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Pseudoprepotherium

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352:. The material documented so far consists mainly of limb bones but includes individual skulls and remains of jaws. Body weight of around 550kg is reconstructed for the smaller relatives using a thigh bone around 42cm long. Large molds with femur lengths of 56 to 59cm are estimated to have weighed between 1.52 and 1.86 tons  A surviving skull has a length of 43cm, but is partly deformed laterally, which means that only a few features are recognizable. A bent profile line was characteristic. Because of this, the rostrum and the cranium were at an angle of 130° from each other. At the occipital bone, the articular processes for the cervical spine protruded with little prominence. The alveoli of the five teeth per row of teeth typical of mylodon can be seen on the upper jaw, but the two front teeth are poorly preserved. From the alveoli, it can be seen that the rearmost tooth was the smallest and possibly had two lobes (bilobate). The fourth and third teeth were each elongated. 762:. The main components are different layers of sand, clay, and/or silt and limestone in which individual coal seams are embedded, at least in the Urumaco Formation. The rock strata were formed in what was originally a coastal area under the influence of a river delta. From the entire Urumaco sequence, many sites are documented, the exploration of which began as early as the 1950s. They are distributed over a good 60 different stratigraphic levels. The find material consists mainly of fish, especially sharks and rays. In addition, there are also reptiles such as turtles, crocodilians, and isolated snakes, as well as mammals appearing with rodents, South American ungulates, 125: 372:. The lower end of the joint was partially rotated out of the shaft axis. The inner articulated roller became larger than the outer. The lower one was somewhat narrower than the upper end of the joint. A tibia associated with a femur about 56cm long measured 29cm in length. This corresponds to the ratio known in other mylodons of extremely short lower sections of the hindlimbs compared to the upper ones. As a result, the tibia was only half the length of the femur. The proportions are broadly similar to 701:. It is a femur found in the first third of the 20th century. The site was then inaccessible for a long time, since it was created during the construction of a reservoir (the Virgen de la Coromoto reservoir) sank. However, a new fossil locality on the lake's eastern shore largely corresponds spatially and temporally to the original site. This was announced in a publication in 2016. The site contained a small collection of vertebrates, such as remains of 146: 498:) possess chewing surface. This particular tooth structure is widely thought to reflect a greater adaptation to grassy diets. The posterior teeth are round, oval or more complex in cross-section and correspond to molar-like teeth. The foremost tooth is designed like a canine. The rear foot also clearly shows twists so that the sole points inwards.  The mylodonts can be detected for the first time in the Oligocene. One of the earliest forms is 516:
negatively.  However, other lineages associated with the Nematheriinae, the Octomylodontinae, or the Urumacotheriinae, depending on the author, are more controversial. The latter, in particular, summarizes the late Miocene representatives of northern South America. In principle, many researchers urge a revision for the entire family since many higher taxonomic units have no formal diagnosis.  The position of
2544: 423: 3300: 617:. Carlini and colleagues had defined this using numerous finds from the Urumaco sequence in the FalcĂłn Basin of north-western Venezuela, the chronological range of which includes the Middle and Upper Miocene. However, a 2020 revision of the fossil remains by Ascanio D. RincĂłn and H. Gregory McDonald led to the dissolution of the genus and relegation to the genus 551:, which also occurs there and is closely related. Both had already been described by Florentino Ameghino at the end of the 19th century using finds from the Santa Cruz Formation. In 1934, R. Lee Collins referred a femur from the RĂ­o Yuca Formation on the RĂ­o Tucupido near Guanare in the Venezuelan state of Portuguesa to 841:
come from the western Amazon basin and date to the Middle to Upper Miocene. They are part of many fossil remains from numerous local sites spread over a large area. Significant find areas are on the RĂ­o Sepa and the RĂ­o Inuya, which flow into the RĂ­o Urubamba in central Peru. From here, the find area
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to the Mylodontidae. Their assessment was based on extensive finds from the important Middle Miocene fossil site of La Venta in Colombia in connection with Collins' thigh find.  In hindsight, Hirschfeld's characterization diagnosis turned out to be incorrect since Collins' find from the RĂ­o Yuca
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lived in a tropical climate with a water-rich environment. Their known remains are limited to limb bones, except for a few skulls and teeth. Based on these remains, they were most likely medium to large-sized mylodontid. The genus was described in 1961 and currently contains three species, which were
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As is usual in the Mylodontidae, the femur stood out due to its flat, board-like shape in front and behind. The shaft was slightly curved at the side. There was only a shallow indentation between the spherical condyle and the greater trochanter. The great calculus was massive but not very elevated.
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The internal organization of the Mylodontidae is complex and currently under discussion. A relatively wide recognition usually only finds the late development lines with the Mylodontinae and Lestodontinae, as several studies have shown since 2004,  but they are sometimes also discussed
494:) also includes one of the two sloth genera that still exist today. The Mylodontidae form one of the most diverse groups within the sloths. Characteristic features can be found in the high-crowned teeth, which, unlike those of the Megatherioidea and Megalocnoidea, have a rather flat ( 486:. In a classic system based on skeletal anatomical features, the Mylodontoidea form a sister group to the Megatherioidea and thus one of the two major lineages of sloths. Molecular genetic analyses and protein studies also differentiate a third large lineage from the 1594:
Tejada-Lara, Julia V.; Salas-Gismondi, Rodolfo; Pujos, François; Baby, Patrice; Benammi, Mouloud; Brusset, Stéphane; De Franceschi, Dario; Espurt, Nicolas; Urbina, Mario; Antoine, Pierre-Olivier (March 2015). Goswami, Anjali (ed.).
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Antoine, Pierre Olivier; Abello, MarĂ­a Alejandra; Adnet, Sylvain; Altamirano Sierra, Ali J.; Baby, Patrice; Billet, Guillaume; Boivin, Myriam; CalderĂłn, Ysabel; Candela, Adriana Magdalena; Chabain, Jules; Corfu, Fernando (2016).
766:, and minor jointed animals, among others. The secondary articulated animals show a high diversity, almost reaching that of the contemporary fauna of southern South America in the Pampas region or Mesopotamia. Armadillos, the 1143:
Varela, Luciano; Tambusso, P Sebastián; McDonald, H Gregory; Fariña, Richard A (2018-09-15). "Phylogeny, Macroevolutionary Trends and Historical Biogeography of Sloths: Insights From a Bayesian Morphological Clock Analysis".
