232:
31:
428:. The hair on the dorsal region is of intense red coloring, and a large portion of the tail hair is black (basal), but the last two-fifths of the tail are white. Its ears are small and light brown, and feature tufts of long hair on the inner surface, but lack hair on the outer surface. Between the eye and the ear, there is a tuft of long black hairs. The thin whiskers can reach up to 13 cm (5 in) long, and has a strip of red fur around its neck.
83:
489:
58:
357:, Colombia. The specimen was of undetermined sex, and as all specimens of Smith's collections were collected by local hunters, there is no specific information regarding the location where the specimen was discovered. A second specimen, also of undetermined sex, was discovered in around 1913 in the
471:
is a nocturnal rodent, and its diet is unknown, but it is assumed to feed on plant matter such as fruits or seeds, like similar species in family
Echimyidae. It is able to easily climb a vertical wooden surface. The specimen observed in 2011 made no sounds. It is endemic to Colombia, and has only
440:
of the skull is large, and the interorbital region is very broad with nearly parallel sides. The facial portion of the skull is very short, and the distance between the incisors and the molars is slightly less than the length of the coronary surface of its upper row of teeth.
423:
specimens measure between 51 cm (20 in) from head to the tip of their tail, with their tails measuring between 18 and 28 cm (7.1 and 11.0 in). The species can weigh up to 500 g (18 oz), and has a woolly, soft, and long coat. The species is mainly
320:
in 1913 at the same location, and a further specimen observed in the same location in 2011. Found at altitudes of 700 to 2,000 meters, the species is endemic to
Colombia in an isolated area with high levels of biodiversity. The species was initially identified as
1113:, a protected area of Sierra Nevada de Santa Marta. Set up on 31 March 2006, this reserve covers 1,024 ha (2,530 acres) at an altitude between 950 and 2,600 m (3,120 and 8,530 ft), and contains a large number of endemic or endangered species.
431:
The upper surfaces of the front and rear legs are covered in a pale gray coat, and the hind legs are very short and wide. The feet lack small tubercles and the thumb is covered by a nail. Its skull is short and wide, and it has a heavy, large, and uncurved
1733:
Galewski, Thomas; Mauffrey, Jean-François; Leite, Yuri L. R.; Patton, James L.; Douzery, Emmanuel J. P. (2005). "Ecomorphological diversification among South
American spiny rats (Rodentia; Echimyidae): a phylogenetic and chronological approach".
472:
been found in Sierra Nevada de Santa Marta at altitudes of 700 to 2,000 m (2,300 to 6,600 ft). Due to the location's isolation and specific geological and climatic conditions, this mountainous region has high levels of biodiversity and
595:, reflecting the fact that these taxa have once been classified in the same genus. These two genera constitute the sister group of the "Dactylomyines", a clade of South American bamboo rats. All these taxa are closely related to the genera
1136:", with much of the forest in the potential range of the species having been cleared or degraded. Much of its potential area is infested with feral cats (who feed on fauna), and climate change is a long-term threat to the species.
361:
by
Carriker, but there is not much information regarding its location either, or the date of discovery. It is assumed that the specimen was obtained through a gift or was purchased. It was recorded as a
1804:
Fabre, Pierre-Henri; Galewski, Thomas; Tilak, Marie-ka; Douzery, Emmanuel J. P. (2013-03-01). "Diversification of South
American spiny rats (Echimyidae): a multigene phylogenetic approach".
1934:
Upham, Nathan S.; Patterson, Bruce D. (2015). "Evolution of
Caviomorph rodents: a complete phylogeny and timetree for living genera". In Vassallo, Aldo Ivan; Antenucci, Daniel (eds.).
3876:
1568:; Leite, Yuri L. R. (2016). "Family Echimyidae (hutias, South American spiny-rats and coypu)". In Wilson, Don E.; Lacher, Thomas E. Jr; Mittermeier, Russell A. (eds.).
1484:
1125:
1769:
Upham, Nathan S.; Patterson, Bruce D. (2012). "Diversification and biogeography of the
Neotropical caviomorph lineage Octodontoidea (Rodentia: Hystricognathi)".
3825:
1996:
1132:", given the lack of information regarding the distribution, ecology and habitat of this rodent. Following a reassessment in 2011, the species was listed as "
1877:
Fabre, Pierre-Henri; Vilstrup, Julia T.; Raghavan, Maanasa; Der
Sarkissian, Clio; Willerslev, Eske; Douzery, Emmanuel J. P.; Orlando, Ludovic (2014-07-01).
1655:; da Silva, Maria Nazareth F. (1996). "The Simultaneous Diversification of South American Echimyid Rodents (Hystricognathi) Based on Complete Cytochrome
1464:
1602:; Justy, Fabienne; Leite, Yuri L. R.; Loss, Ana Carolina; Orlando, Ludovic; Tilak, Marie-Ka; Patterson, Bruce D.; Douzery, Emmanuel J. P. (2017-03-01).
3773:
2018:
3911:
380:
1388:
537:, and identified a number of unique features of this species, including the differences between the teeth of this species and others in genus
3901:
1579:
1540:
1569:
3906:
3886:
367:
1252:
Noble, Elizabeth; McKeown, Simon; Sechrest, Wes (31 October 2011). "Rediscovery of
Santamartamys rufodorsalis (Rodent: Echimyidae)".
3896:
1433:
1369:
1950:
1420:(2005). "A revision of the genera of arboreal Echimyidae (Rodentia: Echimyidae, Echimyinae), with descriptions of two new genera".
3812:
3398:
3371:
3344:
3308:
2255:
2173:
1325:
3615:
3317:
2630:
2594:
2531:
1938:. Buenos Aires: SAREM Series A, Mammalogical Research — Sociedad Argentina para el Estudio de los MamĂferos. pp. 63–120.
1493:
1425:
1361:
3362:
3353:
3326:
2603:
2585:
2576:
62:
1300:
3891:
2489:
2423:
2414:
2405:
2363:
2182:
2011:
457:
358:
317:
309:
82:
3760:
3752:
3021:
2567:
2549:
3830:
1518:
3335:
2612:
2330:
2282:
2264:
2206:
1848:
692:
522:
1698:(2002). "Evolution of South American spiny rats (Rodentia, Echimyidae): the star-phylogeny hypothesis revisited".
