352:. The material documented so far consists mainly of limb bones but includes individual skulls and remains of jaws. Body weight of around 550kg is reconstructed for the smaller relatives using a thigh bone around 42cm long. Large molds with femur lengths of 56 to 59cm are estimated to have weighed between 1.52 and 1.86 tons A surviving skull has a length of 43cm, but is partly deformed laterally, which means that only a few features are recognizable. A bent profile line was characteristic. Because of this, the rostrum and the cranium were at an angle of 130° from each other. At the occipital bone, the articular processes for the cervical spine protruded with little prominence. The alveoli of the five teeth per row of teeth typical of mylodon can be seen on the upper jaw, but the two front teeth are poorly preserved. From the alveoli, it can be seen that the rearmost tooth was the smallest and possibly had two lobes (bilobate). The fourth and third teeth were each elongated.
762:. The main components are different layers of sand, clay, and/or silt and limestone in which individual coal seams are embedded, at least in the Urumaco Formation. The rock strata were formed in what was originally a coastal area under the influence of a river delta. From the entire Urumaco sequence, many sites are documented, the exploration of which began as early as the 1950s. They are distributed over a good 60 different stratigraphic levels. The find material consists mainly of fish, especially sharks and rays. In addition, there are also reptiles such as turtles, crocodilians, and isolated snakes, as well as mammals appearing with rodents, South American ungulates,
125:
372:. The lower end of the joint was partially rotated out of the shaft axis. The inner articulated roller became larger than the outer. The lower one was somewhat narrower than the upper end of the joint. A tibia associated with a femur about 56cm long measured 29cm in length. This corresponds to the ratio known in other mylodons of extremely short lower sections of the hindlimbs compared to the upper ones. As a result, the tibia was only half the length of the femur. The proportions are broadly similar to
701:. It is a femur found in the first third of the 20th century. The site was then inaccessible for a long time, since it was created during the construction of a reservoir (the Virgen de la Coromoto reservoir) sank. However, a new fossil locality on the lake's eastern shore largely corresponds spatially and temporally to the original site. This was announced in a publication in 2016. The site contained a small collection of vertebrates, such as remains of
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498:) possess chewing surface. This particular tooth structure is widely thought to reflect a greater adaptation to grassy diets. The posterior teeth are round, oval or more complex in cross-section and correspond to molar-like teeth. The foremost tooth is designed like a canine. The rear foot also clearly shows twists so that the sole points inwards. The mylodonts can be detected for the first time in the Oligocene. One of the earliest forms is
516:
negatively. However, other lineages associated with the
Nematheriinae, the Octomylodontinae, or the Urumacotheriinae, depending on the author, are more controversial. The latter, in particular, summarizes the late Miocene representatives of northern South America. In principle, many researchers urge a revision for the entire family since many higher taxonomic units have no formal diagnosis. The position of
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617:. Carlini and colleagues had defined this using numerous finds from the Urumaco sequence in the FalcĂłn Basin of north-western Venezuela, the chronological range of which includes the Middle and Upper Miocene. However, a 2020 revision of the fossil remains by Ascanio D. RincĂłn and H. Gregory McDonald led to the dissolution of the genus and relegation to the genus
551:, which also occurs there and is closely related. Both had already been described by Florentino Ameghino at the end of the 19th century using finds from the Santa Cruz Formation. In 1934, R. Lee Collins referred a femur from the RĂo Yuca Formation on the RĂo Tucupido near Guanare in the Venezuelan state of Portuguesa to
841:
come from the western Amazon basin and date to the Middle to Upper
Miocene. They are part of many fossil remains from numerous local sites spread over a large area. Significant find areas are on the RĂo Sepa and the RĂo Inuya, which flow into the RĂo Urubamba in central Peru. From here, the find area
571:
to the
Mylodontidae. Their assessment was based on extensive finds from the important Middle Miocene fossil site of La Venta in Colombia in connection with Collins' thigh find. In hindsight, Hirschfeld's characterization diagnosis turned out to be incorrect since Collins' find from the RĂo Yuca
330:
lived in a tropical climate with a water-rich environment. Their known remains are limited to limb bones, except for a few skulls and teeth. Based on these remains, they were most likely medium to large-sized mylodontid. The genus was described in 1961 and currently contains three species, which were
355:
As is usual in the
Mylodontidae, the femur stood out due to its flat, board-like shape in front and behind. The shaft was slightly curved at the side. There was only a shallow indentation between the spherical condyle and the greater trochanter. The great calculus was massive but not very elevated.
515:
The internal organization of the
Mylodontidae is complex and currently under discussion. A relatively wide recognition usually only finds the late development lines with the Mylodontinae and Lestodontinae, as several studies have shown since 2004, but they are sometimes also discussed
494:) also includes one of the two sloth genera that still exist today. The Mylodontidae form one of the most diverse groups within the sloths. Characteristic features can be found in the high-crowned teeth, which, unlike those of the Megatherioidea and Megalocnoidea, have a rather flat (
486:. In a classic system based on skeletal anatomical features, the Mylodontoidea form a sister group to the Megatherioidea and thus one of the two major lineages of sloths. Molecular genetic analyses and protein studies also differentiate a third large lineage from the
1594:
Tejada-Lara, Julia V.; Salas-Gismondi, Rodolfo; Pujos, François; Baby, Patrice; Benammi, Mouloud; Brusset, Stéphane; De
Franceschi, Dario; Espurt, Nicolas; Urbina, Mario; Antoine, Pierre-Olivier (March 2015). Goswami, Anjali (ed.).
1644:
Antoine, Pierre
Olivier; Abello, MarĂa Alejandra; Adnet, Sylvain; Altamirano Sierra, Ali J.; Baby, Patrice; Billet, Guillaume; Boivin, Myriam; CalderĂłn, Ysabel; Candela, Adriana Magdalena; Chabain, Jules; Corfu, Fernando (2016).