405:. The size of the bite marks, from three to 15 mm in diameter, allows the length of the attacker to be reconstructed to be around 4m. It caught its prey with its front teeth. Attacks on the hind legs are also known by modern 528:, a form widely found in Central and North America. However, the limb bones usually only offer a limited selection of features for determining family relationships. Therefore, investigations on the little skull material see 400:
tibia from the Middle Miocene Pebas Formation near Iquitos in the western Amazon Basin, about 23 cm long, shows more than 60 bite marks, the size and arrangement of which suggest that they were probably caused by a juvenile
362:. The lesser trochanter was only weakly developed. The third trochanter appeared as a slight bulge around the middle of the shaft and continued as an edge to the lower end of the joint. The position roughly matched that of 1263:
Presslee, Samantha; Slater, Graham J.; Pujos, François; Forasiepi, Analía M.; Fischer, Roman; Molloy, Kelly; Mackie, Meaghan; Olsen, Jesper V.; Kramarz, Alejandro; Taglioretti, Matías; Scaglia, Fernando (July 2019).
669:. A skull and several limb bones from La Venta formed this basis.  Referring to deviations in the structure of the femur, RincĂłn and McDonald reclassified the 2020 form as the type species of the new genus 1187:
Delsuc, Frédéric; Kuch, Melanie; Gibb, Gillian C.; Karpinski, Emil; Hackenberger, Dirk; Szpak, Paul; Martínez, Jorge G.; Mead, Jim I.; McDonald, H. Gregory; MacPhee, Ross D. E.; Billet, Guillaume (2019-06-17).
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formations. These go back to a phase when the so-called "Proto-Amazon" existed, a landscape characterized by lakes, swamps and rivers connected to the Caribbean. It can now be described as "the Pebas mega
1540: 723:. The RĂ­o Yuca Formation is composed largely of limestone and sandstone with discrete interstices of the conglomerate. It originated in a freshwater environment with a formation period possibly in the 1702:
Antoine, Pierre-Olivier; Salas-Gismondi, Rodolfo; Pujos, François; Ganerød, Morgan; Marivaux, Laurent (2017-03-01). "Western Amazonia as a Hotspot of Mammalian Biodiversity Throughout the Cenozoic".
613:. The other two species, which are significantly larger with femur lengths of 56 to 59 cm, were recognized in 2006 by a work team led by Alfredo A. Carlini but were originally included in the genus 955:"A new Miocene vertebrate assemblage from the RĂ­o Yuca Formation (Venezuela) and the northernmost record of typical Miocene mammals of high latitude (Patagonian) affinities in South America" 830:
from the lineage Mylodontidae and their immediate relatives. As a special circumstance of taphonomy, the frequent tradition of limb elements in sloths is to be evaluated, however, from
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The most extensive fossil material to date belongs to the Urumaco sequence, a complex depositional unit that is predominantly exposed in the approximately 36,000 km² large
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Carlini, Alfredo A.; Scillato-Yané, Gustavo J.; Sánchez, Rodolfo (2006-01-01). "New Mylodontoidea (Xenarthra, Phyllophaga) from the Middle Miocene-Pliocene of Venezuela".
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within the Mylodontidae is, therefore, ambiguous since the genus is largely defined by the limb bones. Based on their characteristics, phylogenetic analyzes indicate that
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was probably more adapted to tropical climate conditions. The find locations in the deposit units of the Urumaco sequence also speak in favor of this. A possibly related
3364: 1065:"Reexamination of the Relationship of Pseudoprepotherium Hoffstetter, 1961, to the Mylodont Ground Sloths (Xenarthra) from the Miocene of Northern South America" 867:
finds from the region belong to the transition to the Upper Miocene and include a mandible. At the same time, the Pebas mega wetland also includes the related
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Sánchez-Villagra, Marcelo R.; Aguilera, Orangel A. (January 2006). "Neogene vertebrates from Urumaco, Falcón State, Venezuela: Diversity and significance".
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is the type species and the smallest representative with a femur length of around 42 cm. It is based on the form introduced by R. Lee Collins in 1934 as
1476:"First Megatheriines (Xenarthra, Phyllophaga, Megatheriidae) from the Urumaco (Late Miocene) and Codore (Pliocene) Formations, Estado FalcĂłn, Venezuela" 3430: 3420: 3351: 1773: 1331:
Stratigraphy and sedimentary environments of Miocene shallow to marginal marine deposits in the theUrumaco trough, FalcĂłn Basin, Western Venezuela.
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extends eastward to western Brazil and northern Bolivia and northward to the Iquitos region of northern Peru. The found material belongs to the
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possessed even shorter lower leg sections. The shaft of the tibia narrowed sharply in the middle while the ends of the joints protruded far.
633:, also from the Urumaco sequence, is considered a nomen dubium because its femur does not show the corresponding diagnostic features of 440: 572:
Formation and the La Venta material is from today's perspective assigned to different genera, but their assessment of the position of
547:, a member of the Megatheriidae from the Santa Cruz Formation of Patagonia's Early and Middle Miocene. He also referred to the genus 478:
is a member of the Mylodontidae family within the suborder (Folivora). The Mylodontidae, in turn, are often placed together with the
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being limited to the two lower and first-mentioned sequences. The Urumaco sequence covers the period from the Middle Miocene to the
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Morfología sistemática y paleobiología de los perezosos gigantes del género Lestodon Gervais 1855 (Mammalia, Xenarthra, Tardigrada)
1541:"Two new megalonychid sloths (Mammalia: Xenarthra) from the Urumaco Formation (late Miocene), and their phylogenetic affinities" 124: 720: 145: 543:
was scientifically introduced in 1961 by Robert Hoffstetter. He mentioned them in a publication of a skeletal description of
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Hastings, Alexander K. (2012-06-14). "The Incredible Fossils of Urumaco and Beyond: Exploring Venezuela's Geologic Past".
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during the Early to Late Miocene epoch around 21 to 5.3 million years ago. Fossils of the animal have been found in
3460: 3435: 1348: 1096:"Predation of the giant Miocene caiman Purussaurus on a mylodontid ground sloth in the wetlands of proto-Amazonia" 433: 685:
has only been found in the northern part of South America. The genus defining fossil remains came to Taqe in the
657:, to which they attributed further skeletal material;  RincĂłn and McDonald, in turn, kept only the skull at 3450: 778:, have been proven. Mainly in the late 20th and early 21st century, numerous new forms were described, such as 649:,  which did not occur in northern South America; Carlini et al. 2006 simultaneously removed the femur of 926: 3455: 1759: 953:
Rincón, Ascanio D.; Solórzano, Andrés; Macsotay, Oliver; McDonald, H. Gregory; Núñez-Flores, Mónica (2016).