3725:
3380:
3194:
3389:
2558:
2522:
2354:
342:
313:
1422:
Mammalian
Diversification: From Chromosomes to phylogeography (A Celebration of the Career of James L. Patton)
231:
3623:
2621:
2498:
2456:
2249:
2004:
534:
3687:
3251:
2980:
2957:
2866:
2540:
2432:
2273:
377:
photographed one individual at 1,958 m (6,424 ft) above sea level in El Dorado ProAves Reserve (
3765:
3176:
3130:
2998:
2843:
1879:"Rodents of the Caribbean: origin and diversification of hutias unravelled by next-generation museomics"
1133:
294:
204:
30:
2989:
2925:
2907:
2884:
3734:
3498:
3489:
3203:
3053:
2372:
1979:
1278:
374:
354:
3881:
3284:
3185:
3112:
3085:
3030:
2797:
2306:
2297:
2215:
2131:
976:
47:
1207:
3568:
3225:
3076:
2149:
1829:
437:
239:
77:
3607:
3552:
3448:
3439:
3275:
552:
comes from "Santa Marta", part of the name of the location where the specimens were found, and
3739:
3659:
3103:
2948:
2820:
2465:
2381:
2140:
1916:
1898:
1821:
1786:
1751:
1715:
1676:
1633:
1625:
1575:
1546:
1536:
1429:
1365:
1257:
1145:
1121:
1110:
456:
have a grey coat. During the transition to the adult intense red coat, molting begins at the
3853:
3584:
3167:
3094:
2875:
2715:
2706:
1906:
1890:
1813:
1778:
1743:
1707:
1668:
1615:
1599:
1530:
1460:
1417:
1326:"Espectacular mamĂfero redescubierto despuĂ©s de 113 años en la Sierra Nevada de Santa Marta"
1202:
530:
512:
492:
488:
221:
181:
3838:
3804:
3139:
3121:
2224:
1695:
1652:
1565:
278:
1128:(IUCN) described the species as "Vulnerable" in 1996. In 2008, IUCN re-classified it as "
1954:
3529:
3521:
3421:
2916:
2678:
2087:
2081:
1911:
1878:
1604:"Mitogenomic Phylogeny, Diversification, and Biogeography of South American Spiny Rats"
1129:
782:
433:
1711:
3870:
3799:
3640:
2645:
2035:
1817:
1526:
1193:
1150:
445:
has large eyes, which is consistent with its nocturnal behavior. It has two pairs of
370:
in New York. Despite several organized searches, no other specimens were discovered.
67:
1833:
1492:(in Spanish), Sistema de informaciĂłn sobre biodiversidad in Colombia, archived from
1329:
1184:
1095:
The cladogram has been reconstructed from mitochondrial and nuclear DNA characters.
3650:
3543:
2696:
409:
3817:
3786:
3719:
3576:
3560:
3512:
3470:
2739:
2730:
2321:
2093:
2042:
1782:
1747:
615:. In turn, these genera share phylogenetic affinities with the clade containing
350:
568:
meaning "red" and "back", respectively, referring to the color of this rodent.
301:, climate change, and the clearing of forest in its potential range in coastal
3158:
2778:
2480:
2122:
2103:
2028:
1005:
765:
526:
286:
154:
144:
3710:
1902:
1825:
1629:
1261:
395:
382:
3598:
2754:
2513:
2447:
2112:
1620:
1603:
1550:
1023:
863:
643:
580:
298:
164:
94:
1920:
1894:
1790:
1755:
1719:
1672:
1637:
3843:
1680:
3704:
3299:
2345:
2239:
2164:
2063:
922:
820:
660:
584:
517:
504:
473:
302:
289:. It is nocturnal and is believed to feed on plant matter, and is mainly
114:
1389:"Rediscovered in Santa Marta - rodent that has not been seen since 1898"
3778:
3242:
2669:
2396:
2197:
845:
713:
316:, a specimen identified by the American ornithologist and entomologist
308:
It is known only from three specimens, a specimen collected in 1898 in
3791:
3747:
3266:
2763:
2075:
2069:
2057:
1975:
1571:
Handbook of the Mammals of the World. Vol 6. Lagomorphs and Rodents I
1392:
425:
290:
134:
124:
104:
3681:
293:, with young specimens having a grey coat. IUCN list the species as
1472:. Vol. 12. Bulletin of the American Museum of Natural History.
3479:
487:
446:
1532:
Mammal Species of the World: A Taxonomic and Geographic Reference
3685:
2000:
1849:"Unexpected phylogenetic relationships of the painted tree rat
1951:"Lost & Found - Once upon a time, there was an adventurer"
1535:(3rd ed.). Johns Hopkins University Press. p. 1577.
1178:
1176:
1174:
1172:
1170:
1168:
1166:
1328:(in Spanish). ProAves Colombia. 19 May 2011. Archived from
541:. This resulted in the establishment of monotypic genus
1847:
Loss, Ana; Moura, Raquel T.; Leite, Yuri L. R. (2014).
992:
814:
759:
751:
743:
735:
654:
1936:
Biology of caviomorph rodents: diversity and evolution
503:
Known as the "red-crested tree rat", this species was
1301:"Presumed Extinct, The Red-Crested Tree Rat Returns"
3694:
3648:
3639:
3596:
3541:
3510:
3487:
3478:
3469:
3437:
3430:
3419:
3297:
3264:
3240:
3233:
3224:
3156:
3066:
3043:
3011:
2970:
2938:
2897:
2856:
2833:
2810:
2787:
2776:
2752:
2728:
2704:
2695:
2667:
2643:
2511:
2478:
2445:
2394:
2343:
2319:
2295:
2237:
2195:
2162:
2120:
2111:
2102:
1279:"Spectacular mammal rediscovered after 113 years!"
449:on the lateral edge of the abdominal side coat.
1574:. Barcelona: Lynx Edicions. pp. 552–641.
1126:International Union for Conservation of Nature
2012:
1208:10.2305/IUCN.UK.2017-2.RLTS.T6664A22210948.en
8:
1593:
1591:
587:Echimyidae rodents. The closest relative of
373:However, on 4 May 2011, two volunteers from
3877:IUCN Red List critically endangered species
3682:
3645:
3484:
3475:
3434:
3427:
3237:
3230:
2784:
2701:
2117:
2108:
2019:
2005:
1997:
1273:
1271:
230:
56:
29:
20:
1910:
1619:
1320:
1318:
1206:
16:Mammal in the spiny rat family of rodents
1412:
1410:
1408:
1406:
1404:
1402:
1383:
1381:
1247:
1245:
964: "Dactylomyines"
633:Genus-level cladogram of the Echimyini.
277:) is a species of tree-rat found in the
1598:Fabre, Pierre-Henri; Upham, Nathan S.;
1512:
1510:
1466:New Rodents from Colombia and Venezuela
1455:
1453:
1451:
1449:
1447:
1445:
1351:
1349:
1347:
1243:
1241:
1239:
1237:
1235:
1233:
1231:
1229:
1227:
1225:
1162:
1517:Woods, C.A.; Kilpatrick, C.W. (2005).
680: "Eumysopines"
515:in 1899, and was transferred to genus
408:) that was subsequently identified by
1771:Molecular Phylogenetics and Evolution
1736:Molecular Phylogenetics and Evolution
1700:Molecular Phylogenetics and Evolution
1661:Molecular Phylogenetics and Evolution
476:. It is believed that the habitat of
333:was created in 2005 for the species.