766:, and minor jointed animals, among others. The secondary articulated animals show a high diversity, almost reaching that of the contemporary fauna of southern South America in the Pampas region or Mesopotamia. Armadillos, the
1143:
Varela, Luciano; Tambusso, P Sebastián; McDonald, H Gregory; Fariña, Richard A (2018-09-15). "Phylogeny, Macroevolutionary Trends and
Historical Biogeography of Sloths: Insights From a Bayesian Morphological Clock Analysis".
405:. The size of the bite marks, from three to 15 mm in diameter, allows the length of the attacker to be reconstructed to be around 4m. It caught its prey with its front teeth. Attacks on the hind legs are also known by modern
528:, a form widely found in Central and North America. However, the limb bones usually only offer a limited selection of features for determining family relationships. Therefore, investigations on the little skull material see
400:
tibia from the Middle
Miocene Pebas Formation near Iquitos in the western Amazon Basin, about 23 cm long, shows more than 60 bite marks, the size and arrangement of which suggest that they were probably caused by a juvenile
362:. The lesser trochanter was only weakly developed. The third trochanter appeared as a slight bulge around the middle of the shaft and continued as an edge to the lower end of the joint. The position roughly matched that of
1263:
Presslee, Samantha; Slater, Graham J.; Pujos, François; Forasiepi, AnalĂa M.; Fischer, Roman; Molloy, Kelly; Mackie, Meaghan; Olsen, Jesper V.; Kramarz, Alejandro; Taglioretti, MatĂas; Scaglia, Fernando (July 2019).
669:. A skull and several limb bones from La Venta formed this basis. Referring to deviations in the structure of the femur, RincĂłn and McDonald reclassified the 2020 form as the type species of the new genus
1187:
Delsuc, FrĂ©dĂ©ric; Kuch, Melanie; Gibb, Gillian C.; Karpinski, Emil; Hackenberger, Dirk; Szpak, Paul; MartĂnez, Jorge G.; Mead, Jim I.; McDonald, H. Gregory; MacPhee, Ross D. E.; Billet, Guillaume (2019-06-17).
850:
formations. These go back to a phase when the so-called "Proto-Amazon" existed, a landscape characterized by lakes, swamps and rivers connected to the
Caribbean. It can now be described as "the Pebas mega
1540:
723:. The RĂo Yuca Formation is composed largely of limestone and sandstone with discrete interstices of the conglomerate. It originated in a freshwater environment with a formation period possibly in the
1702:
Antoine, Pierre-Olivier; Salas-Gismondi, Rodolfo; Pujos, François; Ganerød, Morgan; Marivaux, Laurent (2017-03-01). "Western Amazonia as a Hotspot of Mammalian Biodiversity Throughout the Cenozoic".
613:. The other two species, which are significantly larger with femur lengths of 56 to 59 cm, were recognized in 2006 by a work team led by Alfredo A. Carlini but were originally included in the genus
955:"A new Miocene vertebrate assemblage from the RĂo Yuca Formation (Venezuela) and the northernmost record of typical Miocene mammals of high latitude (Patagonian) affinities in South America"
830:
from the lineage Mylodontidae and their immediate relatives. As a special circumstance of taphonomy, the frequent tradition of limb elements in sloths is to be evaluated, however, from
734:
The most extensive fossil material to date belongs to the Urumaco sequence, a complex depositional unit that is predominantly exposed in the approximately 36,000 km² large
1005:
Carlini, Alfredo A.; Scillato-Yané, Gustavo J.; Sánchez, Rodolfo (2006-01-01). "New Mylodontoidea (Xenarthra, Phyllophaga) from the Middle Miocene-Pliocene of Venezuela".
520:
within the Mylodontidae is, therefore, ambiguous since the genus is largely defined by the limb bones. Based on their characteristics, phylogenetic analyzes indicate that
396:
was probably more adapted to tropical climate conditions. The find locations in the deposit units of the Urumaco sequence also speak in favor of this. A possibly related
3364:
1065:"Reexamination of the Relationship of Pseudoprepotherium Hoffstetter, 1961, to the Mylodont Ground Sloths (Xenarthra) from the Miocene of Northern South America"
867:
finds from the region belong to the transition to the Upper Miocene and include a mandible. At the same time, the Pebas mega wetland also includes the related
1377:
Sánchez-Villagra, Marcelo R.; Aguilera, Orangel A. (January 2006). "Neogene vertebrates from Urumaco, Falcón State, Venezuela: Diversity and significance".
609:
is the type species and the smallest representative with a femur length of around 42 cm. It is based on the form introduced by R. Lee Collins in 1934 as
1476:"First Megatheriines (Xenarthra, Phyllophaga, Megatheriidae) from the Urumaco (Late Miocene) and Codore (Pliocene) Formations, Estado FalcĂłn, Venezuela"
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Stratigraphy and sedimentary environments of Miocene shallow to marginal marine deposits in the theUrumaco trough, FalcĂłn Basin, Western Venezuela.
842:
extends eastward to western Brazil and northern Bolivia and northward to the Iquitos region of northern Peru. The found material belongs to the
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possessed even shorter lower leg sections. The shaft of the tibia narrowed sharply in the middle while the ends of the joints protruded far.
633:, also from the Urumaco sequence, is considered a nomen dubium because its femur does not show the corresponding diagnostic features of
440:
572:
Formation and the La Venta material is from today's perspective assigned to different genera, but their assessment of the position of
547:, a member of the Megatheriidae from the Santa Cruz Formation of Patagonia's Early and Middle Miocene. He also referred to the genus
478:
is a member of the Mylodontidae family within the suborder (Folivora). The Mylodontidae, in turn, are often placed together with the
758:
being limited to the two lower and first-mentioned sequences. The Urumaco sequence covers the period from the Middle Miocene to the
462:
1350:
MorfologĂa sistemática y paleobiologĂa de los perezosos gigantes del gĂ©nero Lestodon Gervais 1855 (Mammalia, Xenarthra, Tardigrada)
1541:"Two new megalonychid sloths (Mammalia: Xenarthra) from the Urumaco Formation (late Miocene), and their phylogenetic affinities"
124:
720:
145:
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was scientifically introduced in 1961 by Robert Hoffstetter. He mentioned them in a publication of a skeletal description of
444:
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Hastings, Alexander K. (2012-06-14). "The Incredible Fossils of Urumaco and Beyond: Exploring Venezuela's Geologic Past".
