1789:
133:
1807:
112:
497:
individual, possibly a juvenile. The lack of a premaxillary crest has been used as evidence for it being a juvenile, although the absence of a crest has also been interpreted as a distinct characteristic of some species, and more recently as a possible characteristic of
1179:
Stocker, M. R. (2012). "A new phytosaur (Archosauriformes, Phytosauria) from the Lot's Wife beds (Sonsela Member) within the Chinle
Formation (Upper Triassic) of Petrified Forest National Park, Arizona".
462:. The first autapomorphy is the flat shape of the basitubera, an area at the base of the skull in front of its attachment point with the neck. Another is that bony projections called
470:, the hole through which the spinal cord enters the skull. A third autapomorphy is a small depression or fossa surrounding the front tip of a hole in the lower jaw called the
1998:
538:, the two most likely genera to which the specimen could be referred. PEFO 34034's placement outside these groups justified the erection of a new genus and species.
1939:
558:
grouping, meaning that they did not represent one coherent genus. Many isolated phytosaur skull fragments from
Petrified Forest have been assigned to
1926:
1859:
400:
has a long, narrow, tubular-shaped snout. It has a partial rostral crest at the back of the snout, a feature which it shares with phytosaurs like
2018:
1233:
1788:
2008:
354:
389:
word for battleship, which is a reference to
Battleship NW, the locality within Petrified Forest where the skull was found.
871:
357:. The skull was discovered in 2004 by Michelle Stocker and Bill Parker and was described by them as a specimen of
2003:
1348:
1226:
1144:
132:
526:
analysis was conducted to determine its evolutionary relationships. The analysis placed PEFO 34034 outside the
1022:
1005:
981:
895:
814:
487:
440:
402:
359:
1827:
675:
2013:
1965:
1885:
716:
270:
502:. Most juvenile phytosaurs are identified on the basis of incomplete fusion of different parts of each
1219:
912:
624:
247:
562:
because they had broad squamosals, but the presence of this type of squamosal in the distinct taxon
346:
1993:
1810:
1321:
831:
471:
463:
434:
1197:
954:
127:
1970:
1890:
474:. These features were used to distinguish PEFO 34034 as a new genus and species, distinct from
1952:
1944:
1872:
1720:
499:
1957:
1877:
1622:
1615:
1568:
1333:
1281:
1189:
763:
547:
350:
200:
187:
1727:
1497:
1426:
967:
1606:
1149:, a New Phytosaur from the Chinle Formation of Petrified Forest National Park, Arizona"
467:
459:
449:
378:
1987:
1771:
1699:
1643:
1629:
1557:
1529:
1515:
1502:
1465:
1201:
692:
661:
424:
316:
69:
30:
452:, which makes up the cheek region and extends behind the main portion of the skull.
1757:
1742:
1713:
1706:
1691:
1667:
1522:
1458:
1445:
555:
523:
1850:
1193:
1750:
1636:
1595:
1588:
1581:
1472:
1433:
777:
648:
600:
585:
543:
414:
408:
365:
111:
44:
1917:
1764:
1684:
1479:
1306:
1290:
1272:
429:
309:
89:
54:
1734:
1412:
1390:
1360:
1242:
571:
566:
suggests that not all of these fragments can be assigned with confidence to
312:
226:
213:
144:
94:
38:
493:
When it was described in 2004, PEFO 34034 was thought to represent a small
1911:
1844:
1550:
1536:
1266:
733:
503:
342:
334:
164:
84:
79:
64:
59:
49:
1931:
1864:
1677:
324:
320:
302:
174:
99:
74:
1486:
1260:
154:
1821:
527:
386:
338:
305:
486:
are similar in that both have broad squamosals and holes called
1825:
1253:
1215:
1211:
458:
was named on the basis of several unique characteristics, or
432:
crest along the midline of the snout that is present in
466:
at the back of the skull make up the top margin of the
1901:
1834:
1665:
1605:
1567:
1496:
1444:
1410:
1359:
1332:
1305:
550:. As was the case in previous studies, species of
418:, but which is not as prominent as it is in other
345:called PEFO 34034 from the Upper Lot's Wife beds,
1227:
381:word for an animal's face. The specific name
8:
1139:
1137:
510:is not known from any postcranial remains.