7:
1194:IUCN Red List of Threatened Species
996:
968:
913:
906:
836:
810:
755:
747:
739:
697: (Tuft-tailed spiny tree-rat)
684:
676:
650:
639:
2595:Blacked-spined Atlantic tree-rat (
1109:was rediscovered on 4 May 2011 in
464:Behavior, distribution and habitat
368:American Museum of Natural History
14:
3372:Mouse-tailed Atlantic spiny rat (
3309:White-spined Atlantic spiny rat (
2256:Yellow-crowned brush-tailed rat (
2174:Colombian soft-furred spiny rat (
345:identified the first specimen of
329:in 1935, and the monotypic genus
3318:Soft-spined Atlantic spiny rat (
2631:Short-furred Atlantic tree-rat (
2532:Orange-brown Atlantic tree-rat (
1818:10.1111/j.1463-6409.2012.00572.x
81:
3616:Isla De La Juventud tree hutia
3399:Yonenaga's Atlantic spiny rat (
3354:Dark-caped Atlantic spiny rat (
2604:Rusty-sided Atlantic tree-rat (
2586:Long-furred Atlantic tree-rat (
2577:Mantiqueira Atlantic tree-rat (
1608:Molecular Biology and Evolution
495:first described the species as
3912:Taxa named by Joel Asaph Allen
3345:Ihering's Atlantic spiny rat (
2490:Bare-tailed armored tree-rat (
2424:Tufted-tailed spiny tree-rat (
2415:Woolly-headed spiny tree-rat (
2364:Long-tailed armored tree-rat (
2183:Rufous soft-furred spiny rat (
1426:University of California Press
1183:Lacher, T.; Roach, N. (2017).
1116:In 1990, Emmons believed that
1:
3363:Moojen's Atlantic spiny rat (
1712:10.1016/s1055-7903(02)00279-8
1358:Walker's Mammals of the World
944: (Red-crested tree-rat)
3902:Sierra Nevada de Santa Marta
3336:Gracile Atlantic spiny rat (
3327:Elias's Atlantic spiny rat (
2613:Southern Atlantic tree-rat (
2355:Red-nosed armored tree-rat (
2331:Tuft-tailed spiny tree-rat (
2283:Sinnamary brush-tailed rat (
2265:Rio Negro brush-tailed rat (
2207:White-faced spiny tree-rat (
981: (Atlantic bamboo rat)
359:Sierra Nevada de Santa Marta
318:Melbourne Armstrong Carriker
310:Sierra Nevada de Santa Marta
3381:Spiked Atlantic spiny rat (
2406:Ferreira's spiny tree-rat (
2382:Peruvian armored tree-rat (
1976:"Reserva ProAves El Dorado"
1783:10.1016/j.ympev.2012.01.020
1748:10.1016/j.ympev.2004.11.015
460:region and moves backward.
3928:
3753:santamartamys-rufodorsalis
3696:Santamartamys rufodorsalis
3390:Hairy Atlantic spiny rat (
2568:Lund's Atlantic tree-rat (
2559:Pallid Atlantic tree-rat (
2550:Kerr's Atlantic tree-rat (
2523:Golden Atlantic tree-rat (
1187:Santamartamys rufodorsalis
523:George Henry Hamilton Tate
480:is suffering degradation.
325:in 1899, re-classified as
274:Santamartamys rufodorsalis
214:Santamartamys rufodorsalis
40:Santamartamys rufodorsalis
3907:Mammals described in 1899
3887:Endemic fauna of Colombia
2622:Giant Atlantic tree-rat (
2499:Speckled spiny tree-rat (
2457:White-tailed olalla rat (
2225:Vieira's spiny tree-rat (
2052:
2027:Extant species of family
1356:Nowak, Ronald M. (1999).
1111:El Dorado ProAves Reserve
1019:
1001:
994:
973:
966:
936:
918:
911:
904:
859:
841:
834:
816:
808:
779:
761:
753:
745:
737:
709:
689:
682:
674:
656:
648:
525:. In 2005, its subfamily
396:11.1008139°N 74.0720444°W
245:
238:
229:
210:
203:
78:Scientific classification
76:
54:
45:
37:
28:
23:
3897:Rodents of South America
3624:Prehensile-tailed hutia
3252:Broad-headed spiny rat (
2981:Short-tailed spiny rat (
2541:Drab Atlantic tree-rat (
2274:Plain brush-tailed rat (
366:specimen in 1913 at the
343:Herbert Huntingdon Smith
314:Herbert Huntingdon Smith
3131:Gray-footed spiny rat (
3022:O'Connell's spiny rat (
2999:Long-tailed spiny rat (
2844:Stiff-spine spiny rat (
2655:Red-crested tree-rat (
2373:Dark armored tree-rat (
1853:(Rodentia: Echimyidae)"
1254:ConservaciĂłn Colombiana
787: (Giant tree-rat)
560:comes from Latin, with
535:Smithsonian Institution
401:11.1008139; -74.0720444
1895:10.1098/rsbl.2014.0266
1673:10.1006/mpev.1996.0035
1529:; Reeder, D.M (eds.).
1418:Emmons, Louise Hickock
1120:was one of the rarest
1102:Threats and protection
511:by American biologist
500:
3740:Diplomys_rufodorsalis
3726:Diplomys rufodorsalis
3195:Sao Lourenço punaré (
3113:Magdalena spiny rat (
2958:Roberto's spiny rat (
2867:Gardner's spiny rat (
2798:Colombian spiny rat (
2433:Pará spiny tree-rat (
2307:Atlantic bamboo rat (
2216:Dark spiny tree-rat (
2132:Bolivian bamboo rat (
1621:10.1093/molbev/msw261
1564:Fabre, Pierre-Henri;
1521:Diplomys rufodorsalis
1134:Critically endangered
509:Isothrix rufodorsalis
497:Isothrix rufodorsalis
491:
341:On 24 December 1898,
327:Diplomys rufodorsalis
323:Isothrix rufodorsalis
295:critically endangered
254:Isothrix rufodorsalis
249:Diplomys rufodorsalis
63:Critically Endangered
24:Red-crested tree-rat
3077:Trinidad spiny rat (
2990:Cuvier's spiny rat (
2926:Steere's spiny rat (
2908:Goeldi's spiny rat (
2885:Patton's spiny rat (
2150:Montane bamboo rat (
1428:. pp. 247–310.
1201:: e.T6664A22210948.
505:originally described
297:: it is affected by
265:red-crested tree-rat
196:S. rufodorsalis
3892:Mammals of Colombia
3569:Black-tailed hutia
3204:Paraguayan punaré (
3104:Guyanan spiny rat (
3054:Simon's spiny rat (
2949:Guyenne spiny rat (
2821:Pacific spiny rat (
2466:Greedy olalla rat (
2141:Amazon bamboo rat (
1694:Leite, Yuri L. R.;
1281:. IUCN. 19 May 2011
579:is a member of the
556:, meaning "mouse".
452:Young specimens of
392: /
48:Conservation status
3660:Hispaniolan hutia
3587:(M. sanfelipensis)
3555:(M. angelcabrerai)
3499:Desmarest's hutia
3285:Fischer's guiara (
3226:Euryzygomatomyinae
3186:Highlands punaré (
3095:Guaira spiny rat (
3086:Boyacá spiny rat (
3031:Tome's spiny rat (
2876:Kulina spiny rat (
2716:Painted tree-rat (
2579:P. mantiqueirensis
1949:Lost & Found.