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during the Early to Late Miocene epoch around 21 to 5.3 million years ago. Fossils of the animal have been found in
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1096:"Predation of the giant Miocene caiman Purussaurus on a mylodontid ground sloth in the wetlands of proto-Amazonia"
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has only been found in the northern part of South America. The genus defining fossil remains came to Taqe in the
657:, to which they attributed further skeletal material; RincĂłn and McDonald, in turn, kept only the skull at
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778:, have been proven. Mainly in the late 20th and early 21st century, numerous new forms were described, such as
649:, which did not occur in northern South America; Carlini et al. 2006 simultaneously removed the femur of
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Rincón, Ascanio D.; Solórzano, Andrés; Macsotay, Oliver; McDonald, H. Gregory; Núñez-Flores, Mónica (2016).
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Rincón, Ascanio D.; Solórzano, Andrés; McDonald, H. Gregory; Montellano-Ballesteros, Marisol (2019-03-04).
490:. According to the results of the latter two analysis methods, the Mylodontoidea with the two-toed sloths (
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based on the postcranial skeletal elements ). As early as 1985, Sue Hirschfeld named the species
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1647:"A 60-million-year Cenozoic history of western Amazonian ecosystems in Contamana, eastern Peru"
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1597:"Life in proto-Amazonia: Middle Miocene mammals from the Fitzcarrald Arch (Peruvian Amazonia)"
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At first, he saw their systematic position as ambiguous. In 1985, Sue Hirschfeld moved
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Its apex was at or slightly below the level of the condyle and thus lower as compared to
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In: Marcelo R. Sánchez-Villagra, Orangel A. Aguilera und Alfredo A. Carlini (Hrsg.):
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Urumaco and Venezuelan paleontology, the fossil record of the northern Neotropics.
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clearly more basally embedded in the mylodonts and move the form partly closer to
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1190:"Ancient Mitogenomes Reveal the Evolutionary History and Biogeography of Sloths"
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447: in this section. Unsourced material may be challenged and removed.
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genus based on anatomical differences and established a new one with
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is more closely related to some more modern representatives, such as
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1967:
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is based on the characteristics of the femur divided until today.
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Carlini, Alfredo A.; Brandoni, Diego; Sánchez, Rodolfo (2006).
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559:. In 1961, Hoffstetter classified this femur as part of the
621:, largely based on the design of the femur. A third species
859:, which is now considered the type form of the genus
855:. Some of the finds, however, are assigned to the species
742:. It is composed of the lithostratigraphic units of the
1469:
1467:
1094:
Pujos, François; Salas-Gismondi, Rodolfo (2020-08-26).
1063:
RincĂłn, Ascanio D.; McDonald, H. Gregory (2020-08-07).
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Due to its far northern distribution in South America,
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348:is a medium to a large-sized member of the family
1266:"Palaeoproteomics resolves sloth relationships"
596:( Carlini, Scillato-Yané & Sánchez , 2006)
590:(Carlini , Scillato-Yané & Sánchez , 2006)
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8:
1356:(Thesis). Universidad Nacional de La Plata.
583:are currently considered to be valid today:
306:is an extinct genus of sloths of the family
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1337:Indiana University Press 2010, S. 153–172
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463:Learn how and when to remove this message
1329:Luis I. Quiroz und Carlos A. Jaramillo:
890:
899:"Pseudoprepotherium Hoffstetter, 1961"
834:also documented remains of the skull.
653:and reassigned the rest, a skull, to
645:was originally assigned to the giant
7:
1069:Revista Geológica de América Central
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1000:
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536:, but also to the Scelidotheriidae.
445:adding citations to reliable sources
310:. It was widespread across northern
1545:Journal of Systematic Palaeontology
1480:Journal of Systematic Palaeontology
1379:Journal of Systematic Palaeontology
1007:Journal of Systematic Palaeontology
555:and set up with it the new species
14:
927:"Fossilworks: Pseudoprepotherium"
331:originally assigned to the genus
3431:Taxa named by Robert Hoffstetter
3421:Miocene mammals of South America
3298:
2542:
806:as representatives of the large
421:
144:
432:needs additional citations for
248:Pseudoprepotherium venezuelanum
1704:Journal of Mammalian Evolution
1430:Journal of Mammalian Evolution
1273:Nature Ecology & Evolution
16:Extinct genus of ground sloths
1:
3426:Fossil taxa described in 1961
1565:10.1080/14772019.2018.1427639
979:10.1016/j.geobios.2016.06.005
693:about 11km west-southwest of
366:, but was lower than that of
3411:Prehistoric placental genera
579:A total of three species of
857:Pseudoprepotherium confusum
738:in the Venezuelan state of
697:in the Venezuelan state of
667:Pseudoprepotherium confusum
3477:
1347:Czerwonogora, Ada (2010).
837:Finds possibly related to
482:in the superfamily of the
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3093:
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2815:
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2583:
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2076:
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1988:
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1839:
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1822:
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1716:10.1007/s10914-016-9333-1
1500:10.1017/S1477201906001878
1442:10.1007/s10914-012-9208-z
1399:10.1017/s1477201906001829
1293:10.1038/s41559-019-0909-z
1215:10.1016/j.cub.2019.05.043
1027:10.1017/S147720190600191X
611:Prepotherium venezuelanum
557:Prepotherium venezuelanum
263:
258:
242:
235:
141:Scientific classification
139:
131:
122:
23:
1671:10.1016/j.gr.2015.11.001
721:South American ungulates
1112:10.1098/rsbl.2020.0239
661:in 2020 and split off
3378:Paleobiology Database
1158:10.1093/sysbio/syy058
715:, as well as armored
3441:Fossils of Venezuela
1363:10.35537/10915/80374
1200:(12): 2031–2042.e6.