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Rincón, Ascanio D.; Solórzano, Andrés; McDonald, H. Gregory; Montellano-Ballesteros, Marisol (2019-03-04).
490:. According to the results of the latter two analysis methods, the Mylodontoidea with the two-toed sloths ( 3319: 2904: 3223: 2890: 2264: 500: 3445: 3377: 2754: 1265: 686: 3258: 3230: 843: 743: 665:
based on the postcranial skeletal elements  ). As early as 1985, Sue Hirschfeld named the species
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At first, he saw their systematic position as ambiguous.  In 1985, Sue Hirschfeld moved
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Its apex was at or slightly below the level of the condyle and thus lower as compared to
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In: Marcelo R. Sánchez-Villagra, Orangel A. Aguilera und Alfredo A. Carlini (Hrsg.):
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Urumaco and Venezuelan paleontology, the fossil record of the northern Neotropics.
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clearly more basally embedded in the mylodonts and move the form partly closer to
3265: 3209: 3110: 3014: 2897: 2862: 2827: 2761: 2713: 2484: 2470: 2433: 2394: 2345: 2338: 2250: 2229: 2125: 1940: 1930: 1860: 1646: 1190:"Ancient Mitogenomes Reveal the Evolutionary History and Biogeography of Sloths" 780: 711: 422: 57: 42: 3342: 3021: 2931: 2848: 2820: 2747: 2630: 2147: 2019: 2005: 1715: 1499: 1441: 1398: 1292: 1214: 1189: 1026: 771: 706: 406: 102: 67: 1733: 1688: 1670: 1572: 1517: 1449: 1406: 1300: 1233: 1165: 1080: 1034: 986: 3042: 2841: 2257: 2012: 1813: 1157: 319: 157: 107: 51: 1362: 1308: 1241: 1173: 1129: 1111: 739: 3336: 2855: 2676: 2602: 2401: 2182: 2047: 1947: 1844: 1801: 759: 374: 177: 97: 92: 77: 72: 62: 1224: 3356: 2463: 2111: 2040: 2033: 1724: 1679: 1621: 1596: 1508: 898: 763: 694: 509: 447: in this section. Unsourced material may be challenged and removed. 112: 87: 30: 1828: 1807: 1795: 1782: 775: 563:
genus based on anatomical differences and established a new one with
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is more closely related to some more modern representatives, such as
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is based on the characteristics of the femur divided until today.
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Carlini, Alfredo A.; Brandoni, Diego; Sánchez, Rodolfo (2006).
416: 559:. In 1961, Hoffstetter classified this femur as part of the 621:, largely based on the design of the femur. A third species 859:, which is now considered the type form of the genus 855:. Some of the finds, however, are assigned to the species 742:. It is composed of the lithostratigraphic units of the 1469: 1467: 1094:
Pujos, François; Salas-Gismondi, Rodolfo (2020-08-26).
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RincĂłn, Ascanio D.; McDonald, H. Gregory (2020-08-07).
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Due to its far northern distribution in South America,
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Jaramillo: 890: 899:"Pseudoprepotherium Hoffstetter, 1961" 834:also documented remains of the skull. 653:and reassigned the rest, a skull, to 645:was originally assigned to the giant 7: 1069:Revista GeolĂłgica de AmĂ©rica Central 1058: 1056: 1054: 1052: 1000: 998: 996: 948: 946: 536:, but also to the Scelidotheriidae. 445:adding citations to reliable sources 310:. It was widespread across northern 1545:Journal of Systematic Palaeontology 1480:Journal of Systematic Palaeontology 1379:Journal of Systematic Palaeontology 1007:Journal of Systematic Palaeontology 555:and set up with it the new species 14: 927:"Fossilworks: Pseudoprepotherium" 331:originally assigned to the genus 3431:Taxa named by Robert Hoffstetter 3421:Miocene mammals of South America 3298: 2542: 806:as representatives of the large 421: 144: 432:needs additional citations for 248:Pseudoprepotherium venezuelanum 1704:Journal of Mammalian Evolution 1430:Journal of Mammalian Evolution 1273:Nature Ecology & Evolution 16:Extinct genus of ground sloths 1: 3426:Fossil taxa described in 1961 1565:10.1080/14772019.2018.1427639 979:10.1016/j.geobios.2016.06.005 693:about 11km west-southwest of 366:, but was lower than that of 3411:Prehistoric placental genera 579:A total of three species of 857:Pseudoprepotherium confusum 738:in the Venezuelan state of 697:in the Venezuelan state of 667:Pseudoprepotherium confusum 3477: 1347:Czerwonogora, Ada (2010). 