1851:Callistomys pictus
1496:on January 1, 2013
1303:. NPR. 19 May 2011
501:
438:supraorbital ridge
312:and identified by
3864:
3863:
3688:Taxon identifiers
3679:
3678:
3675:
3674:
3671:
3670:
3635:
3634:
3618:(M. meridionalis)
3585:San Felipe hutia
3465:
3464:
3461:
3460:
3449:Owl's spiny rat (
3415:
3414:
3411:
3410:
3276:Brandt's guiara (
3220:
3219:
3216:
3215:
3177:Foster's punaré (
3152:
3151:
3140:Sucre spiny rat (
3122:Minca spiny rat (
2919:P. quadruplicatus
2691:
2690:
2285:I. sinnamariensis
2250:I. barbarabrownae
2046:
2039:
2032:
1980:FundaciĂłn ProAves
1806:Zoologica Scripta
1651:Lara, Márcia C.;
1600:Emmons, Louise H.
1581:978-84-941892-3-4
1542:978-0-8018-8221-0
1461:Allen, Joel Asaph
1424:. Vol. 133.
1146:Fauna of Colombia
1099:
1098:
1089:
1088:
1080:
1079:
1071:
1070:
1062:
1061:
1053:
1052:
1044:
1043:
1035:
1034:
954:
953:
893:
892:
884:
883:
875:
874:
797:
796:
725:
724:
375:FundaciĂłn ProAves
261:
260:
185:
71:
3919:
3857:
3856:
3847:
3846:
3834:
3833:
3821:
3820:
3808:
3807:
3795:
3794:
3782:
3781:
3769:
3768:
3756:
3755:
3743:
3742:
3730:
3729:
3728:
3715:
3714:
3713:
3683:
3646:
3626:(M. prehensilis)
3608:Garrido's hutia
3553:Cabrera's hutia
3485:
3476:
3435:
3428:
3238:
3231:
3001:P. longicaudatus
2917:Napo spiny rat (
2785:
2702:
2679:Giant tree-rat (
2118:
2109:
2044:
2037:
2030:
2021:
2014:
2007:
1998:
1991:
1990:
1988:
1986:
1972:
1966:
1965:
1963:
1962:
1953:. Archived from
1946:
1940:
1939:
1931:
1925:
1924:
1914:
1874:
1868:
1867:
1860:Natureza on Line
1857:
1844:
1838:
1837:
1801:
1795:
1794:
1766:
1760:
1759:
1730:
1724:
1723:
1696:Patton, James L.
1691:
1685:
1684:
1653:Patton, James L.
1648:
1642:
1641:
1623:
1595:
1586:
1585:
1566:Patton, James L.
1561:
1555:
1554:
1514:
1505:
1504:
1503:
1501:
1480:
1474:
1473:
1471:
1457:
1440:
1439:
1414:
1397:
1396:
1385:
1376:
1375:
1364:. p. 1696.
1353:
1342:
1341:
1339:
1337:
1322:
1313:
1312:
1310:
1308:
1297:
1291:
1290:
1288:
1286:
1275:
1266:
1265:
1249:
1220:
1219:
1217:
1215:
1210:
1180:
997:
969:
914:
907:
837:
811:
756:
748:
740:
685:
677:
651:
640:
630:
629:
513:Joel Asaph Allen
493:Joel Asaph Allen
407:
406:
404:
403:
402:
397:
393:
390:
389:
388:
385:
269:Santa Marta toro
234:
216:
180:
86:
85:
65:
60:
59:
38:Illustration of
33:
21:
3927:
3926:
3922:
3921:
3920:
3918:
3917:
3916:
3867:
3866:
3865:
3860:
3852:
3850:
3842:
3839:Observation.org
3837:
3829:
3824:
3816:
3811:
3803:
3798:
3790:
3785:
3777:
3772:
3764:
3759:
3751:
3746:
3738:
3733:
3724:
3723:
3718:
3709:
3708:
3703:
3690:
3680:
3667:
3631:
3592:
3537:
3530:Bahamian hutia
3522:Jamaican coney
3506:
3457:
3407:
3293:
3267:Euryzygomatomys
3260:
3212:
3168:Common punaré (
3148:
3106:P. hoplomyoides
3062:
3039:
3033:P. semispinosus
3007:
2966:
2934:
2893:
2852:
2829:
2806:
2772:
2748:
2724:
2687:
2663:
2657:S. rufodorsalis
2639:
2534:P. brasiliensis
2507:
2501:P. semivillosus
2474:
2441:
2390:
2357:M. didelphoides
2339:
2315:
2291:
2233:
2191:
2158:
2098:
2048:
2025:
1995:
1994:
1984:
1982:
1974:
1973:
1969:
1960:
1958:
1948:
1947:
1943:
1933:
1932:
1928:
1889:(7): 20140266.
1883:Biology Letters
1876:
1875:
1871:
1855:
1846:
1845:
1841:
1803:
1802:
1798:
1768:
1767:
1763:
1732:
1731:
1727:
1693:
1692:
1688:
1650:
1649:
1645:
1597:
1596:
1589:
1582:
1563:
1562:
1558:
1543:
1516:
1515:
1508:
1499:
1497:
1482:
1481:
1477:
1469:
1459:
1458:
1443:
1436:
1416:
1415:
1400:
1387:
1386:
1379:
1372:
1360:. Vol. 1.