683:, Pseudoprepotherium
651:Mirandabradys zabasi
643:Mirandabradys zabasi
627:Mirandabradys zabasi
565:Pseudoprepotherium .
441:improve this article
3416:Miocene xenarthrans
3224:Perezfontanatherium
2265:Paroctodontotherium
1663:2016GondR..31...30A
1613:2015Palgy..58..341T
1557:2019JSPal..17..409R
1492:2006JSPal...4..269C
1391:2006JSPal...4..213S
1285:2019NatEE...3.1121P
1206:2019CBio...29E2031D
1019:2006JSPal...4..255C
971:2016Geobi..49..395R
931:www.fossilworks.org
501:Paroctodontotherium
3406:Prehistoric sloths
3328:Pseudoprepotherium
2755:Megalonychotherium
2272:Pseudoprepotherium
1622:10.1111/pala.12147
1146:Systematic Biology
865:Pseudoprepotherium
839:Pseudoprepotherium
832:Pseudoprepotherium
756:Pseudoprepotherium
754:, with remains of
687:RĂo Yuca Formation
639:Pseudoprepotherium
619:Pseudoprepotherium
581:Pseudoprepotherium
574:Pseudoprepotherium
569:Pseudoprepotherium
541:Pseudoprepotherium
530:Pseudoprepotherium
522:Pseudoprepotherium
518:Pseudoprepotherium
476:Pseudoprepotherium
398:Pseudoprepotherium
394:Pseudoprepotherium
346:Pseudoprepotherium
328:Pseudoprepotherium
303:Pseudoprepotherium
225:Pseudoprepotherium
132:Holotype femur of
25:Pseudoprepotherium
3461:Fossils of Brazil
3436:Neogene Venezuela
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3392:
3320:Taxon identifiers
3311:
3310:
3307:
3306:
3294:
3293:
3290:
3289:
3286:
3285:
3282:
3281:
3259:Pyramiodontherium
3231:Plesiomegatherium
3080:
3079:
2922:
2921:
2918:
2917:
2914:
2913:
2783:Proschismotherium
2551:
2550:
2538:
2537:
2534:
2533:
2530:
2529:
2502:
2501:
2498:
2497:
2494:
2493:
2192:
2191:
2057:
2056:
1961:
1960:
1957:
1956:
1651:Gondwana Research
790:from the family
752:Codore Formations
473:
472:
465:
299:
298:
293:
278:
231:
3468:
3386:
3385:
3373:
3372:
3360:
3359:
3347:
3346:
3345:
3315:
3302:
3091:
3064:Pronothrotherium
2958:
2949:
2930:
2813:
2790:Pseudortotherium
2721:
2659:
2620:
2610:
2600:
2585:
2581:
2570:
2557:
2546:
2508:
2318:Glossotheriopsis
2295:
2244:Octodontotherium
2208:
2200:
2155:Scelidotheridium
2096:Scelidotheriidae
2094:
2074:
2070:
1995:
1990:
1986:
1938:
1928:
1898:
1868:
1858:
1850:
1841:
1837:
1824:
1776:
1769:
1762:
1753:
1746:
1745:
1727:
1699:
1693:
1692:
1682:
1641:
1635:
1634:
1624:
1591:
1585:
1584:
1536:
1530:
1529:
1511:
1471:
1462:
1461:
1425:
1419:
1418:
1374:
1368:
1367:
1365:
1355:
1344:
1338:
1327:
1321:
1320:
1279:(7): 1121–1130.
1270:
1260:
1254:
1253:
1227:
1217:
1184:
1178:
1177:
1140:
1134:
1133:
1123:
1091:
1085:
1084:
1060:
1047:
1046:
1002:
991:
990:
950:
941:
940:
938:
937:
923:
917:
916:
914:
913:
895:
602:( Collins, 1934)
480:Scelidotheriidae
468:
461:
457:
454:
448:
425:
417:
287:
272:
253:Hofstetter, 1961
245:
230:Hofstetter, 1961
229:
222:
209:
149:
148:
127:
117:
54:
29:Temporal range:
21:
3476:
3475:
3471:
3470:
3469:
3467:
3466:
3465:
3451:Fossils of Peru
3396:
3395:
3394:
3389:
3381:
3376:
3368:
3363:
3355:
3350:
3341:
3340:
3335:
3322:
3312:
3303:
3278:
3245:Proeremotherium
3238:Pliomegatherium
3161:Anisodontherium
3146:
3141:Proprepotherium
3105:
3076:
2985:Nothrotheriinae
2979:
2953:Nothrotheriidae
2947:
2934:
2910:
2802:
2719:
2707:Proplatyarthrus
2652:
2577:
2564:
2547:
2526:
2490:
2443:Glossotheridium
2428:
2409:Magdalenabradys
2373:
2284:
2237:Octodontobradys
2204:
2188:
2162:Scelidotheriops
2087:
2066:
2053:
1984:
1979:Pseudoglyptodon
1970:
1953:
1912:Myrmecophagidae
1906:
1876:
1831:
1818:
1786:
1780:
1750:
1749:
1701:
1700:
1696:
1643:
1642:
1638:
1593:
1592:
1588:
1538:
1537:
1533:
1473:
1472:
1465:
1427:
1426:
1422:
1376:
1375:
1371:
1353:
1346:
1345:
1341:
1328:
1324:
1268:
1262:
1261:
1257:
1194:Current Biology
1186:
1185:
1181:
1142:
1141:
1137:
1106:(8): 20200239.
