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over a distance of less than two centimetres. This crest is at its midline extremely thickened, resulting at the position of the rear frontals bones it overgrowths, in a pointed strongly triangular transverse cross-section, the upper angle of which in rear view is less than 130°. Additionally, these
507:
is a large hadrosaurid. Its holotype is the largest brachysaurolophin specimen known. Because of this size the fossil has been nicknamed "Super-Duck". Its size has been estimated at 10 metres (33 ft) in length and 5 metric tons (5.5 short tons) in body mass. Recent body mass estimate suggests
416:
secured the remainder of a hadrosaur skeleton, among which the skull. The fossil was donated to the Museum of the
Rockies by land owners Nolan and Cheryl Fladstol; and John and Claire Wendland. In 2009, 2010 and 2011, the find was reported in the scientific literature as a possible new species of
1184:
research of the bone structure of the left shinbone of the type specimen. This bone showed fourteen LAGs, lines of arrested growth, that likely represented yearly seasons of low food intake. The age of the holotype would thus be about fourteen years. The distance between the lines indicated this
469:, was found in a layer of the Judith River Formation dating from the Campanian and being between 79.8 and 79.5 million years old, plus or minus two hundred thousand years. It consists of a partial skeleton with skull, of an adult individual. It contains the right premaxilla, both
1327:
Freedman, E.A., and Fowler, D.W., 2010, "Stratigraphic correlation of Judith River
Formation (Campanian, Upper Cretaceous) exposures in Kennedy Coulee (northcentral Montana) to the Foremost Formation (Alberta): implications for anagenesis in hadrosaurid dinosaurs",
1421:"Testing Size–frequency Distributions As a Method of Ontogenetic Aging: A Life-history Assessment of Hadrosaurid Dinosaurs from the Dinosaur Park Formation of Alberta, Canada, with Implications for Hadrosaurid Paleoecology"
1192:
The lines also showed a general slowing of the growth after the fifth year. This could have been an indication sexual maturity was reached at that age. If so, the onset of maturity was about two years later than with
1292:
Fowler, Elizabeth A. Freedman, and John R. Horner. "A New
Brachylophosaurin Hadrosaur (Dinosauria: Ornithischia) with an Intermediate Nasal Crest from the Campanian Judith River Formation of Northcentral Montana."
473:, the left jugal, a part of the right lacrimal, the rear of a left nasal bone, the middle part of a right nasal bone, the skull roof from the frontal bones to the exoccipitals, both
3924:
405:
485:, nineteen ribs, the entire pelvis, both lower legs, a right second metatarsal and a right fourth metatarsal. The bones have not been found in articulation.
1359:
488:
Specimen MOR 1097, a fragmentary skull of a subadult individual, was referred to the species. It had been found at a kilometre distance from the holotype.
481:, the predentary of the lower jaws, both dentaries, a right surangular, eleven neck vertebrae, eleven back vertebrae, twenty-nine tail vertebrae, nineteen
1311:(Dinosauria: Hadrosauridae): Ontogenetic and stratigraphic implications of a large new specimen from the Judith River Formation of northcentral Montana",
3949:
527:, unique derived characters. The skull crest is made of massive bone and is entirely formed by the nasal bones which, in adult individuals, overhang the
384:
evolving from a crestless ancestor (the former genus) to its crested descendant (the latter genus) while changing the morphology of its nasal bones.
3422:
412:
in the north of
Montana, at a site originally discovered by Kyoko Kishi. After a school class found some more bones, in 2007 and 2008 a team of the
2507:
3934:
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1395:
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432:
1313:
Abstracts of papers, Sixty-ninth Annual
Meeting, Society of Vertebrate Paleontology. Journal of Vertebrate Paleontology
3428:
447:
1345:(Ornithischia: Hadrosauridae) from the Judith River Formation (Late Cretaceous: Campanian) of northcentral Montana",
1185:
individual had not yet reached its maximum size, but closely approached it, proof that it was not simply a subadult
3914:
2514:
1487:
1216:
Thompson, Michael G.W.; Bedek, Fern V.; Schröder-Adams, Claudia; Evans, David C.; Ryan, Michael J. (2021-10-01).
546:. Of the front branch of the jugal, the lower corner is positioned behind the level of the upper corner, as with
1363:
1594:
1291:
455:
372:
The significance of this particular hadrosaur taxon is that it is a transitional species between the genera
665:
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1953:
1805:
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511:
354:
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3763:
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was one of eighteen dinosaur taxa from 2015 to be described in open access or free-to-read journals.
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autapomorphies form a unique combination with traits that in themselves are not unique. The rear
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1932:
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analyses, resulted in slightly conflicting positions. In one analysis it was recovered as the
578:
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1347:
International
Hadrosaur Symposium, Royal Tyrrell Museum, Drumheller, Alberta. Abstract Volume
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2919:
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1961:
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1719:
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562:. The nasal bone skull crest is massive and almost horizontally oriented to the rear as in
458:
honours Sam Berge, one of the landowners, and his friends who have supported the research.
17:
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and the latter, in view of its intermediate smaller crest form, again an ancestor of
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104:
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523:
In 2015, several distinguishing traits were established. Two of these are
3858:
3619:
3288:
3176:
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2947:
2874:
2799:
2764:
2665:
2651:
2609:
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109:
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168:
3835:
1218:"The oldest occurrence of brachylophosaurin hadrosaurids in Canada"
2623:
2153:
1330:
Journal of
Vertebrate Paleontology, SVP Program and Abstracts Book
1171:
632:
595:
587:
577:
510:
495:
446:
and refers to the genus being situated in a lower position in the
436:
391:
3878:
1991:
3839:
3444:
2537:
1672:
1514:
1476:
554:. The squamosals touch each other at the skull midline as with
1472:
508:
that it reached more than 4.3 metric tons (4.7 short tons).
