1800:
144:
1818:
123:
508:
individual, possibly a juvenile. The lack of a premaxillary crest has been used as evidence for it being a juvenile, although the absence of a crest has also been interpreted as a distinct characteristic of some species, and more recently as a possible characteristic of
1190:
Stocker, M. R. (2012). "A new phytosaur (Archosauriformes, Phytosauria) from the Lot's Wife beds (Sonsela Member) within the Chinle
Formation (Upper Triassic) of Petrified Forest National Park, Arizona".
473:. The first autapomorphy is the flat shape of the basitubera, an area at the base of the skull in front of its attachment point with the neck. Another is that bony projections called
481:, the hole through which the spinal cord enters the skull. A third autapomorphy is a small depression or fossa surrounding the front tip of a hole in the lower jaw called the
2009:
549:, the two most likely genera to which the specimen could be referred. PEFO 34034's placement outside these groups justified the erection of a new genus and species.
1950:
569:
grouping, meaning that they did not represent one coherent genus. Many isolated phytosaur skull fragments from
Petrified Forest have been assigned to
1937:
1870:
411:
has a long, narrow, tubular-shaped snout. It has a partial rostral crest at the back of the snout, a feature which it shares with phytosaurs like
2029:
1244:
1799:
2019:
365:
400:
word for battleship, which is a reference to
Battleship NW, the locality within Petrified Forest where the skull was found.
882:
368:. The skull was discovered in 2004 by Michelle Stocker and Bill Parker and was described by them as a specimen of
2014:
1359:
1237:
1155:
143:
537:
analysis was conducted to determine its evolutionary relationships. The analysis placed PEFO 34034 outside the
1033:
1016:
992:
906:
825:
498:
451:
413:
370:
1838:
686:
2024:
1976:
1896:
727:
281:
513:. Most juvenile phytosaurs are identified on the basis of incomplete fusion of different parts of each
1230:
923:
635:
258:
573:
because they had broad squamosals, but the presence of this type of squamosal in the distinct taxon
357:
2004:
1821:
1332:
842:
482:
474:
445:
1208:
965:
138:
1981:
1901:
485:. These features were used to distinguish PEFO 34034 as a new genus and species, distinct from
1963:
1955:
1883:
1731:
510:
1968:
1888:
1633:
1626:
1579:
1344:
1292:
1200:
774:
558:
361:
211:
198:
1738:
1508:
1437:
978:
1617:
1160:, a New Phytosaur from the Chinle Formation of Petrified Forest National Park, Arizona"
478:
470:
460:
389:
1998:
1782:
1710:
1654:
1640:
1568:
1540:
1526:
1513:
1476:
1212:
703:
672:
435:
327:
80:
41:
463:, which makes up the cheek region and extends behind the main portion of the skull.
1768:
1753:
1724:
1717:
1702:
1678:
1533:
1469:
1456:
566:
534:
1861:
1204:
17:
1761:
1647:
1606:
1599:
1592:
1483:
1444:
788:
659:
611:
596:
554:
425:
419:
376:
122:
55:
1928:
1775:
1695:
1490:
1317:
1301:
1283:
440:
320:
100:
65:
1745:
1423:
1401:
1371:
1253:
582:
577:
suggests that not all of these fragments can be assigned with confidence to
323:
237:
224:
155:
105:
49:
504:
When it was described in 2004, PEFO 34034 was thought to represent a small
1922:
1855:
1561:
1547:
1277:
744:
514:
353:
345:
175:
95:
90:
75:
70:
60:
1942:
1875:
1688:
335:
331:
313:
185:
110:
85:
1497:
1271:
165:
1832:
538:
397:
349:
316:
497:
are similar in that both have broad squamosals and holes called
1836:
1264:
1226:
1222:
469:
was named on the basis of several unique characteristics, or
443:
crest along the midline of the snout that is present in
477:
at the back of the skull make up the top margin of the
1912:
1845:
1676:
1616:
1578:
1507:
1455:
1421:
1370:
1343:
1316:
561:. As was the case in previous studies, species of
429:, but which is not as prominent as it is in other
356:called PEFO 34034 from the Upper Lot's Wife beds,
1238:
392:word for an animal's face. The specific name
8:
1150:
1148:
521:is not known from any postcranial remains.