1100:Biology Letters
1093:
1092:
1088:
1062:
1061:
1050:
1004:
1003:
994:
952:
951:
944:
935:
933:
925:
924:
920:
911:
909:
897:
896:
892:
887:
861:Magdalenabradys
812:Magdalenabradys
803:Proeremotherium
679:
671:Magdalenabradys
663:Magdalenabradys
607:P. venezuelanum
600:P. venezuelanum
594:P. urumaquensis
469:
458:
452:
449:
438:
426:
415:
390:
364:Magdalenabradys
359:Magdalenabradys
343:
284:P. urumaquensis
254:
251:
228:
220:
207:
143:
118:
116:
115:
110:
105:
100:
95:
90:
85:
80:
75:
70:
65:
60:
49:
48:
46:
27:
17:
12:
11:
5:
3474:
3472:
3464:
3463:
3458:
3456:Neogene Brazil
3453:
3448:
3443:
3438:
3433:
3428:
3423:
3418:
3413:
3408:
3398:
3397:
3391:
3390:
3388:
3387:
3374:
3361:
3348:
3332:
3330:
3324:
3323:
3318:
3309:
3308:
3305:
3304:
3297:
3295:
3292:
3291:
3288:
3287:
3284:
3283:
3280:
3279:
3277:
3276:
3269:
3262:
3255:
3252:Promegatherium
3248:
3241:
3234:
3227:
3220:
3213:
3206:
3199:
3192:
3189:Megathericulus
3185:
3178:
3171:
3168:Diabolotherium
3164:
3156:
3154:
3148:
3147:
3145:
3144:
3137:
3130:
3123:
3115:
3113:
3107:
3106:
3104:
3103:
3094:
3088:
3082:
3081:
3078:
3077:
3075:
3074:
3067:
3060:
3053:
3050:Nothrotheriops
3046:
3039:
3032:
3025:
3018:
3011:
3004:
2997:
2989:
2987:
2981:
2980:
2978:
2977:
2969:
2961:
2955:
2946:
2945:
2937:
2935:
2927:
2924:
2923:
2920:
2919:
2916:
2915:
2912:
2911:
2909:
2908:
2901:
2894:
2887:
2884:Protomegalonyx
2880:
2877:Pliometanastes
2873:
2870:Pattersonocnus
2866:
2859:
2852:
2845:
2838:
2831:
2824:
2816:
2810:
2808:Megalonychinae
2804:
2803:
2801:
2800:
2793:
2786:
2779:
2772:
2765:
2758:
2751:
2744:
2737:
2729:
2727:
2718:
2717:
2710:
2703:
2696:
2693:Mesopotamocnus
2689:
2681:
2673:
2670:Deseadognathus
2665:
2663:
2661:Megalonychidae
2654:
2653:
2651:
2650:
2642:
2639:Schismotherium
2634:
2626:
2616:
2606:
2596:
2588:
2578:
2575:Megatherioidea
2573:
2566:
2565:
2562:Megatherioidea
2560:
2553:
2552:
2549:
2548:
2541:
2539:
2536:
2535:
2532:
2531:
2528:
2527:
2525:
2524:
2523:
2522:
2516:
2514:
2512:Megatherioidea
2504:
2503:
2500:
2499:
2496:
2495:
2492:
2491:
2489:
2488:
2481:
2478:Pleurolestodon
2474:
2467:
2460:
2453:
2446:
2438:
2436:
2430:
2429:
2427:
2426:
2419:
2412:
2405:
2398:
2391:
2388:Bolivartherium
2383:
2381:
2375:
2374:
2372:
2371:
2364:
2357:
2350:
2342:
2335:
2328:
2321:
2314:
2306:
2303:Archaeomylodon
2298:
2292:
2286:
2285:
2283:
2282:
2279:Urumacotherium
2275:
2268:
2261:
2254:
2247:
2240:
2233:
2226:
2223:Eionaletherium
2219:
2216:Baraguatherium
2211:
2205:
2197:
2194:
2193:
2190:
2189:
2187:
2186:
2179:
2176:Sibyllotherium
2172:
2169:Scelidotherium
2165:
2158:
2151:
2144:
2137:
2134:Neonematherium
2130:
2122:
2115:
2108:
2100:
2098:
2089:
2088:
2086:
2085:
2077:
2067:
2062:
2059:
2058:
2055:
2054:
2052:
2051:
2044:
2037:
2030:
2023:
2016:
2009:
2001:
1999:
1983:
1982:
1973:
1971:
1966:
1963:
1962:
1959:
1958:
1955:
1954:
1952:
1951:
1944:
1934:
1924:
1916:
1914:
1908:
1907:
1905:
1904:
1901:Palaeomyrmidon
1894:
1886:
1884:
1878:
1877:
1875:
1874:
1864:
1853:
1847:
1833:
1832:
1827:
1820:
1819:
1817:
1816:
1810:
1804:
1798:
1791:
1788:
1787:
1781:
1779:
1778:
1771:
1764:
1756:
1748:
1747:
1694:
1636:
1607:(2): 341–378.
1586:
1551:(5): 409–421.
1531:
1486:(3): 269–278.
1463:
1436:(2): 147–148.
1420:
1385:(3): 213–220.
1369:
1339:
1322:
1255:
1179:
1152:(2): 204–218.
1135:
1086:
1048:
1013:(3): 255–267.
992:
965:(5): 395–405.
