42:
65:
292:, another member of Frankiales. Additionally, five conserved signature proteins have been identified which are found only in the orders Pseudonocardiales and Corynebacteriales. Homologs of the proteins are generally found in
277:
are closely related. Several conserved signature indels have been identified which are found in both
Pseudonocardiales and Corynebacteriales, including a three-amino-acid insertion in a conserved region of
1906:
1689:
718:
1880:
1919:
1854:
1893:
300:, providing additional evidence of these two species being closely related to the orders Pseudonocardiales and Corynebacteriales.
1955:
1712:
Nouioui I, Carro L, García-López M, Meier-Kolthoff JP, Woyke T, Kyrpides NC, Pukall R, Klenk HP, Goodfellow M, Göker M (2018).
64:
1898:
41:
1983:
279:
254:
250:
1924:
288:
241:
1806:
407:
1768:
50:
1450:
497:
1374:
597:
671:
1960:
1934:
1828:
587:
1283:
1082:
681:
617:
487:
1629:"Phylogenetic Framework and Molecular Signatures for the Main Clades of the Phylum Actinobacteria"
808:
691:
647:
637:
447:
977:
657:
143:
59:
1058:
701:
273:
is closely related to this group. Some evidence also suggests the orders
Pseudonocardiales and
1942:
1815:
1749:
1658:
1193:
887:
846:
758:
607:
517:
347:
317:
274:
236:
224:
1885:
1573:
Embley MT, Smida J, Stackebrandt E (1988). "The phylogeny of mycolate-less wall chemotype IV
1947:
1739:
1729:
1648:
1640:
1612:
ord. nov.". In
Goodfellow M, Kämpfer P, Trujillo ME, Suzuki K, Ludwig W, Whitman WB (eds.).
1590:
1234:
377:
337:
1457:
1300:
1217:
1034:
870:
477:
467:
457:
367:
357:
220:
1744:
1713:
1653:
1628:
775:
577:
266:
86:
1594:
1977:
1333:
1099:
960:
557:
507:
417:
397:
387:
327:
96:
261:
1911:
1820:
1680:
17:
1867:
1800:
1357:
627:
567:
1791:
437:
262:
1734:
1841:
929:
527:
427:
1753:
1662:
1644:
235:
1785:
537:
249:, also formed a clade with the Pseudonocardiaceae species in 100% of the
76:
1010:
547:
246:
1872:
1859:
1762:
1846:
1766:
1616:. Vol. 5 (2nd ed.). New York: Springer. p. 1301.
1833:
721:(LPSN). The phylogeny is based on whole-genome analysis.
1690:
List of
Prokaryotic names with Standing in Nomenclature
719:
List of
Prokaryotic names with Standing in Nomenclature
239:
based on concatenated protein sequences. Additionally,
269:
1714:"Genome-Based Taxonomic Classification of the Phylum
1775:
717:The currently accepted taxonomy is based on the
612:Lacey and Goodfellow 1975 (Approved Lists 1980)
1608:Labeda DP, Goodfellow M (2012). "Order XIII.
602:Nonomura and Ohara 1971 (Approved Lists 1980)
8:
1553:are not included in this phylogenetic tree.
372:Gochnauer et al. 1975 (Approved Lists 1980)
1763:
1633:Microbiology and Molecular Biology Reviews
1614:Bergey's Manual of Systematic Bacteriology
392:Hasegawa et al. 1978 (Approved Lists 1980)
253:replications of the phylogenetic trees. A
40:
31:
1743:
1733:
1652:
1565:
1487:
1674:
1672:
666:Tomita et al. 1978) Tomita et al. 1987
257:has been identified which is found in
219:are a family of bacteria in the order
7:
1935:4af443d7-4708-4069-9c3f-e789fb62261c
282:. This insertion is also present in
223:and the only member of the suborder
1349:
1325:
1275:
1268:
1209:
1185:
1178:
1074:
1050:
1026:
1002:
952:
945:
921:
862:
838:
831:
824:
800:
750:
743:
731:
724:
582:Henssen 1957 (Approved Lists 1980)
25:
311:comprises the following genera:
63:
1627:Gao, B.; Gupta, R. S. (2012).