1822:
1573:
1507:
1450:
1418:
1407:
1338:
1311:
1302:
1250:
1234:
1220:
1212:
574:showing the phylogenetic relationships of
448:with other phytosaurs can be found in its
110:
20:
1999:Late Triassic reptiles of North America
1174:
1172:
1170:
1168:
1166:
1133:
490:that are located on top of the skull.
444:. Many characteristics that associate
337:incomplete, partially disarticulated
7:
997:
973:
946:
887:
863:
856:
806:
769:
708:
667:
640:
616:
592:
581:
1182:Journal of Vertebrate Paleontology
369:). It was placed in the new genus
14:
1806:
1805:
1787:
131:
355:Petrified Forest National Park
1:
2019:Fossil taxa described in 2012
1194:10.1080/02724634.2012.649815
333:. It is known from a single
2035:
872:Smilosuchus lithodendrorum
323:, represented by a single
1801:
1784:
1576:
1510:
1453:
1421:
1406:
1387:
1341:
1314:
1301:
1249:
1019:
1002:
995:
978:
971:
951:
944:
909:
892:
885:
868:
861:
854:
828:
811:
804:
791:
774:
767:
747:
730:
713:
706:
689:
672:
665:
645:
638:
621:
614:
601:"Paleorhinus" scurriensis
597:
590:
276:
269:
128:Scientific classification
126:
118:
109:
23:
16:Extinct genus of reptiles
734:"Machaeroprosopus" zunii
373:in 2012. The genus name
2009:Paleontology in Arizona
1023:Pseudopalatus pristinus
1006:Mystriosuchus westphali
982:Pseudopalatus mccauleyi
896:Smilosuchus adamanensis
815:Leptosuchus crosbiensis
546:just outside the clade
488:supratemporal fenestrae
476:Leptosuchus adamanensis
441:Mystriosuchus westphali
403:Smilosuchus adamanensis
360:Smilosuchus adamanensis
778:"Phytosaurus" doughtyi
676:Brachysuchus megalodon
530:containing species of
428:species. It lacks the
1966:Paleobiology Database
1886:Paleobiology Database
717:Rutiodon carolinensis
578:from Stocker (2012):
522:was named in 2012, a
913:Smilosuchus gregorii
649:"Paleorhinus" sawini
625:Paleorhinus bransoni
119:Life restoration of
832:Leptosuchus studeri
472:mandibular fenestra
435:Leptosuchus studeri
363:(then a species of
955:Pravusuchus hortus
542:was found to be a
464:exoccipital prongs
1981:
1980:
1953:Open Tree of Life
1873:Open Tree of Life
1836:Protome batalaria
1828:Taxon identifiers
1819:
1818:
1797:
1796:
1782:
1781:
1721:Heterodontosuchus
1661:
1660:
1657:
1656:
1653:
1652:
1402:
1401:
1398:
1397:
1383:
1382:
1379:
1378:
1147:Protome batalaria
1124:
1123:
1115:
1114:
1106:
1105:
1097:
1096:
1088:
1087:
1079:
1078:
1070:
1069:
1061:
1060:
1052:
1051:
1043:
1042:
1034:
1033:
933:
932:
924:
923:
843:
842:
751:Protome batalaria
500:sexual dimorphism
422:species and some
330:Protome batalaria
294:
293:
283:Protome batalaria
262:P. batalaria
250:
121:Protome batalaria
37:221.5โ205.6
25:Protome batalaria
2026:
2004:Triassic Arizona
1974:
1973:
1961:
1960:
1948:
1947:
1935:
1934:
1922:
1921:
1920:
1894:
1893:
1881:
1880:
1868:
1867:
1855:
1854:
1853:
1823:
1809:
1808:
1792:
1791:
1623:Machaeroprosopus
1616:Angistorhinopsis
1574:
1569:Leptosuchomorpha
1508:
1451:
1419:
1408:
1349:Archosauromorpha
1339:
1334:Archosauromorpha
1312:
1303:
1296:
1295:
1282:Archosauromorpha
1251:
1236:
1229:
1222:
1213:
1206:
1205:
1176:
1161:
1160:
1158:
1156:
1141:
998:
974:
947:
888:
864:
857:
807:
770:
764:Leptosuchomorpha
709:
668:
641:
617:
593:
582:
548:Leptosuchomorpha
351:Chinle Formation
285:
281:
245:
238:
225:
212:
201:Archosauriformes
188:Archosauromorpha
136:
135:
114:
104:
41:
29:Temporal range:
21:
2034:
2033:
2029:
2028:
2027:
2025:
2024:
2023:
1984:
1983:
1982:
1977:
1969:
1964:
1956:
1951:
1943:
1938:
1930:
1925:
1916:
1915:
1910:
1897:
1889:
1884:
1876:
1871:
1863:
1858:
1849:
1848:
1843:
1830:
1820:
1815:
1793:
1786:
1778:
1728:Mesorhinosuchus
1649:
1601:
1563:
1500:
1498:Mystriosuchinae
1492:
1440:
1427:Diandongosuchus
1415:
1394:
1375:
1355:
1328:
1297:
1256:
1255:
1245:
1240:
1210:
1209:
1178:
1177:
1164:
1154:
1152:
1143:
1142:
1135:
1130:
1125:
1116:
1107:
1098:
1089:
1080:
1071:
1062:
1053:
1044:
1035:
968:Pseudopalatinae
934:
925:
844:
516:
395:
290:
287:
279:
278:
265:
244:
236:
223:
210:
130:
105:
103:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
36:
35:
34:
27:
17:
12:
11:
5:
2032:
2030:
2022:
2021:
2016:
2011:
2006:
2001:
1996:
1986:
1985:
1979:
1978:
1976:
1975:
1962:
1949:
1936:
1923:
1907:
1905:
1899:
1898:
1896:
1895:
1882:
1869:
1856:
1840:
1838:
1832:
1831:
1826:
1817:
1816:
1814:
1813:
1802:
1799:
1798:
1795:
1794:
1785:
1783:
1780:
1779:
1777:
1776:
1768:
1761:
1754:
1747:
1739:
1731:
1724:
1717:
1710:
1703:
1696:
1688:
1681:
1673:
1671:
1663:
1662:
1659:
1658:
1655:
1654:
1651:
1650:
1648:
1647:
1640:
1633:
1626:
1619:
1611:
1609:
1607:Mystriosuchini
1603:
1602:
1600:
1599:
1592:
1585:
1577:
1571:
1565:
1564:
1562:
1561:
1554:
1547:
1540:
1533:
1526:
1519:
1511:
1505:
1494:
1493:
1491:
1490:
1483:
1476:
1469:
1462:
1454:
1448:
1442:
1441:
1439:
1438:
1430:
1422:
1416:
1411:
1404:
1403:
1400:
1399:
1396:
1395:
1388:
1385:
1384:
1381:
1380:
1377:
1376:
1374:
1373:
1372:
1371:
1365:
1363:
1357:
1356:
1354:
1353:
1352:
1351:
1342:
1336:
1330:
1329:
1327:
1326:
1325:
1324:
1315:
1309:
1299:
1298:
1294:
1293:
1284:
1275:
1269:
1263:
1254:
1247:
1246:
1241:
1239:
1238:
1231:
1224:
1216:
1208:
1207:
1188:(3): 573โ586.
1162:
1132:
1131:
1129:
1126:
1122:
1121:
1118:
1117:
1113:
1112:
1109:
1108:
1104:
1103:
1100:
1099:
1095:
1094:
1091:
1090:
1086:
1085:
1082:
1081:
1077:
1076:
1073:
1072:
1068:
1067:
1064:
1063:
1059:
1058:
1055:
1054:
1050:
1049:
1046:
1045:
1041:
1040:
1037:
1036:
1032:
1031:
1028:
1027:
1018:
1015:
1014:
1011:
1010:
1001:
996:
994:
991:
990:
987:
986:
977:
972:
970:
964:
963:
960:
959:
950:
945:
943:
940:
939:
936:
935:
931:
930:
927:
926:
922:
921:
918:
917:
908:
905:
904:
901:
900:
891:
886:
884:
881:
880:
877:
876:
867:
862:
860:
855:
853:
850:
849:
846:
845:
841:
840:
837:
836:
827:
824:
823:
820:
819:
810:
805:
803:
800:
799:
796:
795:
793:TMM 31173-120
790:
787:
786:
783:
782:
773:
768:
766:
760:
759:
756:
755:
746:
743:
742:
739:
738:
729:
726:
725:
722:
721:
712:
707:
705:
702:
701:
698:
697:
688:
685:
684:
681:
680:
671:
666:
664:
658:
657:
654:
653:
644:
639:
637:
634:
633:
630:
629:
620:
615:
613:
610:
609:
606:
605:
596:
591:
589:
580:
515:
512:
468:foramen magnum
460:autapomorphies
450:squamosal bone
394:
391:
347:Sonsela Member
292:
291:
288:
274:
273:
267:
266:
258:
256:
252:
251:
234:
230:
229:
221:
217:
216:
208:
204:
203:
198:
191:
190:
185:
178:
177:
172:
168:
167:
162:
158:
157:
152:
148:
147:
142:
138:
137:
124:
123:
116:
115:
107:
106:
98:
93:
88:
83:
78:
73:
68:
63:
58:
53:
48:
43:
42:
28:
15:
13:
10:
9:
6:
4:
3:
2:
2031:
2020:
2017:
2015:
2012:
2010:
2007:
2005:
2002:
2000:
1997:
1995:
1992:
1991:
1989:
1972:
1967:
1963:
1959:
1954:
1950:
1946:
1941:
1937:
1933:
1928:
1924:
1919:
1913:
1909:
1908:
1906:
1904:
1900:
1892:
1887:
1883:
1879:
1874:
1870:
1866:
1861:
1857:
1852:
1846:
1842:
1841:
1839:
1837:
1833:
1829:
1824:
1812:
1804:
1803:
1800:
1790:
1774:
1773:
1772:Termatosaurus
1769:
1767:
1766:
1762:
1760:
1759:
1755:
1753:
1752:
1748:
1745:
1744:
1740:
1737:
1736:
1732:
1730:
1729:
1725:
1723:
1722:
1718:
1716:
1715:
1711:
1709:
1708:
1704:
1702:
1701:
1700:Coburgosuchus
1697:
1694:
1693:
1689:
1687:
1686:
1682:
1680:
1679:
1675:
1674:
1672:
1670:
1669:
1664:
1646:
1645:
1644:Redondasaurus
1641:
1639:
1638:
1634:
1632:
1631:
1630:Mystriosuchus
1627:
1625:
1624:
1620:
1618:
1617:
1613:
1612:
1610:
1608:
1604:
1598:
1597:
1593:
1591:
1590:
1586:
1584:
1583:
1579:
1578:
1575:
1572:
1570:
1566:
1560:
1559:
1558:Volcanosuchus
1555:
1553:
1552:
1548:
1546:
1545:
1541:
1539:
1538:
1534:
1532:
1531:
1530:Colossosuchus
1527:
1525:
1524:
1520:
1518:
1517:
1516:Angistorhinus
1513:
1512:
1509:
1506:
1504:
1503:Phytosauridae
1499:
1495:
1489:
1488:
1484:
1482:
1481:
1477:
1475:
1474:
1470:
1468:
1467:
1466:Ebrachosuchus
1463:
1461:
1460:
1456:
1455:
1452:
1449:
1447:
1443:
1436:
1435:
1431:
1429:
1428:
1424:
1423:
1420:
1417:
1414:
1409:
1405:
1393:
1392:
1386:
1369:
1368:
1367:
1366:
1364:
1362:
1358:
1350:
1346:
1345:
1344:
1343:
1340:
1337:
1335:
1331:
1323:
1319:
1318:
1317:
1316:
1313:
1310:
1308:
1304:
1300:
1292:
1288:
1285:
1283:
1279:
1276:
1274:
1270:
1268:
1264:
1262:
1258:
1257:
1252:
1248:
1244:
1237:
1232:
1230:
1225:
1223:
1218:
1217:
1214:
1203:
1199:
1195:
1191:
1187:
1183:
1175:
1173:
1171:
1169:
1167:
1163:
1150:
1148:
1145:"Introducing
1140:
1138:
1134:
1127:
1120:
1119:
1111:
1110:
1102:
1101:
1093:
1092:
1084:
1083:
1075:
1074:
1066:
1065:
1057:
1056:
1048:
1047:
1039:
1038:
1030:
1029:
1026:
1025:
1024:
1017:
1016:
1013:
1012:
1009:
1008:
1007:
1000:
999:
993:
992:
989:
988:
985:
984:
983:
976:
975:
969:
966:
965:
962:
961:
958:
957:
956:
949:
948:
942:
941:
938:
937:
929:
928:
920:
919:
916:
915:
914:
907:
906:
903:
902:
899:
898:
897:
890:
889:
883:
882:
879:
878:
875:
874:
873:
866:
865:
859:
858:
852:
851:
848:
847:
839:
838:
835:
834:
833:
826:
825:
822:
821:
818:
817:
816:
809:
808:
802:
801:
798:
797:
794:
789:
788:
785:
784:
781:
780:
779:
772:
771:
765:
762:
761:
758:
757:
754:
753:
752:
745:
744:
741:
740:
737:
736:
735:
728:
727:
724:
723:
720:
719:
718:
711:
710:
704:
703:
700:
699:
696:
695:
694:
693:Angistorhinus
687:
686:
683:
682:
679:
678:
677:
670:
669:
663:
662:Phytosauridae
660:
659:
656:
655:
652:
651:
650:
643:
642:
636:
635:
632:
631:
628:
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425:Pseudopalatus
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317:Late Triassic
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289:Stocker, 2012
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271:Binomial name
