
317:
312:
306:
302:
297:
291:
290:
288:
283:
279:
273:
272:
265:
262:
261:
258:
252:
249:
248:
245:
244:Poposauroidea
239:
236:
233:
232:
229:
226:
223:
220:
219:
216:
213:
210:
207:
206:
203:
200:
197:
196:
193:
190:
187:
186:
183:
180:
177:
176:
173:
170:
167:
166:
161:
156:
152:
148:
144:
140:
135:
131:
126:
119:
114:
109:
104:
99:
94:
89:
84:
79:
74:
69:
64:
58:
55:237–216
51:
47:
43:
42:Late Triassic
37:
33:
30:
19:
4780:Chinle fauna
4770:Poposauroids
4678:
4634:Sphenosuchia
4591:
4554:crocodilians
4525:notosuchians
4479:
4474:Nominosuchus
4472:
4465:
4458:
4451:
4431:
4424:
4417:
4412:Hadongsuchus
4410:
4382:
4372:
4361:
4351:
4341:
4331:
4321:
4311:
4278:
4272:Lisboasaurus
4270:
4263:
4258:Cassissuchus
4256:
4232:
4225:
4202:
4194:
4186:
4179:
4172:
4165:
4160:Edentosuchus
4158:
4151:
4144:
4123:Tagarosuchus
4121:
4111:
4101:
4093:Microchampsa
4091:
4081:
4071:
4061:
3998:
3991:
3986:Almadasuchus
3984:
3977:Hallopodidae
3961:
3954:
3947:
3924:
3913:
3905:Sphenosuchus
3903:
3893:
3883:
3873:
3863:
3853:
3843:
3832:
3822:
3812:
3802:
3792:
3722:
3671:
3663:
3656:
3648:
3643:Teratosaurus
3641:
3635:Scythosuchus
3633:
3626:
3619:
3611:
3604:
3599:Polonosuchus
3597:
3589:
3583:Energosuchus
3581:
3575:Arganasuchus
3573:
3566:Rauisuchidae
3552:Youngosuchus
3550:
3540:
3530:
3520:
3512:Prestosuchus
3510:
3499:
3491:Luperosuchus
3489:
3478:
3467:
3457:
3449:Fasolasuchus
3447:
3437:
3427:
3417:
3407:
3399:Arganasuchus
3397:
3387:
3356:
3349:
3342:
3323:
3322:
3315:
3308:Poposauridae
3295:
3289:Tsylmosuchus
3287:
3280:
3273:
3266:
3261:Bromsgroveia
3259:
3252:
3232:
3224:
3217:
3210:
3183:
3126:
3078:Yonghesuchus
3076:
3069:
3062:
3054:
3047:
2997:
2990:
2985:Acaenasuchus
2983:
2956:
2951:Parringtonia
2949:
2944:Pagosvenator
2942:
2937:Erpetosuchus
2935:
2927:
2919:
2898:Ticinosuchus
2896:
2885:
2875:
2864:
2853:
2820:
2813:
2806:
2801:Dynamosuchus
2799:
2793:Aenigmaspina
2791:
2767:
2757:
2747:
2728:Pseudosuchia
2715:Pseudosuchia
2674:Pseudosuchia
2599:
2565:Pseudosuchia
2509:Poposauridae
2498:
2485:
2458:. Retrieved
2452:
2448:
2405:
2401:
2397:
2349:
2345:
2339:
2331:
2306:
2302:
2293:
2283:
2276:
2266:
2241:
2237:
2232:
2224:
2215:
2211:
2201:
2176:
2172:
2168:
2162:
2153:
2149:
2143:
2097:
2093:
2087:
2035:
2031:
2025:
2017:
1992:
1988:
1984:
1944:
1940:
1938:
1920:
1916:
1912:
1908:
1904:
1900:
1896:
1891:
1889:
1884:
1879:
1867:crocodilians
1850:
1848:
1838:
1834:
1830:
1826:
1824:
1819:
1815:
1811:
1799:
1794:
1792:
1786:
1772:Paleobiology
1666:
1632:
1631:
1617:Polonosuchus
1615:
1614:
1591:
1590:
1562:Fasolasuchus
1560:
1559:
1536:
1535:
1512:
1511:
1490:Prestosuchus
1488:
1487:
1410:
1409:
1393:
1392:
1369:
1368:
1345:
1344:
1322:
1321:
1320:
1288:
1287:
1271:
1270:
1240:
1239:
1201:Ticinosuchus
1199:
1198:
1167:
1151:
1150:
1127:
1126:
1110:
1109:
1082:
1070:Pseudosuchia
1055:
1031:
1017:
1003:
995:
991:
985:
982:sister taxon
977:
974:phylogenetic
969:
965:
961:
958:shuvosaurids
946:Poposauridae
941:
940:
932:
914:
910:
898:
894:
892:
886:
864:
852:
850:
845:
841:
829:
821:
818:P. langstoni
817:
814:P. langstoni
813:
809:
805:
793:
791:
777:
767:Rauisuchidae
762:
759:rauisuchians
755:Lythrosuchus
754:
751:Bromsgroveia
750:
746:
742:
738:
730:
710:
706:
704:
699:
695:Teratosaurus
693:
687:
683:Bromsgroveia
681:
675:
671:
667:
659:
656:Peter Galton
653:
648:
641:Dockum Group
636:
628:
624:
618:
597:
589:
578:Poposauridae
562:Franz Nopcsa
557:
555:
549:
545:
541:
529:
523:
519:
515:
511:
506:
500:
481:
479:
473:
467:
447:
444:crocodilians
439:
433:
416:
413:Lythrosuchus
412:
409:P. langstoni
408:
400:
397:type species
349:
348:
347:
336:Lythrosuchus
335:
311:P. langstoni
310:
299:Mehl, 1915 (
295:
270:
269:
257:Poposauridae
234:
228:Pseudosuchia
221:
208:
147:Field Museum
142:
141:specimen of
35:
29:
4639:Protosuchia
4519:Ziphosuchia
4508:Ziphosuchia
4453:Adzhosuchus
4323:Hsisosuchus
4280:Zaraasuchus
4265:Gobiosuchus
4234:Orthosuchus
4227:Notochampsa
4204:Protosuchus
4188:Orthosuchus
4181:Hoplosuchus
4153:Dianosuchus
3834:Heteropelta
3606:Postosuchus
3532:Saurosuchus
3501:Mandasuchus
3469:Heteropelta
3459:Heptasuchus
3389:Apatosuchus
3358:Sillosuchus
3351:Shuvosaurus
3297:Xilousuchus
3234:Qianosuchus
3185:Mambawakale
3064:Parvosuchus
2921:Archeopelta
2887:Nundasuchus
2877:Mandasuchus
2855:Heteropelta
2815:Riojasuchus
2769:Phytosauria
2759:Nundasuchus
2604:Archosauria
2513:Palaeos.com
2309:(7): 1–53.
2156:(4): 59–78.