431:
was named and described by
Elizabeth A. Freedman Fowler and
649:
was in a more basal position. The authors suggested that
1419:
Wosik, M.; Chiba, K.; Therrien, F.; Evans, D.C. (2020).
1307:
Freedman, E.A., 2009, "Variation in nasal crest size of
1287:
1285:
1283:
1281:
1279:
1277:
1275:
1273:
1271:
1269:
1267:
1180:
The describing article also published the result of
3848:
3749:
3600:
3580:
3455:
3332:
3298:
3242:
3201:
3145:
3121:
3048:
3007:
2891:
2830:
2550:
2466:
2423:
2357:
2183:
2115:
2071:
2008:
1972:
1931:
1767:
1730:
1685:
1632:
1605:
1578:
615:by the describing authors. Adding the traits of
406:University of California Museum of Paleontology
1488:
668:resulted in the following evolutionary tree:
8:
1390:. Princeton University Press. p. 336.
3925:Late Cretaceous dinosaurs of North America
3836:
3452:
3441:
3127:
2897:
2838:
2558:
2547:
2534:
2363:
2191:
2083:
2079:
1937:
1697:
1693:
1682:
1669:
1611:
1584:
1575:
1511:
1495:
1481:
1473:
661:that lived about 1.5 million years later.
127:
31:
1444:
408:excavated limb bones and a vertebra near
1341:Freedman, E.A., 2011, "A new species of
631:; in the other as a sister species of a
1208:
435:. The generic name is a combination of
1387:The Princeton Field Guide to Dinosaurs
404:In 1981 and 1994, Mark Goodwin of the
653:might have been a direct ancestor of
7:
1360:"The Open Access Dinosaurs of 2015"
1055:
1031:
963:
939:
915:
908:
844:
820:
813:
767:
738:
709:
685:
671:
1222:Canadian Journal of Earth Sciences
337:is a species of large herbivorous
325:Freedman Fowler & Horner, 2015
285:Freedman Fowler & Horner, 2015
25:
3950:Campanian genus first appearances
3427:
3421:
3416:
3410:
3403:
2520:
2513:
2506:
1459:
145:
3784:List of informal dinosaur names
396:Brachylophosaurini localities,
3794:Timeline of hadrosaur research
1:
3935:Fossil taxa described in 2015
1189:specimen with a small crest.
645:. In both evolutionary trees
592:Cervical and dorsal vertebrae
1362:. PLOS Paleo. Archived from
1309:Brachylophosaurus canadensis
1176:Tibial histology of MOR 2919
3955:Campanian genus extinctions
619:to two earlier datasets of
536:is transversely wide as in
429:Probrachylophosaurus bergei
334:Probrachylophosaurus bergei
319:Probrachylophosaurus bergei
18:Probrachylophosaurus bergei
3976:
27:Extinct genus of dinosaurs
3940:Taxa named by Jack Horner
3451:
3440:
3401:
3130:
2900:
2841:
2561:
2546:
2533:
2504:
2366:
2194:
2086:
2082:
1940:
1700:
1696:
1681:
1668:
1614:
1587:
1574:
1510:
1384:Paul, Gregory S. (2016).
1077:
1060:
1053:
1036:
1029:
985:
968:
961:
944:
937:
920:
913:
906:
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842:
825:
818:
811:
789:
772:
765:
743:
736:
714:
707:
690:
683:
312:
305:
142:Scientific classification
140:
135:
126:
34:
664:The revised analysis of
3945:Paleontology in Montana
3920:Paleontology in Alberta
1297:10.11 (2015): e0141304.
529:supratemporal fenestrae
1242:10.1139/cjes-2020-0007
1177:
601:
593:
585:
520:
515:Braincase of subadult
501:
401:
355:Judith River Formation
3887:Paleobiology Database
3764:Basturs Poble bonebed
1175:
666:Albert Prieto-Márquez
599:
591:
581:
514:
499:
414:Museum of the Rockies
395:
3960:Ornithischian genera
3850:Probrachylophosaurus
3786:(contains hadrosaur
3184:Probrachylophosaurus
948:Probrachylophosaurus
655:Probrachylophosaurus
617:Probrachylophosaurus
605:Probrachylophosaurus
505:Probrachylophosaurus
460:Probrachylophosaurus
400:marked with red star
398:Probrachylophosaurus
388:Discovery and naming
280:Probrachylophosaurus
36:Probrachylophosaurus
1437:2020Pbio...46..379W
1234:2021CaJES..58..993T
566:but different from
3536:Ornithopodichnites
3147:Brachylophosaurini
1732:Hypsilophodontidae