942:
918:
889:
888:
886:
883:
874:Megathericulus
827:Urumacotherium
822:Eionaletherium
817:Bolivartherium
792:Megalonychidae
787:Pattersonocnus
772:Glyptodontinae
768:Pampatheriidae
760:Early Pliocene
678:
675:
659:Bolivartherium
655:Bolivartherium
604:
603:
597:
591:
588:P. socorrensis
534:Urumacotherium
471:
470:
429:
427:
420:
414:
413:Classification
411:
389:
386:
369:Eionaletherium
342:
339:
297:
296:
295:
294:
279:
269:P. socorrensis
261:
260:
259:Other species
256:
255:
252:
240:
239:
233:
232:
218:
214:
213:
205:
201:
200:
195:
191:
190:
185:
181:
180:
175:
171:
170:
165:
161:
160:
155:
151:
150:
137:
136:
129:
128:
120:
119:
111:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
55:
50:21.0–5.3
28:
15:
13:
10:
9:
6:
4:
3:
2:
3473:
3462:
3459:
3457:
3454:
3452:
3449:
3447:
3444:
3442:
3439:
3437:
3434:
3432:
3429:
3427:
3424:
3422:
3419:
3417:
3414:
3412:
3409:
3407:
3404:
3403:
3401:
3384:
3379:
3375:
3371:
3366:
3362:
3358:
3353:
3349:
3344:
3338:
3334:
3333:
3331:
3329:
3325:
3321:
3316:
3301:
3275:
3274:
3270:
3268:
3267:
3263:
3261:
3260:
3256:
3254:
3253:
3249:
3247:
3246:
3242:
3240:
3239:
3235:
3233:
3232:
3228:
3226:
3225:
3221:
3219:
3218:
3214:
3212:
3211:
3207:
3205:
3204:
3200:
3198:
3197:
3196:Megatheridium
3193:
3191:
3190:
3186:
3184:
3183:
3179:
3177:
3176:
3175:Eomegatherium
3172:
3170:
3169:
3165:
3163:
3162:
3158:
3157:
3155:
3153:
3152:Megatheriinae
3149:
3143:
3142:
3138:
3136:
3135:
3131:
3129:
3128:
3124:
3122:
3121:
3117:
3116:
3114:
3112:
3108:
3101:
3100:
3096:
3095:
3092:
3089:
3087:
3086:Megatheriidae
3083:
3073:
3072:
3068:
3066:
3065:
3061:
3059:
3058:
3057:Nothrotherium
3054:
3052:
3051:
3047:
3045:
3044:
3040:
3038:
3037:
3033:
3031:
3030:
3029:Mcdonaldocnus
3026:
3024:
3023:
3019:
3017:
3016:
3012:
3010:
3009:
3008:Chasicobradys
3005:
3003:
3002:
3001:Aymaratherium
2998:
2996:
2995:
2991:
2990:
2988:
2986:
2982:
2975:
2974:
2970:
2968:
2967:
2963:
2962:
2959:
2956:
2954:
2950:
2944:
2943:
2942:Analcimorphus
2939:
2938:
2936:
2933:
2925:
2907:
2906:
2902:
2900:
2899:
2895:
2893:
2892:
2888:
2886:
2885:
2881:
2879:
2878:
2874:
2872:
2871:
2867:
2865:
2864:
2860:
2858:
2857:
2853:
2851:
2850:
2846:
2844:
2843:
2839:
2837:
2836:
2835:Megalonychops
2832:
2830:
2829:
2825:
2823:
2822:
2818:
2817:
2814:
2811:
2809:
2805:
2799:
2798:
2794:
2792:
2791:
2787:
2785:
2784:
2780:
2778:
2777:
2773:
2771:
2770:
2766:
2764:
2763:
2759:
2757:
2756:
2752:
2750:
2749:
2745:
2743:
2742:
2738:
2736:
2735:
2731:
2730:
2728:
2726:
2725:Ortotheriinae
2722:
2716:
2715:
2711:
2709:
2708:
2704:
2702:
2701:
2697:
2695:
2694:
2690:
2687:
2686:
2682:
2679:
2678:
2674:
2672:
2671:
2667:
2666:
2664:
2662:
2655:
2649:
2648:
2647:Similhapalops
2643:
2641:
2640:
2635:
2633:
2632:
2627:
2625:
2624:
2617:
2615:
2614:
2607:
2605:
2604:
2597:
2595:
2594:
2590:
2589:
2586:
2582:
2579:
2576:
2571:
2567:
2563:
2558:
2554:
2545:
2520:
2519:
2518:
2517:
2515:
2513:
2509:
2505:
2487:
2486:
2482:
2480:
2479:
2475:
2473:
2472:
2468:
2466:
2465:
2461:
2459:
2458:
2454:
2452:
2451:
2450:Glossotherium
2447:
2445:
2444:
2440:
2439:
2437:
2435:
2431:
2425:
2424:
2423:Thinobadistes
2420:
2418:
2417:
2413:
2411:
2410:
2406:
2404:
2403:
2399:
2397:
2396:
2392:
2390:
2389:
2385:
2384:
2382:
2380:
2376:
2370:
2369:
2365:
2363:
2362:
2358:
2356:
2355:
2351:
2348:
2347:
2343:
2341:
2340:
2336:
2334:
2333:
2329:
2327:
2326:
2322:
2320:
2319:
2315:
2312:
2311:
2307:
2305:
2304:
2300:
2299:
2296:
2293:
2291:
2287:
2281:
2280:
2276:
2274:
2273:
2269:
2267:
2266:
2262:
2260:
2259:
2255:
2253:
2252:
2248:
2246:
2245:
2241:
2239:
2238:
2234:
2232:
2231:
2227:
2225:
2224:
2220:
2218:
2217:
2213:
2212:
2209:
2206:
2203:
2195:
2185:
2184:
2180:
2178:
2177:
2173:
2171:
2170:
2166:
2164:
2163:
2159:
2157:
2156:
2152:
2150:
2149:
2145:
2143:
2142:
2141:Proscelidodon
2138:
2136:
2135:
2131:
2128:
2127:
2123:
2121:
2120:
2116:
2114:
2113:
2109:
2107:
2106:
2105:Analcitherium
2102:
2101:
2099:
2097:
2090:
2084:
2083:
2079:
2078:
2075:
2071:
2068:
2065:
2064:Mylodontoidea
2060:
2050:
2049:
2045:
2043:
2042:
2038:
2036:
2035:
2031:
2029:
2028:
2024:
2022:
2021:
2017:
2015:
2014:
2010:
2008:
2007:
2003:
2002:
2000:
1998:
1991:
1987:
1981:
1980:
1975:
1974:
1972:
1969:
1964:
1950:
1949:
1945:
1943:
1942:
1935:
1933:
1932:
1925:
1923:
1922:
1918:
1917:
1915:
1913:
1909:
1903:
1902:
1895:
1893:
1892:
1888:
1887:
1885:
1883:
1879:
1873:
1872:
1865:
1863:
1862:
1855:
1854:
1851:
1848:
1846:
1842:
1838:
1834:
1830:
1825:
1821:
1815:
1811:
1809:
1805:
1803:
1799:
1797:
1793:
1792:
1789:
1784:
1777:
1772:
1770:
1765:
1763:
1758:
1757:
1754:
1743:
1739:
1735:
1731:
1726:
1721:
1717:
1713:
1709:
1705:
1698:
1695:
1690:
1686:
1681:
1676:
1672:
1668:
1664:
1660:
1656:
1652:
1648:
1640:
1637:
1632:
1628:
1623:
1618:
1614:
1610:
1606:
1602:
1601:Palaeontology
1598:
1590:
1587:
1582:
1578:
1574:
1570:
1566:
1562:
1558:
1554:
1550:
1546:
1542:
1535:
1532:
1527:
1523:
1519:
1515:
1510:
1505:
1501:
1497:
1493:
1489:
1485:
1481:
1477:
1470:
1468:
1464:
1459:
1455:
1451:
1447:
1443:
1439:
1435:
1431:
1424:
1421:
1416:
1412:
1408:
1404:
1400:
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919:
908:
904:
900:
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891:
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879:Megatheriidae
876:
875:
870:
866:
862:
858:
854:
849:
845:
840:
835:
833:
829:
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823:
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813:
809:
808:Megatheriidae
805:
804:
799:
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789:
788:
783:
782:
777:
774:, as well as
773:
769:
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745:
741:
737:
732:
730:
726:
722:
718:
717:Peltephilidae
714:
713:
708:
704:
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696:
692:
688:
684:
676:
674:
672:
668:
664:
660:
656:
652:
648:
644:
640:
636:
635:Mirandabradys
632:
628:
624:
620:
616:
615:Mirandabradys
612:
608:
601:
598:
595:
592:
589:
586:
585:
584:
582:
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570:
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546:
542:
537:
535:
531:
527:
526:Thinobadistes
523:
519:
513:
511:
507:
506:Salla-Luribay
503:
502:
497:
493:
489:
488:Megalocnoidea
485:
484:Mylodontoidea
481:
477:
467:
464:
456:
446:
442:
436:
435:
430:This section
428:
424:
419:
418:
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410:
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404:
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395:
387:
385:
383:
382:
381:Glossotherium
377:
376:
371:
370:
365:
361:
360:
353:
351:
347:
340:
338:
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335:
329:
325:
321:
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313:
312:South America
309:
305:
304:
291:
286:
285:
280:
276:
271:
270:
265:
264:
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227:
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135:
134:P. venezuelam
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109:
104:
99:
94:
89:
84:
79:
74:
69:
64:
59:
53:
44:
40:
39:Colhuehuapian
36:
32:
31:Early Miocene
26:
22:
19:
3446:Neogene Peru
3327:
3271:
3264:
3257:
3250:
3243:
3236:
3229:
3222:
3215:
3208:
3203:Megatheriops
3201:
3194:
3187:
3182:Eremotherium
3180:
3173:
3166:
3159:
3139:
3134:Prepoplanops
3132:
3127:Prepotherium
3125:
3118:
3099:Thalassocnus
3097:
3069:
3062:
3055:
3048:
3041:
3036:Mionothropus
3034:
3027:
3020:
3013:
3006:
2999:
2992:
2973:Thalassocnus
2971:
2964:
2940:
2903:
2896:
2889:
2882:
2875:
2868:
2861:
2854:
2847:
2840:
2833:
2826:
2819:
2795:
2788:
2781:
2774:
2769:Paranabradys
2767:
2760:
2753:
2746:
2739:
2732:
2712:
2705:
2698:
2691:
2683:
2675:
2668:
2645:
2637:
2629:
2621:
2613:Hiskatherium
2611:
2601:
2591:
2483:
2476:
2469:
2462:
2455:
2448:
2441:
2421:
2416:Sphenotherus
2414:
2407:
2400:
2393:
2386:
2379:Lestodontini
2366:
2359:
2352:
2344:
2337:
2332:Mylodonopsis
2330:
2323:
2316:
2310:Brievabradys
2308:
2301:
2290:Mylodontinae
2277:
2271:
2270:
2263:
2256:
2249:
2242:
2235:
2228:
2221:
2214:
2202:Mylodontidae
2181:
2174:
2167:
2160:
2153:
2146:
2139:
2132:
2124:
2119:Chubutherium
2117:
2110:
2103:
2080:
2046:
2039:
2032:
2025:
2018:
2011:
2004:
1997:Megalocnidae
1977:
1946:
1939:
1929:
1921:Myrmecophaga
1919:
1899:
1889:
1882:Cyclopedidae
1871:Orthoarthrus
1869:
1859:
1812:Superorder:
1707:
1703:
1697:
1654:
1650:
1639:
1604:
1600:
1589:
1548:
1544:
1534:
1483:
1479:
1433:
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1423:
1382:
1378:
1372:
1349:
1342:
1334:
1330:
1325:
1276:
1272:
1258:
1225:11336/136908
1197:
1193:
1182:
1149:
1145:
1138:
1103:
1099:
1089:
1072:
1068:
1010:
1006:
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958:
934:. Retrieved
930:
921:
910:. Retrieved
903:www.gbif.org
902:
893:
872:
868:
864:
860:
856:
852:
838:
836:
831:
825:
821:
815:
811:
801:
795:
785:
779:
755:
736:FalcĂłn Basin
733:
729:Late Miocene
710:
707:crocodilians
691:RĂo Tucupido
682:
680:
677:Paleoecology
670:
666:
662:
658:
654:
650:
646:
642:
641:(a femur of
638:
634:
630:
626:
622:
618:
614:
610:
606:
605:
599:
593:
587:
580:
578:
573:
568:
564:
561:Prepotherium
560:
556:
553:Prepotherium
552:
549:Prepotherium
548:
544:
540:
538:
533:
529:
525:
521:
517:
514:
499:
495:
491:
475:
474:
459:
450:
439:Please help
434:verification
431:
402:
397:
393:
391:
388:Paleobiology
379:
373:
367:
363:
357:
354:
350:Mylodontidae
345:
344:
334:Prepotherium
332:
327:
308:Mylodontidae
302:
301:
300:
289:
283:
282:
274:
268:
267:
247:
246:
237:Type species
224:
223:
211:Mylodontidae
133:
35:Late Miocene
24:
18:
3266:Sibotherium
3210:Megatherium
3111:Planopsinae
3015:Huilabradys
2994:Amphibradys
2966:Neohapalops
2905:Zacatzontli
2898:Xibalbaonyx
2863:Nohochichak
2828:Australonyx
2776:Pliomorphus
2762:Ortotherium
2741:Eucholoeops
2714:Urumacocnus
2685:Hyperleptus
2623:Hyperleptus
2485:Simomylodon
2471:Paramylodon
2457:Kiyumylodon
2434:Mylodontini
2395:Lestobradys
2368:Strabosodon
2346:Oreomylodon
2339:Ocnotherium
2251:Octomylodon
2230:Nematherium
2126:Nematherium
1941:Protamandua
1931:Neotamandua
1861:Argyromanis
1845:Vermilingua
1725:11336/63465
1710:(1): 5–17.