2026:Paleorhinus
1859:crown group
1804:rauisuchids
1787:P. gracilis
1634:Postosuchus
1514:Saurosuchus
1395:Shuvosaurus
1371:Sillosuchus
1290:Xilousuchus
1242:Qianosuchus
1043:Archosauria
1032:P. gracilis
1005:Qianosuchus
924:metatarsals
887:P. gracilis
877:Description
871:rauisuchian
853:P. gracilis
830:P. gracilis
822:P. gracilis
776:Fossils of
763:Postosuchus
739:Postosuchus
731:Postosuchus
689:Postosuchus
649:P. gracilis
633:Carnosauria
588:classified
574:camptosaurs
570:iguanodonts
438:dinosaurs,
429:rauisuchian
401:P. gracilis
296:P. gracilis
215:Archosauria
4764:Categories
4679:Poposaurus
4644:Mesosuchia
4629:Bathyotica
4619:Rauisuchia
4617:See also:
4496:Metasuchia
4333:Induszalim
3926:Trialestes
3694:see below↓
3650:Tikisuchus
3621:Rauisuchus
3591:Jushatyria
3439:Etjosuchus
3419:Dagasuchus
3324:Poposaurus
3219:Lotosaurus
3098:see below↓
3021:Aetosauria
3010:Aetosauria
2683:see below↓
2635:Sauropsida
2620:Sauropsida
2595:Sauropsida
2499:Poposaurus
2486:Poposaurus
2460:January 1,
2144:Poposaurus
1959:References
1945:Poposaurus
1941:Poposaurus
1921:Poposaurus
1917:Poposaurus
1913:Poposaurus
1909:Poposaurus
1905:Poposaurus
1901:Poposaurus
1897:Poposaurus
1892:Poposaurus
1885:Poposaurus
1880:Poposaurus
1875:homologous
1851:Poposaurus
1843:acetabulum
1839:Poposaurus
1835:Poposaurus
1831:Poposaurus
1827:Poposaurus
1820:Poposaurus
1816:Poposaurus
1812:Poposaurus
1800:Poposaurus
1795:Poposaurus
1777:Locomotion
1593:Rauisuchus
1347:Lotosaurus
1323:Poposaurus
1169:Aetosauria
996:Lotosaurus
992:Poposaurus
987:Lotosaurus
978:Poposaurus
976:analysis.
970:Poposaurus
962:Poposaurus
942:Poposaurus
915:Poposaurus
911:Poposaurus
899:Poposaurus
895:Poposaurus
846:Poposaurus
842:Poposaurus
810:Poposaurus
794:Poposaurus
747:Poposaurus
743:Poposaurus
707:Poposaurus
700:Poposaurus
672:Poposaurus
668:Poposaurus
660:Poposaurus
637:Poposaurus
629:Poposaurus
625:Poposaurus
598:Poposaurus
590:Poposaurus
586:Oskar Kuhn
558:Poposaurus
550:Poposaurus
542:Poposaurus
520:Poposaurus
516:Poposaurus
512:Poposaurus
507:Poposaurus
482:Poposaurus
448:Poposaurus
440:Poposaurus
417:Poposaurus
350:Poposaurus
271:Poposaurus
36:Poposaurus
4552:includes
4548:Neosuchia
4537:Neosuchia
4523:includes
4467:Kyasuchus
3865:Parrishia
2581:Kingdom:
2352:. e4235.
2193:129438498
2068:129850970
1949:gastralia
1935:Breathing
1871:high walk
1825:Although
1014:cladogram
968:in 2011,
928:calcaneum
917:has five
873:remains.
866:Doswellia
832:known as
727:calcaneum
711:Placerias
709:from the
664:thecodont
616:in 1950.
614:stegosaur
498:phytosaur
452:high walk
427:, and a "
425:phytosaur
364:from the
362:archosaur
178:Kingdom:
172:Eukaryota
4688:Wikidata
4654:Category
4433:Zosuchus
4083:Kemkemia
3993:Hallopus
3794:Carnufex
3373:Loricata
2958:Tarjadia
2589:Chordata
2587:Phylum:
2583:Animalia
2422:21800358
2378:29472991
2258:84822632
2218:: 1–254.
2124:30067173
2060:30056366
2009:86687464
1020:(2011):
885:Size of
436:theropod
421:dinosaur
393:Virginia
327:Synonyms
285:Species
250:Family:
202:Reptilia
192:Chordata
188:Phylum:
182:Animalia
168:Domain:
139:holotype
4734:4130206
4721:1129969
4708:4819188
4694:Q135726
3658:Vivaron
3344:Effigia
3056:Maehary
2929:Dyoplax
2593:Class:
2502:in the
2430:4651357
2369:5816584
2323:1006472
2102:Bibcode
2040:Bibcode
1412:Effigia
1229:
1217:
1099:
1072:
1045:
960:. Like
926:. The
907:ischium
857:Carnian
826:ischium
788:, Utah.