1355:
1354:
1345:
1335:
1333:
1324:
1323:
1316:
1306:
1304:
1299:
1298:
1294:
1284:
1282:
1277:
1276:
1269:
1251:
1250:
1223:
1213:
1211:
1182:
1181:
1164:
1159:
1142:
1104:
1090:
1081:
1072:
1063:
1054:
1045:
1036:
980:
955:
943:
894:
885:
876:
798:
786:
726:
696:
574:
548:The genus name
529:was revised by
486:
466:
418:
400:
398:
394:
391:
386:
383:
381:
379:
378:
339:
225:
218:
212:
199:
179:
80:
72:
61:
57:
50:
17:
12:
11:
5:
3925:
3923:
3915:
3914:
3909:
3904:
3899:
3894:
3889:
3884:
3879:
3869:
3868:
3862:
3861:
3859:
3858:
3848:
3835:
3822:
3809:
3796:
3783:
3770:
3757:
3744:
3731:
3716:
3700:
3698:
3692:
3691:
3686:
3677:
3676:
3673:
3672:
3669:
3668:
3666:
3665:
3656:
3654:
3643:
3637:
3636:
3633:
3632:
3630:
3629:
3621:
3613:
3604:
3602:
3594:
3593:
3591:
3590:
3582:
3574:
3571:(M. melanurus)
3566:
3558:
3549:
3547:
3539:
3538:
3536:
3535:
3532:(G. ingrahami)
3527:
3518:
3516:
3508:
3507:
3505:
3504:
3501:(C. pilorides)
3495:
3493:
3482:
3473:
3467:
3466:
3463:
3462:
3459:
3458:
3456:
3455:
3445:
3443:
3432:
3425:
3422:Incertae sedis
3417:
3416:
3413:
3412:
3409:
3408:
3406:
3405:
3396:
3387:
3378:
3369:
3360:
3356:T. mirapitanga
3351:
3342:
3333:
3324:
3315:
3305:
3303:
3295:
3294:
3292:
3291:
3282:
3272:
3270:
3262:
3261:
3259:
3258:
3248:
3246:
3235:
3228:
3222:
3221:
3218:
3217:
3214:
3213:
3211:
3210:
3201:
3192:
3183:
3174:
3164:
3162:
3154:
3153:
3150:
3149:
3147:
3146:
3137:
3128:
3119:
3110:
3101:
3092:
3088:P. chrysaeolus
3083:
3073:
3071:
3064:
3063:
3061:
3060:
3050:
3048:
3041:
3040:
3038:
3037:
3028:
3018:
3016:
3009:
3008:
3006:
3005:
2996:
2987:
2977:
2975:
2968:
2967:
2965:
2964:
2955:
2951:P. guyannensis
2945:
2943:
2936:
2935:
2933:
2932:
2923:
2914:
2904:
2902:
2895:
2894:
2892:
2891:
2882:
2873:
2863:
2861:
2854:
2853:
2851:
2850:
2846:P. echinothrix
2840:
2838:
2831:
2830:
2828:
2827:
2817:
2815:
2808:
2807:
2805:
2804:
2794:
2792:
2782:
2774:
2773:
2771:
2770:
2760:
2758:
2750:
2749:
2747:
2746:
2736:
2734:
2726:
2725:
2723:
2722:
2712:
2710:
2699:
2693:
2692:
2689:
2688:
2686:
2685:
2675:
2673:
2665:
2664:
2662:
2661:
2651:
2649:
2641:
2640:
2638:
2637:
2628:
2619:
2610:
2601:
2597:P. nigrispinus
2592:
2583:
2574:
2565:
2556:
2547:
2538:
2529:
2525:P. blainvillii
2519:
2517:
2509:
2508:
2506:
2505:
2496:
2486:
2484:
2476:
2475:
2473:
2472:
2463:
2453:
2451:
2443:
2442:
2440:
2439:
2430:
2421:
2412:
2402:
2400:
2392:
2391:
2389:
2388:
2379:
2370:
2361:
2351:
2349:
2341:
2340:
2338:
2337:
2327:
2325:
2317:
2316:
2314:
2313:
2303:
2301:
2293:
2292:
2290:
2289:
2280:
2271:
2262:
2253:
2245:
2243:
2235:
2234:
2232:
2231:
2222:
2213:
2203:
2201:
2193:
2192:
2190:
2189:
2180:
2170:
2168:
2160:
2159:
2157:
2156:
2147:
2138:
2134:D. boliviensis
2128:
2126:
2115:
2106:
2100:
2099:
2097:
2096:
2090:
2088:Hystricognathi
2084:
2082:Hystricomorpha
2078:
2072:
2066:
2060:
2053:
2050:
2049:
2036:Myocastoridae
2026:
2024:
2023:
2016:
2009:
2001:
1993:
1992:
1978:(in Spanish).
1967:
1941:
1926:
1869:
1839:
1812:(2): 117–134.
1796:
1777:(2): 417–429.
1761:
1742:(3): 601–615.
1725:
1706:(3): 455–464.
1686:
1667:(2): 403–413.
1643:
1614:(3): 613–633.
1587:
1580:
1556:
1541:
1506:
1475:
1441:
1434:
1398:
1395:. 18 May 2011.
1391:(in Spanish).
1377:
1370:
1343:
1314:
1292:
1267:
1221:
1161:
1160:
1158:
1155:
1154:
1153:
1148:
1141:
1138:
1130:Data deficient
1103:
1100:
1097:
1096:
1092:
1091:
1087:
1086:
1083:
1082:
1078:
1077:
1074:
1073:
1069:
1068:
1065:
1064:
1060:
1059:
1056:
1055:
1051:
1050:
1047:
1046:
1042:
1041:
1038:
1037:
1033:
1032:
1029:
1028:
1018:
1015:
1014:
1011:
1010:
1000:
995:
993:
989:
988:
985:
984:
972:
967:
965:
961:
960:
957:
956:
952:
951:
948:
947:
935:
932:
931:
928:
927:
917:
912:
910:
905:
903:
900:
899:
896:
895:
891:
890:
887:
886:
882:
881:
878:
877:
873:
872:
869:
868:
858:
855:
854:
851:
850:
840:
835:
833:
830:
829:
826:
825:
815:
809:
807:
804:
803:
800:
799:
795:
794:
791:
790:
778:
775:
774:
771:
770:
760:
754:
752:
746:
744:
738:
736:
732:
731:
728:
727:
723:
722:
719:
718:
708:
705:
704:
701:
700:
688:
683:
681:
675:
673:
670:
669:
666:
665:
655:
649:
647:
638:
635:
634:
573:
570:
485:
484:Classification
482:
465:
462:
434:zygomatic bone
417:
414:
338:
335:
285:in the family
259:
258:
257:
256:
251:
243:
242:
236:
235:
227:
226:
219:
208:
207:
201:
200:
193:
191:
187:
186:
172:
168:
167:
162:
158:
157:
152:
148:
147:
142:
138:
137:
132:
128:
127:
122:
118:
117:
112:
108:
107:
102:
98:
97:
92:
88:
87:
74:
73:
55:
52:
51:
46:
43:
42:
35:
34:
26:
25:
15:
13:
10:
9:
6:
4:
3:
2:
3924:
3913:
3910:
3908:
3905:
3903:
3900:
3898:
3895:
3893:
3890:
3888:
3885:
3883:
3880:
3878:
3875:
3874:
3872:
3855:
3849:
3845:
3840:
3836:
3832:
3827:
3823:
3819:
3814:
3810:
3806:
3801:
3797:
3793:
3788:
3784:
3780:
3775:
3771:
3767:
3762:
3758:
3754:
3749:
3745:
3741:
3736:
3732:
3727:
3721:
3717:
3712:
3706:
3702:
3701:
3699:
3697:
3693:
3689:
3684:
3664:
3663:
3658:
3657:
3655:
3653:
3652:
3647:
3644:
3642:
3641:Plagiodontini
3638:
3628:
3627:
3622:
3620:
3619:
3614:
3612:
3611:
3610:(M. garridoi)
3606:
3605:
3603:
3601:
3600:
3595:
3589:
3588:
3583:
3581:
3580:
3575:
3573:
3572:
3567:
3565:
3564:
3559:
3557:
3556:
3551:
3550:
3548:
3546:
3545:
3540:
3534:
3533:
3528:
3526:
3525:
3520:
3519:
3517:
3515:
3514:
3509:
3503:
3502:
3497:
3496:
3494:
3492:
3491:
3486:
3483:
3481:
3477:
3474:
3472:
3468:
3454:
3452:
3447:
3446:
3444:
3442:
3441:
3436:
3433:
3429:
3426:
3424:
3423:
3418:
3404:
3402:
3397:
3395:
3393:
3388:
3386:
3384:
3379:
3377:
3375:
3370:
3368:
3366:
3361:
3359:
3357:
3352:
3350:
3348:
3343:
3341:
3339:
3334:
3332:
3330:
3325:
3323:
3321:
3320:T. dimidiatus
3316:
3314:
3312:
3311:T. albispinus
3307:
3306:
3304:
3302:
3301:
3296:
3290:
3288:
3283:
3281:
3279:
3274:
3273:
3271:
3269:
3268:
3263:
3257:
3255:
3250:
3249:
3247:
3245:
3244:
3239:
3236:
3232:
3229:
3227:
3223:
3209:
3207:
3202:
3200:
3198:
3197:T. laurentius
3193:
3191:
3189:
3184:
3182:
3180:
3175:
3173:
3171:
3170:T. apereoides
3166:
3165:
3163:
3161:
3160:
3155:
3145:
3143:
3138:
3136:
3134:
3129:
3127:
3125:
3120:
3118:
3116:
3115:P. magdalenae
3111:
3109:
3107:
3102:
3100:
3098:
3093:
3091:
3089:
3084:
3082:
3080:
3079:P. trinitatus
3075:
3074:
3072:
3070:
3065:
3059:
3057:
3052:
3051:
3049:
3047:
3042:
3036:
3034:
3029:
3027:
3025:
3020:
3019:
3017:
3015:
3010:
3004:
3002:
2997:
2995:
2993:
2988:
2986:
2984:
2983:P. brevicauda
2979:
2978:
2976:
2974:
2973:longicaudatus
2969:
2963:
2961:
2956:
2954:
2952:
2947:
2946:
2944:
2942:
2937:
2931:
2929:
2924:
2922:
2920:
2915:
2913:
2911:
2906:
2905:
2903:
2901:
2896:
2890:
2888:
2883:
2881:
2879:
2874:
2872:
2870:
2865:
2864:
2862:
2860:
2855:
2849:
2847:
2842:
2841:
2839:
2837:
2832:
2826:
2824:
2819:
2818:
2816:
2814:
2809:
2803:
2801:
2800:P. canicollis
2796:
2795:
2793:
2791:
2786:
2783:
2781:
2780:
2775:
2769:
2767:
2762:
2761:
2759:
2757:
2756:
2751:
2745:
2743:
2740:Armored rat (
2738:
2737:
2735:
2733:
2732:
2727:
2721:
2719:
2714:
2713:
2711:
2709:
2708:
2703:
2700:
2698:
2694:
2684:
2682:
2677:
2676:
2674:
2672:
2671:
2666:
2660:
2658:
2653:
2652:
2650:
2648:
2647:
2646:Santamartamys
2642:
2636:
2634:
2629:
2627:
2625:
2620:
2618:
2616:
2611:
2609:
2607:
2602:
2600:
2598:
2593:
2591:
2589:
2584:
2582:
2580:
2575:
2573:
2571:
2566:
2564:
2562:
2557:
2555:
2553:
2548:
2546:
2544:
2539:
2537:
2535:
2530:
2528:
2526:
2521:
2520:
2518:
2516:
2515:
2510:
2504:
2502:
2497:
2495:
2493:
2488:
2487:
2485:
2483:
2482:
2477:
2471:
2469:
2464:
2462:
2460:
2455:
2454:
2452:
2450:
2449:
2444:
2438:
2436:
2431:
2429:
2427:
2422:
2420:
2418:
2413:
2411:
2409:
2404:
2403:
2401:
2399:
2398:
2393:
2387:
2385:
2380:
2378:
2376:
2371:
2369:
2367:
2362:
2360:
2358:
2353:
2352:
2350:
2348:
2347:
2342:
2336:
2334:
2329:
2328:
2326:
2324:
2323:
2318:
2312:
2310:
2305:
2304:
2302:
2300:
2299:
2298:Kannabateomys
2294:
2288:
2286:
2281:
2279:
2277:
2272:
2270:
2268:
2263:
2261:
2259:
2254:
2252:
2251:
2247:
2246:
2244:
2242:
2241:
2236:
2230:
2228:
2223:
2221:
2219:
2214:
2212:
2210:
2205:
2204:
2202:
2200:
2199:
2194:
2188:
2186:
2181:
2179:
2177:
2172:
2171:
2169:
2167:
2166:
2161:
2155:
2153:
2148:
2146:
2144:
2143:D. dactylinus
2139:
2137:
2135:
2130:
2129:
2127:
2125:
2124:
2119:
2116:
2114:
2110:
2107:
2105:
2101:
2095:
2091:
2089:
2085:
2083:
2079:
2077:
2073:
2071:
2067:
2065:
2061:
2059:
2055:
2054:
2051:
2047:
2040:
2033:
2022:
2017:
2015:
2010:
2008:
2003:
2002:
1999:
1981:
1977:
1971:
1968:
1957:on 2017-04-28
1956:
1952:
1945:
1942:
1937:
1930:
1927:
1922:
1918:
1913:
1908:
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1900:
1896:
1892:
1888:
1884:
1880:
1873:
1870:
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1861:
1854:
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1827:
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1815:
1811:
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1800:
1797:
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1788:
1784:
1780:
1776:
1772:
1765:
1762:
1757:
1753:
1749:
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1741:
1737:
1729:
1726:
1721:
1717:
1713:
1709:
1705:
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1697:
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1658:
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1647:
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1639:
1635:
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1613:
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1594:
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1577:
1573:
1572:
1567:
1560:
1557:
1552:
1548:
1544:
1538:
1534:
1533:
1528:
1524:
1522:
1513:
1511:
1507:
1495:
1491:
1490:
1489:(Allen, 1899)
1487:Santamartamys
1486:
1479:
1476:
1468:
1467:
1462:
1456:
1454:
1452:
1450:
1448:
1446:
1442:
1437:
1435:9780520098534
1431:
1427:
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1413:
1411:
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1405:
1403:
1399:
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1390:
1384:
1382:
1378:
1373:
1371:9780801857898
1367:
1363:
1359:
1352:
1350:
1348:
1344:
1332:on 2011-08-23
1331:
1327:
1321:
1319:
1315:
1302:
1296:
1293:
1280:
1274:
1272:
1268:
1263:
1259:
1256:(15): 40–43.