837:Finds possibly related to 482:in the superfamily of the 3296: 3093: 2960: 2815: 2587: 2583: 2568: 2555: 2540: 2506: 2297: 2210: 2076: 2072: 1988: 1852: 1839: 1835: 1822: 1790: 1716:10.1007/s10914-016-9333-1 1500:10.1017/S1477201906001878 1442:10.1007/s10914-012-9208-z 1399:10.1017/s1477201906001829 1293:10.1038/s41559-019-0909-z 1215:10.1016/j.cub.2019.05.043 1027:10.1017/S147720190600191X 611:Prepotherium venezuelanum 557:Prepotherium venezuelanum 263: 258: 242: 235: 141:Scientific classification 139: 131: 122: 23: 1671:10.1016/j.gr.2015.11.001 721:South American ungulates 1112:10.1098/rsbl.2020.0239 661:in 2020 and split off 3378:Paleobiology Database 1158:10.1093/sysbio/syy058 715:, as well as armored 3441:Fossils of Venezuela 1363:10.35537/10915/80374 1200:(12): 2031–2042.e6. 683:, Pseudoprepotherium 651:Mirandabradys zabasi 643:Mirandabradys zabasi 627:Mirandabradys zabasi 565:Pseudoprepotherium . 441:improve this article 3416:Miocene xenarthrans 3224:Perezfontanatherium 2265:Paroctodontotherium 1663:2016GondR..31...30A 1613:2015Palgy..58..341T 1557:2019JSPal..17..409R 1492:2006JSPal...4..269C 1391:2006JSPal...4..213S 1285:2019NatEE...3.1121P 1206:2019CBio...29E2031D 1019:2006JSPal...4..255C 971:2016Geobi..49..395R 931:www.fossilworks.org 501:Paroctodontotherium 3406:Prehistoric sloths 3328:Pseudoprepotherium 2755:Megalonychotherium 2272:Pseudoprepotherium 1622:10.1111/pala.12147 1146:Systematic Biology 865:Pseudoprepotherium 839:Pseudoprepotherium 832:Pseudoprepotherium 756:Pseudoprepotherium 754:, with remains of 687:RĂ­o Yuca Formation 639:Pseudoprepotherium 619:Pseudoprepotherium 581:Pseudoprepotherium 574:Pseudoprepotherium 569:Pseudoprepotherium 541:Pseudoprepotherium 530:Pseudoprepotherium 522:Pseudoprepotherium 518:Pseudoprepotherium 476:Pseudoprepotherium 398:Pseudoprepotherium 394:Pseudoprepotherium 346:Pseudoprepotherium 328:Pseudoprepotherium 303:Pseudoprepotherium 225:Pseudoprepotherium 132:Holotype femur of 25:Pseudoprepotherium 3461:Fossils of Brazil 3436:Neogene Venezuela 3393: 3392: 3320:Taxon identifiers 3311: 3310: 3307: 3306: 3294: 3293: 3290: 3289: 3286: 3285: 3282: 3281: 3259:Pyramiodontherium 3231:Plesiomegatherium 3080: 3079: 2922: 2921: 2918: 2917: 2914: 2913: 2783:Proschismotherium 2551: 2550: 2538: 2537: 2534: 2533: 2530: 2529: 2502: 2501: 2498: 2497: 2494: 2493: 2192: 2191: 2057: 2056: 1961: 1960: 1957: 1956: 1651:Gondwana Research 790:from the family 752:Codore Formations 473: 472: 465: 299: 298: 293: 278: 231: 3468: 3386: 3385: 3373: 3372: 3360: 3359: 3347: 3346: 3345: 3315: 3302: 3091: 3064:Pronothrotherium 2958: 2949: 2930: 2813: 2790:Pseudortotherium 2721: 2659: 2620: 2610: 2600: 2585: 2581: 2570: 2557: 2546: 2508: 2318:Glossotheriopsis 2295: 2244:Octodontotherium 2208: 2200: 2155:Scelidotheridium 2096:Scelidotheriidae 2094: 2074: 2070: 1995: 1990: 1986: 1938: 1928: 1898: 1868: 1858: 1850: 1841: 1837: 1824: 1776: 1769: 1762: 1753: 1746: 1745: 1727: 1699: 1693: 1692: 1682: 1641: 1635: 1634: 1624: 1591: 1585: 1584: 1536: 1530: 1529: 1511: 1471: 1462: 1461: 1425: 1419: 1418: 1374: 1368: 1367: 1365: 1355: 1344: 1338: 1327: 1321: 1320: 1279:(7): 1121–1130. 1270: 1260: 1254: 1253: 1227: 1217: 1184: 1178: 1177: 1140: 1134: 1133: 1123: 1091: 1085: 1084: 1060: 1047: 1046: 1002: 991: 990: 950: 941: 940: 938: 937: 923: 917: 916: 914: 913: 895: 602:( Collins, 1934) 480:Scelidotheriidae 468: 461: 457: 454: 448: 425: 417: 287: 272: 253:Hofstetter, 1961 245: 230:Hofstetter, 1961 229: 222: 209: 149: 148: 127: 117: 54: 29:Temporal range: 21: 3476: 3475: 3471: 3470: 3469: 3467: 3466: 3465: 3451:Fossils of Peru 3396: 3395: 3394: 3389: 3381: 3376: 3368: 3363: 3355: 3350: 3341: 3340: 3335: 3322: 3312: 3303: 3278: 3245:Proeremotherium 3238:Pliomegatherium 3161:Anisodontherium 3146: 3141:Proprepotherium 3105: 3076: 2985:Nothrotheriinae 2979: 2953:Nothrotheriidae 2947: 2934: 2910: 2802: 2719: 2707:Proplatyarthrus 2652: 2577: 2564: 2547: 2526: 2490: 2443:Glossotheridium 2428: 2409:Magdalenabradys 2373: 2284: 2237:Octodontobradys 2204: 2188: 2162:Scelidotheriops 2087: 2066: 2053: 1984: 1979:Pseudoglyptodon 1970: 1953: 1912:Myrmecophagidae 1906: 1876: 1831: 1818: 1786: 1780: 1750: 1749: 1701: 1700: 1696: 1643: 1642: 1638: 1593: 1592: 1588: 1538: 1537: 1533: 1473: 1472: 1465: 1427: 1426: 1422: 1376: 1375: 1371: 1353: 1346: 1345: 1341: 1328: 1324: 1268: 1262: 1261: 1257: 1194:Current Biology 1186: 1185: 1181: 1142: 1141: 1137: 1106:(8): 20200239. 