735:chimera
719:Arizona
596:placed
534:humerus
458:History
385:Arizona
377:Wyoming
263:Genus:
198:Class:
46:Carnian
2838:Suchia
2428:
2420:
2376:
2366:
2321:
2256:
2191:
2122:
2066:
2058:
2007:
1225:
1213:
1097:Suchia
1095:
1068:
1018:et al.
966:et al.
919:sacral
859:-aged
692:, and
391:, and
276:Mehl,
50:Norian
4747:38263
4716:IRMNG
2600:Clade
2426:S2CID
2346:PeerJ
2319:JSTOR
2254:S2CID
2189:S2CID
2120:JSTOR
2064:S2CID
2056:JSTOR
2005:S2CID
1929:femur
1863:birds
1010:basal
903:pubis
761:like
757:from
723:tibia
662:as a
490:ilium
434:Like
389:Texas
370:clade
355:genus
235:Clade
222:Clade
209:Clade
4703:GBIF
4546:see
4517:see
4023:see
3019:see
2660:see
2633:see
2462:2010
2418:PMID
2374:PMID
1865:and
1806:and
956:and
905:and
725:and
572:and
544:and
423:, a
381:Utah
316:1995
301:type
278:1915
63:PreꞒ
2511:at
2410:doi
2406:272
2364:PMC
2354:doi
2311:doi
2271:pp.
2246:doi
2181:doi
2110:doi
2048:doi
1997:doi
1030:of
834:YPM
784:in
717:of
643:of
600:in
431:".
357:of
52:),
48:to
4766::
4744::
4731::
4718::
4705::
4690::
2602::
2438:^
2424:.
2416:.
2404:.
2386:^
2372:.
2362:.
2348:.
2344:.
2317:.
2307:70
2252:.
2242:81
2240:.
2214:.
2210:.
2187:.
2177:51
2175:.
2154:14
2152:.
2148:.
2132:^
2118:.
2108:.
2098:23
2096:.
2092:.
2076:^
2062:.
2054:.
2046:.
2036:15
2034:.
2030:.
2003:.
1993:52
1991:.
1967:^
1931:.
816:.
812:,
769:.
749:,
686:,
680:,
674:,
651:.
454:.
399:,
387:,
383:,
379:,
237::
224::
211::
149:.
113:Pg
57:Ma
4505:†
4396:†
4387:?
4380:†
4370:†
4366:?
4359:†
4349:†
4339:†
4329:†
4319:†
4309:†
4275:?
4246:†
4237:?
4215:†
4199:?
4191:?
4134:†
4119:†
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4099:†
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3911:†
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3830:†
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3790:†
3676:?
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3616:?
3594:?
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3563:†
3548:†
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3528:†
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3508:†
3504:?
3497:†
3487:†
3483:?
3476:†
3472:?
3465:†
3455:†
3445:†
3435:†
3425:†
3415:†
3405:†
3395:†
3385:†
3292:?
3229:?
3196:†
3181:†
3059:?
3037:†
2969:†
2932:?
2924:?
2909:†
2894:†
2890:?
2883:†
2873:†
2869:?
2862:†
2858:?
2851:†
2796:?
2781:†
2772:?
2765:†
2755:†
2745:†
2557:e
2550:t
2543:v
2464:.
2432:.
2412::
2380:.
2356::
2350:6
2325:.
2313::
2260:.
2248::
2216:4
2195:.
2183::
2146:"
2126:.
2112::
2104::
2086:"
2070:.
2050::
2042::
2011:.
1999::
1034:.
476:.
318:)
308:†
303:)
293:†
267:†
254:†
241:†
118:N
108:K
103:J
98:T
93:P
88:C
83:D
78:S
73:O
68:Ꞓ
44:(
20:)
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