1255:
1248:
1246:
1244:
1242:
1240:
1238:
1236:
1234:
1232:
1230:
1228:
1226:
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1209:
1204:
1200:
1196:
1195:
1190:
1188:
1179:
1177:
1175:
1173:
1171:
1169:
1167:
1163:
1156:
1152:
1151:IUCN Red List
1149:
1147:
1144:
1143:
1139:
1137:
1135:
1131:
1127:
1124:mammals. The
1123:
1119:
1118:Santamartamys
1114:
1112:
1108:
1107:Santamartamys
1101:
1094:
1093:
1085:
1084:
1076:
1075:
1067:
1066:
1058:
1057:
1049:
1048:
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1025:
1017:
1016:
1013:
1012:
1009:
1008:
1007:
999:
998:
991:
990:
987:
986:
983:
982:
979:
978:Kannabateomys
971:
970:
963:
962:
959:
958:
950:
949:
946:
945:
942:
941:Santamartamys
934:
933:
930:
929:
926:
925:
924:
916:
915:
909:
908:
902:
901:
898:
897:
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888:
880:
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871:
870:
867:
866:
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857:
856:
853:
852:
849:
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847:
839:
838:
832:
831:
828:
827:
824:
823:
822:
813:
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806:
805:
802:
801:
793:
792:
789:
788:
785:
777:
776:
773:
772:
769:
768:
767:
758:
757:
750:
749:
742:
741:
734:
733:
730:
729:
721:
720:
717:
716:
715:
707:
706:
703:
702:
699:
698:
695:
687:
686:
679:
678:
672:
671:
668:
667:
664:
663:
662:
653:
652:
645:
642:
641:
637:
636:
632:
631:
628:
626:
622:
618:
614:
610:
606:
602:
598:
594:
590:
589:Santamartamys
586:
582:
578:
577:Santamartamys
571:
569:
567:
563:
559:
555:
551:
550:Santamartamys
546:
544:
543:Santamartamys
540:
536:
532:
531:Louise Emmons
528:
524:
520:
519:
514:
510:
506:
498:
494:
490:
483:
481:
479:
478:Santamartamys
475:
470:
469:Santamartamys
463:
461:
459:
455:
454:Santamartamys
450:
448:
444:
443:Santamartamys
439:
435:
429:
427:
422:
421:Santamartamys
415:
413:
411:
405:
387:74°04′19.36″W
384:11°06′02.93″N
376:
371:
369:
365:
364:Santamartamys
360:
356:
352:
348:
347:Santamartamys
344:
336:
334:
332:
331:Santamartamys
328:
324:
319:
315:
311:
306:
304:
300:
296:
292:
288:
284:
283:Santamartamys
280:
276:
275:
270:
266:
255:
252:
250:
247:
246:
244:
241:
237:
233:
228:
223:
217:
215:
209:
206:
205:Binomial name
202:
198:
197:
192:
189:
188:
183:
178:
177:
176:Santamartamys
173:
170:
169:
166:
163:
160:
159:
156:
153:
150:
149:
146:
143:
140:
139:
136:
133:
130:
129:
126:
123:
120:
119:
116:
113:
110:
109:
106:
103:
100:
99:
96:
93:
90:
89:
84:
79:
75:
69:
64:
53:
49:
44:
41:
36:
32:
27:
22:
19:
3695:
3661:
3651:Plagiodontia
3649:
3625:
3617:
3609:
3597:
3586:
3578:
3577:Dwarf hutia
3570:
3563:(M. auritus)
3562:
3561:Eared hutia
3554:
3544:Mesocapromys
3542:
3531:
3524:(G. brownii)
3523:
3511:
3500:
3488:
3451:C. sulcidens
3450:
3438:
3420:
3401:T. yonenagae
3400:
3391:
3382:
3373:
3364:
3355:
3346:
3338:T. gratiosus
3337:
3328:
3319:
3310:
3298:
3286:
3277:
3265:
3253:
3241:
3206:T. pachyurus
3205:
3196:
3187:
3178:
3169:
3157:
3141:
3132:
3123:
3114:
3105:
3096:
3087:
3078:
3068:
3055:
3045:
3032:
3024:P. oconnelli
3023:
3014:semispinosus
3013:
3000:
2991:
2982:
2972:
2959:
2950:
2940:
2927:
2918:
2909:
2899:
2886:
2877:
2868:
2858:
2845:
2835:
2823:P. decumanus
2822:
2812:
2799:
2789:
2777:
2765:
2753:
2741:
2729:
2717:
2705:
2697:Myocastorini
2680:
2668:
2656:
2654:
2644:
2632:
2623:
2614:
2605:
2596:
2587:
2578:
2569:
2560:
2551:
2543:P. dasythrix
2542:
2533:
2524:
2512:
2500:
2491:
2479:
2467:
2459:O. albicauda
2458:
2446:
2434:
2425:
2416:
2407:
2395:
2384:M. rhipidura
2383:
2374:
2365:
2356:
2344:
2332:
2320:
2309:K. amblyonyx
2308:
2296:
2284:
2275:
2267:I. negrensis
2266:
2258:I. bistriata
2257:
2248:
2238:
2226:
2217:
2209:E. chrysurus
2208:
2196:
2184:
2175:
2163:
2151:
2142:
2133:
2121:
2086:Infraorder:
2043:Capromyidae
2034:, including
2031:(Spiny rats)
1983:. Retrieved
1970:
1959:. Retrieved
1955:the original
1944:
1935:
1929:
1886:
1882:
1872:
1863:
1859:
1850:
1842:
1809:
1805:
1799:
1774:
1770:
1764:
1739:
1735:
1728:
1703:
1699:
1689:
1664:
1660:
1659:Sequences".
1656:
1646:
1611:
1607:
1570:
1559:
1531:
1527:Wilson, D.E.
1520:
1498:, retrieved
1494:the original
1488:
1485:
1483:BSi (2012),
1478:
1465:
1421:
1357:
1334:. Retrieved
1330:the original
1305:. Retrieved
1295:
1283:. Retrieved
1253:
1212:. Retrieved
1198:
1192:
1186:
1117:
1115:
1106:
1105:
1022:
1020:
1004:
1002:
977:
974:
940:
939:
937:
921:
919:
862:
860:
844:
842:
819:
817:
783:
780:
764:
762:
712:
710:
693:
690:
659:
657:
624:
620:
616:
612:
608:
604:
600:
596:
592:
588:
576:
575:
565:
561:
558:Rufodorsalis
557:
553:
549:
547:
542:
538:
516:
508:
502:
496:
477:
468:
467:
453:
451:
442:
430:
420:
419:
410:Paul Salaman
372:
363:
346:
340:
330:
326:
322:
307:
282:
273:
272:
268:
264:
262:
253:
248:
213:
211:
195:
194:
175:
174:
39:
18:
3787:iNaturalist
3720:Wikispecies
3662:(P. aedium)
3513:Geocapromys
3471:Capromyinae
3374:T. myosuros
3347:T. iheringi
3287:E. spinosus
3254:C. laticeps
3133:P. poliopus
2941:guyannensis
2869:P. gardneri
2836:echinothrix
2742:H. gymnurus
2707:Callistomys
2633:P. unicolor
2492:P. occasius
2435:M. stimulax
2426:M. occultus
2417:M. leniceps
2408:M. hispidus
2322:Lonchothrix
2218:E. saturnus
2176:D. caniceps
2152:D. peruanus
2094:Caviomorpha
2092:Parvorder:
2029:Echimyidae
1500:28 November
1336:28 November
1307:18 November
1285:28 November
1214:17 November
1122:Neotropical
694:Lonchothrix
623:, and with
617:Lonchothrix
521:in 1935 by
416:Description
399: /
351:Santa Marta
222:J. A. Allen
151:Subfamily:
3882:Echimyidae
3871:Categories
3579:(M. nanus)
3480:Capromyini
3440:Carterodon
3392:T. setosus
3383:T. paratus
3365:T. moojeni
3188:T. inermis
3179:T. fosteri
3159:Thrichomys
3097:P. guairae
3069:trinitatus
3056:P. simonsi
2992:P. cuvieri
2960:P. roberti
2928:P. steerei
2910:P. goeldii
2887:P. pattoni
2878:P. kulinae
2790:canicollis
2779:Proechimys
2681:T. grandis
2624:P. thomasi
2615:P. sulinus
2606:P. pattoni
2561:P. lamarum
2481:Pattonomys
2375:M. obscura
2366:M. macrura
2333:L. emiliae
2276:I. pagurus
2227:E. vieirai
2185:D. labilis
2123:Dactylomys
2104:Echimyinae
2080:Suborder:
1961:2018-04-08
1866:: 132–136.