1833:
1584:
1518:
1461:
1429:
1418:
1349:
1322:
1313:
1261:
1245:
1231:
1223:
585:showing the phylogenetic relationships of
459:with other phytosaurs can be found in its
121:
31:
2010:Late Triassic reptiles of North America
1185:
1183:
1181:
1179:
1177:
1144:
501:that are located on top of the skull.
455:. Many characteristics that associate
348:incomplete, partially disarticulated
7:
1008:
984:
957:
898:
874:
867:
817:
780:
719:
678:
651:
627:
603:
592:
1193:Journal of Vertebrate Paleontology
380:). It was placed in the new genus
25:
1817:
1816:
1798:
142:
366:Petrified Forest National Park
1:
2030:Fossil taxa described in 2012
1205:10.1080/02724634.2012.649815
344:. It is known from a single
2046:
883:Smilosuchus lithodendrorum
334:, represented by a single
1812:
1795:
1587:
1521:
1464:
1432:
1417:
1398:
1352:
1325:
1312:
1260:
1030:
1013:
1006:
989:
982:
962:
955:
920:
903:
896:
879:
872:
865:
839:
822:
815:
802:
785:
778:
758:
741:
724:
717:
700:
683:
676:
656:
649:
632:
625:
612:"Paleorhinus" scurriensis
608:
601:
287:
280:
139:Scientific classification
137:
129:
120:
34:
27:Extinct genus of reptiles
745:"Machaeroprosopus" zunii
384:in 2012. The genus name
2020:Paleontology in Arizona
1034:Pseudopalatus pristinus
1017:Mystriosuchus westphali
993:Pseudopalatus mccauleyi
907:Smilosuchus adamanensis
826:Leptosuchus crosbiensis
557:just outside the clade
499:supratemporal fenestrae
487:Leptosuchus adamanensis
452:Mystriosuchus westphali
414:Smilosuchus adamanensis
371:Smilosuchus adamanensis
789:"Phytosaurus" doughtyi
687:Brachysuchus megalodon
541:containing species of
439:species. It lacks the
1977:Paleobiology Database
1897:Paleobiology Database
728:Rutiodon carolinensis
589:from Stocker (2012):
533:was named in 2012, a
924:Smilosuchus gregorii
660:"Paleorhinus" sawini
636:Paleorhinus bransoni
130:Life restoration of
843:Leptosuchus studeri
483:mandibular fenestra
446:Leptosuchus studeri
374:(then a species of
966:Pravusuchus hortus
553:was found to be a
475:exoccipital prongs
1992:
1991:
1964:Open Tree of Life
1884:Open Tree of Life
1847:Protome batalaria
1839:Taxon identifiers
1830:
1829:
1808:
1807:
1793:
1792:
1732:Heterodontosuchus
1672:
1671:
1668:
1667:
1664:
1663:
1413:
1412:
1409:
1408:
1394:
1393:
1390:
1389:
1158:Protome batalaria
1135:
1134:
1126:
1125:
1117:
1116:
1108:
1107:
1099:
1098:
1090:
1089:
1081:
1080:
1072:
1071:
1063:
1062:
1054:
1053:
1045:
1044:
944:
943:
935:
934:
854:
853:
762:Protome batalaria
511:sexual dimorphism
433:species and some
341:Protome batalaria
305:
304:
294:Protome batalaria
273:P. batalaria
261:
132:Protome batalaria
48:221.5โ205.6
36:Protome batalaria
18:Protome (reptile)
16:(Redirected from
2037:
2015:Triassic Arizona
1985:
1984:
1972:
1971:
1959:
1958:
1946:
1945:
1933:
1932:
1931:
1905:
1904:
1892:
1891:
1879:
1878:
1866:
1865:
1864:
1834:
1820:
1819:
1803:
1802:
1634:Machaeroprosopus
1627:Angistorhinopsis
1585:
1580:Leptosuchomorpha
1519:
1462:
1430:
1419:
1360:Archosauromorpha
1350:
1345:Archosauromorpha
1323:
1314:
1307:
1306:
1293:Archosauromorpha
1262:
1247:
1240:
1233:
1224:
1217:
1216:
1187:
1172:
1171:
1169:
1167:
1152:
1009:
985:
958:
899:
875:
868:
818:
781:
775:Leptosuchomorpha
720:
679:
652:
628:
604:
593:
559:Leptosuchomorpha
362:Chinle Formation
296:
292:
256:
249:
236:
223:
212:Archosauriformes
199:Archosauromorpha
147:
146:
125:
115:
52:
40:Temporal range:
32:
21:
2045:
2044:
2040:
2039:
2038:
2036:
2035:
2034:
1995:
1994:
1993:
1988:
1980:
1975:
1967:
1962:
1954:
1949:
1941:
1936:
1927:
1926:
1921:
1908:
1900:
1895:
1887:
1882:
1874:
1869:
1860:
1859:
1854:
1841:
1831:
1826:
1804:
1797:
1789:
1739:Mesorhinosuchus
1660:
1612:
1574:
1511:
1509:Mystriosuchinae
1503:
1451:
1438:Diandongosuchus
1426:
1405:
1386:
1366:
1339:
1308:
1267:
1266:
1256:
1251:
1221:
1220:
1189:
1188:
1175:
1165:
1163:
1154:
1153:
1146:
1141:
1136:
1127:
1118:
1109:
1100:
1091:
1082:
1073:
1064:
1055:
1046:
979:Pseudopalatinae
945:
936:
855:
527:
406:
301:
298:
290:
289:
276:
255:
247:
234:
221:
141:
116:
114:
113:
108:
103:
98:
93:
88:
83:
78:
73:
68:
63:
58:
47:
46:
45:
38:
28:
23:
22:
15:
12:
11:
5:
2043:
2041:
2033:
2032:
2027:
2022:
2017:
2012:
2007:
1997:
1996:
1990:
1989:
1987:
1986:
1973:
1960:
1947:
1934:
1918:
1916:
1910:
1909:
1907:
1906:
1893:
1880:
1867:
1851:
1849:
1843:
1842:
1837:
1828:
1827:
1825:
1824:
1813:
1810:
1809:
1806:
1805:
1796:
1794:
1791:
1790:
1788:
1787:
1779:
1772:
1765:
1758:
1750:
1742:
1735:
1728:
1721:
1714:
1707:
1699:
1692:
1684:
1682:
1674:
1673:
1670:
1669:
1666:
1665:
1662:
1661:
1659:
1658:
1651:
1644:
1637:
1630:
1622:
1620:
1618:Mystriosuchini
1614:
1613:
1611:
1610:
1603:
1596:
1588:
1582:
1576:
1575:
1573:
1572:
1565:
1558:
1551:
1544:
1537:
1530:
1522:
1516:
1505:
1504:
1502:
1501:
1494:
1487:
1480:
1473:
1465:
1459:
1453:
1452:
1450:
1449:
1441:
1433:
1427:
1422:
1415:
1414:
1411:
1410:
1407:
1406:
1399:
1396:
1395:
1392:
1391:
1388:
1387:
1385:
1384:
1383:
1382:
1376:
1374:
1368:
1367:
1365:
1364:
1363:
1362:
1353:
1347:
1341:
1340:
1338:
1337:
1336:
1335:
1326:
1320:
1310:
1309:
1305:
1304:
1295:
1286:
1280:
1274:
1265:
1258:
1257:
1252:
1250:
1249:
1242:
1235:
1227:
1219:
1218:
1199:(3): 573โ586.
1173:
1143:
1142:
1140:
1137:
1133:
1132:
1129:
1128:
1124:
1123:
1120:
1119:
1115:
1114:
1111:
1110:
1106:
1105:
1102:
1101:
1097:
1096:
1093:
1092:
1088:
1087:
1084:
1083:
1079:
1078:
1075:
1074:
1070:
1069:
1066:
1065:
1061:
1060:
1057:
1056:
1052:
1051:
1048:
1047:
1043:
1042:
1039:
1038:
1029:
1026:
1025:
1022:
1021:
1012:
1007:
1005:
1002:
1001:
998:
997:
988:
983:
981:
975:
974:
971:
970:
961:
956:
954:
951:
950:
947:
946:
942:
941:
938:
937:
933:
932:
929:
928:
919:
916:
915:
912:
911:
902:
897:
895:
892:
891:
888:
887:
878:
873:
871:
866:
864:
861:
860:
857:
856:
852:
851:
848:
847:
838:
835:
834:
831:
830:
821:
816:
814:
811:
810:
807:
806:
804:TMM 31173-120
801:
798:
797:
794:
793:
784:
779:
777:
771:
770:
767:
766:
757:
754:
753:
750:
749:
740:
737:
736:
733:
732:
723:
718:
716:
713:
712:
709:
708:
699:
696:
695:
692:
691:
682:
677:
675:
669:
668:
665:
664:
655:
650:
648:
645:
644:
641:
640:
631:
626:
624:
621:
620:
617:
616:
607:
602:
600:
591:
526:
523:
479:foramen magnum
471:autapomorphies
461:squamosal bone
405:
402:
358:Sonsela Member
303:
302:
299:
285:
284:
278:
277:
269:
267:
263:
262:
245:
241:
240:
232:
228:
227:
219:
215:
214:
209:
202:
201:
196:
189:
188:
183:
179:
178:
173:
169:
168:
163:
159:
158:
153:
149:
148:
135:
134:
127:
126:
118:
117:
109:
104:
99:
94:
89:
84:
79:
74:
69:
64:
59:
54:
53:
39:
26:
24:
14:
13:
10:
9:
6:
4:
3:
2:
2042:
2031:
2028:
2026:
2023:
2021:
2018:
2016:
2013:
2011:
2008:
2006:
2003:
2002:
2000:
1983:
1978:
1974:
1970:
1965:
1961:
1957:
1952:
1948:
1944:
1939:
1935:
1930:
1924:
1920:
1919:
1917:
1915:
1911:
1903:
1898:
1894:
1890:
1885:
1881:
1877:
1872:
1868:
1863:
1857:
1853:
1852:
1850:
1848:
1844:
1840:
1835:
1823:
1815:
1814:
1811:
1801:
1785:
1784:
1783:Termatosaurus
1780:
1778:
1777:
1773:
1771:
1770:
1766:
1764:
1763:
1759:
1756:
1755:
1751:
1748:
1747:
1743:
1741:
1740:
1736:
1734:
1733:
1729:
1727:
1726:
1722:
1720:
1719:
1715:
1713:
1712:
1711:Coburgosuchus
1708:
1705:
1704:
1700:
1698:
1697:
1693:
1691:
1690:
1686:
1685:
1683:
1681:
1680:
1675:
1657:
1656:
1655:Redondasaurus
1652:
1650:
1649:
1645:
1643:
1642:
1641:Mystriosuchus
1638:
1636:
1635:
1631:
1629:
1628:
1624:
1623:
1621:
1619:
1615:
1609:
1608:
1604:
1602:
1601:
1597:
1595:
1594:
1590:
1589:
1586:
1583:
1581:
1577:
1571:
1570:
1569:Volcanosuchus
1566:
1564:
1563:
1559:
1557:
1556:
1552:
1550:
1549:
1545:
1543:
1542:
1541:Colossosuchus
1538:
1536:
1535:
1531:
1529:
1528:
1527:Angistorhinus
1524:
1523:
1520:
1517:
1515:
1514:Phytosauridae
1510:
1506:
1500:
1499:
1495:
1493:
1492:
1488:
1486:
1485:
1481:
1479:
1478:
1477:Ebrachosuchus
1474:
1472:
1471:
1467:
1466:
1463:
1460:
1458:
1454:
1447:
1446:
1442:
1440:
1439:
1435:
1434:
1431:
1428:
1425:
1420:
1416:
1404:
1403:
1397:
1380:
1379:
1378:
1377:
1375:
1373:
1369:
1361:
1357:
1356:
1355:
1354:
1351:
1348:
1346:
1342:
1334:
1330:
1329:
1328:
1327:
1324:
1321:
1319:
1315:
1311:
1303:
1299:
1296:
1294:
1290:
1287:
1285:
1281:
1279:
1275:
1273:
1269:
1268:
1263:
1259:
1255:
1248:
1243:
1241:
1236:
1234:
1229:
1228:
1225:
1214:
1210:
1206:
1202:
1198:
1194:
1186:
1184:
1182:
1180:
1178:
1174:
1161:
1159:
1156:"Introducing
1151:
1149:
1145:
1138:
1131:
1130:
1122:
1121:
1113:
1112:
1104:
1103:
1095:
1094:
1086:
1085:
1077:
1076:
1068:
1067:
1059:
1058:
1050:
1049:
1041:
1040:
1037:
1036:
1035:
1028:
1027:
1024:
1023:
1020:
1019:
1018:
1011:
1010:
1004:
1003:
1000:
999:
996:
995:
994:
987:
986:
980:
977:
976:
973:
972:
969:
968:
967:
960:
959:
953:
952:
949:
948:
940:
939:
931:
930:
927:
926:
925:
918:
917:
914:
913:
910:
909:
908:
901:
900:
894:
893:
890:
889:
886:
885:
884:
877:
876:
870:
869:
863:
862:
859:
858:
850:
849:
846:
845:
844:
837:
836:
833:
832:
829:
828:
827:
820:
819:
813:
812:
809:
808:
805:
800:
799:
796:
795:
792:
791:
790:
783:
782:
776:
773:
772:
769:
768:
765:
764:
763:
756:
755:
752:
751:
748:
747:
746:
739:
738:
735:
734:
731:
730:
729:
722:
721:
715:
714:
711:
710:
707:
706:
705:
704:Angistorhinus
698:
697:
694:
693:
690:
689:
688:
681:
680:
674:
673:Phytosauridae
671:
670:
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581:. Below is a
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328:Late Triassic
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42:Late Triassic