1446:10.1017/pab.2020.2
1178:
902:Brachylophosaurini
613:Brachylophosaurini
602:
594:
586:
521:
502:
450:than its relative
402:
363:Foremost Formation
3915:Fossils of Canada
3902:
3901:
3842:Taxon identifiers
3833:
3832:
3829:
3828:
3825:
3824:
3543:Shiraminesauropus
3436:
3435:
3399:
3398:
3395:
3394:
3391:
3390:
3387:
3386:
3282:Naashoibitosaurus
3203:Austrokritosauria
3163:Brachylophosaurus
3117:
3116:
2529:
2528:
2502:
2501:
2498:
2497:
2494:
2493:
2490:
2489:
2486:
2485:
2179:
2178:
2067:
2066:
2063:
2062:
2047:Transylvanosaurus
1933:Rhabdodontomorpha
1879:Notohypsilophodon
1850:Macrogryphosaurus
1664:
1663:
1660:
1659:
1656:
1655:
1652:
1651:
1397:978-1-78684-190-2
1343:Brachylophosaurus
1187:Brachylophosaurus
1164:
1163:
1155:
1154:
1146:
1145:
1137:
1136:
1128:
1127:
1119:
1118:
1110:
1109:
1101:
1100:
1092:
1091:
1018:
1017:
1009:
1008:
1000:
999:
972:Brachylophosaurus
890:
889:
881:
880:
659:Brachylophosaurus
637:Brachylophosaurus
629:Brachylophosaurus
564:Brachylophosaurus
560:Brachylophosaurus
552:Brachylophosaurus
544:Brachylophosaurus
517:Brachylophosaurus
452:Brachylophosaurus
444:Brachylophosaurus
419:Brachylophosaurus
381:Brachylophosaurus
339:brachylophosaurin
330:
329:
286:
16:(Redirected from
3967:
3895:
3894:
3882:
3881:
3869:
3868:
3867:
3837:
3810:mummy S.M. R4036
3684:Microhadrosaurus
3677:Mandschurosaurus
3487:Camptosaurichnus
3480:Bonaparteichnium
3453:
3446:Related articles
3442:
3431:
3425:
3420:
3415:
3414:
3407:
3128:
3009:Parasaurolophini
2920:Angulomastacator
2898:
2839:
2758:Shuangmiaosaurus
2659:Huehuecanauhtlus
2645:Gongpoquansaurus
2559:
2548:
2535:
2524:
2518:
2517:
2510:
2364:
2359:Hadrosauriformes
2327:Ratchasimasaurus
2192:
2084:
2080:
1962:Muttaburrasaurus
1938:
1871:Muttaburrasaurus
1785:Atlascopcosaurus
1720:Thescelosauridae
1698:
1694:
1683:
1670:
1612:
1585:
1576:
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1568:
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1490:
1483:
1474:
1469:
1467:Dinosaurs portal
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1371:
1356:
1350:
1339:
1333:
1325:
1319:
1305:
1299:
1289:
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1261:
1228:(10): 993–1004.
1213:
1056:
1032:
964:
940:
916:
909:
845:
821:
814:
768:
739:
710:
686:
672:
611:, placed in the
607:was, within the
583:Life restoration
519:specimen MOR 940
442:, "before", and
321:
317:
284:
277:
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40:Temporal range:
32:
21:
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3970:
3969:
3968:
3966:
3965:
3964:
3905:
3904:
3903:
3898:
3890:
3885:
3877:
3872:
3863:
3862:
3857:
3844:
3834:
3821:
3802:mummy AMNH 5060
3769:Dakota (fossil)
3745:
3596:
3576:
3564:Tetrapodosaurus
3515:Hadrosaurichnus
3447:
3432:
3409:
3383:
3378:Shantungosaurus
3328:
3309:Augustynolophus
3294:
3238:
3212:Bonapartesaurus
3197:
3141:
3113:
3044:
3032:Parasaurolophus
3003:
2887:
2835:
2826:
2751:Qianjiangsaurus
2737:Probactrosaurus
2716:Penelopognathus
2555:
2542:
2525:
2512:
2482:
2462:
2419:
2353:
2334:Riabininohadros
2207:Bayannurosaurus
2188:
2175:
2111:
2076:
2059:
2004:
1974:Tenontosauridae
1968:
1927:
1763:
1742:Gideonmantellia
1726:
1690:
1677:
1648:
1628:
1601:
1595:Avemetatarsalia
1580:Avemetatarsalia
1570:
1517:
1516:
1506:
1501:
1465:
1460:
1458:
1455:
1454:
1418:
1417:
1413:
1398:
1383:
1382:
1378:
1369:
1367:
1358:
1357:
1353:
1340:
1336:
1326:
1322:
1306:
1302:
1290:
1265:
1215:
1214:
1210:
1205:
1170:
1165:
1156:
1147:
1138:
1129:
1120:
1111:
1102:
1093:
1019:
1010:
1001:
891:
882:
576:
494:
390:
349:Late Cretaceous
347:known from the
326:
323:
315:
314:
301:
283:
275:
262:
249:
236:
220:
204:
144:
122:
120:
119:
118:
117:
112:
107:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
46:
45:
42:Late Cretaceous
38:
28:
23:
22:
15:
12:
11:
5:
3973:
3971:
3963:
3962:
3957:
3952:
3947:
3942:
3937:
3932:
3927:
3922:
3917:
3907:
3906:
3900:
3899:
3897:
3896:
3883:
3870:
3854:
3852:
3846:
3845:
3840:
3831:
3830:
3827:
3826:
3823:
3822:
3820:
3819:
3812:
3804:
3796:
3791:
3781:
3779:Hypsilophodont
3776:
3774:Hadrosaur diet
3771:
3766:
3761:
3753:
3751:
3750:Other articles
3747:
3746:
3744:
3743:
3736:
3729:
3722:
3715:
3712:Stephanosaurus
3708:
3701:
3694:
3687:
3680:
3673:
3666:
3659:
3652:
3649:Fulgurotherium
3645:
3638:
3631:
3623:
3616:
3608:
3606:
3598:
3597:
3595:
3594:
3586:
3584:
3578:
3577:
3575:
3574:
3567:
3560:
3553:
3546:
3539:
3532:
3525:
3522:Iguanodontipus
3518:
3511:
3504:
3497:
3490:
3483:
3476:
3469:
3461:
3459:
3449:
3448:
3445:
3438:
3437:
3434:
3433:
3402:
3400:
3397:
3396:
3393:
3392:
3389:
3388:
3385:
3384:
3382:
3381:
3374:
3371:Laiyangosaurus
3367:
3360:
3353:
3346:
3338:
3336:
3334:Edmontosaurini
3330:
3329:
3327:
3326:
3319:
3316:Prosaurolophus
3312:
3304:
3302:
3296:
3295:
3293:
3292:
3285:
3278:
3271:
3264:
3261:Coahuilasaurus
3257:
3248:
3246:
3240:
3239:
3237:
3236:
3229:
3222:
3215:
3207:
3205:
3199:
3198:
3196:
3195:
3187:
3180:
3173:
3166:
3159:
3151:
3149:
3143:
3142:
3140:
3139:
3131:
3125:
3119:
3118:
3115:
3114:
3112:
3111:
3104:
3097:
3090:
3083:
3076:
3069:
3062:
3054:
3052:
3046:
3045:
3043:
3042:
3035:
3028:
3021:
3013:
3011:
3005:
3004:
3002:
3001:
2994:
2987:
2980:
2973:
2965:
2958:
2951:
2944:
2937:
2930:
2923:
2916:
2909:
2901:
2895:
2893:Lambeosaurinae
2889:
2888:
2886:
2885:
2878:
2871:
2864:
2857:
2850:
2842:
2836:
2831:
2828:
2827:
2825:
2824:
2817:
2810:
2803:
2796:
2789:
2782:
2775:
2768:
2761:
2754:
2747:
2740:
2733:
2726:
2719:
2712:
2709:Nanyangosaurus
2705:
2702:Nanningosaurus
2698:
2691:
2684:
2676:
2669:
2662:
2655:
2648:
2641:
2634:
2631:Gilmoreosaurus
2627:
2620:
2613:
2606:
2599:
2592:
2585:
2577:
2570:
2562:
2556:
2553:Hadrosauroidea
2551:
2544:
2543:
2540:Hadrosauroidea
2538:
2531:
2530:
2527:
2526:
2505:
2503:
2500:
2499:
2496:
2495:
2492:
2491:
2488:
2487:
2484:
2483:
2481:
2480:
2479:
2478:
2472:
2470:
2468:Hadrosauroidea
2464:
2463:
2461:
2460:
2456:Mantellisaurus
2452:
2445:
2438:
2429:
2427:
2425:Iguanodontidae
2421:
2420:
2418:
2417:
2410:
2403:
2400:Mantellisaurus
2396:
2389:
2382:
2375:
2367:
2361:
2355:
2354:
2352:
2351:
2344:
2337:
2330:
2323:
2316:
2309:
2302:
2295:
2288:
2281:
2274:
2267:
2264:Iguanacolossus
2260:
2253:
2246:
2239:
2231:
2224:
2217:
2210:
2203:
2195:
2189:
2186:Ankylopollexia
2184:
2181:
2180:
2177:
2176:
2174:
2173:
2166:
2158:
2150:
2147:Eousdryosaurus
2143:
2136:
2133:Dysalotosaurus
2129:
2121:
2119:
2113:
2112:
2110:
2109:
2102:
2095:
2087:
2077:
2072:
2069:
2068:
2065:
2064:
2061:
2060:
2058:
2057:
2050:
2043:
2036:
2029:
2022:
2014:
2012:
2010:Rhabdodontidae
2006:
2005:
2003:
2002:
1995:
1988:
1979:
1977:
1970:
1969:
1967:
1966:
1958:
1950:
1941:
1935:
1929:
1928:
1926:
1925:
1921:Weewarrasaurus
1917:
1910:
1903:
1896:
1889:
1882:
1875:
1867:
1860:
1853:
1846:
1843:Leaellynasaura
1839:
1831:
1824:
1821:Gasparinisaura
1817:
1814:Galleonosaurus
1810:
1802:
1795:
1788:
1781:
1773:
1771:
1765:
1764:
1762:
1761:
1753:
1746:
1737:
1735:
1728:
1727:
1725:
1724:
1716:
1709:
1701:
1691:
1686:
1679:
1678:
1673:
1666:
1665:
1662:
1661:
1658:
1657:
1654:
1653:
1650:
1649:
1647:
1646:
1645:
1644:
1638:
1636:
1630:
1629:
1627:
1626:
1625:
1624:
1615:
1609:
1603:
1602:
1600:
1599:
1598:
1597:
1588:
1582:
1572:
1571:
1567:
1566:
1557:
1555:Neornithischia
1548:
1539:
1530:
1524:
1515:
1508:
1507:
1502:
1500:
1499:
1492:
1485:
1477:
1471:
1470:
1453:
1452:
1431:(3): 379–404.