1100:Biology Letters 1093: 1092: 1088: 1062: 1061: 1050: 1004: 1003: 994: 952: 951: 944: 935: 933: 925: 924: 920: 911: 909: 897: 896: 892: 887: 861:Magdalenabradys 812:Magdalenabradys 803:Proeremotherium 679: 671:Magdalenabradys 663:Magdalenabradys 607:P. venezuelanum 600:P. venezuelanum 594:P. urumaquensis 469: 458: 452: 449: 438: 426: 415: 390: 364:Magdalenabradys 359:Magdalenabradys 343: 284:P. urumaquensis 254: 251: 228: 220: 207: 143: 118: 116: 115: 110: 105: 100: 95: 90: 85: 80: 75: 70: 65: 60: 49: 48: 46: 27: 17: 12: 11: 5: 3474: 3472: 3464: 3463: 3458: 3456:Neogene Brazil 3453: 3448: 3443: 3438: 3433: 3428: 3423: 3418: 3413: 3408: 3398: 3397: 3391: 3390: 3388: 3387: 3374: 3361: 3348: 3332: 3330: 3324: 3323: 3318: 3309: 3308: 3305: 3304: 3297: 3295: 3292: 3291: 3288: 3287: 3284: 3283: 3280: 3279: 3277: 3276: 3269: 3262: 3255: 3252:Promegatherium 3248: 3241: 3234: 3227: 3220: 3213: 3206: 3199: 3192: 3189:Megathericulus 3185: 3178: 3171: 3168:Diabolotherium 3164: 3156: 3154: 3148: 3147: 3145: 3144: 3137: 3130: 3123: 3115: 3113: 3107: 3106: 3104: 3103: 3094: 3088: 3082: 3081: 3078: 3077: 3075: 3074: 3067: 3060: 3053: 3050:Nothrotheriops 3046: 3039: 3032: 3025: 3018: 3011: 3004: 2997: 2989: 2987: 2981: 2980: 2978: 2977: 2969: 2961: 2955: 2946: 2945: 2937: 2935: 2927: 2924: 2923: 2920: 2919: 2916: 2915: 2912: 2911: 2909: 2908: 2901: 2894: 2887: 2884:Protomegalonyx 2880: 2877:Pliometanastes 2873: 2870:Pattersonocnus 2866: 2859: 2852: 2845: 2838: 2831: 2824: 2816: 2810: 2808:Megalonychinae 2804: 2803: 2801: 2800: 2793: 2786: 2779: 2772: 2765: 2758: 2751: 2744: 2737: 2729: 2727: 2718: 2717: 2710: 2703: 2696: 2693:Mesopotamocnus 2689: 2681: 2673: 2670:Deseadognathus 2665: 2663: 2661:Megalonychidae 2654: 2653: 2651: 2650: 2642: 2639:Schismotherium 2634: 2626: 2616: 2606: 2596: 2588: 2578: 2575:Megatherioidea 2573: 2566: 2565: 2562:Megatherioidea 2560: 2553: 2552: 2549: 2548: 2541: 2539: 2536: 2535: 2532: 2531: 2528: 2527: 2525: 2524: 2523: 2522: 2516: 2514: 2512:Megatherioidea 2504: 2503: 2500: 2499: 2496: 2495: 2492: 2491: 2489: 2488: 2481: 2478:Pleurolestodon 2474: 2467: 2460: 2453: 2446: 2438: 2436: 2430: 2429: 2427: 2426: 2419: 2412: 2405: 2398: 2391: 2388:Bolivartherium 2383: 2381: 2375: 2374: 2372: 2371: 2364: 2357: 2350: 2342: 2335: 2328: 2321: 2314: 2306: 2303:Archaeomylodon 2298: 2292: 2286: 2285: 2283: 2282: 2279:Urumacotherium 2275: 2268: 2261: 2254: 2247: 2240: 2233: 2226: 2223:Eionaletherium 2219: 2216:Baraguatherium 2211: 2205: 2197: 2194: 2193: 2190: 2189: 2187: 2186: 2179: 2176:Sibyllotherium 2172: 2169:Scelidotherium 2165: 2158: 2151: 2144: 2137: 2134:Neonematherium 2130: 2122: 2115: 2108: 2100: 2098: 2089: 2088: 2086: 2085: 2077: 2067: 2062: 2059: 2058: 2055: 2054: 2052: 2051: 2044: 2037: 2030: 2023: 2016: 2009: 2001: 1999: 1983: 1982: 1973: 1971: 1966: 1963: 1962: 1959: 1958: 1955: 1954: 1952: 1951: 1944: 1934: 1924: 1916: 1914: 1908: 1907: 1905: 1904: 1901:Palaeomyrmidon 1894: 1886: 1884: 1878: 1877: 1875: 1874: 1864: 1853: 1847: 1833: 1832: 1827: 1820: 1819: 1817: 1816: 1810: 1804: 1798: 1791: 1788: 1787: 1781: 1779: 1778: 1771: 1764: 1756: 1748: 1747: 1694: 1636: 1607:(2): 341–378. 1586: 1551:(5): 409–421. 1531: 1486:(3): 269–278. 1463: 1436:(2): 147–148. 1420: 1385:(3): 213–220. 1369: 1339: 1322: 1255: 1179: 1152:(2): 204–218. 1135: 1086: 1048: 1013:(3): 255–267. 992: 965:(5): 395–405. 942: 918: 889: 888: 886: 883: 874:Megathericulus 827:Urumacotherium 822:Eionaletherium 817:Bolivartherium 792:Megalonychidae 787:Pattersonocnus 772:Glyptodontinae 768:Pampatheriidae 760:Early Pliocene 678: 675: 659:Bolivartherium 655:Bolivartherium 604: 603: 597: 591: 588:P. socorrensis 534:Urumacotherium 471: 470: 429: 427: 420: 414: 413:Classification 411: 389: 386: 369:Eionaletherium 342: 339: 297: 296: 295: 294: 279: 269:P. socorrensis 261: 260: 259:Other species 256: 255: 252: 240: 239: 233: 232: 218: 214: 213: 205: 201: 200: 195: 191: 190: 185: 181: 180: 175: 171: 170: 165: 161: 160: 155: 151: 150: 137: 136: 129: 128: 120: 119: 111: 106: 101: 96: 91: 86: 81: 76: 71: 66: 61: 56: 55: 50:21.0–5.3  28: 15: 13: 10: 9: 6: 4: 3: 2: 3473: 3462: 3459: 3457: 3454: 3452: 3449: 3447: 3444: 3442: 3439: 3437: 3434: 3432: 3429: 3427: 3424: 3422: 3419: 3417: 3414: 3412: 3409: 3407: 3404: 3403: 3401: 3384: 3379: 3375: 3371: 3366: 3362: 3358: 3353: 3349: 3344: 3338: 3334: 3333: 3331: 3329: 3325: 3321: 3316: 3301: 3275: 3274: 3270: 3268: 3267: 3263: 3261: 3260: 3256: 3254: 3253: 3249: 3247: 3246: 3242: 3240: 3239: 3235: 3233: 3232: 3228: 3226: 3225: 3221: 3219: 3218: 3214: 