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31:Late Triassic
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2014:Chinle fauna
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1758:Rileyasuchus
1756:
1749:
1743:Palaeoctonus
1741:
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1726:
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1714:Francosuchus
1712:
1707:Compsosaurus
1705:
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1692:Clepsysaurus
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1668:Nomina dubia
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1523:Brachysuchus
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1459:Arganarhinus
1457:
1446:Parasuchidae
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1153:. Retrieved
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227:Parasuchidae
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24:
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1751:Phytosaurus
1637:Nicrosaurus
1596:Smilosuchus
1589:Pravusuchus
1582:Leptosuchus
1473:Paleorhinus
1434:Heteropelta
1413:Phytosauria
1391:Phytosauria
1361:Phytosauria
1243:Phytosauria
1151:. Chinleana
586:Phytosauria
568:Leptosuchus
560:Leptosuchus
552:Leptosuchus
544:phytosaurid
536:Smilosuchus
532:Leptosuchus
495:Leptosuchus
484:Leptosuchus
420:Smilosuchus
415:Leptosuchus
409:Pravusuchus
393:Description
366:Leptosuchus
214:Phytosauria
1994:Phytosaurs
1988:Categories
1765:Suchoprion
1685:Centemodon
1480:Parasuchus
1370:see belowโ
1322:Sauropsida
1307:Sauropsida
1291:Crurotarsi
1273:Sauropsida
1128:References
310:parasuchid
1851:Q28598688
1735:Omosaurus
1259:Kingdom:
1202:129527672
572:cladogram
554:formed a
514:Phylogeny
383:batalaria
343:lower jaw
341:and left
315:from the
313:phytosaur
255:Species:
246:Stocker,
151:Kingdom:
145:Eukaryota
1918:Q3280960
1912:Wikidata
1845:Wikidata
1811:Category
1551:Rutiodon
1537:Jupijkam
1267:Chordata
1265:Phylum:
1261:Animalia
1155:20 March
504:vertebra
335:holotype
220:Family:
175:Reptilia
165:Chordata
161:Phylum:
155:Animalia
141:Domain:
1958:6149768
1945:1483727
1932:8481116
1903:Protome
1878:6149769
1865:8590893
1678:Belodon
1544:Protome
1271:Class:
588:
576:Protome
564:Protome
540:Protome
520:Protome
508:Protome
480:Protome
456:Protome
446:Protome
398:Protome
385:is the
377:is the
375:Protome
371:Protome
349:of the
325:species
321:Arizona
303:extinct
298:Protome
280:†
259:†
241:Protome
233:Genus:
207:Order:
171:Class:
1971:242994
1891:242995
1487:Wannia
1200:
528:clades
506:, but
412:, and
301:is an
1940:IRMNG
1287:Clade
1278:Clade
1198:S2CID
518:When
387:Latin
379:Greek
339:skull
306:genus
195:Clade
182:Clade
1927:GBIF
1860:GBIF
1347:see
1320:see
1157:2021
534:and
482:and
438:and
248:2012
45:Pre๊
1190:doi
353:in
319:of
308:of
1990::
1968::
1955::
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1501:/
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1196:.
1186:32
1184:.
1165:^
1136:^
478:.
406:,
327:,
197::
184::
95:Pg
39:Ma
33:,
1775:?
1746:?
1738:?
1695:?
1437:?
1235:e
1228:t
1221:v
1204:.
1192::
1159:.
237:โ
224:โ
211:โ
100:N
90:K
85:J
80:T
75:P
70:C
65:D
60:S
55:O
50:๊
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