1411:
1396:
1376:
1351:
1334:
1320:
1300:
1263:
1207:
1206:
1204:
1201:
1169:
1166:
1162:
1161:
1158:
1157:
1153:
1152:
1149:
1148:
1144:
1143:
1140:
1139:
1135:
1134:
1131:
1130:
1126:
1125:
1122:
1121:
1117:
1116:
1113:
1112:
1108:
1107:
1104:
1103:
1099:
1098:
1095:
1094:
1090:
1089:
1086:
1085:
1076:
1073:
1072:
1069:
1068:
1064:Prosaurolophus
1059:
1054:
1052:
1049:
1048:
1045:
1044:
1035:
1030:
1028:
1025:
1024:
1021:
1020:
1016:
1015:
1012:
1011:
1007:
1006:
1003:
1002:
998:
997:
994:
993:
984:
981:
980:
977:
976:
967:
962:
960:
957:
956:
953:
952:
943:
938:
936:
933:
932:
929:
928:
919:
914:
912:
907:
905:
897:
896:
893:
892:
888:
887:
884:
883:
879:
878:
875:
874:
865:
862:
861:
858:
857:
848:
843:
841:
838:
837:
834:
833:
824:
819:
817:
812:
810:
802:
801:
798:
797:
788:
785:
784:
781:
780:
771:
766:
764:
756:
755:
752:
751:
742:
737:
735:
727:
726:
723:
722:
713:
708:
706:
703:
702:
699:
698:
689:
684:
682:
670:
635:consisting of
625:sister species
575:
574:Classification
572:
525:autapomorphies
493:
490:
479:quadrate bones
465:The holotype,
448:stratification
389:
386:
328:
327:
324:
310:
309:
303:
302:
298:P. bergei
294:
292:
288:
287:
273:
269:
268:
260:
256:
255:
247:
243:
242:
234:
227:
226:
224:Neornithischia
218:
211:
210:
202:
195:
194:
189:
182:
181:
176:
172:
171:
166:
162:
161:
156:
152:
151:
138:
137:
133:
132:
124:
123:
115:
114:
113:
108:
103:
98:
93:
88:
83:
78:
73:
68:
63:
58:
53:
52:
39:
26:
24:
14:
13:
10:
9:
6:
4:
3:
2:
3972:
3961:
3958:
3956:
3953:
3951:
3948:
3946:
3943:
3941:
3938:
3936:
3933:
3931:
3930:Saurolophines
3928:
3926:
3923:
3921:
3918:
3916:
3913:
3912:
3910:
3893:
3888:
3884:
3880:
3875:
3871:
3866:
3860:
3856:
3855:
3853:
3851:
3847:
3843:
3838:
3818:
3817:
3813:
3811:
3809:
3808:Edmontosaurus
3805:
3803:
3801:
3800:Edmontosaurus
3797:
3795:
3792:
3789:
3785:
3782:
3780:
3777:
3775:
3772:
3770:
3767:
3765:
3762:
3760:
3759:
3755:
3754:
3752:
3748:
3742:
3741:
3737:
3735:
3734:
3730:
3728:
3727:
3723:
3721:
3720:
3719:Syngonosaurus
3716:
3714:
3713:
3709:
3707:
3706:
3702:
3700:
3699:
3695:
3693:
3692:
3688:
3686:
3685:
3681:
3679:
3678:
3674:
3672:
3671:
3667:
3665:
3664:
3663:Koutalisaurus
3660:
3658:
3657:
3653:
3651:
3650:
3646:
3644:
3643:
3642:Eucercosaurus
3639:
3637:
3636:
3632:
3629:
3628:
3624:
3622:
3621:
3617:
3615:
3614:
3613:Arstanosaurus
3610:
3609:
3607:
3605:
3604:
3599:
3593:
3592:
3588:
3587:
3585:
3583:
3579:
3573:
3572:
3568:
3566:
3565:
3561:
3559:
3558:
3554:
3552:
3551:
3547:
3545:
3544:
3540:
3538:
3537:
3533:
3531:
3530:
3529:Orcauichnites
3526:
3524:
3523:
3519:
3517:
3516:
3512:
3510:
3509:
3505:
3503:
3502:
3498:
3496:
3495:
3491:
3489:
3488:
3484:
3482:
3481:
3477:
3475:
3474:
3470:
3468:
3467:
3466:Amblydactylus
3463:
3462:
3460:
3458:
3454:
3450:
3443:
3439:
3430:
3426:
3424:
3419:
3413:
3406:
3380:
3379:
3375:
3373:
3372:
3368:
3366:
3365:
3364:Kundurosaurus
3361:
3359:
3358:
3357:Kerberosaurus
3354:
3352:
3351:
3347:
3345:
3344:
3343:Edmontosaurus
3340:
3339:
3337:
3335:
3331:
3325:
3324:
3320:
3318:
3317:
3313:
3311:
3310:
3306:
3305:
3303:
3301:
3297:
3291:
3290:
3286:
3284:
3283:
3279:
3277:
3276:
3272:
3270:
3269:
3265:
3263:
3262:
3258:
3255:
3254:
3253:Anasazisaurus
3250:
3249:
3247:
3245:
3241:
3235:
3234:
3233:Secernosaurus
3230:
3228:
3227:
3226:Kelumapusaura
3223:
3221:
3220:
3216:
3214:
3213:
3209:
3208:
3206:
3204:
3200:
3193:
3192:
3188:
3186:
3185:
3181:
3179:
3178:
3174:
3172:
3171:
3167:
3165:
3164:
3160:
3158:
3157:
3153:
3152:
3150:
3148:
3144:
3138:
3137:
3133:
3132:
3129:
3126:
3124:
3123:Saurolophinae
3120:
3110:
3109:
3105:
3103:
3102:
3098:
3096:
3095:
3091:
3089:
3088:
3084:
3082:
3081:
3077:
3075:
3074:
3073:Hypacrosaurus
3070:
3068:
3067:
3066:Corythosaurus
3063:
3061:
3060:
3056:
3055:
3053:
3051:
3050:Lambeosaurini
3047:
3041:
3040:
3036:
3034:
3033:
3029:
3027:
3026:
3025:Charonosaurus
3022:
3020:
3019:
3015:
3014:
3012:
3010:
3006:
3000:
2999:
2998:Tsintaosaurus
2995:
2993:
2992:
2991:Pararhabdodon
2988:
2986:
2985:
2984:Nipponosaurus
2981:
2979:
2978:
2974:
2971:
2970:
2966:
2964:
2963:
2959:
2957:
2956:
2955:Jaxartosaurus
2952:
2950:
2949:
2945:
2943:
2942:
2938:
2936:
2935:
2931:
2929:
2928:
2924:
2922:
2921:
2917:
2915:
2914:
2910:
2908:
2907:
2906:Adynomosaurus
2903:
2902:
2899:
2896:
2894:
2890:
2884:
2883:
2879:
2877:
2876:
2872:
2870:
2869:
2868:Lapampasaurus
2865:
2863:
2862:
2858:
2856:
2855:
2851:
2849:
2848:
2844:
2843:
2840:
2837:
2834:
2833:Hadrosauridae
2829:
2823:
2822:
2818:
2816:
2815:
2814:Zhanghenglong
2811:
2809:
2808:
2804:
2802:
2801:
2797:
2795:
2794:
2790:
2788:
2787:
2786:Telmatosaurus
2783:
2781:
2780:
2776:
2774:
2773:
2769:
2767:
2766:
2762:
2760:
2759:
2755:
2753:
2752:
2748:
2746:
2745:
2741:
2739:
2738:
2734:
2732:
2731:
2730:Portellsaurus
2727:
2725:
2724:
2720:
2718:
2717:
2713:
2711:
2710:
2706:
2704:
2703:
2699:
2697:
2696:
2692:
2690:
2689:
2685:
2682:
2681:
2677:
2675:
2674:
2670:
2668:
2667:
2663:
2661:
2660:
2656:
2654:
2653:
2649:
2647:
2646:
2642:
2640:
2639:
2635:
2633:
2632:
2628:
2626:
2625:
2621:
2619:
2618:
2614:
2612:
2611:
2607:
2605:
2604:
2600:
2598:
2597:
2593:
2591:
2590:
2586:
2583:
2582:
2578:
2576:
2575:
2571:
2569:
2568:
2564:
2563:
2560:
2557:
2554:
2549:
2545:
2541:
2536:
2532:
2523:
2519:
2516:
2509:
2476:
2475:
2474:
2473:
2471:
2469:
2465:
2458:
2457:
2453:
2451:
2450:
2446:
2444:
2443:
2439:
2436:
2435:
2431:
2430:
2428:
2426:
2422:
2416:
2415:
2411:
2409:
2408:
2404:
2402:
2401:
2397:
2395:
2394:
2393:Jinzhousaurus
2390:
2388:
2387:
2386:Hypselospinus
2383:
2381:
2380:
2376:
2374:
2373:
2369:
2368:
2365:
2362:
2360:
2356:
2350:
2349:
2345:
2343:
2342:
2341:Theiophytalia
2338:
2336:
2335:
2331:
2329:
2328:
2324:
2322:
2321:
2317:
2315:
2314:
2310:
2308:
2307:
2303:
2301:
2300:
2296:
2294:
2293:
2289:
2287:
2286:
2282:
2280:
2279:
2275:
2273:
2272:
2271:Lanzhousaurus
2268:
2266:
2265:
2261:
2259:
2258:
2254:
2252:
2251:
2247:
2245:
2244:
2240:
2237:
2236:
2232:
2230:
2229:
2225:
2223:
2222:
2218:
2216:
2215:
2211:
2209:
2208:
2204:
2202:
2201:
2197:
2196:
2193:
2190:
2187:
2182:
2172:
2171:
2167:
2164:
2163:
2159:
2156:
2155:
2151:
2149:
2148:
2144:
2142:
2141:
2140:Elrhazosaurus
2137:
2135:
2134:
2130:
2128:
2127:
2123:
2122:
2120:
2118:
2114:
2108:
2107:
2103:
2101:
2100:
2096:
2094:
2093:
2089:
2088:
2085:
2081:
2078:
2075:
2070:
2056:
2055:
2051:
2049:
2048:
2044:
2042:
2041:
2037:
2035:
2034:
2030:
2028:
2027:
2023:
2021:
2020:
2016:
2015:
2013:
2011:
2007:
2001:
2000:
1999:Tenontosaurus
1996:
1994:
1993:
1989:
1986:
1985:
1984:Convolosaurus
1981:
1980:
1978:
1975:
1971:
1964:
1963:
1959:
1956:
1955:
1951:
1948:
1947:
1946:Ampelognathus
1943:
1942:
1939:
1936:
1934:
1930:
1923:
1922:
1918:
1916:
1915:
1911:
1909:
1908:
1904:
1902:
1901:
1897:
1895:
1894:
1890:
1888:
1887:
1883:
1881:
1880:
1876:
1873:
1872:
1868:
1866:
1865:
1861:
1859:
1858:
1857:Mahuidacursor
1854:
1852:
1851:
1847:
1845:
1844:
1840:
1837:
1836:
1832:
1830:
1829:
1825:
1823:
1822:
1818:
1816:
1815:
1811:
1808:
1807:
1803:
1801:
1800:
1796:
1794:
1793:
1789:
1787:
1786:
1782:
1780:
1779:
1775:
1774:
1772:
1770:
1766:
1759:
1758:
1754:
1752:
1751:
1750:Hypsilophodon
1747:
1744:
1743:
1739:
1738:
1736:
1733:
1729:
1722:
1721:
1717:
1715:
1714:
1713:Callovosaurus
1710:
1708:
1707:
1706:Burianosaurus
1703:
1702:
1699:
1695:
1692:
1689:
1684:
1680:
1676:
1671:
1667:
1642:
1641:
1640:
1639:
1637:
1635:
1631:
1623:
1619:
1618:
1617:
1616:
1613:
1610:
1608:
1604:
1596:
1592:
1591:
1590:
1589:
1586:
1583:
1581:
1577:
1573:
1565:
1561:
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1422:
1415:
1412:
1407:
1403:
1399:
1393:
1389:
1388:
1380:
1377:
1366:on 2017-07-15
1365:
1361:
1355:
1352:
1348:
1344:
1338:
1335:
1331:
1324:
1321:
1318:(3): 99A-100A
1317:
1314:
1310:
1304:
1301:
1298:
1296:
1288:
1286:
1284:
1282:
1280:
1278:
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1123:
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1067:
1066:
1065:
1058:
1057:
1051:
1050:
1047:
1046:
1043:
1042:
1041:
1040:Edmontosaurus
1034:
1033:
1027:
1026:
1023:
1022:
1014:
1013:
1005:
1004:
996:
995:
992:
991:
990:
983:
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979:
978:
975:
974:
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966:
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927:
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918:
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903:
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877:
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864:
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860:
859:
856:
855:
854:
847:
846:
840:
839:
836:
835:
832:
831:
830:
829:Secernosaurus
823:
822:
816:
815:
808:
807:Saurolophinae
804:
803:
800:
799:
796:
795:
794:
793:Corythosaurus
787:
786:
783:
782:
779:
778:
777:
770:
769:
762:
761:Saurolophidae
758:
757:
754:
753:
750:
749:
748:
741:
740:
733:
732:Hadrosauridae
729:
728:
725:
724:
721:
720:
719:
712:
711:
705:
704:
701:
700:
697:
696:
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688:
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674:
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669:
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660:
656:
652:
648:
644:
643:
638:
634:
630:
626:
622:
618:
614:
610:
609:Hadrosaurinae
606:
598:
590:
584:
580:
573:
571:
569:
565:
561:
557:
553:
549:
545:
541:
540:
535:
534:lacrimal bone
530:
526:
518:
513:
509:
506:
498:
491:
489:
486:
484:
483:chevron bones
480:
476:
472:
468:
463:
461:
457:
456:specific name
453:
449:
445:
441:
438:
434:
430:
427:
424:In 2015, the
422:
420:
415:
411:
407:
399:
394:
387:
385:
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377:
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368:
364:
360:
356:
353:
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346:
343:
340:
336:
335:
322:
320:
311:
308:
307:Binomial name
304:
300:
299:
293:
290:
289:
282:
281:
274:
271:
270:
267:
266:Saurolophinae
261:
258:
257:
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253:Hadrosauridae
248:
245:
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232:
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228:
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130:
125:
111:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
50:
43:
37:
33:
30:
19:
3849:
3814:
3807:
3799:
3787:
3756:
3738:
3731:
3724:
3717:
3710:
3703:
3696:
3689:
3682:
3675:
3668:
3661:
3654:
3647:
3640:
3633:
3627:Claorhynchus
3625:
3618:
3611:
3603:Nomina dubia
3601:
3591:Guegoolithus
3589:
3569:
3562:
3557:Staurichnium
3555:
3550:Sousaichnium
3548:
3541:
3534:
3527:
3520:
3513:
3506:
3501:Dakotasaurus
3499:
3494:Caririchnium
3492:
3485:
3478:
3471:
3464:
3408:
3376:
3369:
3362:
3355:
3348:
3341:
3321:
3314:
3307:
3300:Saurolophini
3287:
3280:
3273:
3266:
3259:
3251:
3244:Kritosaurini
3231:
3224:
3219:Huallasaurus
3217:
3210:
3191:Wulagasaurus
3189:
3183:
3182:
3175:
3168:
3161:
3154:
3134:
3106:
3099:
3092:
3085:
3080:Lambeosaurus
3078:
3071:
3064:
3057:
3037:
3030:
3023:
3016:
2996:
2989:
2982:
2975:
2967:
2960:
2953:
2946:
2939:
2932:
2925:
2918:
2911:
2904:
2882:Yamatosaurus
2880:
2873:
2866:
2859:
2852:
2847:Aquilarhinus
2845:
2819:
2812:
2805:
2798:
2793:Tethyshadros
2791:
2784:
2777:
2770:
2763:
2756:
2749:
2742:
2735:
2728:
2723:Plesiohadros
2721:
2714:
2707:
2700:
2695:Lophorhothon
2693:
2686:
2678:
2671:
2664:
2657:
2650:
2643:
2636:
2629:
2622:
2615:
2608:
2601:
2594:
2587:
2581:Batyrosaurus
2579:
2574:Bactrosaurus
2572:
2565:
2511:
2454:
2447:
2440:
2432:
2414:Ouranosaurus
2412:
2405:
2398:
2391:
2384:
2379:Brighstoneus
2377:
2370:
2346:
2339:
2332:
2325:
2318:
2311:
2306:Osmakasaurus
2304:
2299:Oblitosaurus
2297:
2290:
2283:
2276:
2269:
2262:
2255:
2248:
2241:
2233:
2226:
2221:Camptosaurus
2219:
2212:
2205:
2198:
2168:
2162:Kangnasaurus
2160:
2152:
2145:
2138:
2131:
2124:
2117:Dryosauridae
2104:
2097:
2090:
2052:
2045:
2038:
2031:
2024:
2019:Matheronodon
2017:
1997:
1990:
1982:
1960:
1952:
1944:
1919:
1912:
1905:
1898:
1893:Sektensaurus
1891:
1886:Qantassaurus
1884:
1877:
1869:
1862:
1855:
1848:
1841:
1835:Kangnasaurus
1833:
1826:
1819:
1812:
1804:
1799:Diluvicursor
1797:
1790:
1783:
1776:
1757:Vectidromeus
1755:
1748:
1740:
1718:
1711:
1704:
1622:Ornithischia
1607:Ornithischia
1559:
1550:
1546:Ornithischia
1541:
1532:
1428:
1425:Paleobiology
1424:
1414:
1386:
1379:
1368:. Retrieved
1364:the original
1354:
1346:
1342:
1337:
1329:
1323:
1315:
1312:
1308:
1303:
1294:
1225:
1221:
1211:
1194:
1191:
1186:
1182:histological
1179:
1168:Paleobiology