3212: 3211: 3207: 3205: 3204: 3200: 3198: 3197: 3196:Megatheridium 3193: 3191: 3190: 3186: 3184: 3183: 3179: 3177: 3176: 3175:Eomegatherium 3172: 3170: 3169: 3165: 3163: 3162: 3158: 3157: 3155: 3153: 3152:Megatheriinae 3149: 3143: 3142: 3138: 3136: 3135: 3131: 3129: 3128: 3124: 3122: 3121: 3117: 3116: 3114: 3112: 3108: 3101: 3100: 3096: 3095: 3092: 3089: 3087: 3086:Megatheriidae 3083: 3073: 3072: 3068: 3066: 3065: 3061: 3059: 3058: 3057:Nothrotherium 3054: 3052: 3051: 3047: 3045: 3044: 3040: 3038: 3037: 3033: 3031: 3030: 3029:Mcdonaldocnus 3026: 3024: 3023: 3019: 3017: 3016: 3012: 3010: 3009: 3008:Chasicobradys 3005: 3003: 3002: 3001:Aymaratherium 2998: 2996: 2995: 2991: 2990: 2988: 2986: 2982: 2975: 2974: 2970: 2968: 2967: 2963: 2962: 2959: 2956: 2954: 2950: 2944: 2943: 2942:Analcimorphus 2939: 2938: 2936: 2933: 2925: 2907: 2906: 2902: 2900: 2899: 2895: 2893: 2892: 2888: 2886: 2885: 2881: 2879: 2878: 2874: 2872: 2871: 2867: 2865: 2864: 2860: 2858: 2857: 2853: 2851: 2850: 2846: 2844: 2843: 2839: 2837: 2836: 2835:Megalonychops 2832: 2830: 2829: 2825: 2823: 2822: 2818: 2817: 2814: 2811: 2809: 2805: 2799: 2798: 2794: 2792: 2791: 2787: 2785: 2784: 2780: 2778: 2777: 2773: 2771: 2770: 2766: 2764: 2763: 2759: 2757: 2756: 2752: 2750: 2749: 2745: 2743: 2742: 2738: 2736: 2735: 2731: 2730: 2728: 2726: 2725:Ortotheriinae 2722: 2716: 2715: 2711: 2709: 2708: 2704: 2702: 2701: 2697: 2695: 2694: 2690: 2687: 2686: 2682: 2679: 2678: 2674: 2672: 2671: 2667: 2666: 2664: 2662: 2655: 2649: 2648: 2647:Similhapalops 2643: 2641: 2640: 2635: 2633: 2632: 2627: 2625: 2624: 2617: 2615: 2614: 2607: 2605: 2604: 2597: 2595: 2594: 2590: 2589: 2586: 2582: 2579: 2576: 2571: 2567: 2563: 2558: 2554: 2545: 2520: 2519: 2518: 2517: 2515: 2513: 2509: 2505: 2487: 2486: 2482: 2480: 2479: 2475: 2473: 2472: 2468: 2466: 2465: 2461: 2459: 2458: 2454: 2452: 2451: 2450:Glossotherium 2447: 2445: 2444: 2440: 2439: 2437: 2435: 2431: 2425: 2424: 2423:Thinobadistes 2420: 2418: 2417: 2413: 2411: 2410: 2406: 2404: 2403: 2399: 2397: 2396: 2392: 2390: 2389: 2385: 2384: 2382: 2380: 2376: 2370: 2369: 2365: 2363: 2362: 2358: 2356: 2355: 2351: 2348: 2347: 2343: 2341: 2340: 2336: 2334: 2333: 2329: 2327: 2326: 2322: 2320: 2319: 2315: 2312: 2311: 2307: 2305: 2304: 2300: 2299: 2296: 2293: 2291: 2287: 2281: 2280: 2276: 2274: 2273: 2269: 2267: 2266: 2262: 2260: 2259: 2255: 2253: 2252: 2248: 2246: 2245: 2241: 2239: 2238: 2234: 2232: 2231: 2227: 2225: 2224: 2220: 2218: 2217: 2213: 2212: 2209: 2206: 2203: 2195: 2185: 2184: 2180: 2178: 2177: 2173: 2171: 2170: 2166: 2164: 2163: 2159: 2157: 2156: 2152: 2150: 2149: 2145: 2143: 2142: 2141:Proscelidodon 2138: 2136: 2135: 2131: 2128: 2127: 2123: 2121: 2120: 2116: 2114: 2113: 2109: 2107: 2106: 2105:Analcitherium 2102: 2101: 2099: 2097: 2090: 2084: 2083: 2079: 2078: 2075: 2071: 2068: 2065: 2064:Mylodontoidea 2060: 2050: 2049: 2045: 2043: 2042: 2038: 2036: 2035: 2031: 2029: 2028: 2024: 2022: 2021: 2017: 2015: 2014: 2010: 2008: 2007: 2003: 2002: 2000: 1998: 1991: 1987: 1981: 1980: 1975: 1974: 1972: 1969: 1964: 1950: 1949: 1945: 1943: 1942: 1935: 1933: 1932: 1925: 1923: 1922: 1918: 1917: 1915: 1913: 1909: 1903: 1902: 1895: 1893: 1892: 1888: 1887: 1885: 1883: 1879: 1873: 1872: 1865: 1863: 1862: 1855: 1854: 1851: 1848: 1846: 1842: 1838: 1834: 1830: 1825: 1821: 1815: 1811: 1809: 1805: 1803: 1799: 1797: 1793: 1792: 1789: 1784: 1777: 1772: 1770: 1765: 1763: 1758: 1757: 1754: 1743: 1739: 1735: 1731: 1726: 1721: 1717: 1713: 1709: 1705: 1698: 1695: 1690: 1686: 1681: 1676: 1672: 1668: 1664: 1660: 1656: 1652: 1648: 1640: 1637: 1632: 1628: 1623: 1618: 1614: 1610: 1606: 1602: 1601:Palaeontology 1598: 1590: 1587: 1582: 1578: 1574: 1570: 1566: 1562: 1558: 1554: 1550: 1546: 1542: 1535: 1532: 1527: 1523: 1519: 1515: 1510: 1505: 1501: 1497: 1493: 1489: 1485: 1481: 1477: 1470: 1468: 1464: 1459: 1455: 1451: 1447: 1443: 1439: 1435: 1431: 1424: 1421: 1416: 1412: 1408: 1404: 1400: 1396: 1392: 1388: 1384: 1380: 1373: 1370: 1364: 1359: 1352: 1351: 1343: 1340: 1336: 1332: 1326: 1323: 1318: 1314: 1310: 1306: 1302: 1298: 1294: 1290: 1286: 1282: 1278: 1274: 1267: 1259: 1256: 1251: 1247: 1243: 1239: 1235: 1231: 1226: 1221: 1216: 1211: 1207: 1203: 1199: 1195: 1191: 1183: 1180: 1175: 1171: 1167: 1163: 1159: 1155: 1151: 1147: 1139: 1136: 1131: 1127: 1122: 1117: 1113: 1109: 1105: 1101: 1097: 1090: 1087: 1082: 1078: 1074: 1070: 1066: 1059: 1057: 1055: 1053: 1049: 1044: 1040: 1036: 1032: 1028: 1024: 1020: 1016: 1012: 1008: 1001: 999: 997: 993: 988: 984: 980: 976: 972: 968: 964: 960: 