1680:11336/99030
1509:11336/80745
781:Urumacocnus
712:Purussaurus
629:by Carlini
403:Purussaurus
341:Description
43:Huayquerian
3400:Categories
3022:Lakukullus
2932:Megatheria
2891:Sinclairia
2849:Megistonyx
2821:Ahytherium
2748:Hapaloides
2631:Pelecyodon
2521:see below↓
2354:Promylodon
2325:Megabradys
2148:Scelidodon
2020:Megalocnus
2006:Acratocnus
936:2022-05-11
912:2022-05-11
885:References
709:; such as
699:Portuguesa
407:crocodiles
3273:Urumaquia
3071:Xyophorus
3043:Nothropus
2842:Megalonyx
2797:Torcellia
2700:Paulocnus
2361:Ranculcus
2258:Orophodon
2082:Choloepus
2013:Imagocnus
1814:Xenarthra
1794:Kingdom:
1742:254699242
1734:1573-7055
1689:1342-937X
1657:: 30–59.
1631:129225428
1573:1477-2019
1526:129207595
1518:1477-2019
1458:254688963
1450:1064-7554
1407:1477-2019
1317:174813630
1301:2397-334X
1250:177661447
1234:0960-9822
1166:1063-5157
1081:2215-261X
1035:1477-2019
987:0016-6995
877:from the
869:Urumaquia
797:Urumaquia
539:The term
492:Choloepus
453:June 2022
320:Venezuela
164:Kingdom:
158:Eukaryota
3343:Q6090179
3337:Wikidata
2856:Meizonyx
2734:Diodomus
2677:Hapalops
2603:Hapalops
2593:Bradypus
2402:Lestodon
2183:Valgipes
2048:Parocnus
2027:Mesocnus
1968:Folivora
1948:Tamandua
1891:Cyclopes
1808:Mammalia
1802:Chordata
1800:Phylum:
1796:Animalia
1581:90207481
1415:84357359
1309:31171860
1242:31178321
1174:30239971
1130:32842894
1043:86701294
881:family.
863:. Other
764:manatees
647:Lestodon
378:, while
375:Lestodon
288:Carlini
273:Carlini
204:Family:
188:Mammalia
178:Chordata
174:Phylum:
168:Animalia
154:Domain:
3370:1367752
3357:4834501
3217:Ocnopus
3120:Planops
2464:Mylodon
2112:Catonyx
2041:Neocnus
2034:Miocnus
1806:Class:
1783:Pilosan
1659:Bibcode
1609:Bibcode
1553:Bibcode
1488:Bibcode
1387:Bibcode
1281:Bibcode
1202:Bibcode
1121:7480153
1015:Bibcode
967:Bibcode
959:Geobios
851:wetland
844:Ipururo
748:Urumaco
744:Socorro
695:Guanare
689:on the
545:Planops
510:Bolivia
292:., 2006
277:., 2006
217:Genus:
194:Order:
184:Class:
1829:Pilosa
1785:genera
1740:
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776:sloths
740:FalcĂłn
725:Middle
681:So far
625:named
496:lobate
322:, and
316:Brazil
198:Pilosa
3383:43644
3365:IRMNG
1738:S2CID
1627:S2CID
1577:S2CID
1522:S2CID
1454:S2CID
1411:S2CID
1354:(PDF)
1313:S2CID
1269:(PDF)
1246:S2CID
1039:S2CID
848:Pebas
631:et al
504:from
290:et al
275:et al
3352:GBIF
1730:ISSN
1685:ISSN
1569:ISSN
1514:ISSN
1446:ISSN
1403:ISSN
1305:PMID
1297:ISSN
1238:PMID
1230:ISSN
1170:PMID
1162:ISSN
1126:PMID
1077:ISSN
1031:ISSN
983:ISSN
907:GBIF
871:and
846:and
824:and
810:and
800:and
784:and
770:and
750:and
727:and
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705:and
703:fish
324:Peru
58:Preęž’
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1712:doi
1675:hdl
1667:doi
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1561:doi
1504:hdl
1496:doi
1438:doi
1395:doi
1358:doi
1289:doi
1220:hdl
1210:doi
1154:doi
1116:PMC
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