1079:
1078:
1062:
1061:
1038:
1037:
987:
986:
970:
969:
947:
946:
945:
922:
921:
868:
867:
851:
850:
827:
826:
791:
790:
776:Wulagasaurus
774:
773:
745:
744:
718:Bactrosaurus
716:
715:
692:
691:
678:Iguanodontia
676:
663:
658:
654:
650:
646:
640:
636:
628:
616:
604:
603:
567:
563:
559:
555:
551:
547:
543:
537:
522:
516:
504:
503:
500:Nasal crests
487:
466:
464:
459:
451:
443:
439:
428:
426:type species
423:
418:
403:
397:
379:
373:
371:
333:
332:
331:
318:
313:
297:
296:
279:
278:
230:
214:
208:Ornithischia
198:
185:
35:
29:
3816:Willinakaqe
3788:nomina nuda
3705:Siluosaurus
3670:Loncosaurus
3508:Dinehichnus
3350:Kamuysaurus
3323:Saurolophus
3275:Kritosaurus
3268:Gryposaurus
3101:Sahaliyania
3087:Magnapaulia
3059:Amurosaurus
3039:Tlatolophus
3018:Adelolophus
2962:Kazaklambia
2941:Blasisaurus
2934:Arenysaurus
2927:Aralosaurus
2861:Hadrosaurus
2854:Eotrachodon
2807:Yunganglong
2772:Sirindhorna
2744:Protohadros
2680:Koshisaurus
2673:Jintasaurus
2442:Comptonatus
2292:Napaisaurus
2278:Lurdusaurus
2250:Fukuisaurus
2228:Cedrorestes
2170:Valdosaurus
2033:Pareisactus
1864:Morrosaurus
1828:Isasicursor
1792:Chakisaurus
1688:Ornithopoda
1675:Ornithopoda
1634:Ornithopoda
1504:Ornithopoda
1081:Saurolophus
870:Gryposaurus
853:Kritosaurus
747:Hadrosaurus
492:Description
433:Jack Horner
342:hadrosaurid
259:Subfamily:
240:Ornithopoda
3909:Categories
3758:Arkharavia
3733:Tichosteus
3726:Thespesius
3698:Pteropelyx
3691:Orthomerus
3571:Wintonopus
3156:Acristavus
3136:Barsboldia
3094:Olorotitan
2969:Latirhinus
2821:Zuoyunlong
2688:Levnesovia
2638:Gobihadros
2603:Datonglong
2596:Claosaurus
2567:Altirhinus
2477:see below↓
2407:Morelladon
2285:Magnamanus
2257:Hippodraco
2126:Dryosaurus
2099:Hesperonyx
2074:Dryomorpha
1914:Trinisaura
1907:Tietasaura
1900:Talenkauen
1778:Anabisetia
1643:see below↓
1537:Dinosauria
1370:2016-01-30
1203:References
924:Acristavus
651:Acristavus
647:Acristavus
600:Skull cast
568:Acristavus
556:Acristavus
548:Acristavus
539:Acristavus
475:squamosals
375:Acristavus
192:Dinosauria
3865:Q21494374
3740:Trachodon
3656:Glishades
3635:Diclonius
3473:Anomoepus
3457:Ichnotaxa
3170:Maiasaura
3108:Velafrons
2617:Equijubus
2589:Choyrodon
2449:Iguanodon
2313:Planicoxa
2243:Dakotadon
2214:Calvarius
2040:Rhabdodon
2026:Mochlodon
1769:Elasmaria
1520:Kingdom:
1406:985402380
1258:238720084
1250:0008-4077
1196:Maiasaura
989:Maiasaura
694:Iguanodon
642:Maiasaura
621:cladistic
352:Campanian
291:Species:
165:Kingdom:
159:Eukaryota
3879:53430563
3859:Wikidata
3620:Cionodon
3289:Rhinorex
3177:Ornatops
2977:Minqaria
2948:Canardia
2875:Malefica
2800:Xuwulong
2765:Siamodon
2666:Jeyawati
2652:Gonkoken
2610:Eolambia
2434:Barilium
2235:Cumnoria
2200:Barilium
2106:Owenodon
2092:Draconyx
2054:Zalmoxes
1954:Fostoria
1806:Fostoria
1564:Cerapoda
1528:Chordata
1526:Phylum:
1522:Animalia
1295:PLOS One
558:but not
550:but not
542:but not
471:maxillae
467:MOR 2919
361:and the
345:dinosaur
246:Family:
179:Chordata
175:Phylum:
169:Animalia
155:Domain:
47:79
2913:Ajnabia
2348:Uteodon
1433:Bibcode
1349:, 53-54
1230:Bibcode
904:
809:
763:
734:
681:
477:, both
410:Rudyard
367:Alberta
359:Montana
316:†
295:†
272:Genus:
116:↓
3892:372649
3582:Ootaxa
2779:Tanius
2372:Bolong
1404:
1394:
1256:
1248:
900:
805:
759:
730:
675:
454:. The
136:Skull
2624:Fylax
2154:Iyuku
1560:Clade
1551:Clade
1542:Clade
1533:Clade
1254:S2CID
633:clade
437:Latin
357:, of
231:Clade
215:Clade
199:Clade
186:Clade
2320:Proa
1992:Iani
1620:see
1593:see
1402:OCLC
1392:ISBN
1332:, 92
1246:ISSN
639:and
378:and
55:PreꞒ
3874:EoL
1441:doi
1238:doi
627:of
440:pro
365:of
3911::
3889::
3876::
3861::
1562::
1553::
1544::
1535::
1439:.
1429:46
1427:.
1423:.
1400:.
1316:29
1266:^
1252:.
1244:.
1236:.
1226:58
1224:.
1220:.
1199:.
570:.
421:.
369:.
233::
217::
201::
188::
105:Pg
49:Ma
44:,
3790:)
3630:?
3256:?
3194:?
2972:?
2683:?
2584:?
2459:?
2437:?
2238:?
2165:?
2157:?
1987:?
1976:?
1965:?
1957:?
1949:?
1924:?
1874:?
1838:?
1809:?
1760:?
1745:?
1734:?
1723:?
1496:e
1489:t
1482:v
1449:.
1443::
1435::
1408:.
1373:.
1260:.
1240::
1232::
276:†
263:†
250:†
237:†
221:†
205:†
110:N
100:K
95:J
90:T
85:P
80:C
75:D
70:S
65:O
60:Ꞓ
20:)
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