956: 949: 947: 943: 932: 928: 922: 919: 908: 904: 900: 894: 891: 884: 882: 880: 879:Megatheriidae 876: 875: 870: 866: 862: 858: 854: 849: 845: 840: 835: 833: 829: 828: 823: 819: 818: 813: 809: 808:Megatheriidae 805: 804: 799: 798: 793: 789: 788: 783: 782: 777: 774:, as well as 773: 769: 765: 761: 757: 753: 749: 745: 741: 737: 732: 730: 726: 722: 718: 717:Peltephilidae 714: 713: 708: 704: 700: 696: 692: 688: 684: 676: 674: 672: 668: 664: 660: 656: 652: 648: 644: 640: 636: 635:Mirandabradys 632: 628: 624: 620: 616: 615:Mirandabradys 612: 608: 601: 598: 595: 592: 589: 586: 585: 584: 582: 577: 575: 570: 566: 562: 558: 554: 550: 546: 542: 537: 535: 531: 527: 526:Thinobadistes 523: 519: 513: 511: 507: 506:Salla-Luribay 503: 502: 497: 493: 489: 488:Megalocnoidea 485: 484:Mylodontoidea 481: 477: 467: 464: 456: 446: 442: 436: 435: 430:This section 428: 424: 419: 418: 412: 410: 408: 404: 399: 395: 387: 385: 383: 382: 381:Glossotherium 377: 376: 371: 370: 365: 361: 360: 353: 351: 347: 340: 338: 336: 335: 329: 325: 321: 317: 313: 312:South America 309: 305: 304: 291: 286: 285: 280: 276: 271: 270: 265: 264: 262: 257: 250: 249: 241: 238: 234: 227: 226: 219: 216: 215: 212: 206: 203: 202: 199: 196: 193: 192: 189: 186: 183: 182: 179: 176: 173: 172: 169: 166: 163: 162: 159: 156: 153: 152: 147: 142: 138: 135: 134:P. venezuelam 130: 126: 121: 114: 109: 104: 99: 94: 89: 84: 79: 74: 69: 64: 59: 53: 44: 40: 39:Colhuehuapian 36: 32: 31:Early Miocene 26: 22: 19: 3446:Neogene Peru 3327: 3271: 3264: 3257: 3250: 3243: 3236: 3229: 3222: 3215: 3208: 3203:Megatheriops 3201: 3194: 3187: 3182:Eremotherium 3180: 3173: 3166: 3159: 3139: 3134:Prepoplanops 3132: 3127:Prepotherium 3125: 3118: 3099:Thalassocnus 3097: 3069: 3062: 3055: 3048: 3041: 3036:Mionothropus 3034: 3027: 3020: 3013: 3006: 2999: 2992: 2973:Thalassocnus 2971: 2964: 2940: 2903: 2896: 2889: 2882: 2875: 2868: 2861: 2854: 2847: 2840: 2833: 2826: 2819: 2795: 2788: 2781: 2774: 2769:Paranabradys 2767: 2760: 2753: 2746: 2739: 2732: 2712: 2705: 2698: 2691: 2683: 2675: 2668: 2645: 2637: 2629: 2621: 2613:Hiskatherium 2611: 2601: 2591: 2483: 2476: 2469: 2462: 2455: 2448: 2441: 2421: 2416:Sphenotherus 2414: 2407: 2400: 2393: 2386: 2379:Lestodontini 2366: 2359: 2352: 2344: 2337: 2332:Mylodonopsis 2330: 2323: 2316: 2310:Brievabradys 2308: 2301: 2290:Mylodontinae 2277: 2271: 2270: 2263: 2256: 2249: 2242: 2235: 2228: 2221: 2214: 2202:Mylodontidae 2181: 2174: 2167: 2160: 2153: 2146: 2139: 2132: 2124: 2119:Chubutherium 2117: 2110: 2103: 2080: 2046: 2039: 2032: 2025: 2018: 2011: 2004: 1997:Megalocnidae 1977: 1946: 1939: 1929: 1921:Myrmecophaga 1919: 1899: 1889: 1882:Cyclopedidae 1871:Orthoarthrus 1869: 1859: 1812:Superorder: 1707: 1703: 1697: 1654: 1650: 1639: 1604: 1600: 1589: 1548: 1544: 1534: 1483: 1479: 1433: 1429: 1423: 1382: 1378: 1372: 1349: 1342: 1334: 1330: 1325: 1276: 1272: 1258: 1225:11336/136908 1197: 1193: 1182: 1149: 1145: 1138: 1103: 1099: 1089: 1072: 1068: 1010: 1006: 962: 958: 934:. 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Retrieved 903:www.gbif.org 902: 893: 872: 868: 864: 860: 856: 852: 838: 836: 831: 825: 821: 815: 811: 801: 795: 785: 779: 755: 736:FalcĂłn Basin 733: 729:Late Miocene 710: 707:crocodilians 691:RĂ­o Tucupido 682: 680: 677:Paleoecology 670: 666: 662: 658: 654: 650: 646: 642: 641:(a femur of 638: 634: 630: 626: 622: 618: 614: 610: 606: 605: 599: 593: 587: 580: 578: 573: 568: 564: 561:Prepotherium 560: 556: 553:Prepotherium 552: 549:Prepotherium 548: 544: 540: 538: 533: 529: 525: 521: 517: 514: 499: 495: 491: 475: 474: 459: 450: 439:Please help 434:verification 431: 402: 397: 393: 391: 388:Paleobiology 379: 373: 367: 363: 357: 354: 350:Mylodontidae 345: 344: 334:Prepotherium 332: 327: 308:Mylodontidae 302: 301: 300: 289: 283: 282: 274: 268: 267: 247: 246: 237:Type species 224: 223: 211:Mylodontidae 133: 35:Late Miocene 24: 18: 3266:Sibotherium 3210:Megatherium 3111:Planopsinae 3015:Huilabradys 2994:Amphibradys 2966:Neohapalops 2905:Zacatzontli 2898:Xibalbaonyx 2863:Nohochichak 2828:Australonyx 2776:Pliomorphus 2762:Ortotherium 2741:Eucholoeops 2714:Urumacocnus 2685:Hyperleptus 2623:Hyperleptus 2485:Simomylodon 2471:Paramylodon 2457:Kiyumylodon 2434:Mylodontini 2395:Lestobradys 2368:Strabosodon 2346:Oreomylodon 2339:Ocnotherium 2251:Octomylodon 2230:Nematherium 2126:Nematherium 1941:Protamandua 1931:Neotamandua 1861:Argyromanis 1845:Vermilingua 1725:11336/63465 1710:(1): 5–17. 