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19:
2025:Chinle fauna
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1769:Rileyasuchus
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1754:Palaeoctonus
1752:
1744:
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1725:Francosuchus
1723:
1718:Compsosaurus
1716:
1709:
1703:Clepsysaurus
1701:
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1679:Nomina dubia
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1625:
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1534:Brachysuchus
1532:
1525:
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1489:
1482:
1475:
1470:Arganarhinus
1468:
1457:Parasuchidae
1443:
1436:
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1164:. Retrieved
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238:Parasuchidae
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1762:Phytosaurus
1648:Nicrosaurus
1607:Smilosuchus
1600:Pravusuchus
1593:Leptosuchus
1484:Paleorhinus
1445:Heteropelta
1424:Phytosauria
1402:Phytosauria
1372:Phytosauria
1254:Phytosauria
1162:. Chinleana
597:Phytosauria
579:Leptosuchus
571:Leptosuchus
563:Leptosuchus
555:phytosaurid
547:Smilosuchus
543:Leptosuchus
506:Leptosuchus
495:Leptosuchus
431:Smilosuchus
426:Leptosuchus
420:Pravusuchus
404:Description
377:Leptosuchus
225:Phytosauria
2005:Phytosaurs
1999:Categories
1776:Suchoprion
1696:Centemodon
1491:Parasuchus
1381:see belowโ
1333:Sauropsida
1318:Sauropsida
1302:Crurotarsi
1284:Sauropsida
1139:References
321:parasuchid
1862:Q28598688
1746:Omosaurus
1270:Kingdom:
1213:129527672
583:cladogram
565:formed a
525:Phylogeny
394:batalaria
354:lower jaw
352:and left
326:from the
324:phytosaur
266:Species:
257:Stocker,
162:Kingdom:
156:Eukaryota
1929:Q3280960
1923:Wikidata
1856:Wikidata
1822:Category
1562:Rutiodon
1548:Jupijkam
1278:Chordata
1276:Phylum:
1272:Animalia
1166:20 March
515:vertebra
346:holotype
231:Family:
186:Reptilia
176:Chordata
172:Phylum:
166:Animalia
152:Domain:
1969:6149768
1956:1483727
1943:8481116
1914:Protome
1889:6149769
1876:8590893
1689:Belodon
1555:Protome
1282:Class:
599:
587:Protome
575:Protome
551:Protome
531:Protome
519:Protome
491:Protome
467:Protome
457:Protome
409:Protome
396:is the
388:is the
386:Protome
382:Protome
360:of the
336:species
332:Arizona
314:extinct
309:Protome
291:†
270:†
252:Protome
244:Genus:
218:Order:
182:Class:
1982:242994
1902:242995
1498:Wannia
1211:
539:clades
517:, but
423:, and
312:is an
1951:IRMNG
1298:Clade
1289:Clade
1209:S2CID
529:When
398:Latin
390:Greek
350:skull
317:genus
206:Clade
193:Clade
1938:GBIF
1871:GBIF
1358:see
1331:see
1168:2021
545:and
493:and
449:and
259:2012
56:Pre๊
1201:doi
364:in
330:of
319:of
2001::
1979::
1966::
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1512:/
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1207:.
1197:32
1195:.
1176:^
1147:^
489:.
417:,
338:,
208::
195::
106:Pg
50:Ma
44:,
1786:?
1757:?
1749:?
1706:?
1448:?
1246:e
1239:t
1232:v
1215:.
1203::
1170:.
248:โ
235:โ
222:โ
111:N
101:K
96:J
91:T
86:P
81:C
76:D
71:S
66:O
61:๊
20:)
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