245:, currently part of the order
1:
1595:10.1016/S0723-2020(88)80047-X
706:Labeda and Kroppenstedt 2007
402:Labeda and Kroppenstedt 2008
188:Labeda and Kroppenstedt 2000
159:Goodfellow and Trujillo 2015
632:Moshtaghi Nikou et al. 2017
2000:
280:UDP-galactopyranose mutase
111:Labeda and Goodfellow 2015
1454:
1371:
1354:
1347:
1330:
1323:
1297:
1280:
1273:
1266:
1231:
1214:
1207:
1190:
1183:
1176:
1096:
1079:
1072:
1055:
1048:
1031:
1024:
1007:
1000:
974:
957:
950:
943:
926:
919:
884:
867:
860:
843:
836:
829:
822:
805:
798:
772:
755:
748:
741:
729:
686:Korn-Wendisch et al. 1995
289:Geodermatophilus obscurus
265:serves to both provide a
255:conserved signature indel
149:
142:
135:
130:
60:Scientific classification
58:
48:
39:
34:
1735:10.3389/fmicb.2018.02007
532:Stackebrandt et al. 1994
242:Nakamurella multipartite
204:Stackebrandt et al. 1997
572:Kim and Goodfellow 1999
422:Lechevalier et al. 1986
1679:Euzéby JP, Parte AC.
1645:10.1128/MMBR.05011-11
1583:Syst. Appl. Microbiol
1523:Halosaccharopolyspora
498:Halosaccharopolyspora
408:Allosaccharopolyspora
362:Indananda et al. 2010
165:Pseudonocardiaceae:
1577:and description of
522:Shearer et al. 1986
452:Ningsih et al. 2019
185:Actinosynnemataceae
177:Actinopolysporineae
169:Actinopolysporaceae
152:Pseudonocardiales:
51:Pseudonocardia pini
35:Pseudonocardiaceae
1912:pseudonocardiaceae
1807:Pseudonocardiaceae
1777:Pseudonocardiaceae
1683:Pseudonocardiaceae
1579:Pseudonocardiaceae
978:Streptoalloteichus
658:Streptoalloteichus
622:Labeda et al. 1984
552:Yassin et al. 1995
462:Labeda et al. 2008
432:Meklat et al. 2015
382:Quadri et al. 2016
332:Hatano et al. 2016
322:Tamura et al. 2000
309:Pseudonocardiaceae
237:phylogenetic trees
217:Pseudonocardiaceae
156:Actinopolysporales
125:Embley et al. 1989
121:Pseudonocardiaceae
27:Family of bacteria
1984:Pseudonocardiales
1971:
1970:
1943:Open Tree of Life
1769:Taxon identifiers
1610:Pseudonocardiales
1478:
1477:
1469:
1468:
1440:
1439:
1431:
1430:
1422:
1421:
1413:
1412:
1404:
1403:
1395:
1394:
1386:
1385:
1375:Saccharomonospora
1312:
1311:
1255:
1254:
1246:
1245:
1194:Kibdelosporangium
1165:
1164:
1156:
1155:
1147:
1146:
1138:
1137:
1129:
1128:
1120:
1119:
1111:
1110:
989:
988:
908:
907:
899:
898:
888:Saccharopolyspora
847:Actinoalloteichus
787:
786:
759:Actinomycetospora
737:Pseudonocardiales
707:
697:
687:
677:
667:
653:
643:
633:
623:
613:
608:Saccharopolyspora
603:
598:Saccharomonospora
593:
583:
573:
563:
553:
543:
533:
523:
518:Kibdelosporangium
513:
512:Zhang et al. 2016
503:
493:
483:
473:
463:
453:
443:
433:
423:
413:
403:
393:
383:
373:
363:
353:
352:Jiang et al. 2008
348:Actinomycetospora
343:
333:
323:
318:Actinoalloteichus
275:Corynebacteriales
225:Pseudonocardineae
213:
212:
205:
201:Pseudonocardineae
197:
196:Saker et al. 2015
189:
181:
173:
160:
126:
112:
107:Pseudonocardiales
18:Pseudonocardineae
16:(Redirected from
1991:
1964:
1963:
1951:
1950:
1938:
1937:
1928:
1927:
1915:
1914:
1902:
1901:
1889:
1888:
1876:
1875:
1863:
1862:
1850:
1849:
1837:
1836:
1824:
1823:
1811:
1810:
1809:
1796:
1795:
1794:
1764:
1758:
1757:
1747:
1737:
1722:Front. Microbiol
1709:
1703:
1702:
1700:
1698:
1676:
1667:
1666:
1656:
1624:
1618:
1617:
1605:
1599:
1598:
1570:
1554:
1547:Tamaricihabitans
1499:Actinorectispora
1492:
1350:
1326:
1276:
1269:
1235:Actinokineospora
1210:
1186:
1179:
1075:
1051:
1027:
1003:
953:
946:
922:
863:
839:
832:
825:
801:
751:
744:
732:
725:
705:
695:
685:
675:
672:Tamaricihabitans
661:
651:
642:Tian et al. 2009
641:
631:
621:
611:
601:
592:Tian et al. 2022
591:
581:
571:
561:
551:
541:
531:
521:
511:
501:
491:
482:Tang et al. 2010
481:
471:
461:
451:
441:
431:
421:
411:
401:
391:
381:
378:Actinorectispora
371:
361:
351:
341:
338:Actinokineospora
331:
321:
267:molecular marker
203:
195:
187:
179:
171:
158:
124:
110:
68:
67:
44:
32:
21:
1999:
1998:
1994:
1993:
1992:
1990:
1989:
1988:
1974:
1973:
1972:
1967:
1959:
1954:
1946:
1941:
1933:
1931:
1923:
1918:
1910:
1905:
1897:
1892:
1884:
1879:
1871:
1866:
1858:
1853:
1845:
1840:
1832:
1827:
1819:
1814:
1805:
1804:
1799:
1790:
1789:
1784:
1771:
1761:
1711:
1710:
1706:
1696:
1694:
1678:
1677:
1670:
1626:
1625:
1621:
1607:
1606:
1602:
1572:
1571:
1567:
1563:
1558:
1557:
1515:Haloactinomyces
1493:
1489:
1484:
1479:
1470:
1458:Mycobacteriales
1441:
1432:
1423:
1414:
1405:
1396:
1387:
1313:
1301:Haloechinothrix
1256:
1247:
1218:Actinophytocola
1166:
1157:
1148:
1139:
1130:
1121:
1112:
1035:Goodfellowiella
990:
909:
900:
871:Actinopolyspora
788:
715:
710:
676:Qin et al. 2015
652:Jin et al. 2022
588:Qaidamihabitans
562:Xia et al. 2013
502:Teo et al. 2021
492:Lai et al. 2014
478:Haloechinothrix
472:Lai et al. 2017
468:Haloactinomyces
458:Goodfellowiella
412:Teo et al. 2021
368:Actinopolyspora
358:Actinophytocola
306:
294:N. multipartite
284:N. multipartite
271:N. multipartite
259:N. multipartite
233:
221:Actinomycetales
193:Mzabimycetaceae
180:Zhi et al. 2009
172:Zhi et al. 2009
123:
109:
62:
28:
23:
22:
15:
12:
11:
5:
1997:
1995:
1987:
1986:
1976:
1975:
1969:
1968:
1966:
1965:
1952:
1939:
1929:
1916:
1903:
1890:
1877:
1864:
1851:
1838:
1825:
1812:
1797:
1781:
1779:
1773:
1772:
1767:
1760:
1759:
1716:Actinobacteria
1704:
1668:
1619:
1600:
1564:
1562:
1559:
1556:
1555:
1486:
1485:
1483:
1480:
1476:
1475:
1472:
1471:
1467:
1466:
1463:
1462:
1453:
1447:
1446:
1443:
1442:
1438:
1437:
1434:
1433:
1429:
1428:
1425:
1424:
1420:
1419:
1416:
1415:
1411:
1410:
1407:
1406:
1402:
1401:
1398:
1397:
1393:
1392:
1389:
1388:
1384:
1383:
1380:
1379:
1370:
1367:
1366:
1363:
1362:
1353:
1348:
1346:
1343:
1342:
1339:
1338:
1329:
1324:
1322:
1319:
1318:
1315:
1314:
1310:
1309:
1306:
1305:
1296:
1293:
1292:
1289:
1288:
1279:
1274:
1272:
1267:
1265:
1262:
1261:
1258:
1257:
1253:
1252:
1249:
1248:
1244:
1243:
1240:
1239:
1230:
1227:
1226:
1223:
1222:
1213:
1208:
1206:
1203:
1202:
1199:
1198:
1189:
1184:
1182:
1177:
1175:
1172:
1171:
1168:
1167:
1163:
1162:
1159:
1158:
1154:
1153:
1150:
1149:
1145:
1144:
1141:
1140:
1136:
1135:
1132:
1131:
1127:
1126:
1123:
1122:
1118:
1117:
1114:
1113:
1109:
1108:
1105:
1104:
1095:
1092:
1091:
1088:
1087:
1078:
1073:
1071:
1068:
1067:
1064:
1063:
1054:
1049:
1047:
1044:
1043:
1040:
1039:
1030:
1025:
1023:
1020:
1019:
1016:
1015:
1006:
1001:
999:
996:
995:
992:
991:
987:
986:
983:
982:
973:
970:
969:
966:
965:
956:
951:
949:
944:
942:
939:
938:
935:
934:
925:
920:
918:
915:
914:
911:
910:
906:
905:
902:
901:
897:
896:
893:
892:
883:
880:
879:
876:
875:
866:
861:
859:
856:
855:
852:
851:
842:
837:
835:
830:
828:
823:
821:
818:
817:
814:
813:
804:
799:
797:
794:
793:
790:
789:
785:
784:
781:
780:
776:Pseudonocardia
771:
768:
767:
764:
763:
754:
749:
747:
742:
740:
730:
728:
723:
714:
711:
709:
708:
698:
696:Wu et al. 2014
688:
678:
668:
654:
644:
634:
624:
614:
604:
594:
584:
578:Pseudonocardia
574:
564:
554:
544:
534:
524:
514:
504:
494:
484:
474:
464:
454:
444:
434:
424:
414:
404:
394:
384:
374:
364:
354:
344:
334:
324:
313:
305:
302:
232:
229:
211:
210:
209:
208:
207:
206:
198:
190:
182:
174:
163:
162:
161:
147:
146:
140:
139:
133:
132:
128:
127:
118:
114:
113:
104:
100:
99:
94:
90:
89:
87:Actinomycetota
84:
80:
79:
74:
70:
69:
56:
55:
54:on agar plate
46:
45:
37:
36:
26:
24:
14:
13:
10:
9:
6:
4:
3:
2:
1996:
1985:
1982:
1981:
1979:
1962:
1957:
1953:
1949:
1944:
1940:
1936:
1930:
1926:
1921:
1917:
1913:
1908:
1904:
1900:
1895:
1891:
1887:
1882:
1878:
1874:
1869:
1865:
1861:
1856:
1852:
1848:
1843:
1839:
1835:
1830:
1826:
1822:
1817:
1813:
1808:
1802:
1798:
1793:
1787:
1783:
1782:
1780:
1778:
1774:
1770:
1765:
1755:
1751:
1746:
1741:
1736:
1731:
1727:
1723:
1719:
1717:
1708:
1705:
1693:
1691:
1686:
1684:
1675:
1673:
1669:
1664:
1660:
1655:
1650:
1646:
1642:
1639:(1): 66–112.