1680:11336/99030 1509:11336/80745 781:Urumacocnus 712:Purussaurus 629:by Carlini 403:Purussaurus 341:Description 43:Huayquerian 3400:Categories 3022:Lakukullus 2932:Megatheria 2891:Sinclairia 2849:Megistonyx 2821:Ahytherium 2748:Hapaloides 2631:Pelecyodon 2521:see below↓ 2354:Promylodon 2325:Megabradys 2148:Scelidodon 2020:Megalocnus 2006:Acratocnus 936:2022-05-11 912:2022-05-11 885:References 709:; such as 699:Portuguesa 407:crocodiles 3273:Urumaquia 3071:Xyophorus 3043:Nothropus 2842:Megalonyx 2797:Torcellia 2700:Paulocnus 2361:Ranculcus 2258:Orophodon 2082:Choloepus 2013:Imagocnus 1814:Xenarthra 1794:Kingdom: 1742:254699242 1734:1573-7055 1689:1342-937X 1657:: 30–59. 1631:129225428 1573:1477-2019 1526:129207595 1518:1477-2019 1458:254688963 1450:1064-7554 1407:1477-2019 1317:174813630 1301:2397-334X 1250:177661447 1234:0960-9822 1166:1063-5157 1081:2215-261X 1035:1477-2019 987:0016-6995 877:from the 869:Urumaquia 797:Urumaquia 539:The term 492:Choloepus 453:June 2022 320:Venezuela 164:Kingdom: 158:Eukaryota 3343:Q6090179 3337:Wikidata 2856:Meizonyx 2734:Diodomus 2677:Hapalops 2603:Hapalops 2593:Bradypus 2402:Lestodon 2183:Valgipes 2048:Parocnus 2027:Mesocnus 1968:Folivora 1948:Tamandua 1891:Cyclopes 1808:Mammalia 1802:Chordata 1800:Phylum: 1796:Animalia 1581:90207481 1415:84357359 1309:31171860 1242:31178321 1174:30239971 1130:32842894 1043:86701294 881:family. 863:. Other 764:manatees 647:Lestodon 378:, while 375:Lestodon 288:Carlini 273:Carlini 204:Family: 188:Mammalia 178:Chordata 174:Phylum: 168:Animalia 154:Domain: 3370:1367752 3357:4834501 3217:Ocnopus 3120:Planops 2464:Mylodon 2112:Catonyx 2041:Neocnus 2034:Miocnus 1806:Class: 1783:Pilosan 1659:Bibcode 1609:Bibcode 1553:Bibcode 1488:Bibcode 1387:Bibcode 1281:Bibcode 1202:Bibcode 1121:7480153 1015:Bibcode 967:Bibcode 959:Geobios 851:wetland 844:Ipururo 748:Urumaco 744:Socorro 695:Guanare 689:on the 545:Planops 510:Bolivia 292:., 2006 277:., 2006 217:Genus: 194:Order: 184:Class: 1829:Pilosa 1785:genera 1740:  1732:  1687:  1629:  1579:  1571:  1524:  1516:  1456:  1448:  1413:  1405:  1315:  1307:  1299:  1248:  1240:  1232:  1172:  1164:  1128:  1118:  1079:  1075:(63). 1041:  1033:  985:  776:sloths 740:FalcĂłn 725:Middle 681:So far 625:named 496:lobate 322:, and 316:Brazil 198:Pilosa 3383:43644 3365:IRMNG 1738:S2CID 1627:S2CID 1577:S2CID 1522:S2CID 1454:S2CID 1411:S2CID 1354:(PDF) 1313:S2CID 1269:(PDF) 1246:S2CID 1039:S2CID 848:Pebas 631:et al 504:from 290:et al 275:et al 3352:GBIF 1730:ISSN 1685:ISSN 1569:ISSN 1514:ISSN 1446:ISSN 1403:ISSN 1305:PMID 1297:ISSN 1238:PMID 1230:ISSN 1170:PMID 1162:ISSN 1126:PMID 1077:ISSN 1031:ISSN 983:ISSN 907:GBIF 871:and 846:and 824:and 810:and 800:and 784:and 770:and 750:and 727:and 719:and 705:and 703:fish 324:Peru 58:Preęž’ 1720:hdl 1712:doi 1675:hdl 1667:doi 1617:doi 1561:doi 1504:hdl 1496:doi 1438:doi 1395:doi 1358:doi 1289:doi 1220:hdl 1210:doi 1154:doi 1116:PMC 1108:doi 1023:doi 975:doi 637:or 508:in 443:by 3402:: 3380:: 3367:: 3354:: 3339:: 1736:. 1728:. 1718:. 1708:24 1706:. 1683:. 1673:. 1665:. 1655:31 1653:. 1649:. 1625:. 1615:. 1605:58 1603:. 1599:. 1575:. 1567:. 1559:. 1549:17 1547:. 1543:. 1520:. 1512:. 1502:. 1494:. 1482:. 1478:. 1466:^ 1452:. 1444:. 1434:20 1432:. 1409:. 1401:. 1393:. 1381:. 1311:. 1303:. 1295:. 1287:. 1275:. 1271:. 1244:. 1236:. 1228:. 1218:. 1208:. 1198:29 1196:. 1192:. 1168:. 1160:. 1150:68 1148:. 1124:. 1114:. 1104:16 1102:. 1098:. 1073:63 1071:. 1067:. 1051:^ 1037:. 1029:. 1021:. 1009:. 995:^ 981:. 973:. 963:49 961:. 957:. 945:^ 929:. 905:. 901:. 820:, 814:, 794:, 746:, 731:. 673:. 512:. 409:. 337:. 326:. 318:, 108:Pg 52:Ma 3102:? 2976:? 2929:† 2688:? 2680:? 2658:† 2644:† 2636:† 2628:† 2619:† 2609:† 2599:† 2349:? 2313:? 2199:† 2129:? 2093:† 1994:† 1976:† 1937:† 1927:† 1897:† 1867:† 1857:† 1775:e 1768:t 1761:v 1744:. 1722:: 1714:: 1691:. 1677:: 1669:: 1661:: 1633:. 1619:: 1611:: 1583:. 1563:: 1555:: 1528:. 1506:: 1498:: 1490:: 1484:4 1460:. 1440:: 1417:. 1397:: 1389:: 1383:4 1366:. 1360:: 1319:. 1291:: 1283:: 1277:3 1252:. 1222:: 1212:: 1204:: 1176:. 1156:: 1132:. 1110:: 1083:. 1045:. 1025:: 1017:: 1011:4 989:. 977:: 969:: 939:. 915:. 853:" 623:, 466:) 460:( 455:) 451:( 437:. 281:† 266:† 244:† 221:† 208:† 113:N 103:K 98:J 93:T 88:P 83:C 78:D 73:S 68:O 63:ęž’ 47:~ 45:) 41:- 37:( 33:-

Index

Early Miocene
Late Miocene
Colhuehuapian
Huayquerian
Ma
Preęž’
ęž’
O
S
D
C
P
T
J
K
Pg
N

Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Mammalia
Pilosa
Mylodontidae
Pseudoprepotherium
Type species
Mylodontidae
South America

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