1157:References
1006:Dactylomys
766:Pattonomys
609:Pattonomys
527:Echimyinae
349:in Ocana,
299:feral cats
287:Echimyidae
155:Echimyinae
145:Echimyidae
3599:Mysateles
3329:T. eliasi
3278:E. guiara
3142:P. urichi
3124:P. mincae
2813:decumanus
2766:M. coypus
2755:Myocastor
2718:C. pictus
2588:P. medius
2514:Phyllomys
2448:Olallamys
2113:Echimyini
2056:Kingdom:
1903:1744-9561
1826:1463-6409
1630:0737-4038
1519:"Species
1362:JHU Press
1262:1900-1592
1024:Olallamys
864:Phyllomys
644:Echimyini
601:Phyllomys
583:clade of
581:Echimyini
572:Phylogeny
533:from the
355:Magdalena
337:Discovery
279:monotypic
190:Species:
165:Echimyini
101:Kingdom:
95:Eukaryota
3705:Wikidata
3490:Capromys
3300:Trinomys
2859:gardneri
2764:Nutria (
2731:Hoplomys
2570:P. lundi
2552:P. kerri
2346:Makalata
2240:Isothrix
2165:Diplomys
2076:Rodentia
2070:Mammalia
2064:Chordata
2062:Phylum:
2058:Animalia
2045:(Hutias)
2038:(Coypus)
1921:25115033
1834:83639441
1791:22327013
1756:15683932
1720:12450750
1638:28025278
1551:62265494
1463:(1899).
1140:See also
923:Diplomys
821:Makalata
661:Isothrix
625:Isothrix
605:Makalata
593:Diplomys
585:arboreal
566:dorsalis
539:Diplomys
518:Diplomys
499:in 1899.
474:endemism
458:anterior
303:Colombia
240:Synonyms
141:Family:
135:Rodentia
125:Mammalia
115:Chordata
111:Phylum:
105:Animalia
91:Domain:
68:IUCN 3.1
3831:1568970
3818:1001440
3779:5786579
3711:Q301130
3243:Clyomys
3046:simonsi
2900:goeldii
2670:Toromys
2468:O. edax
2397:Mesomys
2198:Echimys
2074:Order:
2068:Class:
1912:4126619
1681:8728398
1021:
1003:
975:
938:
920:
861:
846:Echimys
843:
818:
784:Toromys
781:
763:
714:Mesomys
711:
691:
658:
646:
621:Mesomys
613:Toromys
597:Echimys
224:, 1899)
171:Genus:
161:Tribe:
131:Order:
121:Class:
66: (
3854:106555
3851:uBio:
3844:201231
3766:914739
3748:ARKive
3067:group
3044:group
3012:group
2971:group
2939:group
2898:group
2857:group
2834:group
2811:group
2788:group
1985:16 May
1919:
1909:
1901:
1832:
1824:
1789:
1754:
1718:
1679:
1636:
1628:
1578:
1549:
1539:
1432:
1393:Semana
1368:
1260:
611:, and
447:udders
436:. The
426:rufous
291:rufous
281:genus
184:, 2005
182:Emmons
3792:75061
1856:(PDF)
1830:S2CID
1525:. In
1470:(PDF)
562:rufus
3826:NCBI
3805:6664
3800:IUCN
3774:GBIF
3761:BOLD
2041:and
1987:2020
1917:PMID
1899:ISSN
1822:ISSN
1787:PMID
1752:PMID
1716:PMID
1677:PMID
1634:PMID
1626:ISSN
1576:ISBN
1547:OCLC
1537:ISBN
1502:2014
1430:ISBN
1366:ISBN
1338:2014
1309:2012
1287:2014
1258:ISSN
1216:2021
1199:2017
619:and
564:and
263:The
3813:MDD
3735:ADW
1907:PMC
1891:doi
1814:doi
1779:doi
1744:doi
1708:doi
1669:doi
1616:doi
1203:doi
591:is
554:mys
507:as
267:or
3873::
3841::
3828::
3815::
3802::
3789::
3776::
3763::
3750::
3737::
3722::
3707::
1915:.
1905:.
1897:.
1887:10
1885:.
1881:.
1864:12
1862:.
1858:.
1828:.
1820:.
1810:42
1808:.
1785:.
1775:63
1773:.
1750:.
1740:34
1738:.
1714:.
1704:25
1702:.
1675:.
1663:.
1632:.
1624:.
1612:34
1610:.
1606:.
1590:^
1545:.
1509:^
1444:^
1401:^
1380:^
1346:^
1317:^
1270:^
1224:^
1197:.
1191:.
1165:^
627:.
607:,
603:,
599:,
545:.
412:.
353:,
305:.
3453:)
3431:—
3403:)
3394:)
3385:)
3376:)
3367:)
3358:)
3349:)
3340:)
3331:)
3322:)
3313:)
3289:)
3280:)
3256:)
3234:—
3208:)
3199:)
3190:)
3181:)
3172:)
3144:)
3135:)
3126:)
3117:)
3108:)
3099:)
3090:)
3081:)
3058:)
3035:)
3026:)
3003:)
2994:)
2985:)
2962:)
2953:)
2930:)
2921:)
2912:)
2889:)
2880:)
2871:)
2848:)
2825:)
2802:)
2768:)
2744:)
2720:)
2683:)
2659:)
2635:)
2626:)
2617:)
2608:)
2599:)
2590:)
2581:)
2572:)
2563:)
2554:)
2545:)
2536:)
2527:)
2503:)
2494:)
2470:)
2461:)
2437:)
2428:)
2419:)
2410:)
2386:)
2377:)
2368:)
2359:)
2335:)
2311:)
2287:)
2278:)
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2220:)
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2187:)
2178:)
2154:)
2145:)
2136:)
2020:e
2013:t
2006:v
1989:.
1964:.
1923:.
1893::
1836:.
1816::
1793:.
1781::
1758:.
1746::
1722:.
1710::
1683:.
1671::
1665:5
1657:b
1640:.
1618::
1584:.
1553:.
1523:"
1438:.
1374:.
1340:.
1311:.
1289:.
1264:.
1218:.
1205::
1189:"
1185:"
271:(
220:(
70:)
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