1638:
1634:
1630:
1623:
1620:
1615:
1611:
1604:
1601:
1596:
1592:
1588:
1584:
1580:
1576:
1575:Actinomycetes
1569:
1566:
1560:
1552:
1548:
1544:
1540:
1536:
1535:Longimycelium
1532:
1528:
1527:Herbihabitans
1524:
1520:
1519:Halopolyspora
1516:
1512:
1508:
1504:
1500:
1496:
1495:Actinocrispum
1491:
1488:
1481:
1474:
1473:
1465:
1464:
1461:
1460:
1459:
1452:
1449:
1448:
1445:
1444:
1436:
1435:
1427:
1426:
1418:
1417:
1409:
1408:
1400:
1399:
1391:
1390:
1382:
1381:
1378:
1377:
1376:
1369:
1368:
1365:
1364:
1361:
1360:
1359:
1352:
1351:
1345:
1344:
1341:
1340:
1337:
1336:
1335:
1334:Amycolatopsis
1328:
1327:
1321:
1320:
1317:
1316:
1308:
1307:
1304:
1303:
1302:
1295:
1294:
1291:
1290:
1287:
1286:
1285:
1284:Thermocrispum
1278:
1277:
1271:
1270:
1264:
1263:
1260:
1259:
1251:
1250:
1242:
1241:
1238:
1237:
1236:
1229:
1228:
1225:
1224:
1221:
1220:
1219:
1212:
1211:
1205:
1204:
1201:
1200:
1197:
1196:
1195:
1188:
1187:
1181:
1180:
1174:
1173:
1170:
1169:
1161:
1160:
1152:
1151:
1143:
1142:
1134:
1133:
1125:
1124:
1116:
1115:
1107:
1106:
1103:
1102:
1101:
1100:Actinosynnema
1094:
1093:
1090:
1089:
1086:
1085:
1084:
1083:Saccharothrix
1077:
1076:
1070:
1069:
1066:
1065:
1062:
1061:
1060:
1053:
1052:
1046:
1045:
1042:
1041:
1038:
1037:
1036:
1029:
1028:
1022:
1021:
1018:
1017:
1014:
1013:
1012:
1005:
1004:
998:
997:
994:
993:
985:
984:
981:
980:
979:
972:
971:
968:
967:
964:
963:
962:
961:Allokutzneria
955:
954:
948:
947:
941:
940:
937:
936:
933:
932:
931:
924:
923:
917:
916:
913:
912:
904:
903:
895:
894:
891:
890:
889:
882:
881:
878:
877:
874:
873:
872:
865:
864:
858:
857:
854:
853:
850:
849:
848:
841:
840:
834:
833:
827:
826:
820:
819:
816:
815:
812:
811:
810:
803:
802:
796:
795:
792:
791:
783:
782:
779:
778:
777:
770:
769:
766:
765:
762:
761:
760:
753:
752:
746:
745:
739:
738:
734:
733:
727:
726:
722:
720:
712:
704:
703:
699:
694:
693:
689:
684:
683:
682:Thermocrispum
679:
674:
673:
669:
665:
660:
659:
655:
650:
649:
645:
640:
639:
635:
630:
629:
625:
620:
619:
618:Saccharothrix
615:
610:
609:
605:
600:
599:
595:
590:
589:
585:
580:
579:
575:
570:
569:
565:
560:
559:
558:Longimycelium
555:
550:
549:
545:
540:
539:
535:
530:
529:
525:
520:
519:
515:
510:
509:
508:Herbihabitans
505:
500:
499:
495:
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489:
488:Halopolyspora
485:
480:
479:
475:
470:
469:
465:
460:
459:
455:
450:
449:
445:
440:
439:
435:
430:
429:
425:
420:
419:
418:Amycolatopsis
415:
410:
409:
405:
400:
399:
398:Allokutzneria
395:
390:
389:
388:Actinosynnema
385:
380:
379:
375:
370:
369:
365:
360:
359:
355:
350:
349:
345:
342:Hasegawa 1988
340:
339:
335:
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329:
328:Actinocrispum
325:
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315:
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145:
141:
138:
134:
129:
122:
119:
116:
115:
108:
105:
102:
101:
98:
97:Actinomycetia
95:
92:
91:
88:
85:
82:
81:
78:
75:
72:
71:
66:
61:
57:
53:
52:
47:
43:
38:
33:
30:
19:
1776:
1725:
1721:
1715:
1707:
1695:. Retrieved
1688:
1682:
1636:
1632:
1622:
1613:
1609:
1603:
1586:
1582:
1578:
1574:
1568:
1551:Thermotunica
1550:
1546:
1543:Solihabitans
1542:
1538:
1534:
1530:
1526:
1522:
1518:
1514:
1511:Gandjariella
1510:
1506:
1502:
1498:
1494:
1490:
1456:
1455:
1373:
1372:
1356:
1355:
1332:
1331:
1299:
1298:
1282:
1281:
1233:
1232:
1216:
1215:
1192:
1191:
1098:
1097:
1081:
1080:
1057:
1056:
1033:
1032:
1009:
1008:
976:
975:
959:
958:
928:
927:
886:
885:
869:
868:
845:
844:
809:Sciscionella
807:
806:
774:
773:
757:
756:
736:
735:
716:
700:
692:Thermotunica
690:
680:
670:
663:
656:
648:Solihabitans
646:
638:Sciscionella
636:
626:
616:
606:
596:
586:
576:
566:
556:
546:
536:
526:
516:
506:
496:
486:
476:
466:
456:
448:Gandjariella
446:
436:
426:
416:
406:
396:
386:
376:
366:
356:
346:
336:
326:
316:
308:
307:
297:
293:
287:
283:
270:
258:
240:
234:
216:
214:
200:
192:
184:
176:
168:
155:
136:
120:
106:
49:
29:
1868:iNaturalist
1801:Wikispecies
1581:fam. nov".
1539:Salinifilum
1358:Prauserella
628:Salinifilum
568:Prauserella
442:Labeda 2001
298:G. obscurus
1561:References
1507:Crossiella
438:Crossiella
263:UMP kinase
247:Frankiales
137:See text.
1589:: 16–19.
1503:Bounagaea
1059:Umezawaea
930:Kutzneria
713:Phylogeny
702:Umezawaea
528:Kutzneria
428:Bounagaea
251:bootstrap
1978:Category
1792:Q7255180
1786:Wikidata
1754:30186281
1728:: 2007.
1663:22390973
1531:Labedaea
1451:outgroup
542:Lee 2012
538:Labedaea
231:Genomics
144:Synonyms
117:Family:
83:Phylum:
77:Bacteria
73:Domain:
1745:6113628
1697:June 9,
1654:3294427
1011:Lentzea
548:Lentzea
131:Genera
103:Order:
93:Class:
1961:394436
1948:126141
1932:NZOR:
1899:956501
1886:105655
1873:356578
1847:1PNOCF
1752:
1742:
1692:(LPSN)
1661:
1651:
1549:, and
304:Genera
1956:WoRMS
1881:IRMNG
1482:Notes
1925:2070
1920:NCBI
1907:LPSN
1894:ITIS
1860:3841
1855:GBIF
1842:EPPO
1834:7884
1750:PMID
1699:2021
1659:PMID
296:and
286:and
215:The
1829:EoL
1821:F9N
1816:CoL
1740:PMC
1730:doi
1649:PMC
1641:doi
1591:doi
1980::
1958::
1945::
1922::
1909::
1896::
1883::
1870::
1857::
1844::
1831::
1818::
1803::
1788::
1748:.
1738:.
1724:.
1720:.
1687:.
1671:^
1657:.
1647:.
1637:76
1635:.
1631:.
1587:11
1585:.
1545:,
1541:,
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1517:,
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1505:,
1501:,
1497:,
664:ex
227:.
1756:.
1732::
1726:9
1718:"
1701:.
1685:"
1681:"
1665:.
1643::
1597:.
1593::
662:(
20:)
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